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<document ID-DOI="10.11646/palaeoentomology.6.1.10" ID-ISSN="2624-2834" ID-Zenodo-Dep="7754820" ID-ZooBank="B822ED53-D559-4DD8-8803-3F2850A00445" approvalRequired="1" approvalRequired_for_taxonomicNames="1" checkinTime="1679380739061" checkinUser="plazi" docAuthor="XUAN, QIANG, CAI, CHEN-YANG &amp; HUANG, DI-YING" docDate="2023" docId="F308E17F435FFFF0FF6F486DFD5C1E56" docLanguage="en" docName="palaeoentomology.6.1.10.pdf" docOrigin="Palaeoentomology 6 (1)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.1.10" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.3:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="3" docTitle="Betaburmesebuthus fleissneri Lourenco &amp; Velten 2016" docType="treatment" docVersion="4" lastPageNumber="83" masterDocId="0F3199074350FFE3FFE74E07FFFE192E" masterDocTitle="Revision of palaeoburmesebuthid scorpions in mid-Cretaceous amber from northern Myanmar (Scorpiones: Buthoidea)" masterLastPageNumber="101" masterPageNumber="64" pageNumber="79" updateTime="1679418755468" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of palaeoburmesebuthid scorpions in mid-Cretaceous amber from northern Myanmar (Scorpiones: Buthoidea)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>XUAN, QIANG</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>CAI, CHEN-YANG</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>HUANG, DI-YING</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Palaeoentomology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>6</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>64</mods:start>
<mods:end>101</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/palaeoentomology.6.1.10</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/palaeoentomology.6.1.10</mods:identifier>
<mods:identifier type="ISSN">2624-2834</mods:identifier>
<mods:identifier type="Zenodo-Dep">7754820</mods:identifier>
<mods:identifier type="ZooBank">B822ED53-D559-4DD8-8803-3F2850A00445</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:F308E17F435FFFF0FF6F486DFD5C1E56" httpUri="http://treatment.plazi.org/id/F308E17F435FFFF0FF6F486DFD5C1E56" lastPageId="19" lastPageNumber="83" pageId="15" pageNumber="79">
<subSubSection pageId="15" pageNumber="79" type="nomenclature">
<paragraph blockId="15.[136,777,1642,1746]" pageId="15" pageNumber="79">
<heading bold="true" centered="true" fontSize="10" level="2" pageId="15" pageNumber="79" reason="3">
<taxonomicName authority="Lourenco &amp; Velten, 2016" authorityName="Lourenco &amp; Velten" authorityYear="2016" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="79" phylum="Arthropoda" rank="species" species="fleissneri">
<emphasis bold="true" pageId="15" pageNumber="79">
<emphasis bold="true" box="[136,366,1642,1669]" italics="true" pageId="15" pageNumber="79">Betaburmesebuthus</emphasis>
fleissneri Lourenço &amp; Velten, 2016
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[136,280,1719,1746]" pageId="15" pageNumber="79" type="description">
<paragraph blockId="15.[136,777,1642,1746]" box="[136,280,1719,1746]" pageId="15" pageNumber="79">
(
<figureCitation box="[144,272,1719,1746]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStartId-0="16.[136,235,1557,1581]" captionStartId-1="17.[136,235,1927,1951]" captionStartId-2="18.[136,235,1946,1970]" captionTargetBox-0="[151,1436,189,1534]" captionTargetBox-1="[170,1416,189,1904]" captionTargetBox-2="[151,1436,189,1922]" captionTargetId-0="figure-165@16.[151,1436,189,1534]" captionTargetId-1="figure-28@17.[170,1416,189,1904]" captionTargetId-2="figure-18@18.[151,1436,189,1922]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionText-0="FIGURE 12. General habitus of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Dorsal view. B, Ventral view. Scale bars = 5 mm." captionText-1="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." captionText-2="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi-0="http://doi.org/10.5281/zenodo.7754850" figureDoi-1="http://doi.org/10.5281/zenodo.7754852" figureDoi-2="http://doi.org/10.5281/zenodo.7754856" httpUri-0="https://zenodo.org/record/7754850/files/figure.png" httpUri-1="https://zenodo.org/record/7754852/files/figure.png" httpUri-2="https://zenodo.org/record/7754856/files/figure.png" pageId="15" pageNumber="79">Figs 12–14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="79" type="materials_examined">
