1196 lines
138 KiB
XML
1196 lines
138 KiB
XML
<document id="17E110A63296F2D205678283BBFAC985" ID-CLB-Dataset="38930" ID-DOI="10.11646/zootaxa.4118.1.1" ID-GBIF-Dataset="a4e408c4-2575-4b5c-878b-e1b9010ec619" ID-ISSN="1175-5326" ID-Zenodo-Dep="264597" ID-ZooBank="CFA586DA-10EE-468B-80EE-35351E3845FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465242077321" checkinUser="plazi" docAuthor="C, Lees David" docDate="2016" docId="038747324C6DC6481EB72DBAFBA8218A" docLanguage="en" docName="zootaxa.4118.pdf.imf" docOrigin="Zootaxa 4118 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heteropsis harveyi Lees & Kremen, sp. nov." docType="treatment" docVersion="9" lastPageNumber="31" masterDocId="FFBE3F4A4C76C6571E202D2DFFD32174" masterDocTitle="Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision" masterLastPageNumber="97" masterPageNumber="1" pageNumber="28" updateTime="1698662715872" updateUser="ExternalLinkService">
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<mods:title id="66C351B03350A5E1FAA457AADFED7038">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
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<mods:namePart id="128EF6E932E6E65C10131A2ECB08190B">C, Lees David</mods:namePart>
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<mods:title id="19AEAF32BCBB7DF4A8A58FD14D6B582F">Zootaxa</mods:title>
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<mods:date id="765D0A25B2A872CE2D8A4E1A8C6C2B49">2016</mods:date>
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<mods:number id="C9BC912B61282329742DEF06924C4D27">4118</mods:number>
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<treatment id="038747324C6DC6481EB72DBAFBA8218A" ID-DOI="http://doi.org/10.5281/zenodo.6086418" ID-GBIF-Taxon="120817090" ID-Zenodo-Dep="6086418" LSID="urn:lsid:plazi:treatment:038747324C6DC6481EB72DBAFBA8218A" httpUri="http://treatment.plazi.org/id/038747324C6DC6481EB72DBAFBA8218A" lastPageId="31" lastPageNumber="31" pageId="27" pageNumber="28">
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<subSubSection id="C334A5AF4C6DC64C1EB72DBAFBC621A7" pageId="27" pageNumber="28" type="nomenclature">
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<paragraph id="8B91F6244C6DC64C1EB72DBAFD6121C6" blockId="27.[151,1045,151,211]" box="[151,690,151,178]" pageId="27" pageNumber="28">
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<heading id="D0D941484C6DC64C1EB72DBAFD6121C6" bold="true" box="[151,690,151,178]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
|
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<emphasis id="B95A2A364C6DC64C1EB72DBAFD6121C6" bold="true" box="[151,690,151,178]" pageId="27" pageNumber="28">
|
||
<taxonomicName id="4C2E8DA74C6DC64C1EB72DBAFD9421C6" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[151,583,151,178]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi" status="sp. nov.">
|
||
<emphasis id="B95A2A364C6DC64C1EB72DBAFEAA21C5" bold="true" box="[151,377,151,177]" italics="true" pageId="27" pageNumber="28">Heteropsis harveyi</emphasis>
|
||
Lees & Kremen
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A269974D4C6DC64C1C742DB5FD6121C6" box="[596,690,152,178]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6DC64C1EB72D96FBC621A7" blockId="27.[151,1045,151,211]" box="[151,1045,186,211]" pageId="27" pageNumber="28">LSID: urn:lsid:zoobank.org:act:CD7A88FE-87A6-4324-8404-5EDE47D8E9F2</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C6DC64C1EB72DD2FAEB206C" box="[151,1336,255,280]" pageId="27" pageNumber="28" type="description">
|
||
<paragraph id="8B91F6244C6DC64C1EB72DD2FAEB206C" blockId="27.[151,1437,255,2020]" box="[151,1336,255,280]" pageId="27" pageNumber="28">
|
||
<emphasis id="B95A2A364C6DC64C1EB72DD2FE84206C" bold="true" box="[151,343,255,280]" pageId="27" pageNumber="28">Prior references</emphasis>
|
||
:
|
||
<taxonomicName id="4C2E8DA74C6DC64C1F462DD2FD03206C" authority="Mabille" authorityName="Mabille" box="[358,720,255,280]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="andravahana">
|
||
<emphasis id="B95A2A364C6DC64C1F462DD2FDA1206C" box="[358,626,255,280]" italics="true" pageId="27" pageNumber="28">Mycalesis andravahana</emphasis>
|
||
Mabille
|
||
</taxonomicName>
|
||
♂ “var.” (
|
||
<bibRefCitation id="EFBF8BD54C6DC64C1D642DD2FC21206C" author="Mabille" box="[836,1010,255,280]" pageId="27" pageNumber="28" refString="Mabille, P. (1885 - 1887) Histoire, Physique, Naturelle et Politique de Madagascar. Vol. 18 - 19. Imprimerie Nationale, Paris, 364 pp., 55 pls. [Grandidier, A. [1885 - 1887] (Ed.)] http: // dx. doi. org / 10.5962 / bhl. title. 1599" type="book chapter" year="1885">Mabille [1885]</bibRefCitation>
|
||
: Pl. 5
|
||
<figureCitation id="1315EAA14C6DC64C1A182DD2FBA8206C" box="[1080,1147,255,280]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="27" pageNumber="28">Fig. 8</figureCitation>
|
||
, [1887: 57–59]);
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C6DC64C1EE72C08FAC023BC" pageId="27" pageNumber="28" type="reference_group">
|
||
<paragraph id="8B91F6244C6DC64C1EE72C08FB4120F1" blockId="27.[151,1437,255,2020]" pageId="27" pageNumber="28">
|
||
<taxonomicName id="4C2E8DA74C6DC64C1EE72C08FE072049" ID-CoL="6M28R" box="[199,468,292,317]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="andravahana">
|
||
<emphasis id="B95A2A364C6DC64C1EE72C08FE072049" box="[199,468,292,317]" italics="true" pageId="27" pageNumber="28">Henotesia andravahana</emphasis>
|
||
</taxonomicName>
|
||
ab.
|
||
<emphasis id="B95A2A364C6DC64C1C242C08FD512048" box="[516,642,293,316]" italics="true" pageId="27" pageNumber="28">marmorata</emphasis>
|
||
<bibRefCitation id="EFBF8BD54C6DC64C1CA82C09FC942048" author="Aurivillius" box="[648,839,292,317]" pageId="27" pageNumber="28" refString="Aurivillius, P. O. C. (1911) In: Seitz, A. (Ed.), (1908 - 1925) Die Gross-Schmetterlinge der Erde. (2) 13 - Afrikanischen Faunengebietes, A. Kerner, Stuttgart, pp. 1 - 613, pls. 1 - 80." type="journal article" year="1911">Aurivillius, 1911</bibRefCitation>
|
||
(unavailable name;
|
||
<typeStatus id="549548864C6DC64C1A0A2C09FB5F2049" box="[1066,1164,292,317]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
examined in Stockholm and conspecific); Gaede 1931: 401;
|
||
<bibRefCitation id="EFBF8BD54C6DC64C1C602C6AFCFB2014" author="Ackery" box="[576,808,327,352]" pageId="27" pageNumber="28" refString="Ackery, P. R., Smith, C. R. & Vane-Wright, R. I. (1995) Carcasson's African Butterflies: An annotated Catalogue of the Papilionoidea and Hesperioidea of the Afrotropical Region. British Museum (Natural History), London, xi + 803 pp." type="book" year="1995">
|
||
Ackery
|
||
<emphasis id="B95A2A364C6DC64C1CBE2C64FD322014" box="[670,737,327,352]" italics="true" pageId="27" pageNumber="28">et al.,</emphasis>
|
||
1995
|
||
</bibRefCitation>
|
||
: 297 (297 [raised from synonymy with
|
||
<taxonomicName id="4C2E8DA74C6DC64C1ADF2C64FEE220F0" ID-CoL="3KV9C" authority="Ward, 1871" authorityName="Ward" authorityYear="1871" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="anganavo">
|
||
<emphasis id="B95A2A364C6DC64C1ADF2C64FA482014" box="[1279,1435,329,352]" italics="true" pageId="27" pageNumber="28">Ht. anganavo</emphasis>
|
||
(Ward, 1871)
|
||
</taxonomicName>
|
||
]; the name ‘
|
||
<emphasis id="B95A2A364C6DC64C1F9A2C40FDEB20F0" box="[442,568,365,388]" italics="true" pageId="27" pageNumber="28">marmorata</emphasis>
|
||
’ was incorrectly linked to “sp. 13” in
|
||
<bibRefCitation id="EFBF8BD54C6DC64C1DC42C40FBB020F0" author="Lees" box="[996,1123,364,388]" pageId="27" pageNumber="28" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||
: 64;
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6DC64C1EE72CBCFB3E23D0" blockId="27.[151,1437,255,2020]" pageId="27" pageNumber="28">
|
||
<emphasis id="B95A2A364C6DC64C1EE72CBCFE4B20DC" box="[199,408,399,424]" italics="true" pageId="27" pageNumber="28">
|
||
<taxonomicName id="4C2E8DA74C6DC64C1EE72CBCFE5F20DC" ID-CoL="92BDD" authority="Wardii" authorityName="Wardii" box="[199,396,399,424]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">Henotesia Wardii</taxonomicName>
|
||
/
|
||
</emphasis>
|
||
<taxonomicName id="4C2E8DA74C6DC64C1FB82CA2FC7B20DC" ID-CoL="92BDD" authority="Andravahana (Oberthur, 1916: 205)" authorityName="Andravahana (Oberthur" authorityPageNumber="205" authorityYear="1916" box="[408,936,399,424]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A2A364C6DC64C1FB82CA2FD7620DC" box="[408,677,399,424]" italics="true" pageId="27" pageNumber="28">Henotesia Andravahana</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBF8BD54C6DC64C1C952CA2FC7320DC" author="Oberthur" box="[693,928,399,424]" pageId="27" pageNumber="28" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916: 205</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Charles Oberthür labelled his specimens in his original drawer ambiguously as “
|
||
<emphasis id="B95A2A364C6DC64C1C672C99FCFD20B9" box="[583,814,436,461]" italics="true" pageId="27" pageNumber="28">
|
||
<taxonomicName id="4C2E8DA74C6DC64C1C672C99FD5C20B9" ID-CoL="934V4" box="[583,655,436,461]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="wardii">wardii</taxonomicName>
|
||
-
|
||
<taxonomicName id="4C2E8DA74C6DC64C1CB92C99FCFD20B9" ID-CoL="6M28R" box="[665,814,436,461]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="andravahana">andravahana</taxonomicName>
|
||
</emphasis>
|
||
” (see
|
||
<bibRefCitation id="EFBF8BD54C6DC64C1D5D2C99FBFC20B9" author="Oberthur" box="[893,1071,436,461]" pageId="27" pageNumber="28" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||
: 204–205;
|
||
<bibRefCitation id="EFBF8BD54C6DC64C1A922C98FAE620B8" author="Lees" box="[1202,1333,436,460]" pageId="27" pageNumber="28" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||
: 64). He stated (
|
||
<bibRefCitation id="EFBF8BD54C6DC64C1ECD2CFAFE082084" author="Oberthur" box="[237,475,471,496]" pageId="27" pageNumber="28" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür 1916: 205</bibRefCitation>
|
||
) that he had in his collection “22 ♂♂ et 10 ♀♀ d'
|
||
<taxonomicName id="4C2E8DA74C6DC64C1A7D2CFAFB262084" ID-CoL="6M28R" box="[1117,1269,471,496]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="43" phylum="Arthropoda" rank="species" species="andravahana">
|
||
<emphasis id="B95A2A364C6DC64C1A7D2CFAFB262084" box="[1117,1269,471,496]" italics="true" pageId="27" pageNumber="28">Andravahana</emphasis>
|
||
</taxonomicName>
|
||
provenant de Fianarantsoa et d'Antsianaka” (subsequently known as
|
||
<taxonomicName id="4C2E8DA74C6DC64C1D202CD0FB9B2360" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" box="[768,1096,508,532]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="viettei">
|
||
<emphasis id="B95A2A364C6DC64C1D202CD0FC102360" box="[768,963,509,532]" italics="true" pageId="27" pageNumber="28">Heteropsis viettei</emphasis>
|
||
Lees, 2003
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFBF8BD54C6DC64C1A742CD0FB002360" author="Lees" box="[1108,1235,508,532]" pageId="27" pageNumber="28" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||
: 62; 391–392, but here treated as
|
||
<taxonomicName id="4C2E8DA74C6DC64C1F772F0CFDA0234C" box="[343,627,543,568]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="andravahana">
|
||
<emphasis id="B95A2A364C6DC64C1F772F0CFDA0234C" box="[343,627,543,568]" italics="true" pageId="27" pageNumber="28">Heteropsis andravahana</emphasis>
|
||
</taxonomicName>
|
||
), “et 78 ♂♂ et 45 ♀♀ du supposé
|
||
<taxonomicName id="4C2E8DA74C6DC64C1A612F32FB5A234C" ID-CoL="934V4" box="[1089,1161,543,568]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="36" phylum="Arthropoda" rank="species" species="wardii">
|
||
<emphasis id="B95A2A364C6DC64C1A612F32FB5A234C" box="[1089,1161,543,568]" italics="true" pageId="27" pageNumber="28">Wardii</emphasis>
|
||
</taxonomicName>
|
||
, capturés également à Fianarantsoa et dans l’Antsianaka"; referred to here as
|
||
<taxonomicName id="4C2E8DA74C6DC64C1D322F68FC412329" box="[786,914,580,605]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6DC64C1D322F68FC412329" box="[786,914,580,605]" italics="true" pageId="27" pageNumber="28">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
; All 78 ♂♂ and 13 of the ♀♀ of Oberthür’s “
|
||
<taxonomicName id="4C2E8DA74C6DC64C1E832F4AFF3823F4" ID-CoL="934V4" box="[163,235,615,640]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="wardii">
|
||
<emphasis id="B95A2A364C6DC64C1E832F4AFF3823F4" box="[163,235,615,640]" italics="true" pageId="27" pageNumber="28">wardii</emphasis>
|
||
</taxonomicName>
|
||
” (most of the latter from Antsianaka) have been located (in BMNH). An additional four ♀♀ placed above the "
|
||
<taxonomicName id="4C2E8DA74C6DC64C1EEC2FA1FEC723D1" ID-CoL="934V4" box="[204,276,652,677]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="wardii">
|
||
<emphasis id="B95A2A364C6DC64C1EEC2FA1FEC723D1" box="[204,276,652,677]" italics="true" pageId="27" pageNumber="28">wardii</emphasis>
|
||
</taxonomicName>
|
||
♀” label in the Oberthür collection were not referable to this species (
|
||
<bibRefCitation id="EFBF8BD54C6DC64C1A072FA1FB3323D0" author="Lees" box="[1063,1248,652,676]" pageId="27" pageNumber="28" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997: 391</bibRefCitation>
|
||
);
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6DC64C1EE72F9DFAC023BC" blockId="27.[151,1437,255,2020]" box="[199,1299,687,712]" pageId="27" pageNumber="28">
|
||
sp. 12 (
|
||
<bibRefCitation id="EFBF8BD54C6DC64C1F382F9DFE6D23BC" author="Kremen" box="[280,446,688,712]" pageId="27" pageNumber="28" refString="Kremen, C. (1994) Biological inventory using target taxa: a case study of the butterflies of Madagascar. Ecological Applications, 4, 407 - 422. http: // dx. doi. org / 10.2307 / 1941946" type="journal article" year="1994">Kremen, 1994</bibRefCitation>
|
||
: 417, as “
|
||
<taxonomicName id="4C2E8DA74C6DC64C1C0C2F9CFD4E23BC" ID-CoL="92BDD" box="[556,669,689,712]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A2A364C6DC64C1C0C2F9CFD4E23BC" box="[556,669,689,712]" italics="true" pageId="27" pageNumber="28">Henotesia</emphasis>
|
||
</taxonomicName>
|
||
?