<paragraph blockId="15.[136,778,1796,2093]" pageId="15" pageNumber="79">
<emphasis bold="true" box="[136,247,1796,1822]" pageId="15" pageNumber="79">Material.</emphasis>
<collectionCode box="[263,330,1796,1823]" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="15" pageNumber="79">NIGP</collectionCode>
200653, an adult male, part of right pedipalp missing.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="79" type="diagnosis">
<paragraph blockId="15.[136,778,1796,2093]" lastBlockId="15.[810,1452,1642,2093]" pageId="15" pageNumber="79">
<emphasis bold="true" box="[189,449,1873,1899]" pageId="15" pageNumber="79">Diagnosis (emended).</emphasis>
This species can be clearly distinguished from other
<taxonomicName box="[456,778,1912,1938]" pageId="15" pageNumber="79">
<emphasis box="[456,677,1912,1938]" italics="true" pageId="15" pageNumber="79">Betaburmesebuthus</emphasis>
species
</taxonomicName>
by the following set of characters: 1) anterior margin of carapace with a single moderately marked median concavity (
<figureCitation box="[264,376,2028,2054]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="15" pageNumber="79">Figs 13A</figureCitation>
,
<figureCitation box="[392,444,2028,2054]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="15" pageNumber="79">14A</figureCitation>
;
<bibRefCitation author="Lourenco, W. R. &amp; Velten, J." box="[460,775,2028,2054]" pageId="15" pageNumber="79" pagination="4 - 11" refId="ref13434" refString="Lourenco, W. R. &amp; Velten, J. (2016 a) One more new species of Betaburmesebuthus Lourenco, 2015 (Scorpiones: Palaeoburmesebuthinae) from Cretaceous burmite. Arachnida, Rivista Aracnologica Italiana, 6, 4 - 11." type="journal article" year="2016">Lourenço &amp; Velten, 2016a</bibRefCitation>
: fig. 5); 2) spiracles oval to round (
<bibRefCitation author="Lourenco, W. R. &amp; Velten, J." box="[547,883,1642,2092]" pageId="15" pageNumber="79" pagination="4 - 11" refId="ref13434" refString="Lourenco, W. R. &amp; Velten, J. (2016 a) One more new species of Betaburmesebuthus Lourenco, 2015 (Scorpiones: Palaeoburmesebuthinae) from Cretaceous burmite. Arachnida, Rivista Aracnologica Italiana, 6, 4 - 11." type="journal article" year="2016">Lourenço &amp; Velten, 2016a</bibRefCitation>
: fig. 6); 3) pectines with 19–21 teeth (
<figureCitation box="[1341,1446,1642,1668]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="15" pageNumber="79">Fig. 13B</figureCitation>
;
<bibRefCitation author="Lourenco, W. R. &amp; Velten, J." box="[810,1122,1681,1707]" pageId="15" pageNumber="79" pagination="4 - 11" refId="ref13434" refString="Lourenco, W. R. &amp; Velten, J. (2016 a) One more new species of Betaburmesebuthus Lourenco, 2015 (Scorpiones: Palaeoburmesebuthinae) from Cretaceous burmite. Arachnida, Rivista Aracnologica Italiana, 6, 4 - 11." type="journal article" year="2016">Lourenço &amp; Velten, 2016a</bibRefCitation>
); 4) vesicle pear-shaped and very long, and aculeus very long and strongly curved (
<figureCitation captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="15" pageNumber="79">Figs 13B</figureCitation>
,
<figureCitation box="[871,921,1758,1784]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="15" pageNumber="79">14B</figureCitation>
); 5) dorsal patellar spur carina with four strong tubercles and femur with three to four internal strong tubercles (
<figureCitation box="[930,1073,1835,1862]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="15" pageNumber="79">Fig. 14C, D</figureCitation>
;
<bibRefCitation author="Lourenco, W. R. &amp; Velten, J." box="[1088,1400,1835,1862]" pageId="15" pageNumber="79" pagination="4 - 11" refId="ref13434" refString="Lourenco, W. R. &amp; Velten, J. (2016 a) One more new species of Betaburmesebuthus Lourenco, 2015 (Scorpiones: Palaeoburmesebuthinae) from Cretaceous burmite. Arachnida, Rivista Aracnologica Italiana, 6, 4 - 11." type="journal article" year="2016">Lourenço &amp; Velten, 2016a</bibRefCitation>
: fig. 2); 6) Chela manus with two small tubercles on internal surface (
<bibRefCitation author="Lourenco, W. R. &amp; Velten, J." box="[905,1207,1912,1938]" pageId="15" pageNumber="79" pagination="4 - 11" refId="ref13434" refString="Lourenco, W. R. &amp; Velten, J. (2016 a) One more new species of Betaburmesebuthus Lourenco, 2015 (Scorpiones: Palaeoburmesebuthinae) from Cretaceous burmite. Arachnida, Rivista Aracnologica Italiana, 6, 4 - 11." type="journal article" year="2016">Lourenço &amp; Velten, 2016a</bibRefCitation>
: fig. 10); 7) telotarsus covered by numerous ventrosubmedian setae (
<figureCitation captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="15" pageNumber="79">Fig. 14E– G</figureCitation>
,
<bibRefCitation author="Lourenco, W. R. &amp; Velten, J." box="[844,1145,1989,2016]" pageId="15" pageNumber="79" pagination="4 - 11" refId="ref13434" refString="Lourenco, W. R. &amp; Velten, J. (2016 a) One more new species of Betaburmesebuthus Lourenco, 2015 (Scorpiones: Palaeoburmesebuthinae) from Cretaceous burmite. Arachnida, Rivista Aracnologica Italiana, 6, 4 - 11." type="journal article" year="2016">Lourenço &amp; Velten, 2016a</bibRefCitation>
: fig. 8).
</paragraph>
</subSubSection>
<subSubSection lastPageId="19" lastPageNumber="83" pageId="15" pageNumber="79" type="description">
<paragraph blockId="15.[810,1452,1642,2093]" lastBlockId="16.[136,778,1677,2093]" lastPageId="16" lastPageNumber="80" pageId="15" pageNumber="79">
<emphasis bold="true" box="[863,1009,2028,2054]" pageId="15" pageNumber="79">Description.</emphasis>
Carapace. Covered by coarse granules; a pair of superciliary carinae well developed; median, posterior median and posterior furrows evident (
<figureCitation captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="16" pageNumber="80">Figs 13A</figureCitation>
,
<figureCitation box="[201,252,1717,1743]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="16" pageNumber="80">14A</figureCitation>
); median eyes oval, large sized and separated by one ocular diameter (
<figureCitation box="[403,506,1755,1781]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="16" pageNumber="80">Fig. 14A</figureCitation>
); PDMi posterodorsal to PLMa, ADMi mixed with coarse granules, not clear (
<figureCitation captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="16" pageNumber="80">Fig. 14A</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7754850" ID-Zenodo-Dep="7754850" httpUri="https://zenodo.org/record/7754850/files/figure.png" pageId="16" pageNumber="80" startId="16.[136,235,1557,1581]" targetBox="[151,1436,189,1534]" targetPageId="16">
<paragraph blockId="16.[136,1452,1557,1617]" pageId="16" pageNumber="80">
<emphasis bold="true" box="[136,272,1557,1581]" pageId="16" pageNumber="80">FIGURE 12.</emphasis>
General habitus of
<taxonomicName authority=", NIGP" authorityName="NIGP" box="[467,836,1557,1581]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="80" phylum="Arthropoda" rank="species" species="fleissneri">
<emphasis box="[467,765,1557,1581]" italics="true" pageId="16" pageNumber="80">Betaburmesebuthus fleissneri</emphasis>
, NIGP
</taxonomicName>
201154, under incident light.
<emphasis bold="true" box="[1129,1147,1557,1581]" pageId="16" pageNumber="80">A</emphasis>
, Dorsal view.
<emphasis bold="true" box="[1291,1308,1558,1582]" pageId="16" pageNumber="80">B</emphasis>
, Ventral view. Scale bars = 5 mm.
</paragraph>
</caption>
<paragraph blockId="16.[136,778,1677,2093]" pageId="16" pageNumber="80">
Coxosternal region. Surface covered by impurity, not clear but obscure outline visible; genital operculum longitudinally divided, two valves rounded and full (
<figureCitation captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="16" pageNumber="80">Fig. 13B</figureCitation>
).