|
||
<taxonomicName id="4C2E8DA74C6DC64C1C912F82FCD823BC" ID-CoL="6LQ8P" box="[689,779,687,712]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="obscura">
|
||
<emphasis id="B95A2A364C6DC64C1C912F82FCD823BC" box="[689,779,687,712]" italics="true" pageId="27" pageNumber="28">obscura</emphasis>
|
||
</taxonomicName>
|
||
”;
|
||
<bibRefCitation id="EFBF8BD54C6DC64C1D062F9DFC4D23BC" author="Lees" box="[806,926,688,712]" pageId="27" pageNumber="28" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997</bibRefCitation>
|
||
: 61, 64;
|
||
<bibRefCitation id="EFBF8BD54C6DC64C1DDE2F9DFB1D23BC" author="Torres" box="[1022,1230,687,712]" pageId="27" pageNumber="28" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. & Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
|
||
Torres
|
||
<emphasis id="B95A2A364C6DC64C1A6C2F9CFB5823BC" box="[1100,1163,687,712]" italics="true" pageId="27" pageNumber="28">et al.,</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
: 462);
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C6DC64C1EE72FF9FAED2398" box="[199,1342,724,749]" pageId="27" pageNumber="28" type="description">
|
||
<paragraph id="8B91F6244C6DC64C1EE72FF9FAED2398" blockId="27.[151,1437,255,2020]" box="[199,1342,724,749]" pageId="27" pageNumber="28">
|
||
<treatmentCitationGroup id="AB3ED10A4C6DC64C1EE72FF9FAED2398" box="[199,1342,724,749]" pageId="27" pageNumber="28">
|
||
“
|
||
<taxonomicName id="4C2E8DA74C6DC64C1EF32FF8FDBD2398" authority="Lees" authorityName="Lees" box="[211,622,724,749]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi" subGenus="Henotesia">
|
||
<emphasis id="B95A2A364C6DC64C1EF32FF8FE982398" box="[211,331,725,748]" italics="true" pageId="27" pageNumber="28">Heteropsis</emphasis>
|
||
(
|
||
<emphasis id="B95A2A364C6DC64C1F7C2FF8FE1D2398" box="[348,462,725,748]" italics="true" pageId="27" pageNumber="28">Henotesia</emphasis>
|
||
)
|
||
<emphasis id="B95A2A364C6DC64C1FFE2FF9FDE12399" box="[478,562,724,749]" italics="true" pageId="27" pageNumber="28">harveyi</emphasis>
|
||
Lees
|
||
</taxonomicName>
|
||
(MS), nom.nov.” (ms name in Appendix A of
|
||
<treatmentCitation id="0A8FD0354C6DC64C1A592FF8FAE22398" author="Lees" box="[1145,1329,724,748]" page="391" pageId="27" pageNumber="28" year="1997">
|
||
<bibRefCitation id="EFBF8BD54C6DC64C1A592FF8FB2B2398" author="Lees" box="[1145,1272,724,748]" pageId="27" pageNumber="28" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||
: 391
|
||
</treatmentCitation>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C6DC64C1EE72FDAFBC52546" pageId="27" pageNumber="28" type="materials_examined">
|
||
<paragraph id="8B91F6244C6DC64C1EE72FDAFDE5222F" blockId="27.[151,1437,255,2020]" pageId="27" pageNumber="28">
|
||
<emphasis id="B95A2A364C6DC64C1EE72FDAFCD92264" bold="true" box="[199,778,759,784]" pageId="27" pageNumber="28">
|
||
<typeStatus id="549548864C6DC64C1EE72FDAFED12264" box="[199,258,759,784]" pageId="27" pageNumber="28">Type</typeStatus>
|
||
material., Deposition
|
||
<collectionCode id="ED3F6EE14C6DC64C1C0B2FDAFD552264" box="[555,646,759,784]" pageId="27" pageNumber="28">BMNH</collectionCode>
|
||
:
|
||
<typeStatus id="549548864C6DC64C1CBD2FDAFCD92264" box="[669,778,759,784]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
: ♂ (
|
||
<figureCitation id="1315EAA14C6DC64C1D732FDAFC722264" box="[851,929,759,784]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="27" pageNumber="28">Fig. 7</figureCitation>
|
||
A),
|
||
<collectingCountry id="F339B6B44C6DC64C1DF72FDAFB8D2264" box="[983,1118,759,784]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
|
||
E, Sahavondronina VOI, Ranomafana National Park,
|
||
<geoCoordinate id="EE1A90E34C6DC64C1FFA2E33FDEC2243" box="[474,575,795,823]" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="55" value="-21.276">21.276o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C6DC64C1C6E2E33FD672243" box="[590,692,795,823]" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="55" value="47.325">47.325o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C6DC64C1CC72E33FC942243" box="[743,839,798,823]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="27" pageNumber="28" unit="km" value="0.25">0.25 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C6DC64C1D742E32FC2F2243" box="[852,1020,798,823]" metricMagnitude="3" metricUnit="m" metricValue="1.313" metricValueMax="1.338" metricValueMin="1.288" pageId="27" pageNumber="28" unit="m" value="1313.0" valueMax="1338.0" valueMin="1288.0">1313 +/- 25 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C6DC64C1A2A2E32FBAE2243" box="[1034,1149,799,823]" pageId="27" pageNumber="28" value="2014-11-08">8/11/2014</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C6DC64C1B2E2E32FAE52243" box="[1294,1334,799,823]" pageId="27" pageNumber="28">DL</collectionCode>
|
||
14R-229,
|
||
<collectionCode id="ED3F6EE14C6DC64C1EB72E6EFED3222F" box="[151,256,835,859]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="27" pageNumber="28">NHMUK</collectionCode>
|
||
010289119 [
|
||
<collectingCountry id="F339B6B44C6DC64C1FAB2E6EFE15222F" box="[395,454,835,859]" name="Qatar" pageId="27" pageNumber="28">QTR</collectingCountry>
|
||
barcode].
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6DC64C1EE72E4BFBC52546" blockId="27.[151,1437,255,2020]" pageId="27" pageNumber="28">
|
||
<typeStatus id="549548864C6DC64C1EE72E4BFEEC220B" box="[199,319,870,895]" pageId="27" pageNumber="28" type="paratype">
|
||
<emphasis id="B95A2A364C6DC64C1EE72E4BFEEC220B" bold="true" box="[199,319,870,895]" pageId="27" pageNumber="28">Paratypes</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<emphasis id="B95A2A364C6DC64C1F762E4BFD95220B" bold="true" box="[342,582,870,895]" pageId="27" pageNumber="28">
|
||
Deposition
|
||
<collectionCode id="ED3F6EE14C6DC64C1FC52E4BFD93220B" box="[485,576,870,895]" pageId="27" pageNumber="28">BMNH</collectionCode>
|
||
:
|
||
</emphasis>
|
||
♂, same data as HT but
|
||
<date id="FF90D0E44C6DC64C1DB92E4AFBDE220B" box="[921,1037,871,895]" pageId="27" pageNumber="28" value="2014-11-08">8/11/2014</date>
|
||
: 10:40, A. Allaoui:
|
||
<collectionCode id="ED3F6EE14C6DC64C1B2E2E4AFAE5220B" box="[1294,1334,871,895]" pageId="27" pageNumber="28">DL</collectionCode>
|
||
14R-204,
|
||
<collectionCode id="ED3F6EE14C6DC64C1EB72EA6FED222D7" box="[151,257,907,931]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="27" pageNumber="28">NHMUK</collectionCode>
|
||
010289120 [
|
||
<collectingCountry id="F339B6B44C6DC64C1FAC2EA6FE1422D7" box="[396,455,907,931]" name="Qatar" pageId="27" pageNumber="28">QTR</collectingCountry>
|
||
barcode]; ♀ (
|
||
<figureCitation id="1315EAA14C6DC64C1C452EA7FD7422D7" box="[613,679,906,931]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="27" pageNumber="28">Fig. 7</figureCitation>
|
||
B),
|
||
<date id="FF90D0E44C6DC64C1CF12EA6FC9522D7" box="[721,838,907,931]" pageId="27" pageNumber="28" value="2014-11-08">8/11/2014</date>
|
||
: 10:57, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C6DC64C1A3F2EA6FB9422D7" box="[1055,1095,907,931]" pageId="27" pageNumber="28">DL</collectionCode>
|
||
14R-207,
|
||
<collectionCode id="ED3F6EE14C6DC64C1A942EA6FACC22D7" box="[1204,1311,907,931]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="27" pageNumber="28">NHMUK</collectionCode>
|
||
010289121 [
|
||
<collectingCountry id="F339B6B44C6DC64C1EBD2E82FF0B22B3" box="[157,216,943,967]" name="Qatar" pageId="27" pageNumber="28">QTR</collectingCountry>
|
||
barcode]; ♂, same data as HT but
|
||
<date id="FF90D0E44C6DC64C1C5F2E83FD2622B3" box="[639,757,942,967]" pageId="27" pageNumber="28" value="2014-11-06">6/11/2014</date>
|
||
: 13:03, fruit trap, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C6DC64C1A762E83FBAD22B3" box="[1110,1150,942,967]" pageId="27" pageNumber="28">DL</collectionCode>
|
||
14R-126, IA599 [isotope analysis],
|
||
<collectionCode id="ED3F6EE14C6DC64C1F2E2EFEFEAB229F" box="[270,376,979,1003]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="27" pageNumber="28">NHMUK</collectionCode>
|
||
010289122 [
|
||
<collectingCountry id="F339B6B44C6DC64C1C292EFEFD97229F" box="[521,580,979,1003]" name="Qatar" pageId="27" pageNumber="28">QTR</collectingCountry>
|
||
barcode]; ♂, same data as HT but
|
||
<date id="FF90D0E44C6DC64C1DD92EFEFBBF229F" box="[1017,1132,979,1003]" pageId="27" pageNumber="28" value="2014-11-08">8/11/2014</date>
|
||
, A. Allaoui:
|
||
<collectionCode id="ED3F6EE14C6DC64C1B2E2EFEFAE5229F" box="[1294,1334,979,1003]" pageId="27" pageNumber="28">DL</collectionCode>
|
||
14R-104,
|
||
<collectionCode id="ED3F6EE14C6DC64C1EB72EDAFED2257B" box="[151,257,1015,1039]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="27" pageNumber="28">NHMUK</collectionCode>
|
||
010289123 [
|
||
<collectingCountry id="F339B6B44C6DC64C1FAD2EDAFE1B257B" box="[397,456,1015,1039]" name="Qatar" pageId="27" pageNumber="28">QTR</collectingCountry>
|
||
barcode]; ♀, same data as HT but
|
||
<date id="FF90D0E44C6DC64C1D7A2EDAFC1E257B" box="[858,973,1015,1039]" pageId="27" pageNumber="28" value="2014-11-08">8/11/2014</date>
|
||
, A. Allaoui:
|
||
<collectionCode id="ED3F6EE14C6DC64C1A422EDBFB59257B" box="[1122,1162,1014,1039]" pageId="27" pageNumber="28">DL</collectionCode>
|
||
14R-206,
|
||
<collectionCode id="ED3F6EE14C6DC64C1AD92EDBFA82257B" box="[1273,1361,1014,1039]" pageId="27" pageNumber="28">BMAD</collectionCode>
|
||
217-15 [
|
||
<collectionCode id="ED3F6EE14C6DC64C1EBD2936FF0E2547" box="[157,221,1051,1075]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="27" pageNumber="28">DNA</collectionCode>
|
||
barcode], IA597 [isotope voucher],
|
||
<collectionCode id="ED3F6EE14C6DC64C1C552936FD0C2547" box="[629,735,1051,1075]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="27" pageNumber="28">NHMUK</collectionCode>
|
||
010289180 [
|
||
<collectingCountry id="F339B6B44C6DC64C1D4A2936FC762547" box="[874,933,1051,1075]" name="Qatar" pageId="27" pageNumber="28">QTR</collectingCountry>
|
||
barcode];
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C6DC64C1EE72913FDD02422" pageId="27" pageNumber="28" type="description">
|
||
<paragraph id="8B91F6244C6DC64C1EE72913FAEF25D6" blockId="27.[151,1437,255,2020]" pageId="27" pageNumber="28">
|
||
<emphasis id="B95A2A364C6DC64C1EE72913FE7C2523" bold="true" box="[199,431,1086,1111]" pageId="27" pageNumber="28">Deposition MNHN:</emphasis>
|
||
♂, same data as HT but
|
||
<date id="FF90D0E44C6DC64C1CE12913FCE72523" box="[705,820,1086,1111]" pageId="27" pageNumber="28" value="2014-11-06">6/11/2014</date>
|
||
, A. Allaoui: DL14R-116; ♀, same data as HT but
|
||
<date id="FF90D0E44C6DC64C1B432912FF01250F" pageId="27" pageNumber="28" value="2014-11-07">7/11/ 2014</date>
|
||
, D.C. Lees: DL14R-156, E37; ♂,
|
||
<date id="FF90D0E44C6DC64C1C58294FFD07250F" box="[632,724,1122,1147]" pageId="27" pageNumber="28" value="1969-05-25">25-V-69</date>
|
||
FIANAR: SENTIER FORESTIER A 3 KM AU S.O.
|
||
<collectingCountry id="F339B6B44C6DC64C1B50294EFA4B250E" box="[1392,1432,1123,1146]" name="South Georgia and the South Sandwich Islands" pageId="27" pageNumber="28">DE</collectingCountry>
|
||
, VOHIPARARA, [
|
||
<geoCoordinate id="EE1A90E34C6DC64C1F4029A7FE6825D6" box="[352,443,1158,1186]" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="555" value="-21.24">21.24o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C6DC64C1FE729A7FDCC25D5" box="[455,543,1158,1186]" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="555" value="47.43">47.43o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C6DC64C1C6829A4FDA825D6" box="[584,635,1161,1186]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="28" unit="km" value="1.0">1km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C6DC64C1CA629A7FCF925D5" box="[646,810,1161,1186]" metricMagnitude="3" metricUnit="m" metricValue="1.0290000000000001" metricValueMax="1.0790000000000002" metricValueMin="0.9790000000000001" pageId="27" pageNumber="28" unit="m" value="1029.0" valueMax="1079.0" valueMin="979.0">1029 +/- 50 m</quantity>
|
||
], J. Minet; ♀, same data as above but:
|
||
<date id="FF90D0E44C6DC64C1AFC29A4FAEB25D6" box="[1244,1336,1161,1186]" pageId="27" pageNumber="28" value="1969-03-09">9-III-69</date>
|
||
;
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6DC64C1EE72980FDEC259E" blockId="27.[151,1437,255,2020]" pageId="27" pageNumber="28">
|
||
<emphasis id="B95A2A364C6DC64C1EE72980FE4D25B2" bold="true" box="[199,414,1197,1222]" pageId="27" pageNumber="28">Deposition ABRI:</emphasis>
|
||
♂, same data as HT but
|
||
<date id="FF90D0E44C6DC64C1C9A2983FCFE25B2" box="[698,813,1198,1222]" pageId="27" pageNumber="28" value="2014-11-08">8/11/2014</date>
|
||
, D.C. Lees: DL14R-245; ♀, same data at HT but
|
||
<date id="FF90D0E44C6DC64C1B432983FF07259E" pageId="27" pageNumber="28" value="2014-11-08">8/11/ 2014</date>
|
||
: 10:48, A. Allaoui: DL14R-191;
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6DC64C1EE729D8FB40247A" blockId="27.[151,1437,255,2020]" box="[199,1171,1269,1294]" pageId="27" pageNumber="28">
|
||
<emphasis id="B95A2A364C6DC64C1EE729D8FE4D247A" bold="true" box="[199,414,1269,1294]" pageId="27" pageNumber="28">Deposition PBZT:</emphasis>
|
||
♂, same data as HT but
|
||
<date id="FF90D0E44C6DC64C1C9329DBFCFB247A" box="[691,808,1270,1294]" pageId="27" pageNumber="28" value="2014-11-08">8/11/2014</date>
|
||
: 11:27, A. Allaoui: DL14R-213.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6DC64C1EE72834FDD02422" blockId="27.[151,1437,255,2020]" pageId="27" pageNumber="28">
|
||
<emphasis id="B95A2A364C6DC64C1EE72834FE6C2446" bold="true" box="[199,447,1305,1330]" pageId="27" pageNumber="28">Deposition summary</emphasis>
|
||
: BMNH (HT ♂, 3 PT ♂♂ and 2 PT ♀♀), MNHN (2 PT ♂♂ and 2 PT ♀♀); ABRI (1 PT ♂ and 1 PT ♀); PBZT (1 PT ♂).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C6DC64C1EE72849FEBA24D5" pageId="27" pageNumber="28" type="materials_examined">
|
||
<paragraph id="8B91F6244C6DC64C1EE72849FEBA24D5" blockId="27.[151,1437,255,2020]" pageId="27" pageNumber="28">
|
||
<emphasis id="B95A2A364C6DC64C1EE72849FEB42409" bold="true" box="[199,359,1380,1405]" pageId="27" pageNumber="28">
|
||
<typeStatus id="549548864C6DC64C1EE72849FED12409" box="[199,258,1380,1405]" pageId="27" pageNumber="28">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F339B6B44C6DC64C1F4F2849FE252409" box="[367,502,1380,1405]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
|
||
E, Sahavondronina VOI, Ranomafana National Park,
|
||
<geoCoordinate id="EE1A90E34C6DC64C1A402849FB162409" box="[1120,1221,1377,1405]" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="55" value="-21.276">21.276o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C6DC64C1AF42849FAEA2408" box="[1236,1337,1377,1405]" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="55" value="47.325">47.325o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C6DC64C1B4B2849FF6D24D5" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="27" pageNumber="28" unit="km" value="0.25">0.25 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C6DC64C1EE928A4FEB724D5" box="[201,356,1416,1441]" metricMagnitude="3" metricUnit="m" metricValue="1.313" metricValueMax="1.338" metricValueMin="1.288" pageId="27" pageNumber="28" unit="m" value="1313.0" valueMax="1338.0" valueMin="1288.0">
|
||
1313 +/-
|
||
<specimenCount id="9D283DAD4C6DC64C1F0F28A5FEB724D5" box="[303,356,1416,1441]" pageId="27" pageNumber="28" type="generic">25m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C6DC64C1EE72881FE4C260C" pageId="27" pageNumber="28" type="diagnosis">
|
||
<paragraph id="8B91F6244C6DC64C1EE72881FE4C260C" blockId="27.[151,1437,255,2020]" pageId="27" pageNumber="28">
|
||
<emphasis id="B95A2A364C6DC64C1EE72881FE9224B1" bold="true" box="[199,321,1452,1477]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C2E8DA74C6DC64C1F6D2880FDD224B1" box="[333,513,1452,1477]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||
<emphasis id="B95A2A364C6DC64C1F6D2880FDD224B1" box="[333,513,1452,1477]" italics="true" pageId="27" pageNumber="28">Ht. vanewrighti</emphasis>
|
||
</taxonomicName>
|
||
and an undescribed species (‘
|
||
<emphasis id="B95A2A364C6DC64C1D4A2880FC5524B0" box="[874,902,1453,1476]" italics="true" pageId="27" pageNumber="28">Ht</emphasis>
|
||
. sp. 20’) are both generally smaller than
|
||
<taxonomicName id="4C2E8DA74C6DC64C1B592880FF38249D" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6DC64C1B592880FF38249D" italics="true" pageId="27" pageNumber="28">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
and the former occurs only in NW
|
||
<collectingCountry id="F339B6B44C6DC64C1C5728FDFCD3249D" box="[631,768,1488,1513]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
|
||
, whereas
|
||
<taxonomicName id="4C2E8DA74C6DC64C1D4E28FCFC3F249D" box="[878,1004,1488,1513]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6DC64C1D4E28FCFC3F249D" box="[878,1004,1488,1513]" italics="true" pageId="27" pageNumber="28">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
is generally similar to
|
||
<taxonomicName id="4C2E8DA74C6DC64C1ACC28FCFA4F249D" box="[1260,1436,1488,1513]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||
<emphasis id="B95A2A364C6DC64C1ACC28FCFA4F249D" box="[1260,1436,1488,1513]" italics="true" pageId="27" pageNumber="28">Ht. vanewrighti</emphasis>
|
||
</taxonomicName>
|
||
in wing colouration, but in all known specimens shows a prominent androconial patch on the FWV. As currently recognized,
|
||
<taxonomicName id="4C2E8DA74C6DC64C1F3F2B31FE4E2740" box="[287,413,1563,1588]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6DC64C1F3F2B31FE4E2740" box="[287,413,1563,1588]" italics="true" pageId="27" pageNumber="28">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
occurs in the centre-east of the montane rainforest biome at latitudes south of about
|
||
<geoCoordinate id="EE1A90E34C6DC64C1B6C2B36FA452747" box="[1356,1430,1559,1588]" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="5555" value="-17.5">17.5o S</geoCoordinate>
|
||
, whereas
|
||
<taxonomicName id="4C2E8DA74C6DC64C1EDA2B6DFE78272C" box="[250,427,1599,1624]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||
<emphasis id="B95A2A364C6DC64C1EDA2B6DFE78272C" box="[250,427,1599,1624]" italics="true" pageId="27" pageNumber="28">Ht. vanewrighti</emphasis>
|
||
</taxonomicName>
|
||
is only known from the extreme northwestern part of the rainforest biome, but
|
||
<emphasis id="B95A2A364C6DC64C1B0D2B6DFA9A2723" box="[1325,1353,1600,1623]" italics="true" pageId="27" pageNumber="28">Ht</emphasis>
|
||
. sp. 20 appears to occur partially sympatrically, distributed from the NE to CE of
|
||
<collectingCountry id="F339B6B44C6DC64C1A212B4EFB582708" box="[1025,1163,1635,1660]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C2E8DA74C6DC64C1AB82B49FAE82708" box="[1176,1339,1635,1660]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="westwoodi">
|
||
<emphasis id="B95A2A364C6DC64C1AB82B49FAE82708" box="[1176,1339,1635,1660]" italics="true" pageId="27" pageNumber="28">Ht. westwoodi</emphasis>
|
||
</taxonomicName>
|
||
is quite distinctive from either of these species, having a more uniform colouration on the FWV towards the costa than any of these species, while
|
||
<taxonomicName id="4C2E8DA74C6DC64C1F8F2B81FDF127B7" box="[431,546,1708,1731]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="viettei">
|
||
<emphasis id="B95A2A364C6DC64C1F8F2B81FDF127B7" box="[431,546,1708,1731]" italics="true" pageId="27" pageNumber="28">Ht. viettei</emphasis>
|
||
</taxonomicName>
|
||
also has a prominent pale or whitish highlighting distal of the HWV Mb. Counting from the start of the 658 bp COI-5P barcode, the following four putatively fixed nucleotide differences in the DNA barcode appear (n=4) to be diagnostic within the
|
||
<taxonomicName id="4C2E8DA74C6DC64C1D042BD9FC712678" box="[804,930,1779,1804]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6DC64C1D042BD9FC712678" box="[804,930,1779,1804]" italics="true" pageId="27" pageNumber="28">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
complex (cluster number BOLD:ACW5694) throughout its range, where in parentheses is specified the un-derived state: 274 C(T); 298 T(C); 466 A(G); 498 C(A). DNA barcoding is currently the best means for determining material of
|
||
<taxonomicName id="4C2E8DA74C6DC64C1A0A2A11FB782620" box="[1066,1195,1851,1876]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6DC64C1A0A2A11FB782620" box="[1066,1195,1851,1876]" italics="true" pageId="27" pageNumber="28">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
among material that exhibit the FWV patch.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C6DC64B1EE72AAEFCE625B1" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
|
||
<paragraph id="8B91F6244C6DC64B1EE72AAEFE5122BC" blockId="27.[151,1437,255,2020]" lastBlockId="28.[151,1437,151,2013]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
|
||
<emphasis id="B95A2A364C6DC64C1EE72AAEFE7926E8" bold="true" box="[199,426,1923,1948]" pageId="27" pageNumber="28">Description. Wings</emphasis>
|
||
: dorsal surface uniform very dark chocolate brown (fresh specimen). FW and HW margin strongly crenate. HW has dark brown Sml following the margin, highlighted distad with slightly paler dark brown (also on ventral surface). FWD space-CuA1 ocellus expressed conspicuously, spanning intervein distance CuA1- CuA2 including the narrow concentric deep reddish orange ring, and HWD space-cuA1 ocellus to a lesser extent, spanning about 2/3 of intervein distance CuA1-CuA2, elliptic, also with a narrow deep orange ring. FWV ocellus space-CuA1 is expressed to a similar extent to the FWD, but more conspicuous with a brighter orange ring, while FWV ocelli space-M2 ocellus is about half the diameter (though barely visible on FWD), the orange ring of that of space-M2 slightly compromising space-M1 and -M3. HWV ocelli of space-Rs-CuA2 are expressed as white points lacking strongly discernible rings, except in the case of space-Rs and space-CuA2 which have small black irises and faint orange rings, and that of CuA1 already referred to. HWV space-CuA1 ocellus is elliptic, same size as on HWD, with a quite narrow bright orange ring. The dark Mb of the HWV is strongly and concavely indented proximad of space-CuA1 ocellus and is quite irregular in shape, sharply convex in space-M2 and space-CuA2 and concave-inflexed below vein CuA2 terminating at 1A; above M2 it is again concave to beyond mid-costa but not so distinct from background. A general irroration of darker brown pervades the HWV background and a generally darker band is formed between the Mb and PMb of the central symmetry system, the PMb not so irregular and somewhat convex. This band is highlighted distad and proximad and towards the anal angle by yellowish background colour. (This marbled ventral colour pattern actually befits Aurivillius’ [unavailable] name for his aberration (
|
||
<emphasis id="B95A2A364C6AC64B1F392FBCFE4423DC" box="[281,407,657,680]" italics="true" pageId="28" pageNumber="29">marmorata</emphasis>
|
||
) of ‘
|
||
<taxonomicName id="4C2E8DA74C6AC64B1FF02FBCFD0E23DC" box="[464,733,655,680]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="andravahana">
|
||
<emphasis id="B95A2A364C6AC64B1FF02FBCFD0E23DC" box="[464,733,655,680]" italics="true" pageId="28" pageNumber="29">Henotesia andravahana</emphasis>
|
||
</taxonomicName>
|
||
’). Distad of the central HWV band and highlighting it there is a more obscure dark violet to grey-brown region, and the FWV pattern is generally more dark-obscure and less marbled but with about four ochreous strigulae (Costal bars, henceforth ‘Cbs’) perpendicular to the costa beyond mid-costa.
|
||
<emphasis id="B95A2A364C6AC64B1F352FD1FE592261" bold="true" box="[277,394,764,789]" pageId="28" pageNumber="29">Variation.</emphasis>
|
||
♂♂ similar, but vary in the intensity of the darker brown irroration and Mb on the HWV. Sexually dimorphic, but marginally so. The ♂ and ♀ are one of the most similar of the
|
||
<taxonomicName id="4C2E8DA74C6AC64B1A5C2E0CFAD0224C" box="[1148,1283,799,824]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="drepana">
|
||
<emphasis id="B95A2A364C6AC64B1A5C2E0CFAD0224C" box="[1148,1283,799,824]" italics="true" pageId="28" pageNumber="29">Ht. drepana</emphasis>
|
||
</taxonomicName>
|
||
group and in fact the only pair in the
|
||
<taxonomicName id="4C2E8DA74C6AC64B1F862E68FDFD2229" box="[422,558,836,861]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="drepana">
|
||
<emphasis id="B95A2A364C6AC64B1F862E68FDFD2229" box="[422,558,836,861]" italics="true" pageId="28" pageNumber="29">Ht. drepana</emphasis>
|
||
</taxonomicName>
|
||
group to have been correctly matched by
|
||
<bibRefCitation id="EFBF8BD54C6AC64B1A292E69FB102229" author="Oberthur" box="[1033,1219,836,861]" pageId="28" pageNumber="29" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür (1916)</bibRefCitation>
|
||
, but the ♀ is larger and paler. The ♂ and ♀ have been photographed in copula (
|
||
<figureCitation id="1315EAA14C6AC64B1D0D2E4AFCA322F4" box="[813,880,871,896]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="28" pageNumber="29">Fig. 7</figureCitation>
|
||
D). The ♀ is similar to the ♂, but rather larger and lighter brown on either surface, with a similar ocellus configuration, although the space-CuA1 ocelli tend to be larger in either wing.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6AC64B1EE72EF9FE3A252C" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
|
||
<emphasis id="B95A2A364C6AC64B1EE72EF9FE872299" bold="true" box="[199,340,980,1005]" pageId="28" pageNumber="29">Androconia</emphasis>
|
||
: HWD discocellular brush is bright yellowish fawn (in the HT) to russet brown at tip where it locates on to an androconial patch on the ventral FW towards the base of space-CuA2 just above vein 1A. HWD discocellular patch is dingy orange to brick red, very large and broad lenticular, bisected by R vein near the top of the cell (not observed on HT).