</paragraph>
<paragraph blockId="16.[136,778,1677,2093]" pageId="16" pageNumber="80">
Chelicerae. With numerous setae on internal surface (
<figureCitation box="[144,249,2066,2092]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="16" pageNumber="80">Fig. 14A</figureCitation>
); cheliceral dentition not visible (
<figureCitation box="[624,729,2066,2092]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="16" pageNumber="80">Fig. 14A</figureCitation>
).
</paragraph>
<paragraph blockId="16.[810,1452,1677,2093]" pageId="16" pageNumber="80">
Pedipalps. Femur with five carinae (
<figureCitation box="[1270,1374,1677,1704]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="16" pageNumber="80">Figs 13E</figureCitation>
,
<figureCitation box="[1386,1439,1678,1704]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="16" pageNumber="80">14D</figureCitation>
): internomedian carina well-marked with several strong tubercles; dorsointernal carina crenulate; dorsoexternal, ventroexternal and ventrointernal carinae costate. Patella with seven carinae (
<figureCitation box="[1061,1174,1833,1859]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="16" pageNumber="80">Figs 13E</figureCitation>
,
<figureCitation box="[1195,1246,1833,1859]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="16" pageNumber="80">14C</figureCitation>
): Dorsal Patellar Spur carina (DPSc) and Ventral Patellar Spur carina (VPSc) well-developed; dorsointernal, dorsomedian and ventrointernal carinae costate with several granules; ventroexternal and dorsoexternal carinae smooth and costate. Chela very slender (Cl/Cw = 7.67,
<tableCitation box="[1293,1375,2027,2054]" captionStart="TABLE 1" captionStartId="10.[136,224,496,522]" captionTargetPageId="10" captionText="TABLE 1. Measurements (mm) of studied specimens and corresponding holotype. Question marks signify doubtful data; value in brackets signify re-estimation from the images in original papers." httpUri="http://table.plazi.org/id/2FDE00E1435AFFE9FF6F4FF7FED11B00" pageId="16" pageNumber="80" tableUuid="2FDE00E1435AFFE9FF6F4FF7FED11B00">Table 1</tableCitation>
).
</paragraph>
<paragraph blockId="16.[810,1452,1677,2093]" lastBlockId="19.[136,779,158,1913]" lastPageId="19" lastPageNumber="83" pageId="16" pageNumber="80">
Trichobothrial pattern (
<figureCitation box="[1135,1245,2066,2092]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="16" pageNumber="80">Figs 13E</figureCitation>
,
<figureCitation box="[1262,1353,2066,2092]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="16" pageNumber="80">14C, D</figureCitation>
). Femur with 7 trichobothria observed, 4 dorsal, 1 internal and 2 external trichobothria, trichobothrium
<emphasis box="[609,630,195,227]" italics="true" pageId="19" pageNumber="83">
e
<subScript attach="left" box="[622,630,213,227]" fontSize="6" pageId="19" pageNumber="83">1</subScript>
</emphasis>
proximal to trichobothrium
<emphasis box="[313,335,230,263]" italics="true" pageId="19" pageNumber="83">
d
<subScript attach="left" box="[327,335,249,263]" fontSize="6" pageId="19" pageNumber="83">5</subScript>
</emphasis>
. Patella with 8 trichobothria observed, including 1 dorsal, 1internal and 6 external trichobothria. Chela fixed finger with 2 dorsal trichobothria observed.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7754852" ID-Zenodo-Dep="7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="17" pageNumber="81" startId="17.[136,235,1927,1951]" targetBox="[170,1416,189,1904]" targetPageId="17">
<paragraph blockId="17.[136,1452,1927,2060]" pageId="17" pageNumber="81">
<emphasis bold="true" box="[136,277,1927,1952]" pageId="17" pageNumber="81">FIGURE 13.</emphasis>
Details of
<taxonomicName authority=", NIGP" authorityName="NIGP" box="[400,780,1928,1952]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="81" phylum="Arthropoda" rank="species" species="fleissneri">
<emphasis box="[400,704,1928,1952]" italics="true" pageId="17" pageNumber="81">Betaburmesebuthus fleissneri</emphasis>
, NIGP
</taxonomicName>
201154, under incident light.
<emphasis bold="true" box="[1094,1112,1927,1951]" pageId="17" pageNumber="81">A</emphasis>
, Carapace and tergites I–III.
<emphasis bold="true" box="[1428,1445,1928,1952]" pageId="17" pageNumber="81">B</emphasis>
, Coxosternal region, pectines and sternites.