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6AC64B1EE72949FCE625B1" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
|
||
<emphasis id="B95A2A364C6AC64B1EE72949FEBF2509" bold="true" box="[199,364,1124,1149]" pageId="28" pageNumber="29">Wingspan/fwl</emphasis>
|
||
: range 33–37.1/
|
||
<quantity id="4CD65BC14C6AC64B1C0A2949FD6B2508" box="[554,696,1124,1148]" metricMagnitude="-2" metricUnit="m" metricValue="1.8649999999999998" metricValueMax="1.93" metricValueMin="1.8" pageId="28" pageNumber="29" unit="mm" value="18.65" valueMax="19.3" valueMin="18.0">18–19.3 mm</quantity>
|
||
(n=6 ♂♂), including HT ♂ = 36.3/
|
||
<quantity id="4CD65BC14C6AC64B1A762949FB6D2508" box="[1110,1214,1124,1149]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="28" pageNumber="29" unit="mm" value="19.3">19.3 mm</quantity>
|
||
; mean = 35 +/- 1.7 SD/18.7 +/- 0.5 SD mm (n=6 ♂♂). Range
|
||
<quantity id="4CD65BC14C6AC64B1C5929A5FCCF25EB" box="[633,796,1160,1184]" metricMagnitude="-2" metricUnit="m" metricValue="3.6599999999999993" metricValueMax="3.84" metricValueMin="3.4799999999999995" pageId="28" pageNumber="29" unit="mm" value="36.599999999999994" valueMax="38.4" valueMin="34.8">34.8–38.4 mm</quantity>
|
||
(n=3 ♀♀)/18.7–21.6 (n=5 ♀♀); mean= 36.6 +/- 1.8 SD/
|
||
<quantity id="4CD65BC14C6AC64B1EB72981FEF025B0" box="[151,291,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="28" pageNumber="29" unit="mm" value="20.0" valueMax="21.0" valueMin="19.0">20 +/- 1 mm</quantity>
|
||
SD (n=3 ♀♀); all examples from
|
||
<typeStatus id="549548864C6AC64B1C842980FD0725B1" box="[676,724,1197,1221]" pageId="28" pageNumber="29">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C6AC64B1EE729E2FED92479" pageId="28" pageNumber="29" type="etymology">
|
||
<paragraph id="8B91F6244C6AC64B1EE729E2FED92479" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
|
||
<emphasis id="B95A2A364C6AC64B1EE729E2FE9D259C" bold="true" box="[199,334,1231,1256]" pageId="28" pageNumber="29">Etymology.</emphasis>
|
||
After Don Harvey, who first pointed out (to Claire Kremen) the FWV androconial patch (
|
||
<bibRefCitation id="EFBF8BD54C6AC64B1B4629FDFF2E2479" author="Lees" pageId="28" pageNumber="29" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997: 64</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C6AC64A1EE7283AFC992261" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="discussion">
|
||
<paragraph id="8B91F6244C6AC64B1EE7283AFE3E26E1" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
|
||
<emphasis id="B95A2A364C6AC64B1EE7283AFE9F2444" bold="true" box="[199,332,1303,1328]" pageId="28" pageNumber="29">Discussion.</emphasis>
|
||
Apart from
|
||
<taxonomicName id="4C2E8DA74C6AC64B1FFF2834FD812444" box="[479,594,1305,1328]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="tianae" status="sp. nov.">
|
||
<emphasis id="B95A2A364C6AC64B1FFF2834FD812444" box="[479,594,1305,1328]" italics="true" pageId="28" pageNumber="29">Ht. tianae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A269974D4C6AC64B1C7C283AFD6B2444" box="[604,696,1303,1328]" pageId="28" pageNumber="29" rank="species">
|
||
<emphasis id="B95A2A364C6AC64B1C7C283AFD6B2444" bold="true" box="[604,696,1303,1328]" pageId="28" pageNumber="29">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C2E8DA74C6AC64B1CE92834FC992444" box="[713,842,1303,1328]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6AC64B1CE92834FC992444" box="[713,842,1303,1328]" italics="true" pageId="28" pageNumber="29">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
is the
|
||
<taxonomicName id="4C2E8DA74C6AC64B1DBC2834FBC72444" box="[924,1044,1305,1328]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A2A364C6AC64B1DBC2834FBC72444" box="[924,1044,1305,1328]" italics="true" pageId="28" pageNumber="29">Heteropsis</emphasis>
|
||
</taxonomicName>
|
||
species described here which has longest been known in historical collections (in BMNH, MNHN, and SNHM, at least). The first modern field specimens of this species were recognized by C. Kremen in
|
||
<quantity id="4CD65BC14C6AC64B1D1E284DFC06240C" box="[830,981,1375,1400]" metricMagnitude="1" metricUnit="m" metricValue="5.05333" metricValueMax="5.05714" metricValueMin="5.04952" pageId="28" pageNumber="29" unit="in" value="1989.5" valueMax="1991.0" valueMin="1988.0">
|
||
<date id="FF90D0E44C6AC64B1D1E284DFC64240C" box="[830,951,1376,1400]" pageId="28" pageNumber="29" value="1988" valueMax="1991">1988-1991</date>
|
||
in
|
||
</quantity>
|
||
what is now Ranomafana National Park (
|
||
<bibRefCitation id="EFBF8BD54C6AC64B1EBF28A9FE9924E9" author="Lees" box="[159,330,1412,1437]" pageId="28" pageNumber="29" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997: 64</bibRefCitation>
|
||
). Until now, recognition of this species has been taxonomically very confusing (as detailed above). Compared with closely related species, it can readily be distinguished androconially and allopatrically from
|
||
<taxonomicName id="4C2E8DA74C6AC64B1B592884FECE2491" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||
<emphasis id="B95A2A364C6AC64B1B592884FECE2491" italics="true" pageId="28" pageNumber="29">Ht. vanewrighti</emphasis>
|
||
</taxonomicName>
|
||
. However, as mentioned above, morphological and genetic variation across the potential full range of
|
||
<taxonomicName id="4C2E8DA74C6AC64B1EB728DCFECD277C" box="[151,286,1519,1544]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6AC64B1EB728DCFECD277C" box="[151,286,1519,1544]" italics="true" pageId="28" pageNumber="29">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
needs further examination and this is amenable to a phylogeographic analysis, to see if its circumscription might eventually comprise one or more potentially allopatric or sympatric morphospecies that also have a ventral androconial patch, ‘sp. 20’ (same as ‘sp. 77’?) known from the northeastern montane rainforests of the island (see also
|
||
<bibRefCitation id="EFBF8BD54C6AC64B1FAB2B70FDC32700" author="Lees" box="[395,528,1628,1652]" pageId="28" pageNumber="29" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||
). As mentioned above, the HT ♂ (NHRS-TOBI000000050;
|
||
<figureCitation id="1315EAA14C6AC64B1AC42B71FAFF2700" box="[1252,1324,1628,1653]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="28" pageNumber="29">Fig. 7</figureCitation>
|
||
C) of
|
||
<taxonomicName id="4C2E8DA74C6AC64B1B592B70FEFD27EC" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="andrahahana">
|
||
<emphasis id="B95A2A364C6AC64B1B592B70FEFD27EC" italics="true" pageId="28" pageNumber="29">Ht. andrahahana</emphasis>
|
||
</taxonomicName>
|
||
ab.
|
||
<emphasis id="B95A2A364C6AC64B1F7C2BACFE0927EC" box="[348,474,1665,1688]" italics="true" pageId="28" pageNumber="29">marmorata</emphasis>
|
||
Aurivillius was examined in SNHM and is considered to be conspecific, but the name is not nomenclaturally available. Also, the HT ♂ of
|
||
<taxonomicName id="4C2E8DA74C6AC64B1CF82B88FBFA27C8" authority="Butler, 1879" authorityName="Butler" authorityYear="1879" box="[728,1065,1700,1725]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="wardii">
|
||
<emphasis id="B95A2A364C6AC64B1CF82B88FC4B27C9" box="[728,920,1700,1725]" italics="true" pageId="28" pageNumber="29">Henotesia wardii</emphasis>
|
||
<bibRefCitation id="EFBF8BD54C6AC64B1DBF2B89FBFA27C8" author="Butler" box="[927,1065,1700,1725]" pageId="28" pageNumber="29" refString="Butler, A. G. (1879) Descriptions of new species of Lepidoptera from Madagascar, with notes on some of the forms already described. Annals and Magazine of Natural History, Series 5, 4, 227 - 246. http: // dx. doi. org / 10.1080 / 00222937908679820" type="journal article" year="1879">Butler, 1879</bibRefCitation>
|
||
</taxonomicName>
|
||
(BMNH(E) 697802); now called
|
||
<taxonomicName id="4C2E8DA74C6AC64B1EB72BE4FE352794" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" box="[151,486,1736,1760]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="viettei">
|
||
<emphasis id="B95A2A364C6AC64B1EB72BE4FE8E2794" box="[151,349,1737,1760]" italics="true" pageId="28" pageNumber="29">Heteropsis viettei</emphasis>
|
||
Lees, 2003
|
||
</taxonomicName>
|
||
, a species potentially confused with
|
||
<taxonomicName id="4C2E8DA74C6AC64B1DAE2BE4FBDC2794" box="[910,1039,1735,1760]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6AC64B1DAE2BE4FBDC2794" box="[910,1039,1735,1760]" italics="true" pageId="28" pageNumber="29">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
in Mabille’s (1878) description of
|
||
<taxonomicName id="4C2E8DA74C6AC64B1EB72BC1FE742671" box="[151,423,1772,1797]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="andravahana">
|
||
<emphasis id="B95A2A364C6AC64B1EB72BC1FE742671" box="[151,423,1772,1797]" italics="true" pageId="28" pageNumber="29">Mycalesis andravahana</emphasis>
|
||
</taxonomicName>
|
||
, is not conspecific and the genitalia are different (P. Viette dissection slide no. 4847). Differences in the lateral shape of the tegumen and the extent of the valve relative to the uncus tip in this slide to other preparations of
|
||
<taxonomicName id="4C2E8DA74C6AC64B1FBD2A18FDC02638" box="[413,531,1845,1868]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="viettei">
|
||
<emphasis id="B95A2A364C6AC64B1FBD2A18FDC02638" box="[413,531,1845,1868]" italics="true" pageId="28" pageNumber="29">Ht. viettei</emphasis>
|
||
</taxonomicName>
|
||
is considered to represent distortion towards two dimensions due to slide preparation, with distortion occurring most noticeably in the structures most susceptible to pressing, and not in the aedeagus or its relative length.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6AC64A1EE72A8DFC992261" blockId="28.[151,1437,151,2013]" lastBlockId="29.[151,1437,151,2013]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
|
||
<emphasis id="B95A2A364C6AC64B1EE72A8DFDA026CD" bold="true" box="[199,627,1952,1977]" pageId="28" pageNumber="29">Additional information. ♂ genitalia</emphasis>
|
||
: (
|
||
<bibRefCitation id="EFBF8BD54C6AC64B1CAC2A8DFDB326A9" author="Lees" pageId="28" pageNumber="29" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997" yearSuffix="i">
|
||
Lees, 1997: 109,
|
||
<figureCitation id="1315EAA14C6AC64B1D752A8DFC4926CD" box="[853,922,1952,1977]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="28" pageNumber="29">
|
||
Fig.
|
||
<date id="FF90D0E44C6AC64B1DAB2A8DFC4926CD" box="[907,922,1952,1977]" pageId="28" pageNumber="29">7</date>
|
||
</figureCitation>
|
||
i, “12”, voucher 98DL,
|
||
<figureCitation id="1315EAA14C6AC64B1A8A2A8DFB3E26CC" box="[1194,1261,1952,1977]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="28" pageNumber="29">Fig. 8</figureCitation>
|
||
A, specimen of
|
||
<taxonomicName id="4C2E8DA74C6AC64B1EB72AE8FE4326A9" box="[151,400,1988,2013]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6AC64B1EB72AE8FEDC26A8" box="[151,271,1989,2012]" italics="true" pageId="28" pageNumber="29">Heteropsis</emphasis>
|
||
cf.
|
||
<emphasis id="B95A2A364C6AC64B1F1C2AE9FE4326A9" box="[316,400,1988,2013]" italics="true" pageId="28" pageNumber="29">harveyi</emphasis>
|
||
</taxonomicName>
|
||
from Anjozorobe
|
||
</bibRefCitation>
|
||
). From LV, a very shallowly domed tegumen with a rather straight dorsal edge profile and a distinctly angled brow leads to a hand-scythe shaped uncus whose ventral edge at the base is downbent as if like a ‘dewlap’ and with a rather narrow and pointed hook. Tegumen dorsal profile features a quite narrow v-shaped proximad notch and lateral profile bends round to constriction at division with vinculum, which is over half the maximum dimension of the tegumen (and thus the tegumen does not have a quadrate shape from LV). Valve with a very prominent convex dorsal shoulder and valve base very broad leading abruptly to the narrow quite tapering and sub-sinuate valve arms, which are ventrally elbowed and with only a slight indication of a ventral ‘head’ at the unexpanded club, covered with fine setae and sporting a stout spine, inrecurved from DV, where somewhat serrate on mesad edge, pointing towards the uncus tip, which falls well short of the maximum uncus extension, in fact points to the middle of the ‘hook’. Gnathos from right-angled base pointing in line with the uncus tip from LV where relatively straight blade-like, tapered to tip, recurved inward from DV. Saccus quite extended and inflated proximad, less than a third of total valve length (it is longer in the
|
||
<taxonomicName id="4C2E8DA74C6BC64A1A292F2CFB6B236C" box="[1033,1208,511,536]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||
<emphasis id="B95A2A364C6BC64A1A292F2CFB6B236C" box="[1033,1208,511,536]" italics="true" pageId="29" pageNumber="30">Ht. vanewrighti</emphasis>
|
||
</taxonomicName>
|
||
specimen described below). Aedeagus particularly thin and dorsoventrally flattened, very gently uprecurved distad of the ostium, also quite extended proximad of it, distad tip quite pointed, about 1.75 x total length of valve. Juxta quite prominent proximad and rounded like a small plate from DV. A specimen referred to
|
||
<taxonomicName id="4C2E8DA74C6BC64A1DD42F40FBA723F1" box="[1012,1140,620,645]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6BC64A1DD42F40FBA723F1" box="[1012,1140,620,645]" italics="true" pageId="29" pageNumber="30">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
from Miaranony,
|
||
<quantity id="4CD65BC14C6BC64A1B652F41FA4F23F0" box="[1349,1436,620,644]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="29" pageNumber="30" unit="m" value="1080.0">1080 m</quantity>
|
||
(156DL) is very similar in proportions to 98DL, with valve terminating in an inwardly pointing spine that falls short of uncus tip, but has a saccus about 1/4 longer and the hook of uncus about 1/3 deeper (from LV). A future more satisfactory resolution of genitalic variation within the
|
||
<taxonomicName id="4C2E8DA74C6BC64A1D672FF4FC152384" box="[839,966,727,752]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6BC64A1D672FF4FC152384" box="[839,966,727,752]" italics="true" pageId="29" pageNumber="30">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
complex (currently containing four COI- 5P BIN clusters) should be based on DNA barcoded material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C6BC64A1EE72E32FE042794" pageId="29" pageNumber="30" type="description">
|
||
<paragraph id="8B91F6244C6BC64A1EE72E32FD74259C" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="30">
|
||
<emphasis id="B95A2A364C6BC64A1EE72E32FE49224C" bold="true" box="[199,410,799,824]" pageId="29" pageNumber="30">DNA divergences</emphasis>
|
||
: COI-5P cluster number BOLD:ACW5694 (exemplars BMAD217-15 from
|
||
<typeStatus id="549548864C6BC64A1B2E2E0DFAED224C" box="[1294,1342,800,824]" pageId="29" pageNumber="30">type</typeStatus>
|
||
locality and BMAD270-15, DL-4-963 from Ambondrombe) which is 1.52% divergent to
|
||
<emphasis id="B95A2A364C6BC64A1A082E68FB972228" box="[1064,1092,837,860]" italics="true" pageId="29" pageNumber="30">Ht</emphasis>
|
||
. sp. 20 (exemplar BMAD213- 15 DL14A-0387, Anjanaharibe Sud, cluster number BOLD:ACW5777), 1.67% divergent to
|
||
<emphasis id="B95A2A364C6BC64A1AE92E44FB3622F4" box="[1225,1253,873,896]" italics="true" pageId="29" pageNumber="30">Ht</emphasis>
|
||
. sp.
|
||
<taxonomicName id="4C2E8DA74C6BC64A1B012E4AFA4F22F4" box="[1313,1436,871,896]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||
cf.
|
||
<emphasis id="B95A2A364C6BC64A1B682E4AFA4F22F4" box="[1352,1436,871,896]" italics="true" pageId="29" pageNumber="30">harveyi</emphasis>
|
||
</taxonomicName>
|
||
(BMAD268-15, DL14AM-0025 from Anjozorobe, cluster number BOLD:ACW5778) and 1.98% divergent to
|
||
<taxonomicName id="4C2E8DA74C6BC64A1B592EA0FECE22BC" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||
<emphasis id="B95A2A364C6BC64A1B592EA0FECE22BC" italics="true" pageId="29" pageNumber="30">Ht. vanewrighti</emphasis>
|
||
</taxonomicName>
|
||
(BMAD064-04, Bekolosy, cluster number BOLD:AAE4111). The 357 bp cytochrome b sequence of
|
||
<taxonomicName id="4C2E8DA74C6BC64A1EB72EF8FECA2299" box="[151,281,980,1005]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6BC64A1EB72EF8FECA2299" box="[151,281,980,1005]" italics="true" pageId="29" pageNumber="30">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
(exemplars BMNH(E) 671648
|
||
<accessionNumber id="947D6BC74C6BC64A1CAD2EF9FD2C2299" box="[653,767,980,1005]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819453" pageId="29" pageNumber="30">FJ819453</accessionNumber>
|
||
, DL-05-367 from Ranomafana, and DL-0290, BMNH(E) 697406 from Maromizaha), is about 1.12% divergent both to
|
||
<emphasis id="B95A2A364C6BC64A1D482ED4FC572564" box="[872,900,1017,1040]" italics="true" pageId="29" pageNumber="30">Ht</emphasis>
|
||
. sp. 20 (as
|
||
<emphasis id="B95A2A364C6BC64A1A312ED4FBFE2564" box="[1041,1069,1017,1040]" italics="true" pageId="29" pageNumber="30">Ht</emphasis>
|
||
. sp. 77; exemplar CK678 from Anjanaharibe Sud) and to
|
||
<taxonomicName id="4C2E8DA74C6BC64A1FE62930FD942541" box="[454,583,1052,1077]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6BC64A1FE62930FE312540" box="[454,482,1053,1076]" italics="true" pageId="29" pageNumber="30">Ht</emphasis>
|
||
.
|
||
<emphasis id="B95A2A364C6BC64A1FD32931FD942541" box="[499,583,1052,1077]" italics="true" pageId="29" pageNumber="30">harveyi</emphasis>
|
||
</taxonomicName>
|
||
(exemplar BMNH(E) #676467
|
||
<accessionNumber id="947D6BC74C6BC64A1D982931FBFA2541" box="[952,1065,1052,1077]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819454" pageId="29" pageNumber="30">FJ819454</accessionNumber>
|
||
, DL-05-385 from Ranomafana), and is about 1.56% divergent to
|
||
<taxonomicName id="4C2E8DA74C6BC64A1C29296CFD68252C" box="[521,699,1087,1112]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||
<emphasis id="B95A2A364C6BC64A1C29296CFD68252C" box="[521,699,1087,1112]" italics="true" pageId="29" pageNumber="30">Ht. vanewrighti</emphasis>
|
||
</taxonomicName>
|
||
(exemplar DLBEK94-130, BMNH(E) #697875 from Bekolosy), whereas it is about 4.2% pairwise divergent to sequences of
|
||
<taxonomicName id="4C2E8DA74C6BC64A1D6F2948FBFB2508" box="[847,1064,1125,1148]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="imerina">
|
||
<emphasis id="B95A2A364C6BC64A1D6F2948FBFB2508" box="[847,1064,1125,1148]" italics="true" pageId="29" pageNumber="30">Heteropsis imerina</emphasis>
|
||
</taxonomicName>
|
||
(BMNH(E) #676683
|
||
<accessionNumber id="947D6BC74C6BC64A1B062949FA4B2509" box="[1318,1432,1124,1149]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819266" pageId="29" pageNumber="30">FJ819266</accessionNumber>
|
||
, BMNH(E) #676685
|
||
<accessionNumber id="947D6BC74C6BC64A1FA329AAFE2525D4" box="[387,502,1159,1184]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819267" pageId="29" pageNumber="30">FJ819267</accessionNumber>
|
||
;
|
||
<bibRefCitation id="EFBF8BD54C6BC64A1C2329AAFCD925D4" author="Monaghan" box="[515,778,1159,1184]" pageId="29" pageNumber="30" refString="Monaghan, M. T., Wild, R., Elliot, E., Fujisawa, T., Balke, M, Inward, D. J. G., Lees, D. C., Ranaivosolo, R., Eggleton, P., Barraclough, T. G. & Vogler, A. P. (2009) Accelerated species inventory on Madagascar using coalescent-based models of species delineation. Systematic Biology, 58, 298 - 311. http: // dx. doi. org / 10.1093 / sysbio / syp 027" type="journal article" year="2009">
|
||
Monaghan
|
||
<emphasis id="B95A2A364C6BC64A1CA429A4FD6C25D4" box="[644,703,1159,1184]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
|
||
, 2009
|
||
</bibRefCitation>
|
||
; additional members of the
|
||
<taxonomicName id="4C2E8DA74C6BC64A1A6E29A4FB2D25D4" box="[1102,1278,1159,1184]" class="Insecta" family="Nymphalidae" genus="Erebia" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="passandava">
|
||
<emphasis id="B95A2A364C6BC64A1A6E29A4FB2D25D4" box="[1102,1278,1159,1184]" italics="true" pageId="29" pageNumber="30">Ht. passandava</emphasis>
|
||
</taxonomicName>
|
||
group in that study included
|
||
<emphasis id="B95A2A364C6BC64A1F662980FD5425B0" box="[326,647,1196,1221]" italics="true" pageId="29" pageNumber="30">
|
||
<taxonomicName id="4C2E8DA74C6BC64A1F662980FE2A25B1" box="[326,505,1196,1221]" class="Insecta" family="Nymphalidae" genus="Erebia" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="passandava">Ht. passandava</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2E8DA74C6BC64A1C262980FD5125B0" box="[518,642,1197,1220]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="narova">Ht. narova</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
‘
|
||
<taxonomicName id="4C2E8DA74C6BC64A1CB92980FCF225B1" box="[665,801,1196,1221]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="difficilis">
|
||
<emphasis id="B95A2A364C6BC64A1CB92980FCF225B1" box="[665,801,1196,1221]" italics="true" pageId="29" pageNumber="30">Ht. difficilis</emphasis>
|
||
</taxonomicName>
|
||
’ [
|
||
<taxonomicName id="4C2E8DA74C6BC64A1D1C2980FC2E25B1" box="[828,1021,1196,1221]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="andravahana" status="stat. nov.">
|
||
<emphasis id="B95A2A364C6BC64A1D1C2980FC2E25B1" box="[828,1021,1196,1221]" italics="true" pageId="29" pageNumber="30">Ht. andravahana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A269974D4C6BC64A1A262981FBBE25B1" box="[1030,1133,1196,1221]" pageId="29" pageNumber="30">
|
||
<emphasis id="B95A2A364C6BC64A1A262981FBBE25B1" bold="true" box="[1030,1133,1196,1221]" pageId="29" pageNumber="30">stat. rev.</emphasis>
|
||
</taxonomicNameLabel>
|
||
] and
|
||
<taxonomicName id="4C2E8DA74C6BC64A1A912980FACD25B0" box="[1201,1310,1197,1220]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="strato">
|
||
<emphasis id="B95A2A364C6BC64A1A912980FACD25B0" box="[1201,1310,1197,1220]" italics="true" pageId="29" pageNumber="30">Ht. strato</emphasis>
|
||
</taxonomicName>
|
||
). No COII sequence is available on GenBank (see below).
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6BC64A1EE729D9FB79277C" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="30">
|
||
<emphasis id="B95A2A364C6BC64A1EE729D9FE342479" bold="true" box="[199,487,1268,1293]" pageId="29" pageNumber="30">Phylogeny/sister species</emphasis>
|
||
: closely related to
|
||
<taxonomicName id="4C2E8DA74C6BC64A1CE829D8FCA82479" box="[712,891,1268,1293]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||
<emphasis id="B95A2A364C6BC64A1CE829D8FCA82479" box="[712,891,1268,1293]" italics="true" pageId="29" pageNumber="30">Ht. vanewrighti</emphasis>
|
||
</taxonomicName>
|
||
, and to two (?) still undescribed species in the same cryptic complex. In their cladistic analysis of COII sequences, not including
|
||
<taxonomicName id="4C2E8DA74C6BC64A1A692834FECE2421" authority="Torres et al., (2001: 465)" authorityName="Torres et al." authorityPageNumber="465" authorityYear="2001" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||
<emphasis id="B95A2A364C6BC64A1A692834FB292444" box="[1097,1274,1303,1328]" italics="true" pageId="29" pageNumber="30">Ht. vanewrighti</emphasis>
|
||
,
|
||
<bibRefCitation id="EFBF8BD54C6BC64A1B2B2835FF0F2420" author="Torres" pageId="29" pageNumber="30" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. & Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
|
||
Torres
|
||
<emphasis id="B95A2A364C6BC64A1B7B2834FA4F2444" box="[1371,1436,1303,1328]" italics="true" pageId="29" pageNumber="30">et al.,</emphasis>
|
||
(2001
|
||
</bibRefCitation>
|
||
: 465)
|
||
</taxonomicName>
|
||
had full support for a sister relation of
|
||
<taxonomicName id="4C2E8DA74C6BC64A1CCC2810FCBE2421" box="[748,877,1340,1365]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6BC64A1CCC2810FCBE2421" box="[748,877,1340,1365]" italics="true" pageId="29" pageNumber="30">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
(their “
|
||
<emphasis id="B95A2A364C6BC64A1DE92810FC242420" box="[969,1015,1341,1364]" italics="true" pageId="29" pageNumber="30">Hen</emphasis>
|
||
. sp. 12” from Ranomafana National Park); with their “
|
||
<emphasis id="B95A2A364C6BC64A1F44284CFE40240C" box="[356,403,1377,1400]" italics="true" pageId="29" pageNumber="30">Hen</emphasis>
|
||
. sp. 77” from Anjanaharibe Sud, a morphospecies that
|
||
<bibRefCitation id="EFBF8BD54C6BC64A1A28284DFB6F240C" author="Lees" box="[1032,1212,1375,1400]" pageId="29" pageNumber="30" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997: 64)</bibRefCitation>
|
||
also considered the same as “sp. 20” from Ambohitsitondroina Mahalevona; unfortunately there seems to be no sequence on NCBI for the exemplar of “sp. 12”. The “sp. 77” of
|
||
<bibRefCitation id="EFBF8BD54C6BC64A1CA42885FC8A24B4" author="Torres" box="[644,857,1447,1472]" pageId="29" pageNumber="30" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. & Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
|
||
Torres
|
||
<emphasis id="B95A2A364C6BC64A1CF52884FCC524B4" box="[725,790,1447,1472]" italics="true" pageId="29" pageNumber="30">et al.,</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
[here considered to be the same as “sp. 20” from Masoala] appears from putatively fixed differences in its DNA barcodes (n=10 currently on BOLD; cluster number BOLD:ACW5777) not to be conspecific with
|
||
<taxonomicName id="4C2E8DA74C6BC64A1CBD28DCFCC8277C" box="[669,795,1519,1544]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6BC64A1CBD28DCFCC8277C" box="[669,795,1519,1544]" italics="true" pageId="29" pageNumber="30">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
(as also treated by
|
||
<bibRefCitation id="EFBF8BD54C6BC64A1DD428DDFBA1277C" author="Lees" box="[1012,1138,1520,1544]" pageId="29" pageNumber="30" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||
: 64).
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6BC64A1EE72B39FE252759" blockId="29.[151,1437,151,2013]" box="[199,502,1556,1581]" pageId="29" pageNumber="30">
|
||
<emphasis id="B95A2A364C6BC64A1EE72B39FE252759" bold="true" box="[199,502,1556,1581]" pageId="29" pageNumber="30">Ecology and distribution.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6BC64A1EE72B1AFC722724" blockId="29.[151,1437,151,2013]" box="[199,929,1591,1616]" pageId="29" pageNumber="30">
|
||
<emphasis id="B95A2A364C6BC64A1EE72B1AFEF02724" bold="true" box="[199,291,1591,1616]" pageId="29" pageNumber="30">Habitat</emphasis>
|
||
: montane rainforest with fine-leaved climbing bamboos.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6BC64A1EE72B71FC0427EC" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="30">
|
||
<emphasis id="B95A2A364C6BC64A1EE72B71FE902701" bold="true" box="[199,323,1628,1653]" pageId="29" pageNumber="30">Behaviour</emphasis>
|
||
: ♂♂ fly quite rapidly in the substory in fairly open light areas of the forest. ♀♀ have been found resting quite low on tree trunks and fly less readily except just before rain.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6BC64A1EE72B89FC8927C9" blockId="29.[151,1437,151,2013]" box="[199,858,1700,1725]" pageId="29" pageNumber="30">
|
||
<emphasis id="B95A2A364C6BC64A1EE72B89FEEE27C9" bold="true" box="[199,317,1700,1725]" pageId="29" pageNumber="30">Hostplant</emphasis>
|
||
: suspected to be fine-leaved climbing bamboos.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C6BC64A1EE72BEAFE042794" blockId="29.[151,1437,151,2013]" box="[199,471,1735,1760]" pageId="29" pageNumber="30">
|
||
<emphasis id="B95A2A364C6BC64A1EE72BEAFE8B2794" bold="true" box="[199,344,1735,1760]" pageId="29" pageNumber="30">Early stages</emphasis>
|
||
: unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C6BC6491EE72BC1FDF32069" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="distribution">
|
||
<paragraph id="8B91F6244C6BC6491EE72BC1FCDD21A1" blockId="29.[151,1437,151,2013]" lastBlockId="30.[151,1437,151,2036]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||
<emphasis id="B95A2A364C6BC64A1EE72BC1FE8A2671" bold="true" box="[199,345,1772,1797]" pageId="29" pageNumber="30">Distribution</emphasis>
|
||
: from Fianarantsoa/Ranomafana (including
|
||
<typeStatus id="549548864C6BC64A1D482BC0FC4B2671" box="[872,920,1773,1797]" pageId="29" pageNumber="30">type</typeStatus>
|
||
locality) probably at least as far north as Andasibe-Mantadia in the eastern rainforest biome, south to Ambondrombe (according to DNA barcode) and possibly as far South as Chaines Anosyennes (referred specimen) (
|
||
<figureCitation id="1315EAA14C6BC64A1DB02A19FC372638" box="[912,996,1844,1869]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="29" pageNumber="30">Fig. 30</figureCitation>
|
||
D, lilac dots). Light brown dots in
|
||
<figureCitation id="1315EAA14C6BC64A1B502A19FF602604" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="29" pageNumber="30">Fig. 30</figureCitation>
|
||
D represent localities for specimens morphologically similar to
|
||
<taxonomicName id="4C2E8DA74C6BC64A1DAA2A74FBD92604" box="[906,1034,1879,1904]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C6BC64A1DAA2A74FBD92604" box="[906,1034,1879,1904]" italics="true" pageId="29" pageNumber="30">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
falling outside the known range of this species, and where the presence of cluster BOLD:ACW5694 has not been confirmed. Populations in the more central Angavo Massif (exemplar BMAD268-15, cluster number BOLD:ACW5778, Anjozorobe) might represent a different species. However, data is currently insufficient to indicate whether the illustrated genitalia from Anjozorobe represent
|
||
<taxonomicName id="4C2E8DA74C68C6491FB02DB4FDDD21C4" box="[400,526,151,176]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C68C6491FB02DB4FDDD21C4" box="[400,526,151,176]" italics="true" pageId="30" pageNumber="31">Ht. harveyi</emphasis>
|
||
</taxonomicName>
|
||
or not (hence the labelling ‘
|
||
<taxonomicName id="4C2E8DA74C68C6491D6D2DB4FC2321C4" box="[845,1008,151,176]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B95A2A364C68C6491D6D2DB4FCBA21C4" box="[845,873,153,176]" italics="true" pageId="30" pageNumber="31">Ht</emphasis>
|
||
. cf.