<emphasis bold="true" box="[577,595,1963,1987]" pageId="17" pageNumber="81">C</emphasis>
,
<emphasis bold="true" box="[609,622,1964,1988]" pageId="17" pageNumber="81">J</emphasis>
, Metasoma and telson, lateral view.
<emphasis bold="true" box="[1000,1018,1964,1988]" pageId="17" pageNumber="81">D</emphasis>
, Tergites IV–VII.
<emphasis bold="true" box="[1208,1225,1964,1988]" pageId="17" pageNumber="81">E</emphasis>
, Left pedipalp, dorsal view.
<emphasis bold="true" box="[196,211,2000,2024]" pageId="17" pageNumber="81">F</emphasis>
, Dentition of left chela movable finger.
<emphasis bold="true" box="[620,640,1999,2023]" pageId="17" pageNumber="81">G</emphasis>
, Tibial spur of leg IV.
<emphasis bold="true" box="[869,889,2000,2024]" pageId="17" pageNumber="81">H</emphasis>
, Tibial spur of leg III.
<emphasis bold="true" box="[1120,1130,2000,2024]" pageId="17" pageNumber="81">I</emphasis>
, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in
<emphasis bold="true" box="[657,675,2035,2059]" pageId="17" pageNumber="81">C</emphasis>
,
<emphasis bold="true" box="[688,701,2036,2060]" pageId="17" pageNumber="81">J</emphasis>
; 1 mm in
<emphasis bold="true" box="[805,823,2035,2059]" pageId="17" pageNumber="81">A</emphasis>
,
<emphasis bold="true" box="[836,853,2036,2060]" pageId="17" pageNumber="81">B</emphasis>
,
<emphasis bold="true" box="[865,883,2036,2060]" pageId="17" pageNumber="81">D</emphasis>
,
<emphasis bold="true" box="[896,913,2036,2060]" pageId="17" pageNumber="81">E</emphasis>
; 500 μm in
<emphasis bold="true" box="[1036,1051,2036,2060]" pageId="17" pageNumber="81">F</emphasis>
,
<emphasis bold="true" box="[1065,1085,2035,2059]" pageId="17" pageNumber="81">G</emphasis>
,
<emphasis bold="true" box="[1097,1117,2036,2060]" pageId="17" pageNumber="81">H</emphasis>
,
<emphasis bold="true" box="[1129,1139,2036,2060]" pageId="17" pageNumber="81">I</emphasis>
.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7754856" ID-Zenodo-Dep="7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="18" pageNumber="82" startId="18.[136,235,1946,1970]" targetBox="[151,1436,189,1922]" targetPageId="18">
<paragraph blockId="18.[136,1451,1946,2042]" pageId="18" pageNumber="82">
<emphasis bold="true" box="[136,272,1946,1970]" pageId="18" pageNumber="82">FIGURE 14.</emphasis>
Details of
<taxonomicName authority=", NIGP" authorityName="NIGP" box="[384,755,1946,1970]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="82" phylum="Arthropoda" rank="species" species="fleissneri">
<emphasis box="[384,683,1946,1970]" italics="true" pageId="18" pageNumber="82">Betaburmesebuthus fleissneri</emphasis>
, NIGP
</taxonomicName>
201154, under green epifluorescence.
<emphasis bold="true" box="[1139,1157,1946,1970]" pageId="18" pageNumber="82">A</emphasis>
, Carapace and chelicerae.
<emphasis bold="true" box="[1428,1445,1946,1970]" pageId="18" pageNumber="82">B</emphasis>
, Telson, lateral view.
<emphasis bold="true" box="[345,363,1982,2006]" pageId="18" pageNumber="82">C</emphasis>
, Patella, dorsal view.
<emphasis bold="true" box="[584,602,1982,2006]" pageId="18" pageNumber="82">D</emphasis>
, Femur, dorsal view.
<emphasis bold="true" box="[820,837,1982,2006]" pageId="18" pageNumber="82">E</emphasis>
, Telotarsus of leg IV.
<emphasis bold="true" box="[1059,1074,1982,2006]" pageId="18" pageNumber="82">F</emphasis>
, Telotarsus of leg II.
<emphasis bold="true" box="[1289,1309,1982,2006]" pageId="18" pageNumber="82">G</emphasis>
, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm.