|
||
<emphasis id="B95A2A364C68C6491DBC2DBAFC2321C4" box="[924,1008,151,176]" italics="true" pageId="30" pageNumber="31">harveyi</emphasis>
|
||
</taxonomicName>
|
||
’). Ideally, dissections would be based on barcoded material, a matter to be addressed in future.
|
||
</paragraph>
|
||
<paragraph id="8B91F6244C68C6491EE72DF2FDF32069" blockId="30.[151,1437,151,2036]" pageId="30" pageNumber="31">
|
||
<emphasis id="B95A2A364C68C6491EE72DF2FE4F218C" bold="true" box="[199,412,223,248]" pageId="30" pageNumber="31">Elevational range</emphasis>
|
||
:
|
||
<quantity id="4CD65BC14C68C6491F8F2DCDFD912183" box="[431,578,224,248]" metricMagnitude="3" metricUnit="m" metricValue="1.365" metricValueMax="2.03" metricValueMin="0.7" pageId="30" pageNumber="31" unit="m" value="1365.0" valueMax="2030.0" valueMin="700.0">700–2030 m</quantity>
|
||
. (n=86, including referred specimens and observations;
|
||
<quantity id="4CD65BC14C68C6491AEC2DF2FABE2183" box="[1228,1389,223,248]" metricMagnitude="3" metricUnit="m" metricValue="1.3385" metricValueMax="1.59" metricValueMin="1.087" pageId="30" pageNumber="31" unit="m" value="1338.5" valueMax="1590.0" valueMin="1087.0">1087–1590 m</quantity>
|
||
. for DNA barcode-verified specimens).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334A5AF4C68C6481EE72C0AFBA8218A" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="materials_examined">
|
||
<paragraph id="8B91F6244C68C6481EE72C0AFBA8218A" blockId="30.[151,1437,151,2036]" lastBlockId="31.[151,1436,151,254]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
|
||
<emphasis id="B95A2A364C68C6491EE72C0AFE6F2034" bold="true" box="[199,444,295,320]" pageId="30" pageNumber="31">Referred specimens.</emphasis>
|
||
33 ♂♂,
|
||
<collectingCountry id="F339B6B44C68C6491C062C0AFD7E2034" box="[550,685,295,320]" name="Madagascar" pageId="30" pageNumber="31">Madagascar</collectingCountry>
|
||
Fianarantsoa Perrot Frères 2e Semestre 1892/ Ex Oberthür Coll. Brit. Mus. 1927-3; specimen [
|
||
<collectionCode id="ED3F6EE14C68C6491FCC2C60FD992010" box="[492,586,333,356]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="30" pageNumber="31">MNHN</collectionCode>
|
||
], E, Anjorofozo [Anjororo?], Forêt Tanala, Ranomafana
|
||
<collectingCountry id="F339B6B44C68C6491AF12C61FA892011" box="[1233,1370,332,357]" name="Madagascar" pageId="30" pageNumber="31">Madagascar</collectingCountry>
|
||
, [?ca.
|
||
<geoCoordinate id="EE1A90E34C68C6491EB72C5EFF2E20FF" box="[151,253,367,395]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="55" value="-20.933">20.933o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491F2C2C5FFEA120FE" box="[268,370,367,395]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="55" value="47.367">47.367o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491F862C5FFE2020FF" box="[422,499,370,395]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="30" pageNumber="31" unit="km" value="15.0">15 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491C212C5EFD7E20FE" box="[513,685,371,395]" metricMagnitude="3" metricUnit="m" metricValue="1.304" metricValueMax="1.315" metricValueMin="1.293" pageId="30" pageNumber="31" unit="m" value="1304.0" valueMax="1315.0" valueMin="1293.0">1304 +/- 11 m</quantity>
|
||
], 1901,
|
||
<collectingCountry id="F339B6B44C68C6491D2C2C5FFCE320FF" box="[780,816,370,395]" name="Switzerland" pageId="30" pageNumber="31">Ch</collectingCountry>
|
||
. Allaud; ♂, E, Miaranony,
|
||
<quantity id="4CD65BC14C68C6491A4E2C5FFB6D20FF" box="[1134,1214,370,395]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="km" value="0.5">0.5 km</quantity>
|
||
N of, Ranomafana National Park,
|
||
<quantity id="4CD65BC14C68C6491F642CB7FE4820C5" box="[324,411,410,434]" metricMagnitude="3" metricUnit="m" metricValue="1.079" pageId="30" pageNumber="31" unit="m" value="1079.0">1079 m</quantity>
|
||
[
|
||
<geoCoordinate id="EE1A90E34C68C6491F8A2CB4FDF320C6" box="[426,544,406,434]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-21.1633">21.1633o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491C0F2CB4FD7120C5" box="[559,674,406,434]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="47.5528">47.5528o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491CF52CB7FCF420C6" box="[725,807,409,434]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="km" value="0.3">0.3 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491D152CB7FC4320C5" box="[821,912,410,434]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="30" pageNumber="31" unit="m" value="1080.0">1080 m</quantity>
|
||
], 10 hi, Station 11 [fruit trap],
|
||
<date id="FF90D0E44C68C6491AD32CB7FAA620C6" box="[1267,1397,410,434]" pageId="30" pageNumber="31" value="1992-10-23">23/10/1992</date>
|
||
, C. Kremen and H. Raharatsimba|156
|
||
<collectionCode id="ED3F6EE14C68C6491C392CEDFD9320AD" box="[537,576,448,473]" pageId="30" pageNumber="31">DL</collectionCode>
|
||
; 2 ♂♂, E, Tsitery, Forêt de, Tsarafidy [Vatobe forest lies to W.],
|
||
<geoCoordinate id="EE1A90E34C68C6491B032CECFA4520AD" box="[1315,1430,445,473]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-21.1881">21.1881o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491EB72CCAFED9208B" box="[151,266,484,511]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="47.2132">47.2132o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491F1E2CCAFE732374" box="[318,416,487,512]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="30" pageNumber="31" unit="km" value="0.25">0.25 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491F8D2CCAFD84208B" box="[429,599,487,512]" metricMagnitude="3" metricUnit="m" metricValue="1.425" metricValueMax="1.475" metricValueMin="1.375" pageId="30" pageNumber="31" unit="m" value="1425.0" valueMax="1475.0" valueMin="1375.0">1425 +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491C452CCAFD182374" box="[613,715,487,512]" pageId="30" pageNumber="31" value="1995-03-02">2/3/1995</date>
|
||
, D.C. Lees; ♂, E, Vohiparara, 1.4. km W,
|
||
<quantity id="4CD65BC14C68C6491A9A2CCAFAC7208B" box="[1210,1300,487,511]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="30" pageNumber="31" unit="m" value="1220.0">1220 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491B022CCAFA45208B" box="[1314,1430,484,512]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-21.2346">21.2346o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491EB72F23FED92352" box="[151,266,522,550]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="47.3943">47.3943o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491F192F23FE592352" box="[313,394,525,550]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="30" pageNumber="31" unit="km" value="0.9">0.9 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491FB52F23FE3F2352" box="[405,492,526,550]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="30" pageNumber="31" unit="m" value="1220.0">1220 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491FD82F23FDBF2352" box="[504,620,526,550]" pageId="30" pageNumber="31" value="1991-03-13">13/3/1991</date>
|
||
, C. Kremen et al.: Genitalia 100; ♂, data as above but: Genitalia 102; ♂, E, Vohiparara,
|
||
<geoCoordinate id="EE1A90E34C68C6491F602F19FE672339" box="[320,436,561,589]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-21.2346">21.2346o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491FE12F18FDE62338" box="[449,565,561,589]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="47.3943">47.3943o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491C452F18FD162339" box="[613,709,564,589]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="30" pageNumber="31" unit="km" value="0.92">0.92 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491CF12F18FCB72338" box="[721,868,564,589]" metricMagnitude="3" metricUnit="m" metricValue="1.19" metricValueMax="1.1949999999999998" metricValueMin="1.185" pageId="30" pageNumber="31" unit="m" value="1190.0" valueMax="1195.0" valueMin="1185.0">1190 +/- 5 m</quantity>
|
||
<date id="FF90D0E44C68C6491D4C2F19FC332339" box="[876,992,564,589]" pageId="30" pageNumber="31" value="1995-02-19">19/2/1995</date>
|
||
, 11:00, D.C. Lees; specimen [
|
||
<collectionCode id="ED3F6EE14C68C6491B142F18FA412338" box="[1332,1426,565,588]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="30" pageNumber="31">MNHN</collectionCode>
|
||
], E, Vohiparara, Sentier Forestier a
|
||
<quantity id="4CD65BC14C68C6491C342F71FD9E2300" box="[532,589,603,628]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="km" value="3.0">3 km</quantity>
|
||
au S.O. de, Fianarantsoa [
|
||
<geoCoordinate id="EE1A90E34C68C6491D542F71FC1C2300" box="[884,975,600,628]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="555" value="-21.24">21.24o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491DFC2F71FBE02307" box="[988,1075,600,628]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="555" value="47.43">47.43o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491A432F71FB4D2300" box="[1123,1182,603,628]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="km" value="1.0">1 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491A8A2F71FA9D2307" box="[1194,1358,603,628]" metricMagnitude="3" metricUnit="m" metricValue="1.0290000000000001" metricValueMax="1.0790000000000002" metricValueMin="0.9790000000000001" pageId="30" pageNumber="31" unit="m" value="1029.0" valueMax="1079.0" valueMin="979.0">1029 +/- 50 m</quantity>
|
||
],
|
||
<date id="FF90D0E44C68C6491B422F76FF0023EF" pageId="30" pageNumber="31" value="1969-05-25">25/5/ 1969</date>
|
||
; specimen [
|
||
<collectionCode id="ED3F6EE14C68C6491F7F2FAEFE6E23EE" box="[351,445,643,666]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="30" pageNumber="31">MNHN</collectionCode>
|
||
], data as above but:
|
||
<date id="FF90D0E44C68C6491C982FAFFCCD23EF" box="[696,798,642,667]" pageId="30" pageNumber="31" value="1969-03-09">9/3/1969</date>
|
||
; ♀, E, Vohiparara,
|
||
<quantity id="4CD65BC14C68C6491A282FAFFB8823EF" box="[1032,1115,642,667]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="30" pageNumber="31" unit="km" value="0.6">0.6 km</quantity>
|
||
S[W],
|
||
<quantity id="4CD65BC14C68C6491A962FAEFAC123EE" box="[1206,1298,643,667]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="30" pageNumber="31" unit="m" value="1220.0">1220 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491B032FAEFA4523EF" box="[1315,1430,639,667]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-21.2427">21.2427o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491EB72F84FED923B5" box="[151,266,678,706]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="47.3785">47.3785o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491F1C2F84FEA923B6" box="[316,378,681,706]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="km" value="1.0">1 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491FA72F84FDF323B5" box="[391,544,681,706]" metricMagnitude="3" metricUnit="m" metricValue="1.22" metricValueMax="1.226" metricValueMin="1.214" pageId="30" pageNumber="31" unit="m" value="1220.0" valueMax="1226.0" valueMin="1214.0">1220 +/- 6 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491C0D2F84FD7223B5" box="[557,673,681,705]" pageId="30" pageNumber="31" value="1991-03-11">11/3/1991</date>
|
||
, C. Kremen et al.; specimen, data as above but:
|
||
<date id="FF90D0E44C68C6491AED2F84FA9223B6" box="[1229,1345,681,706]" pageId="30" pageNumber="31" value="1995-02-19">19/2/1995</date>
|
||
, 14:30– 15:09, D.C. Lees; ♂, E, Ranomafana National Park, Sahamalaotra,
|
||
<geoCoordinate id="EE1A90E34C68C6491DAB2FFDFC2C239C" box="[907,1023,716,744]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-21.2437">21.2437o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491A2C2FFDFBAC2393" box="[1036,1151,716,744]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="47.4019">47.4019o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491A8E2FFDFADE239C" box="[1198,1293,719,744]" metricMagnitude="3" metricUnit="m" metricValue="1.77" pageId="30" pageNumber="31" unit="km" value="1.77">1.77 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491B382FFDFF7C2278" metricMagnitude="3" metricUnit="m" metricValue="1.087" metricValueMax="1.137" metricValueMin="1.037" pageId="30" pageNumber="31" unit="m" value="1087.0" valueMax="1137.0" valueMin="1037.0">1087 +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491E9F2FD9FEE02279" box="[191,307,756,781]" pageId="30" pageNumber="31" value="2004-12-05">5/12/2004</date>
|
||
, D.C. Lees: DL-05-385;
|
||
<collectionCode id="ED3F6EE14C68C6491C7C2FD9FD602278" box="[604,691,756,780]" pageId="30" pageNumber="31">BMNH</collectionCode>
|
||
(E) #676467 [
|
||
<collectionCode id="ED3F6EE14C68C6491D732FD9FC402278" box="[851,915,756,780]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="30" pageNumber="31">DNA</collectionCode>
|
||
voucher; cytochrome b sequence
|
||
<accessionNumber id="947D6BC74C68C6491B062FD9FA4A2279" box="[1318,1433,756,781]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819454" pageId="30" pageNumber="31">
|
||
<collectionCode id="ED3F6EE14C68C6491B062FD9FA912279" box="[1318,1346,756,781]" pageId="30" pageNumber="31">FJ</collectionCode>
|
||
819454
|
||
</accessionNumber>
|
||
:
|
||
<bibRefCitation id="EFBF8BD54C68C6491EB72E37FE492247" author="Monaghan" box="[151,410,794,819]" pageId="30" pageNumber="31" refString="Monaghan, M. T., Wild, R., Elliot, E., Fujisawa, T., Balke, M, Inward, D. J. G., Lees, D. C., Ranaivosolo, R., Eggleton, P., Barraclough, T. G. & Vogler, A. P. (2009) Accelerated species inventory on Madagascar using coalescent-based models of species delineation. Systematic Biology, 58, 298 - 311. http: // dx. doi. org / 10.1093 / sysbio / syp 027" type="journal article" year="2009">
|
||
Monaghan
|
||
<emphasis id="B95A2A364C68C6491F362E31FE862247" box="[278,341,794,819]" italics="true" pageId="30" pageNumber="31">et al.,</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
]; ♂, E, Ranomafana National Park, Sahamalaotra,
|
||
<geoCoordinate id="EE1A90E34C68C6491DF52E36FB9A2247" box="[981,1097,791,819]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-21.2437">21.2437o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491A762E36FB192246" box="[1110,1226,791,819]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="47.4019">47.4019o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491AD92E36FA8A2247" box="[1273,1369,794,819]" metricMagnitude="3" metricUnit="m" metricValue="1.77" pageId="30" pageNumber="31" unit="km" value="1.77">1.77 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491B442E36FED32223" metricMagnitude="3" metricUnit="m" metricValue="1.087" metricValueMax="1.137" metricValueMin="1.037" pageId="30" pageNumber="31" unit="m" value="1087.0" valueMax="1137.0" valueMin="1037.0">1087 +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491F2F2E13FE502223" box="[271,387,830,855]" pageId="30" pageNumber="31" value="2004-12-05">5/12/2004</date>