</paragraph>
</caption>
<paragraph blockId="19.[136,779,158,1913]" pageId="19" pageNumber="83">
Legs. Prolateral and retrolateral pedal spurs present on all legs (
<figureCitation box="[279,420,374,400]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="19" pageNumber="83">Fig. 14E–G</figureCitation>
). Ungues long and moderately curved, dactyl pointed (
<figureCitation box="[401,538,410,436]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="19" pageNumber="83">Fig. 14E–G</figureCitation>
).
</paragraph>
<paragraph blockId="19.[136,779,158,1913]" pageId="19" pageNumber="83">
Pectines (
<figureCitation box="[297,399,446,472]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="19" pageNumber="83">Fig. 13B</figureCitation>
). Lamellae not clear, peg sensillae very short and sensory area fully occupied distal end of teeth.
</paragraph>
<paragraph blockId="19.[136,779,158,1913]" pageId="19" pageNumber="83">
Mesosoma. Tergites covered by coarse granules (
<figureCitation box="[144,254,590,616]" captionStart="FIGURE 12" captionStartId="16.[136,235,1557,1581]" captionTargetBox="[151,1436,189,1534]" captionTargetId="figure-165@16.[151,1436,189,1534]" captionTargetPageId="16" captionText="FIGURE 12. General habitus of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Dorsal view. B, Ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7754850" httpUri="https://zenodo.org/record/7754850/files/figure.png" pageId="19" pageNumber="83">Figs 12A</figureCitation>
,
<figureCitation box="[268,356,590,616]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="19" pageNumber="83">13A, D</figureCitation>
), median carina evident on posterior half of all tergites (
<figureCitation box="[347,451,626,652]" captionStart="FIGURE 12" captionStartId="16.[136,235,1557,1581]" captionTargetBox="[151,1436,189,1534]" captionTargetId="figure-165@16.[151,1436,189,1534]" captionTargetPageId="16" captionText="FIGURE 12. General habitus of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Dorsal view. B, Ventral view. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7754850" httpUri="https://zenodo.org/record/7754850/files/figure.png" pageId="19" pageNumber="83">Fig. 12A</figureCitation>
); dorsolateral carinae present on tergites IV–VI (
<figureCitation box="[347,451,662,688]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="19" pageNumber="83">Fig. 13D</figureCitation>
); tergite VII with five costate carinae (
<figureCitation box="[231,335,698,724]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="19" pageNumber="83">Fig. 13D</figureCitation>
): paired dorsolateral and lateral carinae, and one axial carina on anterior half. Sternites covered by impurity, not clear.
</paragraph>
<paragraph blockId="19.[136,779,158,1913]" pageId="19" pageNumber="83">
Metasoma (
<figureCitation box="[326,460,806,832]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="19" pageNumber="83">Fig. 13C, J</figureCitation>
). All segments longer than width and covered by macrosetae ventrally.Segments I–III with distinct depression dorsally; segments I and II with ten well marked carinae (paired ventromedian, ventrolateral, lateral, dorsolateral, and dorsal carinae); segments III and IV with eight well marked carinae (paired ventromedian, ventrolateral, dorsolateral, and dorsal carinae) and lateral carina reduced on segment III; segment V with five well marked carinae (single ventromedian carina, paired ventrolateral and dorsolateral carinae); dorsal carinae on segments I–IV obviously serrated.
</paragraph>
<paragraph blockId="19.[136,779,158,1913]" pageId="19" pageNumber="83">
Telson (
<figureCitation box="[275,377,1202,1228]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="19" pageNumber="83">Figs13C</figureCitation>
,
<figureCitation box="[385,436,1202,1228]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="19" pageNumber="83">14B</figureCitation>
).Vesicle with a well-developed ventromedian carina, ventral surface covered by several macrosetae and subaculear setal pair significantly evident; one or two small subaculear tuberance present on terminal of ventromedian carina; base of aculeus moderately wide and shorter than half of vesicle; aculeus with several microsetae proximally and progressively darker towards distal.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="83" type="discussion">
<paragraph blockId="19.[136,779,158,1913]" pageId="19" pageNumber="83">
<emphasis bold="true" box="[189,305,1490,1516]" pageId="19" pageNumber="83">Remarks.</emphasis>
Lourenço &amp; Velten (2016) regarded the serrated dorsal carinae of metasoma I–IV as a diagnostic character of
<taxonomicName box="[300,643,1562,1588]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="83" phylum="Arthropoda" rank="species" species="fleissneri">
<emphasis box="[300,643,1562,1588]" italics="true" pageId="19" pageNumber="83">Betaburmesebuthus fleissneri</emphasis>
</taxonomicName>
. However, we noticed that this character is shared by all members of
<taxonomicName box="[170,391,1634,1660]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis box="[170,391,1634,1660]" italics="true" pageId="19" pageNumber="83">Betaburmesebuthus</emphasis>
</taxonomicName>
and especially evident in mature individuals (
<figureCitation box="[276,409,1670,1697]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="19" pageNumber="83">Figs 13C, J</figureCitation>
,
<figureCitation box="[422,472,1670,1696]" captionStart="FIGURE 24" captionStartId="30.[136,235,1951,1975]" captionTargetBox="[211,1377,189,1928]" captionTargetId="figure-18@30.[211,1377,189,1928]" captionTargetPageId="30" captionText="FIGURE 24. Details of Betaburmesebuthus villosus sp. nov., holotype, NIGP200652, under green epifluorescence. A, Carapace. B, tergites. C, Sternum and genital operculum. D, Pectines. E, Tergite IV–VI. F, Femur, dorsal view. G, Patella, dorsal view. H, Metasomal segments III–IV, Lateral view. I, Left leg III. J, Right chela. Scale bars = 400 μm in A, B, D–H, J; 200 μm in C, I." figureDoi="http://doi.org/10.5281/zenodo.7754878" httpUri="https://zenodo.org/record/7754878/files/figure.png" pageId="19" pageNumber="83">24H</figureCitation>
,
<figureCitation box="[485,536,1670,1696]" captionStart="FIGURE 29" captionStartId="34.[136,235,1908,1932]" captionTargetBox="[183,1404,189,1884]" captionTargetId="figure-18@34.[183,1404,189,1884]" captionTargetPageId="34" captionText="FIGURE 29. Details of Betaburmesebuthus fuscus sp. nov., holotype, NIGP200653, under incident light. A, Left pedipalp and carapace. B, Pectines. C, Sternites VI–VII and metasomal segments I–III. D, Femur, dorsal view. E, Patella, dorsal view. F, Tarsus of leg IV. G, Tarsus of leg I. H, Tarsus of leg III. I, Tarsus of leg II. J, Spiracle.Abbreviations: sp, spiracle; ts, tibial spur. Scale bars = 1 mm in A, C; 500 μm in B, F–J; 400 μm in D, E." figureDoi="http://doi.org/10.5281/zenodo.7754892" httpUri="https://zenodo.org/record/7754892/files/figure.png" pageId="19" pageNumber="83">29C</figureCitation>
). Our adult specimen provides more reliable features than the immature
<typeStatus box="[136,238,1742,1768]" pageId="19" pageNumber="83">holotype</typeStatus>
.
</paragraph>
<paragraph blockId="19.[136,779,158,1913]" pageId="19" pageNumber="83">
<materialsCitation location="We" municipality="Velten" pageId="19" pageNumber="83" specimenCount="1" typeStatus="holotype">
<location LSID="urn:lsid:plazi:treatment:F308E17F435FFFF0FF6F486DFD5C1E56:7E7E06B24343FFF0FF5A48F4FF1C1E23" box="[189,226,1779,1805]" municipality="Velten" name="We" pageId="19" pageNumber="83">We</location>
remeasured the ratio of chela length and width from figure
<locationDeviation box="[275,436,1814,1841]" location="Lourenco" pageId="19" pageNumber="83">
<quantity box="[275,320,1814,1840]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="19" pageNumber="83" unit="in" value="2.0">2 in</quantity>
Lourenço
</locationDeviation>
&amp;
<collectingMunicipality box="[475,547,1814,1840]" pageId="19" pageNumber="83">Velten</collectingMunicipality>
(2016) and found it is about
<quantity box="[234,314,1850,1877]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" pageId="19" pageNumber="83" unit="in" value="7.5">7.50 in</quantity>
<typeStatus box="[322,420,1850,1876]" pageId="19" pageNumber="83">holotype</typeStatus>
rather than
<quantity box="[560,639,1850,1876]" metricMagnitude="-1" metricUnit="m" metricValue="1.73228" pageId="19" pageNumber="83" unit="in" value="6.82">6.82 in</quantity>
the original paper, close to the value in our specimen (7.67)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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