|
||
, R. Ranaivosolo: DL-05-367;
|
||
<collectionCode id="ED3F6EE14C68C6491CC42E12FCE82223" box="[740,827,831,855]" pageId="30" pageNumber="31">BMNH</collectionCode>
|
||
(E) #671648 [
|
||
<collectionCode id="ED3F6EE14C68C6491DF92E12FBCA2223" box="[985,1049,831,855]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="30" pageNumber="31">DNA</collectionCode>
|
||
voucher; cytochrome b sequence
|
||
<accessionNumber id="947D6BC74C68C6491EB72E48FED9220A" box="[151,266,869,894]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819453" pageId="30" pageNumber="31">
|
||
<collectionCode id="ED3F6EE14C68C6491EB72E48FF60220A" box="[151,179,869,894]" pageId="30" pageNumber="31">FJ</collectionCode>
|
||
819453
|
||
</accessionNumber>
|
||
:
|
||
<bibRefCitation id="EFBF8BD54C68C6491F342E48FDC5220A" author="Monaghan" box="[276,534,869,894]" pageId="30" pageNumber="31" refString="Monaghan, M. T., Wild, R., Elliot, E., Fujisawa, T., Balke, M, Inward, D. J. G., Lees, D. C., Ranaivosolo, R., Eggleton, P., Barraclough, T. G. & Vogler, A. P. (2009) Accelerated species inventory on Madagascar using coalescent-based models of species delineation. Systematic Biology, 58, 298 - 311. http: // dx. doi. org / 10.1093 / sysbio / syp 027" type="journal article" year="2009">Monaghan et al., 2009</bibRefCitation>
|
||
]; ♂, E, Ranomafana, Sahamahalaotra?,
|
||
<geoCoordinate id="EE1A90E34C68C6491DF42E48FBEA220A" box="[980,1081,866,894]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="55" value="-21.262">21.262o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491A662E4BFB692209" box="[1094,1210,866,894]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="47.4212">47.4212o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491AC92E48FA94220A" box="[1257,1351,869,894]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="30" pageNumber="31" unit="km" value="0.15">0.15 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491B732E4BFF3422D5" metricMagnitude="2" metricUnit="m" metricValue="9.91" metricValueMax="10.16" metricValueMin="9.66" pageId="30" pageNumber="31" unit="m" value="991.0" valueMax="1016.0" valueMin="966.0">991 +/ - 25 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491ED52EA4FEB822D6" box="[245,363,905,930]" pageId="30" pageNumber="31" value="2004-12-05">5/12/2004</date>
|
||
: 12:19, Pierre: DL-05-372, IA194 [isotope voucher]; ♂, E, Vatoharanana, Ranomafana,
|
||
<quantity id="4CD65BC14C68C6491B412EA7FA4F22D6" box="[1377,1436,905,930]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="30" pageNumber="31" unit="km" value="7.0">7 km</quantity>
|
||
SW of,
|
||
<quantity id="4CD65BC14C68C6491ECF2E9DFE9522BC" box="[239,326,944,969]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="30" pageNumber="31" unit="m" value="1150.0">1150 m</quantity>
|
||
, VR,
|
||
<geoCoordinate id="EE1A90E34C68C6491FA82E9DFE2F22BD" box="[392,508,941,969]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-21.2667">21.2667o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491C292E9CFDAE22BC" box="[521,637,941,969]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="47.4333">47.4333o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491C8E2E9CFCDD22BD" box="[686,782,944,969]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="30" pageNumber="31" unit="km" value="0.87">0.87 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491D392E9DFCA322BC" box="[793,880,944,969]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="30" pageNumber="31" unit="m" value="1150.0">1150 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491D5D2E9CFC2222BD" box="[893,1009,945,969]" pageId="30" pageNumber="31" value="1991-02-22">22/2/1991</date>
|
||
, C. Kremen
|
||
<emphasis id="B95A2A364C68C6491AA12E9CFB6922BD" box="[1153,1210,944,969]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
|
||
: Genitalia vial 103; ♂, E, Mt. Maharira,
|
||
<geoCoordinate id="EE1A90E34C68C6491FAD2EFAFDD2229B" box="[397,513,980,1007]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-21.3307">21.3307o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491C322EFAFD55229B" box="[530,646,980,1008]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="47.4162">47.4162o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491C9D2EFAFD2D2284" box="[701,766,983,1008]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="km" value="3.0">3 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491D2D2EFAFC02229B" box="[781,977,983,1008]" metricMagnitude="3" metricUnit="m" metricValue="1.288" metricValueMax="1.3755" metricValueMin="1.2005" pageId="30" pageNumber="31" unit="m" value="1288.0" valueMax="1375.5" valueMin="1200.5">1288 +/- 87.5 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491DC12EFAFB84229B" box="[993,1111,983,1007]" pageId="30" pageNumber="31" value="1992-10-12">12-Oct-92</date>
|
||
, C. Kremen: CK822; ♂, E, Andranobazaha forest, Ambondrombe, camp,
|
||
<geoCoordinate id="EE1A90E34C68C6491C902ED0FCE12562" box="[688,818,1018,1046]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="1" value="-21.87816">21.87816o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491D632ED3FC642562" box="[835,951,1018,1046]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="47.2474">47.2474o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491DCF2ED0FB812562" box="[1007,1106,1021,1046]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="30" pageNumber="31" unit="km" value="0.25">0.25 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491A422ED0FAC12562" box="[1122,1298,1021,1046]" metricMagnitude="3" metricUnit="m" metricValue="1.6039999999999999" metricValueMax="1.654" metricValueMin="1.554" pageId="30" pageNumber="31" unit="m" value="1604.0" valueMax="1654.0" valueMin="1554.0">1604 +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491B032ED3FA4B2562" box="[1315,1432,1022,1046]" pageId="30" pageNumber="31" value="2004-03-22">22/3/2004</date>
|
||
: 11:38, R. Ranaivosolo: DL-4-883; ♂, E, Ambondrombe,
|
||
<geoCoordinate id="EE1A90E34C68C6491D332908FCAF2549" box="[787,892,1057,1085]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="55" value="-21.878">21.878 o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491DA82908FC222548" box="[904,1009,1057,1085]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="55" value="47.247">47.247 o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491A3E2909FBBD2549" box="[1054,1134,1060,1085]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="km" value="0.5">0.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491A592909FACA2548" box="[1145,1305,1060,1085]" metricMagnitude="3" metricUnit="m" metricValue="1.59" metricValueMax="1.64" metricValueMin="1.54" pageId="30" pageNumber="31" unit="m" value="1590.0" valueMax="1640.0" valueMin="1540.0">1590 +/- 50 m</quantity>
|
||
, D.C. Lees: DL-4-963,
|
||
<collectionCode id="ED3F6EE14C68C6491F352966FEBF2510" box="[277,364,1099,1124]" pageId="30" pageNumber="31">BMAD</collectionCode>
|
||
270-15 [
|
||
<collectionCode id="ED3F6EE14C68C6491FE52961FDD62510" box="[453,517,1100,1124]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="30" pageNumber="31">DNA</collectionCode>
|
||
barcode, verified]; ♂, E, Ambondrombe,
|
||
<geoCoordinate id="EE1A90E34C68C6491DFB2961FB962510" box="[987,1093,1096,1124]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="55" value="-21.878">21.878 o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491A712961FB682517" box="[1105,1211,1096,1124]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="55" value="47.247">47.247 o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491ACA2966FAE82510" box="[1258,1339,1099,1124]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="km" value="0.5">0.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491B652966FF3225F3" metricMagnitude="3" metricUnit="m" metricValue="1.59" metricValueMax="1.64" metricValueMin="1.54" pageId="30" pageNumber="31" unit="m" value="1590.0" valueMax="1640.0" valueMin="1540.0">1590 +/ - 50 m</quantity>
|
||
, D.C. Lees: DL-0614,
|
||
<collectionCode id="ED3F6EE14C68C6491FFE295DFDE625FC" box="[478,565,1136,1160]" pageId="30" pageNumber="31">BMNH</collectionCode>
|
||
(E) 671979 [
|
||
<collectionCode id="ED3F6EE14C68C6491C9D295DFD2E25FC" box="[701,765,1136,1160]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="30" pageNumber="31">DNA</collectionCode>
|
||
voucher], KA509 [=KA-P509,
|
||
<collectionCode id="ED3F6EE14C68C6491A7F295DFB4F25FC" box="[1119,1180,1136,1160]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="30" pageNumber="31">DNA</collectionCode>
|
||
extract number]; ♀, E, Ambavafatra-Vohitrasiva, Andringitra,
|
||
<geoCoordinate id="EE1A90E34C68C6491C7629BBFD1A25DB" box="[598,713,1171,1199]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-22.1579">22.1579o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491CF929BBFCED25DA" box="[729,830,1171,1199]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="55" value="47.025">47.025o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491D5329BAFC0625DB" box="[883,981,1174,1199]" metricMagnitude="3" metricUnit="m" metricValue="1.24" pageId="30" pageNumber="31" unit="km" value="1.24">1.24 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491DC329BBFBEE25DA" box="[995,1085,1174,1199]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="30" pageNumber="31" unit="m" value="1350.0">1350 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491A6C29BBFB6C25DB" box="[1100,1215,1174,1199]" pageId="30" pageNumber="31" value="1995-02-17">17/2/1995</date>
|
||
, C. Kremen; ♂, E, Vohitrasiva, near ridge,
|
||
<quantity id="4CD65BC14C68C6491F882990FD9425A1" box="[424,583,1213,1238]" metricMagnitude="3" metricUnit="m" metricValue="1.42" metricValueMax="1.45" metricValueMin="1.39" pageId="30" pageNumber="31" unit="m" value="1420.0" valueMax="1450.0" valueMin="1390.0">1390–1450 m</quantity>
|
||
, Andringitra, [mountaintop on way to Vohipia (
|
||
<quantity id="4CD65BC14C68C6491A482990FB1725A1" box="[1128,1220,1213,1238]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="30" pageNumber="31" unit="m" value="1760.0">1760 m</quantity>
|
||
. pass)],
|
||
<geoCoordinate id="EE1A90E34C68C6491B032990FA4525A2" box="[1315,1430,1210,1238]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-22.1684">22.1684o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491EB729C9FED92588" box="[151,266,1248,1276]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="47.0289">47.0289o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491F7029C9FE782589" box="[336,427,1252,1277]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="km" value="0.1">0.1 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491FE229C9FD562588" box="[450,645,1252,1277]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="30" pageNumber="31" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1450 +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491CBC29C9FCC32589" box="[668,784,1252,1277]" pageId="30" pageNumber="31" value="1995-02-17">17/2/1995</date>
|
||
, 11:40–13:15, D.C. Lees:, specimen [
|
||
<collectionCode id="ED3F6EE14C68C6491ACC29C8FA992588" box="[1260,1354,1253,1276]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="30" pageNumber="31">MNHN</collectionCode>
|
||
], forêt d’Anjavidilava, Andringitra oriental,
|
||
<collectingCountry id="F339B6B44C68C6491C1D2827FD172457" box="[573,708,1290,1315]" name="Madagascar" pageId="30" pageNumber="31">Madagascar</collectingCountry>
|
||
Centre,
|
||
<quantity id="4CD65BC14C68C6491D052827FCA82456" box="[805,891,1290,1315]" metricMagnitude="3" metricUnit="m" metricValue="2.005" pageId="30" pageNumber="31" unit="m" value="2005.0">2005 m</quantity>
|
||
[ca.
|
||
<geoCoordinate id="EE1A90E34C68C6491D932826FBD82457" box="[947,1035,1287,1315]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="555" value="-22.21">22.21o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491A382827FBA32456" box="[1048,1136,1287,1315]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="555" value="46.94">46.94o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491A812827FB262457" box="[1185,1269,1290,1315]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="30" pageNumber="31" unit="km" value="7.5">7.5 km</quantity>
|
||
],
|
||
<date id="FF90D0E44C68C6491B272827FA442457" box="[1287,1431,1290,1315]" pageId="30" pageNumber="31" value="1971-01-01" valueMax="1971-01-15" valueMin="1971-01-01">1–15/1/1971</date>
|
||
, P. Griveaud; specimen [
|
||
<collectionCode id="ED3F6EE14C68C6491F82281DFDD32433" box="[418,512,1328,1351]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="30" pageNumber="31">MNHN</collectionCode>
|
||
], forêt d’Imaitso Anjavidilava, Andringitra-Ambalavao,
|
||
<collectingCountry id="F339B6B44C68C6491A5C2803FAD02433" box="[1148,1283,1326,1351]" name="Madagascar" pageId="30" pageNumber="31">Madagascar</collectingCountry>
|
||
Centre,
|
||
<quantity id="4CD65BC14C68C6491B442802FF7E2419" metricMagnitude="3" metricUnit="m" metricValue="2.03" pageId="30" pageNumber="31" unit="m" value="2030.0">2030 m</quantity>
|
||
[ca.
|
||
<geoCoordinate id="EE1A90E34C68C6491EC7287BFEED241A" box="[231,318,1362,1390]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="555" value="-22.21">22.21o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491F6D2878FE762419" box="[333,421,1362,1390]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="555" value="46.94">46.94o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491FF72878FDF8241A" box="[471,555,1365,1390]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="30" pageNumber="31" unit="km" value="7.5">7.5 km</quantity>
|
||
],
|
||
<date id="FF90D0E44C68C6491C1F2878FD60241A" box="[575,691,1365,1390]" pageId="30" pageNumber="31" value="1958-01-20">20/1/1958</date>
|
||
, P. Griveaud; ♂, [
|
||
<collectionCode id="ED3F6EE14C68C6491DA0287BFC0D2419" box="[896,990,1366,1389]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="30" pageNumber="31">MNHN</collectionCode>
|
||
], SE, Chaines Anosynnes, massif nord, N.-O. de Manentenina, [ca.
|
||
<geoCoordinate id="EE1A90E34C68C6491FCF2851FD9524E1" box="[495,582,1401,1429]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="555" value="-24.16">24.16o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491C7B2850FD6124E0" box="[603,690,1401,1429]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="555" value="47.01">47.01o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491CD12850FC9124E1" box="[753,834,1404,1429]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="30" pageNumber="31" unit="km" value="12.0">12 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491D752851FC6524E0" box="[853,950,1404,1429]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="30" pageNumber="31" unit="m" value="1050.0">1050 m</quantity>
|
||
],
|
||
<date id="FF90D0E44C68C6491DF12850FB8124E1" box="[977,1106,1405,1429]" pageId="30" pageNumber="31" value="1971-11-22">22/11/1971</date>
|
||
, P. Griveaud; [treated with reservation; more northerly records]: ♀, E, Maromizaha, near granite quarry, S side R.N.2
|
||
<quantity id="4CD65BC14C68C6491A83288CFB3824CD" box="[1187,1259,1440,1465]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="30" pageNumber="31" unit="km" value="28.0">28 km</quantity>
|
||
E Moramanga,
|
||
<quantity id="4CD65BC14C68C6491EB728EAFF362494" box="[151,229,1479,1504]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="30" pageNumber="31" unit="km" value="3.2">3.2 km</quantity>
|
||
E of turnoff for Perinet, then right on dirt road to quarry, path to S,
|
||
<quantity id="4CD65BC14C68C6491DF928EAFBA624AB" box="[985,1141,1479,1503]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="30" pageNumber="31" unit="m" value="1150.0" valueMax="1200.0" valueMin="1100.0">1100–1200 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491AA028EAFB2724AB" box="[1152,1268,1476,1504]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-18.9545">18.9545o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491B2028EAFAA724AB" box="[1280,1396,1476,1503]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="48.4477">48.4477o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491EB728C3FF002772" box="[151,211,1517,1542]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="km" value="1.0">1 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491EFE28C0FEAC2772" box="[222,383,1517,1542]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="30" pageNumber="31" unit="m" value="1150.0" valueMax="1200.0" valueMin="1100.0">1150 +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491FAB28C0FE2D2772" box="[395,510,1517,1542]" pageId="30" pageNumber="31" value="1995-01-29">29/1/1995</date>
|
||
, C. Kremen: CK215 [
|
||
<collectionCode id="ED3F6EE14C68C6491CD228C3FCE12772" box="[754,818,1518,1542]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="30" pageNumber="31">DNA</collectionCode>
|
||
voucher]; ♂, E, Maromizaha,
|
||
<geoCoordinate id="EE1A90E34C68C6491AAA28C0FB2E2772" box="[1162,1277,1514,1542]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-18.9668">18.9668o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491B2A28C0FAAE2772" box="[1290,1405,1514,1542]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="48.4547">48.4547o E</geoCoordinate>
|
||
+/ -
|
||
<quantity id="4CD65BC14C68C6491E8A2B3FFEDF275F" box="[170,268,1554,1579]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="30" pageNumber="31" unit="km" value="1.49">1.49 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491F3B2B3FFE68275E" box="[283,443,1554,1578]" metricMagnitude="3" metricUnit="m" metricValue="1.109" metricValueMax="1.182" metricValueMin="1.036" pageId="30" pageNumber="31" unit="m" value="1109.0" valueMax="1182.0" valueMin="1036.0">1109 +/-73 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491FEA2B3FFDED275F" box="[458,574,1554,1579]" pageId="30" pageNumber="31" value="2002-02-25">25/2/2002</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C68C6491CF32B3FFD28275F" box="[723,763,1554,1579]" pageId="30" pageNumber="31">DL</collectionCode>
|
||
02-516; ♂, data as above but: D.C. Lees: A71,
|
||
<collectionCode id="ED3F6EE14C68C6491B052B3FFAAF275E" box="[1317,1404,1554,1578]" pageId="30" pageNumber="31">BMNH</collectionCode>
|
||
(E) #697406 [
|
||
<collectionCode id="ED3F6EE14C68C6491F272B1BFE94273A" box="[263,327,1590,1614]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="30" pageNumber="31">DNA</collectionCode>
|
||
voucher; cytochrome b], KAP545 [=KA-P545,
|
||
<collectionCode id="ED3F6EE14C68C6491D4B2B1BFC7B273A" box="[875,936,1590,1614]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="30" pageNumber="31">DNA</collectionCode>
|
||
extract number]; ♀, E, Andasibe-Mantadia, 11/2014, D.C.Lees:
|
||
<collectionCode id="ED3F6EE14C68C6491F5F2B71FE742701" box="[383,423,1628,1653]" pageId="30" pageNumber="31">DL</collectionCode>
|
||
14AM-156, IA598 [isotope voucher]; ♂, E, Ankeniheny,
|
||
<geoCoordinate id="EE1A90E34C68C6491A612B71FB4B2701" box="[1089,1176,1625,1653]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="555" value="-19.15">19.15o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491A882B70FADE2700" box="[1192,1293,1625,1653]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="55" value="48.303">48.303o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491B632B71FA4B2701" box="[1347,1432,1628,1653]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="30" pageNumber="31" unit="km" value="2.5">2.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491EB72BADFE9327EC" box="[151,320,1664,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.068" metricValueMax="1.1179999999999999" metricValueMin="1.018" pageId="30" pageNumber="31" unit="m" value="1068.0" valueMax="1118.0" valueMin="1018.0">1068 +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491F6E2BACFE6727ED" box="[334,436,1665,1689]" pageId="30" pageNumber="31" value="1994-01-07">7/1/1994</date>
|
||
, WG16 [wing image], IA56 [isotope voucher]; ♂, E, near Ankosy summit, Zahamena,
|
||
<geoCoordinate id="EE1A90E34C68C6491EB72B8AFECB27B4" box="[151,280,1700,1728]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="1" value="-17.49405">17.49405o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491F052B8AFE7527CB" box="[293,422,1700,1728]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="1" value="48.73325">48.73325o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491FF52B8AFD9227B4" box="[469,577,1703,1728]" metricMagnitude="1" metricUnit="m" metricValue="6.7" pageId="30" pageNumber="31" unit="km" value="0.067">0.067 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491C6C2B85FD3E27CB" box="[588,749,1703,1728]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.325" metricValueMin="1.275" pageId="30" pageNumber="31" unit="m" value="1300.0" valueMax="1325.0" valueMin="1275.0">1300 +/- 25 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491CD82B8AFCBF27B4" box="[760,876,1703,1728]" pageId="30" pageNumber="31" value="2006-11-07">7/11/2006</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C68C6491DD52B8AFBCD27B4" box="[1013,1054,1703,1728]" pageId="30" pageNumber="31">DL</collectionCode>
|
||
06-157; ♀, E, slope above cascade [Zahamena NW],
|
||
<geoCoordinate id="EE1A90E34C68C6491F462BE3FE6E2793" box="[358,445,1739,1767]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="555" value="-17.53">17.53o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491FEC2BE3FD932792" box="[460,576,1739,1767]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="48.7269">48.7269o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491C532BE3FD162793" box="[627,709,1742,1767]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="30" pageNumber="31" unit="km" value="1.5">1.5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491CF32BE3FCAF2792" box="[723,892,1742,1767]" metricMagnitude="3" metricUnit="m" metricValue="1.064" metricValueMax="1.089" metricValueMin="1.039" pageId="30" pageNumber="31" unit="m" value="1064.0" valueMax="1089.0" valueMin="1039.0">1064 +/- 25 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491DA92BE3FC2E2793" box="[905,1021,1742,1767]" pageId="30" pageNumber="31" value="2006-11-06">6/11/2006</date>
|
||
, 09:28, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C68C6491AFE2BE3FAD52793" box="[1246,1286,1742,1767]" pageId="30" pageNumber="31">DL</collectionCode>
|
||
06-094, IA31 [isotope voucher];
|
||
<specimenCount id="9D283DAD4C68C6491F4A2BD8FE20267A" box="[362,499,1781,1806]" pageId="30" pageNumber="31" type="generic">5 specimens</specimenCount>
|
||
[
|
||
<collectionCode id="ED3F6EE14C68C6491C202BDBFD8F2679" box="[512,604,1782,1805]" pageId="30" pageNumber="31">BMNH</collectionCode>
|
||
], E, Antsianaka [ca.
|
||
<geoCoordinate id="EE1A90E34C68C6491D612BD8FC67267A" box="[833,948,1778,1806]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-17.6182">17.6182o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491DE02BDBFBF52679" box="[960,1062,1778,1806]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="55" value="48.738">48.738o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491A742BDBFB61267A" box="[1108,1202,1781,1806]" metricMagnitude="4" metricUnit="m" metricValue="1.0699999999999998" pageId="30" pageNumber="31" unit="km" value="10.7">10.7 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491A9D2BDBFAC6267A" box="[1213,1301,1782,1806]" metricMagnitude="3" metricUnit="m" metricValue="1.328" pageId="30" pageNumber="31" unit="m" value="1328.0">1328 m</quantity>
|
||
],1892, E. & B. Perrot;
|
||
<specimenCount id="9D283DAD4C68C6491F2C2A31FE4A2641" box="[268,409,1820,1845]" pageId="30" pageNumber="31" type="generic">2 specimens</specimenCount>
|
||
, E, Belongoina, Reserve Nat. III, Andranomalaza [ca.
|
||
<geoCoordinate id="EE1A90E34C68C6491A212A31FB8A2640" box="[1025,1113,1816,1845]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="555" value="-17.65">17.65o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491A462A31FB1F2640" box="[1126,1228,1816,1845]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="55" value="48.617">48.617o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491ADC2A31FAEA2641" box="[1276,1337,1820,1845]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="km" value="5.0">5 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491B642A31FA432640" box="[1348,1424,1820,1845]" metricMagnitude="2" metricUnit="m" metricValue="9.15" pageId="30" pageNumber="31" unit="m" value="915.0">915 m</quantity>
|
||
], 10/1956, P. Soga; ♀, E, Analamay,
|
||
<geoCoordinate id="EE1A90E34C68C6491C142A6FFD66262F" box="[564,693,1855,1883]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="1" value="-18.80965">18.80965o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491CE52A6FFC95262E" box="[709,838,1855,1883]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="1" value="48.34535">48.34535o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491D5B2A6EFC1C262F" box="[891,975,1858,1883]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="km" value="0.3">0.3 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491DFC2A6FFB54262E" box="[988,1159,1858,1883]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="30" pageNumber="31" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">1050 +/- 50 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491AB52A6EFAD9262F" box="[1173,1290,1859,1883]" pageId="30" pageNumber="31" value="2004-03-13">13/3/2004</date>
|
||
| 16:29, D.C. Lees: DL-4-545; specimen [
|
||
<collectionCode id="ED3F6EE14C68C6491FF22A47FDE326F5" box="[466,560,1898,1921]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="30" pageNumber="31">MNHN</collectionCode>
|
||
], E, Moramanga, Reg. [
|
||
<geoCoordinate id="EE1A90E34C68C6491D1B2A47FC7026F6" box="[827,931,1894,1922]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="55" value="-18.933">18.933o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491D902A47FC2926F5" box="[944,1018,1894,1922]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5555" value="48.2">48.2o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491A0A2A44FBA726F6" box="[1066,1140,1897,1922]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="30" pageNumber="31" unit="km" value="25.0">25 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491AA02A47FB1F26F6" box="[1152,1228,1898,1922]" metricMagnitude="2" metricUnit="m" metricValue="9.02" pageId="30" pageNumber="31" unit="m" value="902.0">902 m</quantity>
|
||
]|193- [last digit of year not specified], G. Oulsoufieff;
|
||
<specimenCount id="9D283DAD4C68C6491C032ABCFD7C26DD" box="[547,687,1936,1961]" pageId="30" pageNumber="31" type="generic">2 specimens</specimenCount>
|
||
, E, Analamazaotra Forest Station,
|
||
<geoCoordinate id="EE1A90E34C68C6491A132ABCFB7526DD" box="[1075,1190,1933,1961]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="-18.9384">18.9384o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491A932ABCFAD826DC" box="[1203,1291,1933,1961]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="555" value="48.41">48.41o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491B192ABCFA4B26DD" box="[1337,1432,1936,1961]" metricMagnitude="2" metricUnit="m" metricValue="7.199999999999999" pageId="30" pageNumber="31" unit="km" value="0.72">0.72 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491EB72A98FE9A26B8" box="[151,329,1973,1997]" metricMagnitude="3" metricUnit="m" metricValue="1.043" metricValueMax="1.136" metricValueMin="0.95" pageId="30" pageNumber="31" unit="m" value="1043.0" valueMax="1136.0" valueMin="950.0">1043 +/- 93 m</quantity>
|
||
,
|
||
<date id="FF90D0E44C68C6491F7B2A98FE1C26B9" box="[347,463,1973,1997]" pageId="30" pageNumber="31" value="2002-02-27">27/2/2002</date>
|
||
, D.C. Lees:
|
||
<collectionCode id="ED3F6EE14C68C6491C4A2A99FD4126B9" box="[618,658,1972,1997]" pageId="30" pageNumber="31">DL</collectionCode>
|
||
02-535A,
|
||
<collectionCode id="ED3F6EE14C68C6491D272A99FCFC26B9" box="[775,815,1972,1997]" pageId="30" pageNumber="31">DL</collectionCode>
|
||
02-535B; specimen [
|
||
<collectionCode id="ED3F6EE14C68C6491A3F2A98FBAE26B8" box="[1055,1149,1973,1996]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="30" pageNumber="31">MNHN</collectionCode>
|
||
], Ambodiriana, route de Lakato, km 10,
|
||
<collectingCountry id="F339B6B44C68C6491F712AF6FE0B2680" box="[337,472,2011,2036]" name="Madagascar" pageId="30" pageNumber="31">Madagascar</collectingCountry>
|
||
Est,
|
||
<quantity id="4CD65BC14C68C6491C372AF6FDA32687" box="[535,624,2011,2036]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="30" pageNumber="31" unit="m" value="1050.0">1050 m</quantity>
|
||
[
|
||
<geoCoordinate id="EE1A90E34C68C6491CA02AF6FD3A2687" box="[640,745,2007,2036]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="55" value="-19.025">19.025o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C68C6491CD82AF6FC832687" box="[760,848,2007,2036]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="555" value="48.35">48.35o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C68C6491DA62AF6FC162680" box="[902,965,2011,2036]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="km" value="1.0">1 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C68C6491DF32AF6FBFC2687" box="[979,1071,2011,2036]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="30" pageNumber="31" unit="m" value="1050.0">1050 m</quantity>
|
||
], 3/1957, P. Griveaud, R. Vieu; specimen, E, Sandrangato [
|
||
<geoCoordinate id="EE1A90E34C69C6481FE72DB6FDEE21C7" box="[455,573,151,179]" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="5" value="-19.1083">19.1083o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C69C6481C6A2DB6FD6D21C6" box="[586,702,151,179]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="5" value="48.2417">48.2417o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C69C6481CCD2DB7FCE421C7" box="[749,823,154,179]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="32" unit="km" value="15.0">15 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C69C6481D622DB6FC4F21C6" box="[834,924,155,179]" metricMagnitude="3" metricUnit="m" metricValue="1.033" pageId="31" pageNumber="32" unit="m" value="1033.0">1033 m</quantity>
|
||
]; ♀, E, Sandrangate,
|
||
<geoCoordinate id="EE1A90E34C69C6481AA92DB6FB2E21C7" box="[1161,1277,151,179]" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="5" value="-19.1083">19.1083o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C69C6481B2A2DB6FAAE21C6" box="[1290,1405,151,179]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="5" value="48.2417">48.2417o E</geoCoordinate>
|
||
+/ -
|
||
<quantity id="4CD65BC14C69C6481E872D93FF2221A3" box="[167,241,190,215]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="32" unit="km" value="15.0">15 km</quantity>
|
||
, 1033,
|
||
<date id="FF90D0E44C69C6481F6A2D93FE6E21A3" box="[330,445,190,215]" pageId="31" pageNumber="32" value="1994-11-06">6/11/1994</date>
|
||
, WG41 [wing image], IA215 [isotope voucher]; specimen [
|
||
<collectionCode id="ED3F6EE14C69C6481A792DEDFB6421A3" box="[1113,1207,192,215]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="31" pageNumber="32">MNHN</collectionCode>
|
||
], E, Mahatsinjo pres Tananarive
|
||
<collectingCountry id="F339B6B44C69C6481F3A2DC8FE70218A" box="[282,419,229,254]" name="Madagascar" pageId="31" pageNumber="32">Madagascar</collectingCountry>
|
||
, [?ca.
|
||
<geoCoordinate id="EE1A90E34C69C6481FCB2DC8FD83218A" box="[491,592,226,254]" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="55" value="-19.167">19.167o S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1A90E34C69C6481C7D2DCBFD112189" box="[605,706,226,254]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="55" value="48.033">48.033o E</geoCoordinate>
|
||
+/-
|
||
<quantity id="4CD65BC14C69C6481CD12DC8FCE9218A" box="[753,826,229,254]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="32" unit="km" value="15.0">15 km</quantity>
|
||
,
|
||
<quantity id="4CD65BC14C69C6481D662DCBFC422189" box="[838,913,230,254]" metricMagnitude="2" metricUnit="m" metricValue="9.129999999999999" pageId="31" pageNumber="32" unit="m" value="913.0">913 m</quantity>
|
||
], 1913, H. Donckier.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |