856 lines
134 KiB
XML
856 lines
134 KiB
XML
<document ID-DOI="10.11646/palaeoentomology.6.1.10" ID-ISSN="2624-2834" ID-Zenodo-Dep="7754820" ID-ZooBank="B822ED53-D559-4DD8-8803-3F2850A00445" approvalRequired="1" approvalRequired_for_taxonomicNames="1" checkinTime="1679380739061" checkinUser="plazi" docAuthor="XUAN, QIANG, CAI, CHEN-YANG & HUANG, DI-YING" docDate="2023" docId="F308E17F4351FFEBFCCD48A1FBE41C3A" docLanguage="en" docName="palaeoentomology.6.1.10.pdf" docOrigin="Palaeoentomology 6 (1)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.1.10" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.3:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="3" docTitle="Betaburmesebuthus Lourenco 2015" docType="treatment" docVersion="3" lastPageNumber="72" masterDocId="0F3199074350FFE3FFE74E07FFFE192E" masterDocTitle="Revision of palaeoburmesebuthid scorpions in mid-Cretaceous amber from northern Myanmar (Scorpiones: Buthoidea)" masterLastPageNumber="101" masterPageNumber="64" pageNumber="65" updateTime="1679418454157" updateUser="tatiana">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of palaeoburmesebuthid scorpions in mid-Cretaceous amber from northern Myanmar (Scorpiones: Buthoidea)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>XUAN, QIANG</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>CAI, CHEN-YANG</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>HUANG, DI-YING</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-28</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>6</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>64</mods:start>
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<mods:end>101</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/palaeoentomology.6.1.10</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/palaeoentomology.6.1.10</mods:identifier>
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<mods:identifier type="ISSN">2624-2834</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7754820</mods:identifier>
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<mods:identifier type="ZooBank">B822ED53-D559-4DD8-8803-3F2850A00445</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:F308E17F4351FFEBFCCD48A1FBE41C3A" httpUri="http://treatment.plazi.org/id/F308E17F4351FFEBFCCD48A1FBE41C3A" lastPageId="8" lastPageNumber="72" pageId="1" pageNumber="65">
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<subSubSection box="[810,1315,1701,1728]" pageId="1" pageNumber="65" type="nomenclature">
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||
<paragraph blockId="1.[810,1315,1701,1728]" box="[810,1315,1701,1728]" pageId="1" pageNumber="65">
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||
<heading box="[810,1315,1701,1728]" fontSize="10" level="2" pageId="1" pageNumber="65" reason="2">
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||
<emphasis bold="true" box="[810,1315,1701,1728]" pageId="1" pageNumber="65">
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Genus
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<taxonomicName authority="Lourenco, 2015" authorityName="Lourenco" authorityYear="2015" box="[893,1315,1701,1728]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[893,1123,1701,1728]" italics="true" pageId="1" pageNumber="65">Betaburmesebuthus</emphasis>
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Lourenço, 2015
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection lastPageId="3" lastPageNumber="67" pageId="1" pageNumber="65" type="type_taxon">
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||
<paragraph blockId="1.[810,1451,1775,2093]" pageId="1" pageNumber="65">
|
||
<emphasis bold="true" box="[810,974,1775,1801]" pageId="1" pageNumber="65">
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||
<typeStatus box="[810,869,1775,1801]" pageId="1" pageNumber="65">Type</typeStatus>
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species.
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||
</emphasis>
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<taxonomicName authority="Lourenco, 2015" authorityName="Lourenco & Beigel" authorityYear="2015" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="kobberti">
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<emphasis box="[990,1320,1775,1801]" italics="true" pageId="1" pageNumber="65">Betaburmesebuthus kobberti</emphasis>
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Lourenço, 2015
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</taxonomicName>
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||
(
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||
<emphasis box="[882,904,1812,1837]" italics="true" pageId="1" pageNumber="65">in</emphasis>
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||
<bibRefCitation author="Lourenco, W. R. & Beigel, A." box="[911,1202,1811,1838]" pageId="1" pageNumber="65" pagination="476 - 480" refId="ref13343" refString="Lourenco, W. R. & Beigel, A. (2015) A new genus and species of Palaeoburmesebuthinae Lourenco, 2014 (Scorpiones: Archaeobuthidae:) from Cretaceous amber of Myanmar. Beitrage zur Araneologie, 9, 476 - 480." type="journal article" year="2015">Lourenço & Beigel, 2015</bibRefCitation>
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).
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||
</paragraph>
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<paragraph blockId="1.[810,1451,1775,2093]" lastBlockId="3.[136,778,1904,2093]" lastPageId="3" lastPageNumber="67" pageId="1" pageNumber="65">
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||
<emphasis bold="true" box="[863,1085,1847,1874]" pageId="1" pageNumber="65">Included species.</emphasis>
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<taxonomicName authority="(Lourenco & Beigel, 2015)" baseAuthorityName="Lourenco & Beigel" baseAuthorityYear="2015" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="kobberti">
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<emphasis box="[1111,1451,1848,1874]" italics="true" pageId="1" pageNumber="65">Betaburmesebuthus kobberti</emphasis>
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(
|
||
<bibRefCitation author="Lourenco, W. R. & Beigel, A." box="[817,1143,1884,1911]" pageId="1" pageNumber="65" pagination="476 - 480" refId="ref13343" refString="Lourenco, W. R. & Beigel, A. (2015) A new genus and species of Palaeoburmesebuthinae Lourenco, 2014 (Scorpiones: Archaeobuthidae:) from Cretaceous amber of Myanmar. Beitrage zur Araneologie, 9, 476 - 480." type="journal article" year="2015">Lourenço & Beigel, 2015</bibRefCitation>
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||
)
|
||
</taxonomicName>
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||
,
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<taxonomicName authority="(Lourenco, 2015 a)" baseAuthorityName="Lourenco" baseAuthorityYear="2015" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="muelleri">
|
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<emphasis box="[1174,1191,1885,1910]" italics="true" pageId="1" pageNumber="65">B</emphasis>
|
||
.
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||
<emphasis box="[1216,1309,1884,1910]" italics="true" pageId="1" pageNumber="65">muelleri</emphasis>
|
||
(
|
||
<bibRefCitation author="Lourenco, W. R." pageId="1" pageNumber="65" pagination="27 - 36" refId="ref13090" refString="Lourenco, W. R. (2015 a) A new contribution to the knowledge of Cretaceous Burmese amber scorpions with the description of two new species of Betaburmesebuthus Lourenco, 2015 (Scorpiones: Archaeobuthidae: Palaeoburmesebuthinae). Arachnida, Rivista Aracnologica Italiana, 1 (3), 27 - 36." type="journal article" year="2015">Lourenço, 2015a</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
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||
<taxonomicName authority="(Lourenco, 2015 a)" baseAuthorityName="Lourenco" baseAuthorityYear="2015" box="[914,1317,1920,1947]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis box="[914,931,1921,1946]" italics="true" pageId="1" pageNumber="65">B</emphasis>
|
||
.
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||
<emphasis box="[957,1074,1921,1947]" italics="true" pageId="1" pageNumber="65">bidentatus</emphasis>
|
||
(
|
||
<bibRefCitation author="Lourenco, W. R." box="[1101,1309,1920,1947]" pageId="1" pageNumber="65" pagination="27 - 36" refId="ref13090" refString="Lourenco, W. R. (2015 a) A new contribution to the knowledge of Cretaceous Burmese amber scorpions with the description of two new species of Betaburmesebuthus Lourenco, 2015 (Scorpiones: Archaeobuthidae: Palaeoburmesebuthinae). Arachnida, Rivista Aracnologica Italiana, 1 (3), 27 - 36." type="journal article" year="2015">Lourenço, 2015a</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
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||
<taxonomicName authority="(Lourenco, 2016)" baseAuthorityName="Lourenco" baseAuthorityYear="2016" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="bellus">
|
||
<emphasis box="[1341,1358,1921,1946]" italics="true" pageId="1" pageNumber="65">B</emphasis>
|
||
.
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||
<emphasis box="[1384,1451,1921,1947]" italics="true" pageId="1" pageNumber="65">bellus</emphasis>
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||
(Lourenço, 2016)
|
||
</taxonomicName>
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||
,
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||
<taxonomicName authority="(Lourenco & Velten, 2016 a)" baseAuthorityName="Lourenco & Velten" baseAuthorityYear="2016" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="fleissneri">
|
||
<emphasis box="[1040,1057,1958,1983]" italics="true" pageId="1" pageNumber="65">B</emphasis>
|
||
.
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||
<emphasis box="[1081,1184,1957,1983]" italics="true" pageId="1" pageNumber="65">fleissneri</emphasis>
|
||
(
|
||
<bibRefCitation author="Lourenco, W. R. & Velten, J." pageId="1" pageNumber="65" pagination="4 - 11" refId="ref13434" refString="Lourenco, W. R. & Velten, J. (2016 a) One more new species of Betaburmesebuthus Lourenco, 2015 (Scorpiones: Palaeoburmesebuthinae) from Cretaceous burmite. Arachnida, Rivista Aracnologica Italiana, 6, 4 - 11." type="journal article" year="2016">Lourenço & Velten, 2016a</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(Lourenco & Velten, 2016 b)" baseAuthorityName="Lourenco & Velten" baseAuthorityYear="2016" box="[905,1447,1993,2020]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="larafleissnerae">
|
||
<emphasis box="[905,922,1994,2019]" italics="true" pageId="1" pageNumber="65">B</emphasis>
|
||
.
|
||
<emphasis box="[939,1107,1993,2019]" italics="true" pageId="1" pageNumber="65">larafleissnerae</emphasis>
|
||
(
|
||
<bibRefCitation author="Lourenco, W. R. & Velten, J." box="[1125,1439,1993,2020]" pageId="1" pageNumber="65" pagination="10 - 17" refId="ref13479" refString="Lourenco, W. R. & Velten, J. (2016 b) A sixth new species of Cretaceous Burmese amber scorpion of the genus Betaburmesebuthus Lourenco, 2015 (Scorpiones: Palaeoburmesebuthidae). Arachnida, Rivista Aracnologica Italiana, 10, 10 - 17." type="journal article" year="2016">Lourenço & Velten, 2016b</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(Lourenco & Rossi, 2017)" baseAuthorityName="Lourenco & Rossi" baseAuthorityYear="2017" box="[810,1211,2030,2056]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="joergi">
|
||
<emphasis box="[810,827,2031,2056]" italics="true" pageId="1" pageNumber="65">B</emphasis>
|
||
.
|
||
<emphasis box="[841,907,2031,2056]" italics="true" pageId="1" pageNumber="65">joergi</emphasis>
|
||
(
|
||
<bibRefCitation author="Lourenco, W. R. & Rossi, A." box="[921,1203,2030,2056]" pageId="1" pageNumber="65" pagination="11 - 21" refId="ref13389" refString="Lourenco, W. R. & Rossi, A. (2017) Two more new species of Burmese amber scorpions of the family Palaeoburmesebuthidae Lourenco, 2015 (Scorpiones). Arachnida, Rivista Aracnologica Italiana, 13, 11 - 21." type="journal article" year="2017">Lourenço & Rossi, 2017</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(Xuan, Cai & Huang, 2022)" baseAuthorityName="Xuan, Cai & Huang" baseAuthorityYear="2022" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="spinipedis">
|
||
<emphasis box="[1222,1239,2031,2056]" italics="true" pageId="1" pageNumber="65">B</emphasis>
|
||
.
|
||
<emphasis box="[1254,1368,2030,2056]" italics="true" pageId="1" pageNumber="65">spinipedis</emphasis>
|
||
(
|
||
<bibRefCitation author="Xuan, Q. & Cai, C. Y. & Huang, D. Y." pageId="1" pageNumber="65" pagination="105165" refId="ref14060" refString="Xuan, Q., Cai, C. Y. & Huang, D. Y. (2022) A new Palaeoburmesebuthidae scorpion from mid-Cretaceous Burmese amber (Arachnida: Scorpiones: Buthoidea). Cretaceous Research, 134, 105165. https: // doi. org / 10.1016 / j. cretres. 2022.105165" type="journal article" year="2022">Xuan, Cai & Huang, 2022</bibRefCitation>
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||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="XUAN & CAI & HUANG, 2023" authorityName="XUAN & CAI & HUANG" authorityYear="2023" box="[1064,1185,2066,2092]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="villosus" status="sp. nov.">
|
||
<emphasis box="[1064,1185,2066,2092]" italics="true" pageId="1" pageNumber="65">B. villosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1194,1287,2066,2092]" pageId="1" pageNumber="65">
|
||
<taxonomicNameLabel box="[1194,1287,2066,2092]" pageId="1" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName authority="XUAN & CAI & HUANG, 2023" authorityName="XUAN & CAI & HUANG" authorityYear="2023" box="[1304,1408,2066,2093]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="fuscus" status="sp. nov.">
|
||
<emphasis box="[1304,1321,2067,2092]" italics="true" pageId="1" pageNumber="65">B</emphasis>
|
||
.
|
||
<emphasis box="[1338,1408,2066,2092]" italics="true" pageId="1" pageNumber="65">fuscus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" lastPageId="3" lastPageNumber="67" pageId="1" pageNumber="65">
|
||
<taxonomicNameLabel lastPageId="3" lastPageNumber="67" pageId="1" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName authority="(Lourenco & Velten, 2017)" authorityName="XUAN & CAI & HUANG" authorityYear="2023" baseAuthorityName="Lourenco & Velten" baseAuthorityYear="2017" box="[199,596,1904,1931]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="pohli" status="comb. nov.">
|
||
<emphasis box="[199,216,1905,1930]" italics="true" pageId="3" pageNumber="67">B</emphasis>
|
||
.
|
||
<emphasis box="[230,288,1904,1930]" italics="true" pageId="3" pageNumber="67">pohli</emphasis>
|
||
(
|
||
<bibRefCitation author="Lourenco, W. R. & Velten, J." box="[301,588,1904,1930]" pageId="3" pageNumber="67" pagination="2 - 10" refId="ref13575" refString="Lourenco, W. R. & Velten, J. (2017) One more new genus and species of fossil scorpion from Burmese Cretaceous amber belonging to the family Palaeoburmesebuthidae (Scorpiones). Arachnida, Rivista Aracnologica Italiana, 13, 2 - 10." type="journal article" year="2017">Lourenço & Velten, 2017</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[602,731,1904,1930]" pageId="3" pageNumber="67">
|
||
<taxonomicNameLabel box="[602,731,1904,1930]" pageId="3" pageNumber="67" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName authority="(Lourenco, 2021)" authorityName="XUAN & CAI & HUANG" authorityYear="2023" baseAuthorityName="Lourenco" baseAuthorityYear="2021" box="[136,504,1945,1971]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="knodelorum" status="comb. nov.">
|
||
<emphasis box="[136,153,1946,1971]" italics="true" pageId="3" pageNumber="67">B</emphasis>
|
||
.
|
||
<emphasis box="[167,301,1945,1971]" italics="true" pageId="3" pageNumber="67">knodelorum</emphasis>
|
||
(
|
||
<bibRefCitation author="Lourenco, W. R." box="[316,496,1945,1971]" pageId="3" pageNumber="67" pagination="1 - 6" refId="ref13280" refString="Lourenco, W. R. (2021) Further comments on the elements of the family Palaeoburmesebuthidae Lourenco, 2015 with description of a new species of Spinoburmesebuthus Lourenco, 2017 from Early Cretaceous Burmite amber (Scorpiones). Faunitaxys, 9 (17), 1 - 6. https: // hal. science / hal- 03358766 v 2" type="journal article" year="2021">Lourenço, 2021</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[511,639,1945,1971]" pageId="3" pageNumber="67">
|
||
<taxonomicNameLabel box="[511,639,1945,1971]" pageId="3" pageNumber="67" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7754826" ID-Zenodo-Dep="7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="2" pageNumber="66" startId="2.[136,235,1854,1878]" targetBox="[164,1423,189,1830]" targetPageId="2">
|
||
<paragraph blockId="2.[136,1452,1854,2058]" pageId="2" pageNumber="66">
|
||
<emphasis bold="true" box="[136,259,1854,1878]" pageId="2" pageNumber="66">FIGURE 2.</emphasis>
|
||
Details of
|
||
<taxonomicName authority=", NIGP" authorityName="NIGP" box="[367,705,1854,1878]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="bellus">
|
||
<emphasis box="[367,633,1854,1878]" italics="true" pageId="2" pageNumber="66">Betaburmesebuthus bellus</emphasis>
|
||
, NIGP
|
||
</taxonomicName>
|
||
200648, under confocal microscopy with depth color-coding.
|
||
<emphasis bold="true" box="[1321,1339,1854,1878]" pageId="2" pageNumber="66">A</emphasis>
|
||
, Carapace.
|
||
<emphasis bold="true" box="[136,153,1890,1914]" pageId="2" pageNumber="66">B</emphasis>
|
||
, Lateral ocelli.
|
||
<emphasis bold="true" box="[314,332,1890,1914]" pageId="2" pageNumber="66">C</emphasis>
|
||
, Coxapophysis, coxa and trochanter of leg I.
|
||
<emphasis bold="true" box="[797,815,1890,1914]" pageId="2" pageNumber="66">D</emphasis>
|
||
, Sternum and genital operculum.
|
||
<emphasis bold="true" box="[1161,1178,1890,1914]" pageId="2" pageNumber="66">E</emphasis>
|
||
, Spiracle of sternite VI.
|
||
<emphasis bold="true" box="[1430,1445,1890,1914]" pageId="2" pageNumber="66">F</emphasis>
|
||
, Tergites I–VI.
|
||
<emphasis bold="true" box="[283,303,1926,1950]" pageId="2" pageNumber="66">G</emphasis>
|
||
, Dentition of movable finger.
|
||
<emphasis bold="true" box="[608,628,1926,1950]" pageId="2" pageNumber="66">H</emphasis>
|
||
, Dentition of fixed finger.
|
||
<emphasis bold="true" box="[899,909,1926,1950]" pageId="2" pageNumber="66">I</emphasis>
|
||
, Tibial spur of leg III.
|
||
<emphasis bold="true" box="[1137,1150,1926,1950]" pageId="2" pageNumber="66">J</emphasis>
|
||
, Tibial spur of leg IV.
|
||
<emphasis bold="true" box="[1377,1397,1926,1950]" pageId="2" pageNumber="66">K</emphasis>
|
||
, Left pectine.
|
||
<emphasis bold="true" box="[221,270,1962,1986]" pageId="2" pageNumber="66">L–O</emphasis>
|
||
, Telotarsi of legs I–IV. Abbreviations: ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in
|
||
<emphasis bold="true" box="[376,394,2034,2058]" pageId="2" pageNumber="66">A</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[407,422,2034,2058]" pageId="2" pageNumber="66">F</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[436,456,2034,2058]" pageId="2" pageNumber="66">G</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[468,488,2034,2058]" pageId="2" pageNumber="66">K</emphasis>
|
||
; 100 μm in
|
||
<emphasis bold="true" box="[611,629,2034,2058]" pageId="2" pageNumber="66">C</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[642,660,2034,2058]" pageId="2" pageNumber="66">D</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[673,690,2034,2058]" pageId="2" pageNumber="66">E</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[702,722,2034,2058]" pageId="2" pageNumber="66">H</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[735,745,2034,2058]" pageId="2" pageNumber="66">I</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[757,770,2034,2058]" pageId="2" pageNumber="66">J</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[783,800,2034,2058]" pageId="2" pageNumber="66">L</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[812,836,2034,2058]" pageId="2" pageNumber="66">M</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[849,867,2034,2058]" pageId="2" pageNumber="66">N</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[880,900,2034,2058]" pageId="2" pageNumber="66">O</emphasis>
|
||
; 50 μm in
|
||
<emphasis bold="true" box="[1010,1027,2034,2058]" pageId="2" pageNumber="66">B</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7754828" ID-Zenodo-Dep="7754828" httpUri="https://zenodo.org/record/7754828/files/figure.png" pageId="3" pageNumber="67" startId="3.[136,235,1712,1736]" targetBox="[151,1436,189,1689]" targetPageId="3">
|
||
<paragraph blockId="3.[136,1452,1712,1845]" pageId="3" pageNumber="67">
|
||
<emphasis bold="true" box="[136,259,1712,1737]" pageId="3" pageNumber="67">FIGURE 3.</emphasis>
|
||
Trichobothrial pattern of
|
||
<taxonomicName authority=", NIGP" authorityName="NIGP" box="[519,858,1713,1737]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="bellus">
|
||
<emphasis box="[519,785,1713,1737]" italics="true" pageId="3" pageNumber="67">Betaburmesebuthus bellus</emphasis>
|
||
, NIGP
|
||
</taxonomicName>
|
||
200648, under confocal microscopy (
|
||
<emphasis bold="true" box="[1233,1250,1713,1737]" pageId="3" pageNumber="67">B</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1262,1280,1712,1736]" pageId="3" pageNumber="67">C</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1292,1309,1713,1737]" pageId="3" pageNumber="67">E</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1321,1336,1713,1737]" pageId="3" pageNumber="67">F</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1349,1401,1713,1737]" pageId="3" pageNumber="67">H–K</emphasis>
|
||
) and green epifluorescence (
|
||
<emphasis bold="true" box="[370,388,1748,1772]" pageId="3" pageNumber="67">A</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[401,419,1749,1773]" pageId="3" pageNumber="67">D</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[431,451,1748,1772]" pageId="3" pageNumber="67">G</emphasis>
|
||
).
|
||
<emphasis bold="true" box="[470,488,1748,1772]" pageId="3" pageNumber="67">A</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[500,517,1749,1773]" pageId="3" pageNumber="67">B</emphasis>
|
||
, Left femur, dorsal view.
|
||
<emphasis bold="true" box="[775,793,1748,1772]" pageId="3" pageNumber="67">C</emphasis>
|
||
, Right femur, dorsal view.
|
||
<emphasis bold="true" box="[1064,1082,1749,1773]" pageId="3" pageNumber="67">D</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1094,1111,1749,1773]" pageId="3" pageNumber="67">E</emphasis>
|
||
, left patella, dorsal view.
|
||
<emphasis bold="true" box="[1368,1383,1749,1773]" pageId="3" pageNumber="67">F</emphasis>
|
||
, Right patella, dorsal view.
|
||
<emphasis bold="true" box="[341,361,1784,1808]" pageId="3" pageNumber="67">G</emphasis>
|
||
, Left chela, dorsal view.
|
||
<emphasis bold="true" box="[611,631,1785,1809]" pageId="3" pageNumber="67">H</emphasis>
|
||
, Left fixed finger, dorsal view.
|
||
<emphasis bold="true" box="[945,955,1785,1809]" pageId="3" pageNumber="67">I</emphasis>
|
||
, Right manus, ventral view.
|
||
<emphasis bold="true" box="[1240,1253,1785,1809]" pageId="3" pageNumber="67">J</emphasis>
|
||
, Left manus, dorsal view.
|
||
<emphasis bold="true" box="[196,216,1821,1845]" pageId="3" pageNumber="67">K</emphasis>
|
||
, Left manus, ventral view. Scale bars = 200 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="8" lastPageNumber="72" pageId="3" pageNumber="67" type="diagnosis">
|
||
<paragraph blockId="3.[136,778,1904,2093]" lastBlockId="8.[136,778,158,2092]" lastPageId="8" lastPageNumber="72" pageId="3" pageNumber="67">
|
||
<emphasis bold="true" box="[189,451,1985,2011]" pageId="3" pageNumber="67">Diagnosis (emended).</emphasis>
|
||
Five pairs of lateral ocelli present, including three major lateral ocelli (ALMa, MLMa, PLMa) and two minor lateral ocelli (PDMi,
|
||
<materialsCitation collectionCode="III, IV" lastPageId="8" lastPageNumber="72" pageId="3" pageNumber="67" specimenCount="1">
|
||
ADMi) (
|
||
<figureCitation box="[906,996,1904,1930]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="3" pageNumber="67">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation box="[1006,1041,1904,1930]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="3" pageNumber="67">6B</figureCitation>
|
||
,
|
||
<figureCitation box="[1051,1101,1904,1930]" captionStart="FIGURE 10" captionStartId="14.[136,235,1845,1869]" captionTargetBox="[224,1363,189,1822]" captionTargetId="figure-18@14.[224,1363,189,1822]" captionTargetPageId="14" captionText="FIGURE 10. Details of Betaburmesebuthus kobberti, NIGP200650, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Coxapophyses I–II. D, Tergites I–VI. E, Sternites IV–VII. F, Pectine. G, Tergite VII and Metasomal segments I–II. H, Telson. I, Tibial spur of leg II. J, Tibial spur of leg IV. K, Telotarsi of leg III. L, Telotarsi of leg IV. Abbreviations: ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, D, E, F, G, H; 100 μm in C, I, J, K, L; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754844" httpUri="https://zenodo.org/record/7754844/files/figure.png" pageId="3" pageNumber="67">10B</figureCitation>
|
||
,
|
||
<figureCitation box="[1110,1159,1904,1930]" captionStart="FIGURE 17" captionStartId="22.[136,235,1829,1853]" captionTargetBox="[239,1349,189,1806]" captionTargetId="figure-18@22.[239,1349,189,1806]" captionTargetPageId="22" captionText="FIGURE 17. Details of Betaburmesebuthus larafleissnerae, NIGP201155, A, C–G under green epifluorescence; B under confocal microscopy; H, I under incident light. A, Carapace and chelicerae, dorsal view. B, Lateral ocelli. C, Tarsus of right leg III. D, Tergites. E, Metasomal segments I–IV and telson, lateral view. F, Metasomal segments I–III, ventral view. G, Metasomal segments IV–V and telson, lateral view. H, Pectines. I, Sternite V with right spiracle. Abbreviations: ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 400 μm in A, D, E, F, G, H; 200 μm in C, I; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754864" httpUri="https://zenodo.org/record/7754864/files/figure.png" pageId="3" pageNumber="67">17B</figureCitation>
|
||
,
|
||
<figureCitation box="[1170,1219,1904,1930]" captionStart="FIGURE 20" captionStartId="25.[136,235,1836,1860]" captionTargetBox="[209,1378,189,1813]" captionTargetId="figure-28@25.[209,1378,189,1813]" captionTargetPageId="25" captionText="FIGURE 20. Details of Betaburmesebuthus muelleri NIGP201156, A, D, E, F under green epifluorescence; B, C, F, G, I–M under confocal microscopy. A, Carapace and tergites. B, Chelicerae, dorsal view. C, Lateral ocelli. D, Tergite VII and metasomal segments I–IV, dorsal view. E, Coxosternal region, pectines and sternites. F, Sternum and genital operculum. G, Pectine. H, Tergite VII and metasomal segments I–IV, ventral view. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Tarsus of leg I. M, Tarsus of leg IV.Abbreviations:ADMi, anterodorsal minor ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 400 μm in A, D, E, H; 100 μm in B, F, G, I–M; 50 μm in C." figureDoi="http://doi.org/10.5281/zenodo.7754870" httpUri="https://zenodo.org/record/7754870/files/figure.png" pageId="3" pageNumber="67">20C</figureCitation>
|
||
,
|
||
<figureCitation box="[1229,1280,1904,1931]" captionStart="FIGURE 25" captionStartId="31.[136,235,1885,1909]" captionTargetBox="[164,1423,189,1862]" captionTargetId="figure-28@31.[164,1423,189,1862]" captionTargetPageId="31" captionText="FIGURE 25. Details of Betaburmesebuthus villosus sp. nov., holotype, NIGP200652, under confocal microscopy. A, Lateral ocelli. B, Femur, dorsal view. C, Spiracle. D, Tibial spur of leg III. E, Tibial spur of leg IV. F, Right chela manus, ventral view. G, Left chela manus, ventral view. H, Tarsus of leg III. I, Tarsus of leg IV. J–K, Telson, lateral view. Abbreviations: ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; st, subaculear tubercle; ts, tibial spur. Scale bars = 100 μm in A–C, K; 200 μm in D–J." figureDoi="http://doi.org/10.5281/zenodo.7754880" httpUri="https://zenodo.org/record/7754880/files/figure.png" pageId="3" pageNumber="67">25A</figureCitation>
|
||
,
|
||
<figureCitation box="[1289,1340,1904,1930]" captionStart="FIGURE 28" captionStartId="33.[136,235,1950,1974]" captionTargetBox="[183,1404,1048,1926]" captionTargetId="figure-59@33.[183,1404,1048,1926]" captionTargetPageId="33" captionText="FIGURE 28. Micro-CT reconstruction of Betaburmesebuthus fuscus sp. nov., holotype, NIGP200653. A, Dorsal view. B, Ventral view. C, Lateral ocelli.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus. Scale bars = 3 mm in A, B; 500 μm in C." figureDoi="http://doi.org/10.5281/zenodo.7754888" httpUri="https://zenodo.org/record/7754888/files/figure.png" pageId="3" pageNumber="67">28C</figureCitation>
|
||
).Anterior margin of carapace with a wide weakly to strongly marked median concavity (
|
||
<figureCitation box="[1031,1128,1985,2011]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="3" pageNumber="67">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation box="[1145,1182,1985,2011]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="3" pageNumber="67">6A</figureCitation>
|
||
,
|
||
<figureCitation box="[1197,1247,1985,2011]" captionStart="FIGURE 10" captionStartId="14.[136,235,1845,1869]" captionTargetBox="[224,1363,189,1822]" captionTargetId="figure-18@14.[224,1363,189,1822]" captionTargetPageId="14" captionText="FIGURE 10. Details of Betaburmesebuthus kobberti, NIGP200650, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Coxapophyses I–II. D, Tergites I–VI. E, Sternites IV–VII. F, Pectine. G, Tergite VII and Metasomal segments I–II. H, Telson. I, Tibial spur of leg II. J, Tibial spur of leg IV. K, Telotarsi of leg III. L, Telotarsi of leg IV. Abbreviations: ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, D, E, F, G, H; 100 μm in C, I, J, K, L; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754844" httpUri="https://zenodo.org/record/7754844/files/figure.png" pageId="3" pageNumber="67">10A</figureCitation>
|
||
,
|
||
<figureCitation box="[1263,1314,1985,2011]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="3" pageNumber="67">14A</figureCitation>
|
||
,
|
||
<figureCitation box="[1330,1381,1985,2011]" captionStart="FIGURE 17" captionStartId="22.[136,235,1829,1853]" captionTargetBox="[239,1349,189,1806]" captionTargetId="figure-18@22.[239,1349,189,1806]" captionTargetPageId="22" captionText="FIGURE 17. Details of Betaburmesebuthus larafleissnerae, NIGP201155, A, C–G under green epifluorescence; B under confocal microscopy; H, I under incident light. A, Carapace and chelicerae, dorsal view. B, Lateral ocelli. C, Tarsus of right leg III. D, Tergites. E, Metasomal segments I–IV and telson, lateral view. F, Metasomal segments I–III, ventral view. G, Metasomal segments IV–V and telson, lateral view. H, Pectines. I, Sternite V with right spiracle. Abbreviations: ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 400 μm in A, D, E, F, G, H; 200 μm in C, I; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754864" httpUri="https://zenodo.org/record/7754864/files/figure.png" pageId="3" pageNumber="67">17A</figureCitation>
|
||
,
|
||
<figureCitation box="[1396,1447,1985,2011]" captionStart="FIGURE 20" captionStartId="25.[136,235,1836,1860]" captionTargetBox="[209,1378,189,1813]" captionTargetId="figure-28@25.[209,1378,189,1813]" captionTargetPageId="25" captionText="FIGURE 20. Details of Betaburmesebuthus muelleri NIGP201156, A, D, E, F under green epifluorescence; B, C, F, G, I–M under confocal microscopy. A, Carapace and tergites. B, Chelicerae, dorsal view. C, Lateral ocelli. D, Tergite VII and metasomal segments I–IV, dorsal view. E, Coxosternal region, pectines and sternites. F, Sternum and genital operculum. G, Pectine. H, Tergite VII and metasomal segments I–IV, ventral view. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Tarsus of leg I. M, Tarsus of leg IV.Abbreviations:ADMi, anterodorsal minor ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 400 μm in A, D, E, H; 100 μm in B, F, G, I–M; 50 μm in C." figureDoi="http://doi.org/10.5281/zenodo.7754870" httpUri="https://zenodo.org/record/7754870/files/figure.png" pageId="3" pageNumber="67">20A</figureCitation>
|
||
,
|
||
<figureCitation box="[810,861,2026,2052]" captionStart="FIGURE 24" captionStartId="30.[136,235,1951,1975]" captionTargetBox="[211,1377,189,1928]" captionTargetId="figure-18@30.[211,1377,189,1928]" captionTargetPageId="30" captionText="FIGURE 24. Details of Betaburmesebuthus villosus sp. nov., holotype, NIGP200652, under green epifluorescence. A, Carapace. B, tergites. C, Sternum and genital operculum. D, Pectines. E, Tergite IV–VI. F, Femur, dorsal view. G, Patella, dorsal view. H, Metasomal segments III–IV, Lateral view. I, Left leg III. J, Right chela. Scale bars = 400 μm in A, B, D–H, J; 200 μm in C, I." figureDoi="http://doi.org/10.5281/zenodo.7754878" httpUri="https://zenodo.org/record/7754878/files/figure.png" pageId="3" pageNumber="67">24A</figureCitation>
|
||
,
|
||
<figureCitation box="[870,922,2026,2052]" captionStart="FIGURE 29" captionStartId="34.[136,235,1908,1932]" captionTargetBox="[183,1404,189,1884]" captionTargetId="figure-18@34.[183,1404,189,1884]" captionTargetPageId="34" captionText="FIGURE 29. Details of Betaburmesebuthus fuscus sp. nov., holotype, NIGP200653, under incident light. A, Left pedipalp and carapace. B, Pectines. C, Sternites VI–VII and metasomal segments I–III. D, Femur, dorsal view. E, Patella, dorsal view. F, Tarsus of leg IV. G, Tarsus of leg I. H, Tarsus of leg III. I, Tarsus of leg II. J, Spiracle.Abbreviations: sp, spiracle; ts, tibial spur. Scale bars = 1 mm in A, C; 500 μm in B, F–J; 400 μm in D, E." figureDoi="http://doi.org/10.5281/zenodo.7754892" httpUri="https://zenodo.org/record/7754892/files/figure.png" pageId="3" pageNumber="67">29A</figureCitation>
|
||
); central median, posterior median and posterior marginal furrows of carapace well-marked in adults (
|
||
<figureCitation captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" lastPageId="6" lastPageNumber="70" pageId="3" pageNumber="67">Figs 13A</figureCitation>
|
||
,
|
||
<figureCitation box="[205,255,1922,1948]" captionStart="FIGURE 28" captionStartId="33.[136,235,1950,1974]" captionTargetBox="[183,1404,1048,1926]" captionTargetId="figure-59@33.[183,1404,1048,1926]" captionTargetPageId="33" captionText="FIGURE 28. Micro-CT reconstruction of Betaburmesebuthus fuscus sp. nov., holotype, NIGP200653. A, Dorsal view. B, Ventral view. C, Lateral ocelli.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus. Scale bars = 3 mm in A, B; 500 μm in C." figureDoi="http://doi.org/10.5281/zenodo.7754888" httpUri="https://zenodo.org/record/7754888/files/figure.png" pageId="6" pageNumber="70">28A</figureCitation>
|
||
,
|
||
<figureCitation box="[273,325,1922,1948]" captionStart="FIGURE 29" captionStartId="34.[136,235,1908,1932]" captionTargetBox="[183,1404,189,1884]" captionTargetId="figure-18@34.[183,1404,189,1884]" captionTargetPageId="34" captionText="FIGURE 29. Details of Betaburmesebuthus fuscus sp. nov., holotype, NIGP200653, under incident light. A, Left pedipalp and carapace. B, Pectines. C, Sternites VI–VII and metasomal segments I–III. D, Femur, dorsal view. E, Patella, dorsal view. F, Tarsus of leg IV. G, Tarsus of leg I. H, Tarsus of leg III. I, Tarsus of leg II. J, Spiracle.Abbreviations: sp, spiracle; ts, tibial spur. Scale bars = 1 mm in A, C; 500 μm in B, F–J; 400 μm in D, E." figureDoi="http://doi.org/10.5281/zenodo.7754892" httpUri="https://zenodo.org/record/7754892/files/figure.png" pageId="6" pageNumber="70">29A</figureCitation>
|
||
). Median ocular tubercle well-marked and located on the first anterior third to the first anterior of carapace (
|
||
<figureCitation box="[288,386,1994,2020]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="6" pageNumber="70">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation box="[402,440,1994,2020]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="6" pageNumber="70">6A</figureCitation>
|
||
,
|
||
<figureCitation box="[455,505,1994,2020]" captionStart="FIGURE 10" captionStartId="14.[136,235,1845,1869]" captionTargetBox="[224,1363,189,1822]" captionTargetId="figure-18@14.[224,1363,189,1822]" captionTargetPageId="14" captionText="FIGURE 10. Details of Betaburmesebuthus kobberti, NIGP200650, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Coxapophyses I–II. D, Tergites I–VI. E, Sternites IV–VII. F, Pectine. G, Tergite VII and Metasomal segments I–II. H, Telson. I, Tibial spur of leg II. J, Tibial spur of leg IV. K, Telotarsi of leg III. L, Telotarsi of leg IV. Abbreviations: ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, D, E, F, G, H; 100 μm in C, I, J, K, L; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754844" httpUri="https://zenodo.org/record/7754844/files/figure.png" pageId="6" pageNumber="70">10A</figureCitation>
|
||
,
|
||
<figureCitation box="[522,573,1994,2020]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="6" pageNumber="70">13A</figureCitation>
|
||
,
|
||
<figureCitation box="[589,640,1994,2020]" captionStart="FIGURE 17" captionStartId="22.[136,235,1829,1853]" captionTargetBox="[239,1349,189,1806]" captionTargetId="figure-18@22.[239,1349,189,1806]" captionTargetPageId="22" captionText="FIGURE 17. Details of Betaburmesebuthus larafleissnerae, NIGP201155, A, C–G under green epifluorescence; B under confocal microscopy; H, I under incident light. A, Carapace and chelicerae, dorsal view. B, Lateral ocelli. C, Tarsus of right leg III. D, Tergites. E, Metasomal segments I–IV and telson, lateral view. F, Metasomal segments I–III, ventral view. G, Metasomal segments IV–V and telson, lateral view. H, Pectines. I, Sternite V with right spiracle. Abbreviations: ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 400 μm in A, D, E, F, G, H; 200 μm in C, I; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754864" httpUri="https://zenodo.org/record/7754864/files/figure.png" pageId="6" pageNumber="70">17A</figureCitation>
|
||
,
|
||
<figureCitation box="[656,706,1994,2020]" captionStart="FIGURE 20" captionStartId="25.[136,235,1836,1860]" captionTargetBox="[209,1378,189,1813]" captionTargetId="figure-28@25.[209,1378,189,1813]" captionTargetPageId="25" captionText="FIGURE 20. Details of Betaburmesebuthus muelleri NIGP201156, A, D, E, F under green epifluorescence; B, C, F, G, I–M under confocal microscopy. A, Carapace and tergites. B, Chelicerae, dorsal view. C, Lateral ocelli. D, Tergite VII and metasomal segments I–IV, dorsal view. E, Coxosternal region, pectines and sternites. F, Sternum and genital operculum. G, Pectine. H, Tergite VII and metasomal segments I–IV, ventral view. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Tarsus of leg I. M, Tarsus of leg IV.Abbreviations:ADMi, anterodorsal minor ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 400 μm in A, D, E, H; 100 μm in B, F, G, I–M; 50 μm in C." figureDoi="http://doi.org/10.5281/zenodo.7754870" httpUri="https://zenodo.org/record/7754870/files/figure.png" pageId="6" pageNumber="70">20A</figureCitation>
|
||
,
|
||
<figureCitation box="[722,774,1994,2020]" captionStart="FIGURE 24" captionStartId="30.[136,235,1951,1975]" captionTargetBox="[211,1377,189,1928]" captionTargetId="figure-18@30.[211,1377,189,1928]" captionTargetPageId="30" captionText="FIGURE 24. Details of Betaburmesebuthus villosus sp. nov., holotype, NIGP200652, under green epifluorescence. A, Carapace. B, tergites. C, Sternum and genital operculum. D, Pectines. E, Tergite IV–VI. F, Femur, dorsal view. G, Patella, dorsal view. H, Metasomal segments III–IV, Lateral view. I, Left leg III. J, Right chela. Scale bars = 400 μm in A, B, D–H, J; 200 μm in C, I." figureDoi="http://doi.org/10.5281/zenodo.7754878" httpUri="https://zenodo.org/record/7754878/files/figure.png" pageId="6" pageNumber="70">24A</figureCitation>
|
||
,
|
||
<figureCitation box="[136,188,2030,2056]" captionStart="FIGURE 29" captionStartId="34.[136,235,1908,1932]" captionTargetBox="[183,1404,189,1884]" captionTargetId="figure-18@34.[183,1404,189,1884]" captionTargetPageId="34" captionText="FIGURE 29. Details of Betaburmesebuthus fuscus sp. nov., holotype, NIGP200653, under incident light. A, Left pedipalp and carapace. B, Pectines. C, Sternites VI–VII and metasomal segments I–III. D, Femur, dorsal view. E, Patella, dorsal view. F, Tarsus of leg IV. G, Tarsus of leg I. H, Tarsus of leg III. I, Tarsus of leg II. J, Spiracle.Abbreviations: sp, spiracle; ts, tibial spur. Scale bars = 1 mm in A, C; 500 μm in B, F–J; 400 μm in D, E." figureDoi="http://doi.org/10.5281/zenodo.7754892" httpUri="https://zenodo.org/record/7754892/files/figure.png" pageId="6" pageNumber="70">29A</figureCitation>
|
||
). Sternum elongated pentagonal (
|
||
<figureCitation box="[572,668,2030,2056]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="6" pageNumber="70">Figs 2D</figureCitation>
|
||
,
|
||
<figureCitation box="[682,715,2030,2056]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="6" pageNumber="70">6E</figureCitation>
|
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,
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||
<figureCitation box="[729,774,2030,2056]" captionStart="FIGURE 20" captionStartId="25.[136,235,1836,1860]" captionTargetBox="[209,1378,189,1813]" captionTargetId="figure-28@25.[209,1378,189,1813]" captionTargetPageId="25" captionText="FIGURE 20. Details of Betaburmesebuthus muelleri NIGP201156, A, D, E, F under green epifluorescence; B, C, F, G, I–M under confocal microscopy. A, Carapace and tergites. B, Chelicerae, dorsal view. C, Lateral ocelli. D, Tergite VII and metasomal segments I–IV, dorsal view. E, Coxosternal region, pectines and sternites. F, Sternum and genital operculum. G, Pectine. H, Tergite VII and metasomal segments I–IV, ventral view. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Tarsus of leg I. M, Tarsus of leg IV.Abbreviations:ADMi, anterodorsal minor ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 400 μm in A, D, E, H; 100 μm in B, F, G, I–M; 50 μm in C." figureDoi="http://doi.org/10.5281/zenodo.7754870" httpUri="https://zenodo.org/record/7754870/files/figure.png" pageId="6" pageNumber="70">20F</figureCitation>
|
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,
|
||
<figureCitation box="[136,187,2066,2092]" captionStart="FIGURE 24" captionStartId="30.[136,235,1951,1975]" captionTargetBox="[211,1377,189,1928]" captionTargetId="figure-18@30.[211,1377,189,1928]" captionTargetPageId="30" captionText="FIGURE 24. Details of Betaburmesebuthus villosus sp. nov., holotype, NIGP200652, under green epifluorescence. A, Carapace. B, tergites. C, Sternum and genital operculum. D, Pectines. E, Tergite IV–VI. F, Femur, dorsal view. G, Patella, dorsal view. H, Metasomal segments III–IV, Lateral view. I, Left leg III. J, Right chela. Scale bars = 400 μm in A, B, D–H, J; 200 μm in C, I." figureDoi="http://doi.org/10.5281/zenodo.7754878" httpUri="https://zenodo.org/record/7754878/files/figure.png" pageId="6" pageNumber="70">24C</figureCitation>
|
||
). Pectines with 13–20 teeth and fulcra present (
|
||
<figureCitation box="[728,846,1922,2092]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="6" pageNumber="70">Figs 2K</figureCitation>
|
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,
|
||
<figureCitation box="[860,905,1922,1948]" captionStart="FIGURE 10" captionStartId="14.[136,235,1845,1869]" captionTargetBox="[224,1363,189,1822]" captionTargetId="figure-18@14.[224,1363,189,1822]" captionTargetPageId="14" captionText="FIGURE 10. Details of Betaburmesebuthus kobberti, NIGP200650, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Coxapophyses I–II. D, Tergites I–VI. E, Sternites IV–VII. F, Pectine. G, Tergite VII and Metasomal segments I–II. H, Telson. I, Tibial spur of leg II. J, Tibial spur of leg IV. K, Telotarsi of leg III. L, Telotarsi of leg IV. Abbreviations: ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, D, E, F, G, H; 100 μm in C, I, J, K, L; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754844" httpUri="https://zenodo.org/record/7754844/files/figure.png" pageId="6" pageNumber="70">10F</figureCitation>
|
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,
|
||
<figureCitation box="[918,969,1922,1948]" captionStart="FIGURE 24" captionStartId="30.[136,235,1951,1975]" captionTargetBox="[211,1377,189,1928]" captionTargetId="figure-18@30.[211,1377,189,1928]" captionTargetPageId="30" captionText="FIGURE 24. Details of Betaburmesebuthus villosus sp. nov., holotype, NIGP200652, under green epifluorescence. A, Carapace. B, tergites. C, Sternum and genital operculum. D, Pectines. E, Tergite IV–VI. F, Femur, dorsal view. G, Patella, dorsal view. H, Metasomal segments III–IV, Lateral view. I, Left leg III. J, Right chela. Scale bars = 400 μm in A, B, D–H, J; 200 μm in C, I." figureDoi="http://doi.org/10.5281/zenodo.7754878" httpUri="https://zenodo.org/record/7754878/files/figure.png" pageId="6" pageNumber="70">24D</figureCitation>
|
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,
|
||
<figureCitation box="[983,1033,1922,1948]" captionStart="FIGURE 29" captionStartId="34.[136,235,1908,1932]" captionTargetBox="[183,1404,189,1884]" captionTargetId="figure-18@34.[183,1404,189,1884]" captionTargetPageId="34" captionText="FIGURE 29. Details of Betaburmesebuthus fuscus sp. nov., holotype, NIGP200653, under incident light. A, Left pedipalp and carapace. B, Pectines. C, Sternites VI–VII and metasomal segments I–III. D, Femur, dorsal view. E, Patella, dorsal view. F, Tarsus of leg IV. G, Tarsus of leg I. H, Tarsus of leg III. I, Tarsus of leg II. J, Spiracle.Abbreviations: sp, spiracle; ts, tibial spur. Scale bars = 1 mm in A, C; 500 μm in B, F–J; 400 μm in D, E." figureDoi="http://doi.org/10.5281/zenodo.7754892" httpUri="https://zenodo.org/record/7754892/files/figure.png" pageId="6" pageNumber="70">29B</figureCitation>
|
||
). Metasomal ventral carinae parallel (
|
||
<figureCitation box="[818,912,1958,1985]" captionStart="FIGURE 5" captionStartId="5.[136,235,1934,1958]" captionTargetBox="[174,1413,189,1910]" captionTargetId="figure-28@5.[174,1413,189,1910]" captionTargetPageId="5" captionText="FIGURE 5. Details of Betaburmesebuthus bellus, NIGP200649, under green epifluorescence. A, Carapace and tergites. B, Coxosternal region, pectines and sternites. C, Metasomal segments I–IV, ventral view. D, Metasomal segments I–IV, dorsal view. E, Metasomal segment V and telson, lateral view. F, Tibial spur of leg III. G, Tibial spur of leg IV. H, Telotarsus of leg III. I, Telotarsus of leg IV. Abbreviations: ts, tibial spur. Scale bars = 400 μm in A–E; 100 μm in F–I." figureDoi="http://doi.org/10.5281/zenodo.7754834" httpUri="https://zenodo.org/record/7754834/files/figure.png" pageId="6" pageNumber="70">Figs 5D</figureCitation>
|
||
,
|
||
<figureCitation box="[925,1000,1958,1985]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="6" pageNumber="70">13C, J</figureCitation>
|
||
,
|
||
<figureCitation box="[1013,1070,1958,1984]" captionStart="FIGURE 30" captionStartId="35.[136,235,1690,1714]" captionTargetBox="[275,1312,189,1667]" captionTargetId="figure-123@35.[275,1312,189,1667]" captionTargetPageId="35" captionText="FIGURE 30. Micro-CT reconstruction details of Betaburmesebuthus fuscus sp. nov., holotype, NIGP200653. A, B, Left pedipalp. C–E, Metasomal segments I–III. Scale bars = 1.5 mm in A, B; 1 mm in C–E." figureDoi="http://doi.org/10.5281/zenodo.7754896" httpUri="https://zenodo.org/record/7754896/files/figure.png" pageId="6" pageNumber="70">30 E</figureCitation>
|
||
). Telson with a very long aculeus and a tiny subaculear tubercle (
|
||
<figureCitation box="[1170,1254,1994,2020]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="6" pageNumber="70">Figs 6I</figureCitation>
|
||
,
|
||
<figureCitation box="[1269,1319,1994,2020]" captionStart="FIGURE 10" captionStartId="14.[136,235,1845,1869]" captionTargetBox="[224,1363,189,1822]" captionTargetId="figure-18@14.[224,1363,189,1822]" captionTargetPageId="14" captionText="FIGURE 10. Details of Betaburmesebuthus kobberti, NIGP200650, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Coxapophyses I–II. D, Tergites I–VI. E, Sternites IV–VII. F, Pectine. G, Tergite VII and Metasomal segments I–II. H, Telson. I, Tibial spur of leg II. J, Tibial spur of leg IV. K, Telotarsi of leg III. L, Telotarsi of leg IV. Abbreviations: ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, D, E, F, G, H; 100 μm in C, I, J, K, L; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754844" httpUri="https://zenodo.org/record/7754844/files/figure.png" pageId="6" pageNumber="70">10H</figureCitation>
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||
,
|
||
<figureCitation box="[1333,1383,1994,2020]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="6" pageNumber="70">14B</figureCitation>
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||
,
|
||
<figureCitation box="[1396,1447,1994,2020]" captionStart="FIGURE 17" captionStartId="22.[136,235,1829,1853]" captionTargetBox="[239,1349,189,1806]" captionTargetId="figure-18@22.[239,1349,189,1806]" captionTargetPageId="22" captionText="FIGURE 17. Details of Betaburmesebuthus larafleissnerae, NIGP201155, A, C–G under green epifluorescence; B under confocal microscopy; H, I under incident light. A, Carapace and chelicerae, dorsal view. B, Lateral ocelli. C, Tarsus of right leg III. D, Tergites. E, Metasomal segments I–IV and telson, lateral view. F, Metasomal segments I–III, ventral view. G, Metasomal segments IV–V and telson, lateral view. H, Pectines. I, Sternite V with right spiracle. Abbreviations: ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 400 μm in A, D, E, F, G, H; 200 μm in C, I; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754864" httpUri="https://zenodo.org/record/7754864/files/figure.png" pageId="6" pageNumber="70">17G</figureCitation>
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||
,
|
||
<figureCitation box="[810,850,2030,2056]" captionStart="FIGURE 20" captionStartId="25.[136,235,1836,1860]" captionTargetBox="[209,1378,189,1813]" captionTargetId="figure-28@25.[209,1378,189,1813]" captionTargetPageId="25" captionText="FIGURE 20. Details of Betaburmesebuthus muelleri NIGP201156, A, D, E, F under green epifluorescence; B, C, F, G, I–M under confocal microscopy. A, Carapace and tergites. B, Chelicerae, dorsal view. C, Lateral ocelli. D, Tergite VII and metasomal segments I–IV, dorsal view. E, Coxosternal region, pectines and sternites. F, Sternum and genital operculum. G, Pectine. H, Tergite VII and metasomal segments I–IV, ventral view. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Tarsus of leg I. M, Tarsus of leg IV.Abbreviations:ADMi, anterodorsal minor ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 400 μm in A, D, E, H; 100 μm in B, F, G, I–M; 50 μm in C." figureDoi="http://doi.org/10.5281/zenodo.7754870" httpUri="https://zenodo.org/record/7754870/files/figure.png" pageId="6" pageNumber="70">20I</figureCitation>
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||
,
|
||
<figureCitation box="[865,908,2030,2057]" captionStart="FIGURE 25" captionStartId="31.[136,235,1885,1909]" captionTargetBox="[164,1423,189,1862]" captionTargetId="figure-28@31.[164,1423,189,1862]" captionTargetPageId="31" captionText="FIGURE 25. Details of Betaburmesebuthus villosus sp. nov., holotype, NIGP200652, under confocal microscopy. A, Lateral ocelli. B, Femur, dorsal view. C, Spiracle. D, Tibial spur of leg III. E, Tibial spur of leg IV. F, Right chela manus, ventral view. G, Left chela manus, ventral view. H, Tarsus of leg III. I, Tarsus of leg IV. J–K, Telson, lateral view. Abbreviations: ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; st, subaculear tubercle; ts, tibial spur. Scale bars = 100 μm in A–C, K; 200 μm in D–J." figureDoi="http://doi.org/10.5281/zenodo.7754880" httpUri="https://zenodo.org/record/7754880/files/figure.png" pageId="6" pageNumber="70">25J</figureCitation>
|
||
). Trichobothriotaxic pattern
|
||
<typeStatus box="[1239,1295,2031,2057]" pageId="6" pageNumber="70">Type</typeStatus>
|
||
A with
|
||
<emphasis box="[1394,1442,2030,2056]" italics="true" pageId="6" pageNumber="70">beta</emphasis>
|
||
- configuration (
|
||
<figureCitation box="[975,1046,2066,2092]" captionStart="FIGURE 3" captionStartId="3.[136,235,1712,1736]" captionTargetBox="[151,1436,189,1689]" captionTargetId="figure-114@3.[151,1436,189,1689]" captionTargetPageId="3" captionText="FIGURE 3. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy (B, C, E, F, H–K) and green epifluorescence (A, D, G). A, B, Left femur, dorsal view. C, Right femur, dorsal view. D, E, left patella, dorsal view. F, Right patella, dorsal view. G, Left chela, dorsal view. H, Left fixed finger, dorsal view. I, Right manus, ventral view. J, Left manus, dorsal view. K, Left manus, ventral view. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7754828" httpUri="https://zenodo.org/record/7754828/files/figure.png" pageId="6" pageNumber="70">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation box="[1058,1074,2067,2093]" captionStart="FIGURE 7" captionStartId="7.[136,235,1951,1975]" captionTargetBox="[168,1420,189,1928]" captionTargetId="figure-28@7.[168,1420,189,1928]" captionTargetPageId="7" captionText="FIGURE 7. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy (B, D, F, G) and green epifluorescence (A, C, E). A–B, Right femur, dorsal view. C–D, Right patella, dorsal view. E–F, Right chela, dorsal view. G, Right manus, ventral view. Scale bars = 200 μm in A, B, C, D, G; 400 μm in E, F." figureDoi="http://doi.org/10.5281/zenodo.7754838" httpUri="https://zenodo.org/record/7754838/files/figure.png" pageId="6" pageNumber="70">7</figureCitation>
|
||
,
|
||
<figureCitation box="[1086,1115,2066,2092]" captionStart="FIGURE 11" captionStartId="15.[136,235,1483,1507]" captionTargetBox="[151,1436,189,1460]" captionTargetId="figure-196@15.[151,1436,189,1460]" captionTargetPageId="15" captionText="FIGURE 11. Trichobothrial pattern of Betaburmesebuthus kobberti, NIGP200650, under confocal microscopy. A, Left femur, patella and chela, dorsal view. B, Left femur, patella and chela, ventral view. C. Left femur and fixed finger, dorsal view. D, Right fixed finger, dorsal view. E, Right chela, dorsal view. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7754846" httpUri="https://zenodo.org/record/7754846/files/figure.png" pageId="6" pageNumber="70">11</figureCitation>
|
||
,
|
||
<figureCitation box="[1126,1209,2066,2093]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="6" pageNumber="70">14C, D</figureCitation>
|
||
,
|
||
<figureCitation box="[1220,1250,2066,2092]" captionStart="FIGURE 18" captionStartId="23.[136,235,1597,1621]" captionTargetBox="[151,1436,189,1574]" captionTargetId="figure-148@23.[151,1436,189,1574]" captionTargetPageId="23" captionText="FIGURE 18. Trichobothrial pattern of Betaburmesebuthus larafleissnerae, NIGP201155, A, C, E, F under green epifluorescence; B, D under confocal microscopy. A, B, Right femur, dorsal view. C, D, Left patella, dorsal view. E. Left femur, dorsal view. F, Left chela, dorsal view. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7754866" httpUri="https://zenodo.org/record/7754866/files/figure.png" pageId="6" pageNumber="70">18</figureCitation>
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||
,
|
||
<figureCitation box="[1261,1291,2066,2092]" captionStart="FIGURE 21" captionStartId="26.[136,235,1712,1736]" captionTargetBox="[151,1436,189,1689]" captionTargetId="figure-99@26.[151,1436,189,1689]" captionTargetPageId="26" captionText="FIGURE 21. Trichobothrial pattern of Betaburmesebuthus muelleri NIGP201156, A, B under green epifluorescence; C, D under confocal microscopy. A, Left pedipalp, dorsal view. B, Left pedipalp, ventral view. C–D, Left chela, dorsal view. E. Left femur and patella, dorsal view. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754872" httpUri="https://zenodo.org/record/7754872/files/figure.png" pageId="6" pageNumber="70">21</figureCitation>
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||
,
|
||
<figureCitation box="[1303,1335,2066,2092]" captionStart="FIGURE 26" captionStartId="32.[136,235,1073,1097]" captionTargetBox="[166,1421,201,1036]" captionTargetId="figure-346@32.[151,1436,189,1049]" captionTargetPageId="32" captionText="FIGURE 26. Line drawings of the trichobothrial pattern of Betaburmesebuthus villosus sp. nov., holotype, NIGP200652. A, femur; B, patella; C, chela. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7754882" httpUri="https://zenodo.org/record/7754882/files/figure.png" pageId="6" pageNumber="70">26</figureCitation>
|
||
). Pedipalp patellar dorsomedian carina (DMc) well-developed (
|
||
<figureCitation captionStart="FIGURE 3" captionStartId="3.[136,235,1712,1736]" captionTargetBox="[151,1436,189,1689]" captionTargetId="figure-114@3.[151,1436,189,1689]" captionTargetPageId="3" captionText="FIGURE 3. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy (B, C, E, F, H–K) and green epifluorescence (A, D, G). A, B, Left femur, dorsal view. C, Right femur, dorsal view. D, E, left patella, dorsal view. F, Right patella, dorsal view. G, Left chela, dorsal view. H, Left fixed finger, dorsal view. I, Right manus, ventral view. J, Left manus, dorsal view. K, Left manus, ventral view. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7754828" httpUri="https://zenodo.org/record/7754828/files/figure.png" pageId="8" pageNumber="72">Figs 3E</figureCitation>
|
||
,
|
||
<figureCitation box="[181,217,195,221]" captionStart="FIGURE 7" captionStartId="7.[136,235,1951,1975]" captionTargetBox="[168,1420,189,1928]" captionTargetId="figure-28@7.[168,1420,189,1928]" captionTargetPageId="7" captionText="FIGURE 7. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy (B, D, F, G) and green epifluorescence (A, C, E). A–B, Right femur, dorsal view. C–D, Right patella, dorsal view. E–F, Right chela, dorsal view. G, Right manus, ventral view. Scale bars = 200 μm in A, B, C, D, G; 400 μm in E, F." figureDoi="http://doi.org/10.5281/zenodo.7754838" httpUri="https://zenodo.org/record/7754838/files/figure.png" pageId="8" pageNumber="72">7D</figureCitation>
|
||
,
|
||
<figureCitation box="[229,278,195,221]" captionStart="FIGURE 11" captionStartId="15.[136,235,1483,1507]" captionTargetBox="[151,1436,189,1460]" captionTargetId="figure-196@15.[151,1436,189,1460]" captionTargetPageId="15" captionText="FIGURE 11. Trichobothrial pattern of Betaburmesebuthus kobberti, NIGP200650, under confocal microscopy. A, Left femur, patella and chela, dorsal view. B, Left femur, patella and chela, ventral view. C. Left femur and fixed finger, dorsal view. D, Right fixed finger, dorsal view. E, Right chela, dorsal view. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7754846" httpUri="https://zenodo.org/record/7754846/files/figure.png" pageId="8" pageNumber="72">11A</figureCitation>
|
||
,
|
||
<figureCitation box="[290,340,194,221]" captionStart="FIGURE 15" captionStartId="20.[136,235,1943,1967]" captionTargetBox="[170,1416,189,1920]" captionTargetId="figure-18@20.[170,1416,189,1920]" captionTargetPageId="20" captionText="FIGURE 15. General habitus and details of Betaburmesebuthus sp., NIGP201157, under incident light. A, Ventral view. B, Dorsal view. C, Tarsus of right leg I. D, Tarsus of right leg II. E, Tarsus of right leg III. F, Tarsus of left leg III.Abbreviations: st, subaculear tubercle. Scale bars = 5 mm in A, B; 1 mm in G; 500 μm in C, D, E, F." figureDoi="http://doi.org/10.5281/zenodo.7754858" httpUri="https://zenodo.org/record/7754858/files/figure.png" pageId="8" pageNumber="72">15A</figureCitation>
|
||
,
|
||
<figureCitation box="[352,402,195,221]" captionStart="FIGURE 18" captionStartId="23.[136,235,1597,1621]" captionTargetBox="[151,1436,189,1574]" captionTargetId="figure-148@23.[151,1436,189,1574]" captionTargetPageId="23" captionText="FIGURE 18. Trichobothrial pattern of Betaburmesebuthus larafleissnerae, NIGP201155, A, C, E, F under green epifluorescence; B, D under confocal microscopy. A, B, Right femur, dorsal view. C, D, Left patella, dorsal view. E. Left femur, dorsal view. F, Left chela, dorsal view. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7754866" httpUri="https://zenodo.org/record/7754866/files/figure.png" pageId="8" pageNumber="72">18D</figureCitation>
|
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,
|
||
<figureCitation box="[414,464,195,221]" captionStart="FIGURE 21" captionStartId="26.[136,235,1712,1736]" captionTargetBox="[151,1436,189,1689]" captionTargetId="figure-99@26.[151,1436,189,1689]" captionTargetPageId="26" captionText="FIGURE 21. Trichobothrial pattern of Betaburmesebuthus muelleri NIGP201156, A, B under green epifluorescence; C, D under confocal microscopy. A, Left pedipalp, dorsal view. B, Left pedipalp, ventral view. C–D, Left chela, dorsal view. E. Left femur and patella, dorsal view. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754872" httpUri="https://zenodo.org/record/7754872/files/figure.png" pageId="8" pageNumber="72">21D</figureCitation>
|
||
,
|
||
<figureCitation box="[476,524,195,221]" captionStart="FIGURE 23" captionStartId="29.[136,235,1952,1976]" captionTargetBox="[151,1436,189,1928]" captionTargetId="figure-28@29.[151,1436,189,1928]" captionTargetPageId="29" captionText="FIGURE 23. Details of Betaburmesebuthus villosus sp. nov., holotype, NIGP200652, under incident light. A, Tergites. B, Sternum and genital operculum and pectines. C, Chela, dorsal view. D, Femur, dorsal view. E, Patella, dorsal view. F, Right Leg III. G, Tergite VII and metasomal segments I–II. H, Telson. Abbreviations: ts, tibial spur. Scale bars = 1 mm in A; 500 μm in B–H." figureDoi="http://doi.org/10.5281/zenodo.7754876" httpUri="https://zenodo.org/record/7754876/files/figure.png" pageId="8" pageNumber="72">23E</figureCitation>
|
||
,
|
||
<figureCitation box="[535,587,195,221]" captionStart="FIGURE 29" captionStartId="34.[136,235,1908,1932]" captionTargetBox="[183,1404,189,1884]" captionTargetId="figure-18@34.[183,1404,189,1884]" captionTargetPageId="34" captionText="FIGURE 29. Details of Betaburmesebuthus fuscus sp. nov., holotype, NIGP200653, under incident light. A, Left pedipalp and carapace. B, Pectines. C, Sternites VI–VII and metasomal segments I–III. D, Femur, dorsal view. E, Patella, dorsal view. F, Tarsus of leg IV. G, Tarsus of leg I. H, Tarsus of leg III. I, Tarsus of leg II. J, Spiracle.Abbreviations: sp, spiracle; ts, tibial spur. Scale bars = 1 mm in A, C; 500 μm in B, F–J; 400 μm in D, E." figureDoi="http://doi.org/10.5281/zenodo.7754892" httpUri="https://zenodo.org/record/7754892/files/figure.png" pageId="8" pageNumber="72">29D</figureCitation>
|
||
). Pedipalp finger dentition composed of about ten discrete linear rows of granules; each slightly oblique with proximal ends directed externally, rows slightly imbricated; a smaller spinous external accessory granule and a bigger spinous internal accessory granule present along the base of each denticle row; tip hooked (
|
||
<figureCitation box="[420,546,413,439]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="8" pageNumber="72">Figs 2G, H</figureCitation>
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,
|
||
<figureCitation box="[556,593,413,439]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="8" pageNumber="72">6H</figureCitation>
|
||
,
|
||
<figureCitation box="[604,669,413,439]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="8" pageNumber="72">13F, I</figureCitation>
|
||
,
|
||
<figureCitation box="[680,765,413,439]" captionStart="FIGURE 21" captionStartId="26.[136,235,1712,1736]" captionTargetBox="[151,1436,189,1689]" captionTargetId="figure-99@26.[151,1436,189,1689]" captionTargetPageId="26" captionText="FIGURE 21. Trichobothrial pattern of Betaburmesebuthus muelleri NIGP201156, A, B under green epifluorescence; C, D under confocal microscopy. A, Left pedipalp, dorsal view. B, Left pedipalp, ventral view. C–D, Left chela, dorsal view. E. Left femur and patella, dorsal view. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754872" httpUri="https://zenodo.org/record/7754872/files/figure.png" pageId="8" pageNumber="72">21A–C</figureCitation>
|
||
). Tibial spurs present on legs
|
||
<collectionCode box="[457,485,450,476]" pageId="8" pageNumber="72">III</collectionCode>
|
||
and
|
||
<collectionCode box="[542,571,450,476]" pageId="8" pageNumber="72">IV</collectionCode>
|
||
(
|
||
<figureCitation box="[587,697,449,476]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="8" pageNumber="72">Figs 2I, J</figureCitation>
|
||
,
|
||
<figureCitation box="[710,774,449,476]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="8" pageNumber="72">6J, K</figureCitation>
|
||
,
|
||
<figureCitation box="[136,222,485,512]" captionStart="FIGURE 10" captionStartId="14.[136,235,1845,1869]" captionTargetBox="[224,1363,189,1822]" captionTargetId="figure-18@14.[224,1363,189,1822]" captionTargetPageId="14" captionText="FIGURE 10. Details of Betaburmesebuthus kobberti, NIGP200650, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Coxapophyses I–II. D, Tergites I–VI. E, Sternites IV–VII. F, Pectine. G, Tergite VII and Metasomal segments I–II. H, Telson. I, Tibial spur of leg II. J, Tibial spur of leg IV. K, Telotarsi of leg III. L, Telotarsi of leg IV. Abbreviations: ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, D, E, F, G, H; 100 μm in C, I, J, K, L; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754844" httpUri="https://zenodo.org/record/7754844/files/figure.png" pageId="8" pageNumber="72">10K, L</figureCitation>
|
||
,
|
||
<figureCitation box="[239,328,485,512]" captionStart="FIGURE 13" captionStartId="17.[136,235,1927,1951]" captionTargetBox="[170,1416,189,1904]" captionTargetId="figure-28@17.[170,1416,189,1904]" captionTargetPageId="17" captionText="FIGURE 13. Details of Betaburmesebuthus fleissneri, NIGP201154, under incident light. A, Carapace and tergites I–III. B, Coxosternal region, pectines and sternites. C, J, Metasoma and telson, lateral view. D, Tergites IV–VII. E, Left pedipalp, dorsal view. F, Dentition of left chela movable finger. G, Tibial spur of leg IV. H, Tibial spur of leg III. I, Dentition of right chela finger. Abbreviations: ts, tibial spur. Scale bars = 2 mm in C, J; 1 mm in A, B, D, E; 500 μm in F, G, H, I." figureDoi="http://doi.org/10.5281/zenodo.7754852" httpUri="https://zenodo.org/record/7754852/files/figure.png" pageId="8" pageNumber="72">13G, H</figureCitation>
|
||
,
|
||
<figureCitation box="[344,425,485,512]" captionStart="FIGURE 20" captionStartId="25.[136,235,1836,1860]" captionTargetBox="[209,1378,189,1813]" captionTargetId="figure-28@25.[209,1378,189,1813]" captionTargetPageId="25" captionText="FIGURE 20. Details of Betaburmesebuthus muelleri NIGP201156, A, D, E, F under green epifluorescence; B, C, F, G, I–M under confocal microscopy. A, Carapace and tergites. B, Chelicerae, dorsal view. C, Lateral ocelli. D, Tergite VII and metasomal segments I–IV, dorsal view. E, Coxosternal region, pectines and sternites. F, Sternum and genital operculum. G, Pectine. H, Tergite VII and metasomal segments I–IV, ventral view. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Tarsus of leg I. M, Tarsus of leg IV.Abbreviations:ADMi, anterodorsal minor ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 400 μm in A, D, E, H; 100 μm in B, F, G, I–M; 50 μm in C." figureDoi="http://doi.org/10.5281/zenodo.7754870" httpUri="https://zenodo.org/record/7754870/files/figure.png" pageId="8" pageNumber="72">20J, K</figureCitation>
|
||
,
|
||
<figureCitation box="[440,524,485,511]" captionStart="FIGURE 29" captionStartId="34.[136,235,1908,1932]" captionTargetBox="[183,1404,189,1884]" captionTargetId="figure-18@34.[183,1404,189,1884]" captionTargetPageId="34" captionText="FIGURE 29. Details of Betaburmesebuthus fuscus sp. nov., holotype, NIGP200653, under incident light. A, Left pedipalp and carapace. B, Pectines. C, Sternites VI–VII and metasomal segments I–III. D, Femur, dorsal view. E, Patella, dorsal view. F, Tarsus of leg IV. G, Tarsus of leg I. H, Tarsus of leg III. I, Tarsus of leg II. J, Spiracle.Abbreviations: sp, spiracle; ts, tibial spur. Scale bars = 1 mm in A, C; 500 μm in B, F–J; 400 μm in D, E." figureDoi="http://doi.org/10.5281/zenodo.7754892" httpUri="https://zenodo.org/record/7754892/files/figure.png" pageId="8" pageNumber="72">29F, H</figureCitation>
|
||
). Telotarsus with few ventrosubmedian setae and a ventromedian spinules row (
|
||
<figureCitation box="[144,282,558,585]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="8" pageNumber="72">Figs 2M, N</figureCitation>
|
||
,
|
||
<figureCitation box="[295,370,558,585]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="8" pageNumber="72">6L, M</figureCitation>
|
||
,
|
||
<figureCitation box="[384,451,558,585]" captionStart="FIGURE 10" captionStartId="14.[136,235,1845,1869]" captionTargetBox="[224,1363,189,1822]" captionTargetId="figure-18@14.[224,1363,189,1822]" captionTargetPageId="14" captionText="FIGURE 10. Details of Betaburmesebuthus kobberti, NIGP200650, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Coxapophyses I–II. D, Tergites I–VI. E, Sternites IV–VII. F, Pectine. G, Tergite VII and Metasomal segments I–II. H, Telson. I, Tibial spur of leg II. J, Tibial spur of leg IV. K, Telotarsi of leg III. L, Telotarsi of leg IV. Abbreviations: ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, D, E, F, G, H; 100 μm in C, I, J, K, L; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754844" httpUri="https://zenodo.org/record/7754844/files/figure.png" pageId="8" pageNumber="72">10I, J</figureCitation>
|
||
,
|
||
<figureCitation box="[466,515,558,584]" captionStart="FIGURE 17" captionStartId="22.[136,235,1829,1853]" captionTargetBox="[239,1349,189,1806]" captionTargetId="figure-18@22.[239,1349,189,1806]" captionTargetPageId="22" captionText="FIGURE 17. Details of Betaburmesebuthus larafleissnerae, NIGP201155, A, C–G under green epifluorescence; B under confocal microscopy; H, I under incident light. A, Carapace and chelicerae, dorsal view. B, Lateral ocelli. C, Tarsus of right leg III. D, Tergites. E, Metasomal segments I–IV and telson, lateral view. F, Metasomal segments I–III, ventral view. G, Metasomal segments IV–V and telson, lateral view. H, Pectines. I, Sternite V with right spiracle. Abbreviations: ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 400 μm in A, D, E, F, G, H; 200 μm in C, I; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754864" httpUri="https://zenodo.org/record/7754864/files/figure.png" pageId="8" pageNumber="72">17C</figureCitation>
|
||
,
|
||
<figureCitation box="[529,617,558,584]" captionStart="FIGURE 20" captionStartId="25.[136,235,1836,1860]" captionTargetBox="[209,1378,189,1813]" captionTargetId="figure-28@25.[209,1378,189,1813]" captionTargetPageId="25" captionText="FIGURE 20. Details of Betaburmesebuthus muelleri NIGP201156, A, D, E, F under green epifluorescence; B, C, F, G, I–M under confocal microscopy. A, Carapace and tergites. B, Chelicerae, dorsal view. C, Lateral ocelli. D, Tergite VII and metasomal segments I–IV, dorsal view. E, Coxosternal region, pectines and sternites. F, Sternum and genital operculum. G, Pectine. H, Tergite VII and metasomal segments I–IV, ventral view. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Tarsus of leg I. M, Tarsus of leg IV.Abbreviations:ADMi, anterodorsal minor ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 400 μm in A, D, E, H; 100 μm in B, F, G, I–M; 50 μm in C." figureDoi="http://doi.org/10.5281/zenodo.7754870" httpUri="https://zenodo.org/record/7754870/files/figure.png" pageId="8" pageNumber="72">20L, M</figureCitation>
|
||
) or numerous ventrosubmedian setae (
|
||
<figureCitation box="[418,562,594,621]" captionStart="FIGURE 14" captionStartId="18.[136,235,1946,1970]" captionTargetBox="[151,1436,189,1922]" captionTargetId="figure-18@18.[151,1436,189,1922]" captionTargetPageId="18" captionText="FIGURE 14. Details of Betaburmesebuthus fleissneri, NIGP201154, under green epifluorescence. A, Carapace and chelicerae. B, Telson, lateral view. C, Patella, dorsal view. D, Femur, dorsal view. E, Telotarsus of leg IV. F, Telotarsus of leg II. G, Telotarsus of leg III. Abbreviations: st, subaculear tubercle. Scale bars = 400 μm." figureDoi="http://doi.org/10.5281/zenodo.7754856" httpUri="https://zenodo.org/record/7754856/files/figure.png" pageId="8" pageNumber="72">Figs 14E–G</figureCitation>
|
||
,
|
||
<figureCitation box="[579,656,594,621]" captionStart="FIGURE 15" captionStartId="20.[136,235,1943,1967]" captionTargetBox="[170,1416,189,1920]" captionTargetId="figure-18@20.[170,1416,189,1920]" captionTargetPageId="20" captionText="FIGURE 15. General habitus and details of Betaburmesebuthus sp., NIGP201157, under incident light. A, Ventral view. B, Dorsal view. C, Tarsus of right leg I. D, Tarsus of right leg II. E, Tarsus of right leg III. F, Tarsus of left leg III.Abbreviations: st, subaculear tubercle. Scale bars = 5 mm in A, B; 1 mm in G; 500 μm in C, D, E, F." figureDoi="http://doi.org/10.5281/zenodo.7754858" httpUri="https://zenodo.org/record/7754858/files/figure.png" pageId="8" pageNumber="72">15C–F</figureCitation>
|
||
,
|
||
<figureCitation box="[672,716,595,621]" captionStart="FIGURE 23" captionStartId="29.[136,235,1952,1976]" captionTargetBox="[151,1436,189,1928]" captionTargetId="figure-28@29.[151,1436,189,1928]" captionTargetPageId="29" captionText="FIGURE 23. Details of Betaburmesebuthus villosus sp. nov., holotype, NIGP200652, under incident light. A, Tergites. B, Sternum and genital operculum and pectines. C, Chela, dorsal view. D, Femur, dorsal view. E, Patella, dorsal view. F, Right Leg III. G, Tergite VII and metasomal segments I–II. H, Telson. Abbreviations: ts, tibial spur. Scale bars = 1 mm in A; 500 μm in B–H." figureDoi="http://doi.org/10.5281/zenodo.7754876" httpUri="https://zenodo.org/record/7754876/files/figure.png" pageId="8" pageNumber="72">23F</figureCitation>
|
||
,
|
||
<figureCitation box="[733,774,595,621]" captionStart="FIGURE 24" captionStartId="30.[136,235,1951,1975]" captionTargetBox="[211,1377,189,1928]" captionTargetId="figure-18@30.[211,1377,189,1928]" captionTargetPageId="30" captionText="FIGURE 24. Details of Betaburmesebuthus villosus sp. nov., holotype, NIGP200652, under green epifluorescence. A, Carapace. B, tergites. C, Sternum and genital operculum. D, Pectines. E, Tergite IV–VI. F, Femur, dorsal view. G, Patella, dorsal view. H, Metasomal segments III–IV, Lateral view. I, Left leg III. J, Right chela. Scale bars = 400 μm in A, B, D–H, J; 200 μm in C, I." figureDoi="http://doi.org/10.5281/zenodo.7754878" httpUri="https://zenodo.org/record/7754878/files/figure.png" pageId="8" pageNumber="72">24I</figureCitation>
|
||
,
|
||
<figureCitation box="[136,209,631,658]" captionStart="FIGURE 25" captionStartId="31.[136,235,1885,1909]" captionTargetBox="[164,1423,189,1862]" captionTargetId="figure-28@31.[164,1423,189,1862]" captionTargetPageId="31" captionText="FIGURE 25. Details of Betaburmesebuthus villosus sp. nov., holotype, NIGP200652, under confocal microscopy. A, Lateral ocelli. B, Femur, dorsal view. C, Spiracle. D, Tibial spur of leg III. E, Tibial spur of leg IV. F, Right chela manus, ventral view. G, Left chela manus, ventral view. H, Tarsus of leg III. I, Tarsus of leg IV. J–K, Telson, lateral view. Abbreviations: ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; st, subaculear tubercle; ts, tibial spur. Scale bars = 100 μm in A–C, K; 200 μm in D–J." figureDoi="http://doi.org/10.5281/zenodo.7754880" httpUri="https://zenodo.org/record/7754880/files/figure.png" pageId="8" pageNumber="72">25H, I</figureCitation>
|
||
,
|
||
<figureCitation box="[222,293,631,657]" captionStart="FIGURE 29" captionStartId="34.[136,235,1908,1932]" captionTargetBox="[183,1404,189,1884]" captionTargetId="figure-18@34.[183,1404,189,1884]" captionTargetPageId="34" captionText="FIGURE 29. Details of Betaburmesebuthus fuscus sp. nov., holotype, NIGP200653, under incident light. A, Left pedipalp and carapace. B, Pectines. C, Sternites VI–VII and metasomal segments I–III. D, Femur, dorsal view. E, Patella, dorsal view. F, Tarsus of leg IV. G, Tarsus of leg I. H, Tarsus of leg III. I, Tarsus of leg II. J, Spiracle.Abbreviations: sp, spiracle; ts, tibial spur. Scale bars = 1 mm in A, C; 500 μm in B, F–J; 400 μm in D, E." figureDoi="http://doi.org/10.5281/zenodo.7754892" httpUri="https://zenodo.org/record/7754892/files/figure.png" pageId="8" pageNumber="72">29F–I</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7754830" ID-Zenodo-Dep="7754830" httpUri="https://zenodo.org/record/7754830/files/figure.png" pageId="4" pageNumber="68" startId="4.[136,235,1982,2006]" targetBox="[138,1449,189,1959]" targetPageId="4">
|
||
<paragraph blockId="4.[136,1452,1982,2043]" pageId="4" pageNumber="68">
|
||
<emphasis bold="true" box="[136,261,1982,2007]" pageId="4" pageNumber="68">FIGURE 4.</emphasis>
|
||
General habitus of
|
||
<taxonomicName authority=", NIGP" authorityName="NIGP" box="[468,811,1983,2007]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="bellus">
|
||
<emphasis box="[468,737,1983,2007]" italics="true" pageId="4" pageNumber="68">Betaburmesebuthus bellus</emphasis>
|
||
, NIGP
|
||
</taxonomicName>
|
||
200649, under incident light.
|
||
<emphasis bold="true" box="[1116,1134,1982,2006]" pageId="4" pageNumber="68">A</emphasis>
|
||
, Dorsal view.
|
||
<emphasis bold="true" box="[1286,1303,1983,2007]" pageId="4" pageNumber="68">B</emphasis>
|
||
, Ventral view. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7754834" ID-Zenodo-Dep="7754834" httpUri="https://zenodo.org/record/7754834/files/figure.png" pageId="5" pageNumber="69" startId="5.[136,235,1934,1958]" targetBox="[174,1413,189,1910]" targetPageId="5">
|
||
<paragraph blockId="5.[136,1452,1934,2066]" pageId="5" pageNumber="69">
|
||
<emphasis bold="true" box="[136,265,1934,1958]" pageId="5" pageNumber="69">FIGURE 5.</emphasis>
|
||
Details of
|
||
<taxonomicName authority=", NIGP" authorityName="NIGP" box="[392,742,1934,1958]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="69" phylum="Arthropoda" rank="species" species="bellus">
|
||
<emphasis box="[392,664,1934,1958]" italics="true" pageId="5" pageNumber="69">Betaburmesebuthus bellus</emphasis>
|
||
, NIGP
|
||
</taxonomicName>
|
||
200649, under green epifluorescence.
|
||
<emphasis bold="true" box="[1147,1165,1934,1958]" pageId="5" pageNumber="69">A</emphasis>
|
||
, Carapace and tergites.
|
||
<emphasis bold="true" box="[1428,1445,1934,1958]" pageId="5" pageNumber="69">B</emphasis>
|
||
, Coxosternal region, pectines and sternites.
|
||
<emphasis bold="true" box="[573,591,1970,1994]" pageId="5" pageNumber="69">C</emphasis>
|
||
, Metasomal segments I–IV, ventral view.
|
||
<emphasis bold="true" box="[1020,1038,1970,1994]" pageId="5" pageNumber="69">D</emphasis>
|
||
, Metasomal segments I–IV, dorsal view.
|
||
<emphasis bold="true" box="[136,153,2006,2030]" pageId="5" pageNumber="69">E</emphasis>
|
||
, Metasomal segment V and telson, lateral view.
|
||
<emphasis bold="true" box="[661,676,2006,2030]" pageId="5" pageNumber="69">F</emphasis>
|
||
, Tibial spur of leg III.
|
||
<emphasis bold="true" box="[920,940,2006,2030]" pageId="5" pageNumber="69">G</emphasis>
|
||
, Tibial spur of leg IV.
|
||
<emphasis bold="true" box="[1180,1200,2006,2030]" pageId="5" pageNumber="69">H</emphasis>
|
||
, Telotarsus of leg III.
|
||
<emphasis bold="true" box="[1435,1445,2006,2030]" pageId="5" pageNumber="69">I</emphasis>
|
||
, Telotarsus of leg IV. Abbreviations: ts, tibial spur. Scale bars = 400 μm in
|
||
<emphasis bold="true" box="[886,934,2042,2066]" pageId="5" pageNumber="69">A–E</emphasis>
|
||
; 100 μm in
|
||
<emphasis bold="true" box="[1057,1095,2042,2066]" pageId="5" pageNumber="69">F–I</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7754836" ID-Zenodo-Dep="7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="6" pageNumber="70" startId="6.[136,235,1657,1681]" targetBox="[151,1436,189,1633]" targetPageId="6">
|
||
<paragraph blockId="6.[136,1452,1657,1862]" pageId="6" pageNumber="70">
|
||
<emphasis bold="true" box="[136,261,1657,1681]" pageId="6" pageNumber="70">FIGURE 6.</emphasis>
|
||
Details of
|
||
<taxonomicName authority=", NIGP" authorityName="NIGP" box="[379,723,1657,1681]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="70" phylum="Arthropoda" rank="species" species="bellus">
|
||
<emphasis box="[379,648,1657,1681]" italics="true" pageId="6" pageNumber="70">Betaburmesebuthus bellus</emphasis>
|
||
, NIGP
|
||
</taxonomicName>
|
||
200649, under confocal microscopy.
|
||
<emphasis bold="true" box="[1103,1121,1657,1681]" pageId="6" pageNumber="70">A</emphasis>
|
||
, Carapace.
|
||
<emphasis bold="true" box="[1245,1262,1657,1681]" pageId="6" pageNumber="70">B</emphasis>
|
||
, Lateral ocelli.
|
||
<emphasis bold="true" box="[1427,1445,1657,1681]" pageId="6" pageNumber="70">C</emphasis>
|
||
, Chelicerae, dorsal view.
|
||
<emphasis bold="true" box="[389,407,1693,1717]" pageId="6" pageNumber="70">D</emphasis>
|
||
, Chelicerae, ventral view.
|
||
<emphasis bold="true" box="[682,699,1693,1717]" pageId="6" pageNumber="70">E</emphasis>
|
||
, Coxosternal region.
|
||
<emphasis bold="true" box="[921,936,1693,1717]" pageId="6" pageNumber="70">F</emphasis>
|
||
, Genital operculum and basal piece.
|
||
<emphasis bold="true" box="[1319,1339,1693,1717]" pageId="6" pageNumber="70">G</emphasis>
|
||
, Spiracles.
|
||
<emphasis bold="true" box="[136,156,1729,1753]" pageId="6" pageNumber="70">H</emphasis>
|
||
, Dentition of movable finger.
|
||
<emphasis bold="true" box="[466,476,1729,1753]" pageId="6" pageNumber="70">I</emphasis>
|
||
, Telson.
|
||
<emphasis bold="true" box="[570,583,1729,1753]" pageId="6" pageNumber="70">J</emphasis>
|
||
, Tibial spur of leg III.
|
||
<emphasis bold="true" box="[817,837,1729,1753]" pageId="6" pageNumber="70">K</emphasis>
|
||
, Tibial spur of leg IV.
|
||
<emphasis bold="true" box="[1070,1087,1729,1753]" pageId="6" pageNumber="70">L</emphasis>
|
||
, Telotarsi of leg IV.
|
||
<emphasis bold="true" box="[1298,1322,1729,1753]" pageId="6" pageNumber="70">M</emphasis>
|
||
, Telotarsi of leg III. Abbreviations: ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in
|
||
<emphasis bold="true" box="[315,333,1837,1861]" pageId="6" pageNumber="70">A</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[346,363,1837,1861]" pageId="6" pageNumber="70">E</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[376,396,1837,1861]" pageId="6" pageNumber="70">G</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[408,428,1837,1861]" pageId="6" pageNumber="70">H</emphasis>
|
||
; 100 μm in
|
||
<emphasis bold="true" box="[551,569,1837,1861]" pageId="6" pageNumber="70">C</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[582,600,1837,1861]" pageId="6" pageNumber="70">D</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[613,628,1837,1861]" pageId="6" pageNumber="70">F</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[641,651,1837,1861]" pageId="6" pageNumber="70">I</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[664,677,1837,1861]" pageId="6" pageNumber="70">J</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[689,709,1837,1861]" pageId="6" pageNumber="70">K</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[721,738,1837,1861]" pageId="6" pageNumber="70">L</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[751,775,1837,1861]" pageId="6" pageNumber="70">M</emphasis>
|
||
; 50 μm in
|
||
<emphasis bold="true" box="[886,903,1837,1861]" pageId="6" pageNumber="70">B</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7754838" ID-Zenodo-Dep="7754838" httpUri="https://zenodo.org/record/7754838/files/figure.png" pageId="7" pageNumber="71" startId="7.[136,235,1951,1975]" targetBox="[168,1420,189,1928]" targetPageId="7">
|
||
<paragraph blockId="7.[136,1451,1951,2048]" pageId="7" pageNumber="71">
|
||
<emphasis bold="true" box="[136,259,1951,1975]" pageId="7" pageNumber="71">FIGURE 7.</emphasis>
|
||
Trichobothrial pattern of
|
||
<taxonomicName authority=", NIGP" authorityName="NIGP" box="[518,858,1951,1975]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="71" phylum="Arthropoda" rank="species" species="bellus">
|
||
<emphasis box="[518,785,1951,1975]" italics="true" pageId="7" pageNumber="71">Betaburmesebuthus bellus</emphasis>
|
||
, NIGP
|
||
</taxonomicName>
|
||
200649, under confocal microscopy (
|
||
<emphasis bold="true" box="[1232,1249,1951,1975]" pageId="7" pageNumber="71">B</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1261,1279,1951,1975]" pageId="7" pageNumber="71">D</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1291,1306,1951,1975]" pageId="7" pageNumber="71">F</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1319,1339,1951,1975]" pageId="7" pageNumber="71">G</emphasis>
|
||
) and green epifluorescence (A, C, E).
|
||
<emphasis bold="true" box="[404,452,1987,2011]" pageId="7" pageNumber="71">A–B</emphasis>
|
||
, Right femur, dorsal view.
|
||
<emphasis bold="true" box="[723,772,1987,2011]" pageId="7" pageNumber="71">C–D</emphasis>
|
||
, Right patella, dorsal view.
|
||
<emphasis bold="true" box="[1053,1097,1987,2011]" pageId="7" pageNumber="71">E–F</emphasis>
|
||
, Right chela, dorsal view.
|
||
<emphasis bold="true" box="[1363,1383,1987,2011]" pageId="7" pageNumber="71">G</emphasis>
|
||
, Right manus, ventral view. Scale bars = 200 μm in
|
||
<emphasis bold="true" box="[592,610,2023,2047]" pageId="7" pageNumber="71">A</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[623,640,2023,2047]" pageId="7" pageNumber="71">B</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[653,671,2023,2047]" pageId="7" pageNumber="71">C</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[684,702,2023,2047]" pageId="7" pageNumber="71">D</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[715,735,2023,2047]" pageId="7" pageNumber="71">G</emphasis>
|
||
; 400 μm in
|
||
<emphasis bold="true" box="[858,875,2023,2047]" pageId="7" pageNumber="71">E</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[887,902,2023,2047]" pageId="7" pageNumber="71">F</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="8" pageNumber="72" type="discussion">
|
||
<paragraph blockId="8.[136,778,158,2092]" pageId="8" pageNumber="72">
|
||
<emphasis bold="true" box="[189,305,666,692]" pageId="8" pageNumber="72">Remarks.</emphasis>
|
||
Previous descriptions of
|
||
<taxonomicName box="[136,357,701,727]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[136,357,701,727]" italics="true" pageId="8" pageNumber="72">Betaburmesebuthus</emphasis>
|
||
</taxonomicName>
|
||
indicated that most members of the genus, except for
|
||
<taxonomicName baseAuthorityName="Lourenco" baseAuthorityYear="2015" box="[344,495,736,762]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis box="[344,361,737,762]" italics="true" pageId="8" pageNumber="72">B</emphasis>
|
||
.
|
||
<emphasis box="[378,495,736,762]" italics="true" pageId="8" pageNumber="72">bidentatus</emphasis>
|
||
</taxonomicName>
|
||
, bear three lateral ocelli (
|
||
<bibRefCitation author="Lourenco, W. R. & Beigel, A." box="[143,466,771,798]" pageId="8" pageNumber="72" pagination="476 - 480" refId="ref13343" refString="Lourenco, W. R. & Beigel, A. (2015) A new genus and species of Palaeoburmesebuthinae Lourenco, 2014 (Scorpiones: Archaeobuthidae:) from Cretaceous amber of Myanmar. Beitrage zur Araneologie, 9, 476 - 480." type="journal article" year="2015">Lourenço & Beigel, 2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Lourenco, W. R." box="[489,692,771,798]" pageId="8" pageNumber="72" pagination="27 - 36" refId="ref13090" refString="Lourenco, W. R. (2015 a) A new contribution to the knowledge of Cretaceous Burmese amber scorpions with the description of two new species of Betaburmesebuthus Lourenco, 2015 (Scorpiones: Archaeobuthidae: Palaeoburmesebuthinae). Arachnida, Rivista Aracnologica Italiana, 1 (3), 27 - 36." type="journal article" year="2015">Lourenço, 2015a</bibRefCitation>
|
||
, 2016;
|
||
<bibRefCitation author="Lourenco, W. R. & Velten, J." box="[136,439,806,832]" pageId="8" pageNumber="72" pagination="4 - 11" refId="ref13434" refString="Lourenco, W. R. & Velten, J. (2016 a) One more new species of Betaburmesebuthus Lourenco, 2015 (Scorpiones: Palaeoburmesebuthinae) from Cretaceous burmite. Arachnida, Rivista Aracnologica Italiana, 6, 4 - 11." type="journal article" year="2016">Lourenço & Velten, 2016a</bibRefCitation>
|
||
, b;
|
||
<bibRefCitation author="Lourenco, W. R. & Rossi, A." box="[482,766,806,832]" pageId="8" pageNumber="72" pagination="11 - 21" refId="ref13389" refString="Lourenco, W. R. & Rossi, A. (2017) Two more new species of Burmese amber scorpions of the family Palaeoburmesebuthidae Lourenco, 2015 (Scorpiones). Arachnida, Rivista Aracnologica Italiana, 13, 11 - 21." type="journal article" year="2017">Lourenço & Rossi, 2017</bibRefCitation>
|
||
). Nonetheless, we confirm the presence of five pairs of lateral ocelli in
|
||
<taxonomicName authority="(Xuan et al., 2022)" baseAuthorityName="Xuan" baseAuthorityYear="2022" box="[304,660,876,902]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="spinipedis">
|
||
<emphasis box="[304,321,877,902]" italics="true" pageId="8" pageNumber="72">B</emphasis>
|
||
.
|
||
<emphasis box="[333,447,876,902]" italics="true" pageId="8" pageNumber="72">spinipedis</emphasis>
|
||
(
|
||
<bibRefCitation author="Xuan, Q. & Cai, C. Y. & Huang, D. Y." box="[459,652,876,902]" pageId="8" pageNumber="72" pagination="105165" refId="ref14060" refString="Xuan, Q., Cai, C. Y. & Huang, D. Y. (2022) A new Palaeoburmesebuthidae scorpion from mid-Cretaceous Burmese amber (Arachnida: Scorpiones: Buthoidea). Cretaceous Research, 134, 105165. https: // doi. org / 10.1016 / j. cretres. 2022.105165" type="journal article" year="2022">
|
||
Xuan
|
||
<emphasis box="[526,573,876,902]" italics="true" pageId="8" pageNumber="72">et al</emphasis>
|
||
., 2022
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. We do not regard this as a new feature since the fourth and fifth pairs of minor lateral ocelli are greatly reduced in size and are likely to be ignored by previous authors. Our examination of studied specimens reveals that
|
||
<taxonomicName box="[526,627,1016,1042]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="bellus">
|
||
<emphasis box="[526,543,1017,1042]" italics="true" pageId="8" pageNumber="72">B</emphasis>
|
||
.
|
||
<emphasis box="[560,627,1016,1042]" italics="true" pageId="8" pageNumber="72">bellus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Lourenco & Beigel" baseAuthorityYear="2015" box="[644,770,1016,1042]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="kobberti">
|
||
<emphasis box="[644,661,1017,1042]" italics="true" pageId="8" pageNumber="72">B</emphasis>
|
||
.
|
||
<emphasis box="[677,770,1016,1042]" italics="true" pageId="8" pageNumber="72">kobberti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Lourenco & Velten" baseAuthorityYear="2016" box="[136,270,1051,1077]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="fleissneri">
|
||
<emphasis box="[136,153,1052,1077]" italics="true" pageId="8" pageNumber="72">B</emphasis>
|
||
.
|
||
<emphasis box="[167,270,1051,1077]" italics="true" pageId="8" pageNumber="72">fleissneri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Lourenco & Velten" baseAuthorityYear="2016" box="[283,481,1051,1077]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="larafleissnerae">
|
||
<emphasis box="[283,300,1052,1077]" italics="true" pageId="8" pageNumber="72">B</emphasis>
|
||
.
|
||
<emphasis box="[313,481,1051,1077]" italics="true" pageId="8" pageNumber="72">larafleissnerae</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName baseAuthorityName="Lourenco" baseAuthorityYear="2015" box="[541,665,1051,1077]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="muelleri">
|
||
<emphasis box="[541,558,1052,1077]" italics="true" pageId="8" pageNumber="72">B</emphasis>
|
||
.
|
||
<emphasis box="[572,665,1051,1077]" italics="true" pageId="8" pageNumber="72">muelleri</emphasis>
|
||
</taxonomicName>
|
||
share this character.
|
||
</paragraph>
|
||
<paragraph blockId="8.[136,778,158,2092]" pageId="8" pageNumber="72">
|
||
<bibRefCitation author="Lourenco, W. R. & Velten, J." box="[189,502,1121,1147]" pageId="8" pageNumber="72" pagination="2 - 10" refId="ref13575" refString="Lourenco, W. R. & Velten, J. (2017) One more new genus and species of fossil scorpion from Burmese Cretaceous amber belonging to the family Palaeoburmesebuthidae (Scorpiones). Arachnida, Rivista Aracnologica Italiana, 13, 2 - 10." type="journal article" year="2017">Lourenço & Velten (2017)</bibRefCitation>
|
||
reported another fossil from Burmese amber, which is assigned to a new genus and species,
|
||
<emphasis box="[292,300,1192,1217]" italics="true" pageId="8" pageNumber="72">i</emphasis>
|
||
.
|
||
<emphasis box="[307,319,1192,1217]" italics="true" pageId="8" pageNumber="72">e</emphasis>
|
||
.,
|
||
<taxonomicName authorityName="XUAN & CAI & HUANG" authorityYear="2023" baseAuthorityName="XUAN & CAI & HUANG" baseAuthorityYear="2023" box="[347,653,1191,1217]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Spinoburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="pohli">
|
||
<emphasis box="[347,653,1191,1217]" italics="true" pageId="8" pageNumber="72">Spinoburmesebuthus pohli</emphasis>
|
||
</taxonomicName>
|
||
. Recently,
|
||
<bibRefCitation author="Lourenco, W. R." box="[136,333,1226,1253]" pageId="8" pageNumber="72" pagination="1 - 6" refId="ref13280" refString="Lourenco, W. R. (2021) Further comments on the elements of the family Palaeoburmesebuthidae Lourenco, 2015 with description of a new species of Spinoburmesebuthus Lourenco, 2017 from Early Cretaceous Burmite amber (Scorpiones). Faunitaxys, 9 (17), 1 - 6. https: // hal. science / hal- 03358766 v 2" type="journal article" year="2021">Lourenço (2021)</bibRefCitation>
|
||
recorded the second species of this genus,
|
||
<taxonomicName authorityName="XUAN & CAI & HUANG" authorityYear="2023" baseAuthorityName="Lourenco" baseAuthorityYear="2021" box="[219,385,1261,1288]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="knodelorum">
|
||
<emphasis box="[219,233,1261,1287]" italics="true" pageId="8" pageNumber="72">S</emphasis>
|
||
.
|
||
<emphasis box="[251,385,1261,1287]" italics="true" pageId="8" pageNumber="72">knodelorum</emphasis>
|
||
</taxonomicName>
|
||
. The most important character of
|
||
<taxonomicName box="[136,369,1296,1322]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Spinoburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[136,369,1296,1322]" italics="true" pageId="8" pageNumber="72">Spinoburmesebuthus</emphasis>
|
||
</taxonomicName>
|
||
claimed by
|
||
<bibRefCitation author="Lourenco, W. R." box="[514,706,1296,1323]" pageId="8" pageNumber="72" pagination="1 - 6" refId="ref13280" refString="Lourenco, W. R. (2021) Further comments on the elements of the family Palaeoburmesebuthidae Lourenco, 2015 with description of a new species of Spinoburmesebuthus Lourenco, 2017 from Early Cretaceous Burmite amber (Scorpiones). Faunitaxys, 9 (17), 1 - 6. https: // hal. science / hal- 03358766 v 2" type="journal article" year="2021">Lourenço (2021)</bibRefCitation>
|
||
is the presence of subaculear tubercle. However, in another paper Lourenço presented a line-drawing of the adult male metasoma of
|
||
<taxonomicName baseAuthorityName="Lourenco" baseAuthorityYear="2015" box="[298,641,1401,1427]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Palaeoburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="knodeli">
|
||
<emphasis box="[298,641,1401,1427]" italics="true" pageId="8" pageNumber="72">Palaeoburmesebuthus knodeli</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Lourenco, W. R." pageId="8" pageNumber="72" pagination="51 - 54" refId="ref13243" refString="Lourenco, W. R. (2018) A further new species of Palaeoburmesebuthus Lourenco, 2002 (Scorpiones: Palaeoburmesebuthidae) from Burmite. Revista Iberica de Aracnologia, 32, 51 - 54." type="journal article" year="2018">Lourenço, 2018</bibRefCitation>
|
||
: fig. 4) which clearly shows a subaculear tubercle. In addition, a small subaculear tubercle also found in the juvenile individual of
|
||
<taxonomicName authority="(Xuan, Cai & Huang, 2022)" baseAuthorityName="Xuan, Cai & Huang" baseAuthorityYear="2022" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="spinipedis">
|
||
<emphasis box="[450,467,1507,1532]" italics="true" pageId="8" pageNumber="72">B</emphasis>
|
||
.
|
||
<emphasis box="[487,601,1506,1532]" italics="true" pageId="8" pageNumber="72">spinipedis</emphasis>
|
||
(
|
||
<bibRefCitation author="Xuan, Q. & Cai, C. Y. & Huang, D. Y." pageId="8" pageNumber="72" pagination="105165" refId="ref14060" refString="Xuan, Q., Cai, C. Y. & Huang, D. Y. (2022) A new Palaeoburmesebuthidae scorpion from mid-Cretaceous Burmese amber (Arachnida: Scorpiones: Buthoidea). Cretaceous Research, 134, 105165. https: // doi. org / 10.1016 / j. cretres. 2022.105165" type="journal article" year="2022">Xuan, Cai & Huang, 2022</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Thus, this feature become a contradictory identification key, and we examined the additional materials of
|
||
<taxonomicName box="[170,391,1611,1637]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[170,391,1611,1637]" italics="true" pageId="8" pageNumber="72">Betaburmesebuthus</emphasis>
|
||
</taxonomicName>
|
||
in this paper to contribute to the solution of this taxonomic confusion. We conclude that the subaculear tubercle is present in
|
||
<taxonomicName box="[550,771,1681,1707]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[550,771,1681,1707]" italics="true" pageId="8" pageNumber="72">Betaburmesebuthus</emphasis>
|
||
</taxonomicName>
|
||
, which is difficult to discover in immature individuals but distinct in adult individuals. The two species assigned to alleged
|
||
<taxonomicName box="[263,496,1786,1812]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Spinoburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[263,496,1786,1812]" italics="true" pageId="8" pageNumber="72">Spinoburmesebuthus</emphasis>
|
||
</taxonomicName>
|
||
show many similarities with
|
||
<taxonomicName box="[200,421,1821,1847]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[200,421,1821,1847]" italics="true" pageId="8" pageNumber="72">Betaburmesebuthus</emphasis>
|
||
</taxonomicName>
|
||
and are the large specimens, as our materials,
|
||
<taxonomicName authority="XUAN & CAI & HUANG, 2023" authorityName="XUAN & CAI & HUANG" authorityYear="2023" box="[339,444,1856,1883]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="fuscus" status="sp. nov.">
|
||
<emphasis box="[339,356,1857,1882]" italics="true" pageId="8" pageNumber="72">B</emphasis>
|
||
.
|
||
<emphasis box="[374,444,1856,1882]" italics="true" pageId="8" pageNumber="72">fuscus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[454,548,1856,1882]" pageId="8" pageNumber="72">
|
||
<taxonomicNameLabel box="[454,548,1856,1882]" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(NIGP200653) and
|
||
<emphasis box="[136,153,1892,1917]" italics="true" pageId="8" pageNumber="72">B</emphasis>
|
||
. sp. (NIGP201157), which all exhibit the obvious subaculear tubercle. As a result, we here merge the genus
|
||
<taxonomicName box="[136,369,1961,1987]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Spinoburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[136,369,1961,1987]" italics="true" pageId="8" pageNumber="72">Spinoburmesebuthus</emphasis>
|
||
</taxonomicName>
|
||
into
|
||
<taxonomicName box="[437,658,1961,1987]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="genus" status="syn. nov.">
|
||
<emphasis box="[437,658,1961,1987]" italics="true" pageId="8" pageNumber="72">Betaburmesebuthus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[669,778,1961,1987]" pageId="8" pageNumber="72">
|
||
<taxonomicNameLabel box="[669,778,1961,1987]" pageId="8" pageNumber="72">syn. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and transfer the two species of
|
||
<taxonomicName box="[544,777,1996,2022]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Spinoburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[544,777,1996,2022]" italics="true" pageId="8" pageNumber="72">Spinoburmesebuthus</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName box="[168,389,2031,2057]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[168,389,2031,2057]" italics="true" pageId="8" pageNumber="72">Betaburmesebuthus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis box="[407,415,2032,2057]" italics="true" pageId="8" pageNumber="72">i</emphasis>
|
||
.
|
||
<emphasis box="[421,433,2032,2057]" italics="true" pageId="8" pageNumber="72">e</emphasis>
|
||
.,
|
||
<taxonomicName authorityName="XUAN & CAI & HUANG" authorityYear="2023" baseAuthorityName="XUAN & CAI & HUANG" baseAuthorityYear="2023" box="[458,551,2031,2058]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Spinoburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="pohli" status="comb. nov.">
|
||
<emphasis box="[458,475,2032,2057]" italics="true" pageId="8" pageNumber="72">B</emphasis>
|
||
.
|
||
<emphasis box="[493,551,2031,2057]" italics="true" pageId="8" pageNumber="72">pohli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[561,692,2031,2057]" pageId="8" pageNumber="72">
|
||
<taxonomicNameLabel box="[561,692,2031,2057]" pageId="8" pageNumber="72" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName authorityName="XUAN & CAI & HUANG" authorityYear="2023" baseAuthorityName="Lourenco" baseAuthorityYear="2021" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="knodelorum" status="comb. nov.">
|
||
<emphasis box="[754,771,2032,2057]" italics="true" pageId="8" pageNumber="72">B</emphasis>
|
||
.
|
||
<emphasis box="[136,270,2066,2092]" italics="true" pageId="8" pageNumber="72">knodelorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[277,405,2066,2092]" pageId="8" pageNumber="72">
|
||
<taxonomicNameLabel box="[277,405,2066,2092]" pageId="8" pageNumber="72" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="8.[810,1452,158,1301]" pageId="8" pageNumber="72">
|
||
The description of denticles of pedipalp fingers for
|
||
<taxonomicName box="[810,1031,194,220]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[810,1031,194,220]" italics="true" pageId="8" pageNumber="72">Betaburmesebuthus</emphasis>
|
||
</taxonomicName>
|
||
remains ambiguous, which could be attributed to the fact that the original authors observed this structure only from the lateral view (
|
||
<bibRefCitation author="Lourenco, W. R." box="[1220,1403,266,292]" pageId="8" pageNumber="72" pagination="1 - 6" refId="ref13280" refString="Lourenco, W. R. (2021) Further comments on the elements of the family Palaeoburmesebuthidae Lourenco, 2015 with description of a new species of Spinoburmesebuthus Lourenco, 2017 from Early Cretaceous Burmite amber (Scorpiones). Faunitaxys, 9 (17), 1 - 6. https: // hal. science / hal- 03358766 v 2" type="journal article" year="2021">Lourenço, 2021</bibRefCitation>
|
||
: fig. 16). We provide clear fluorescence and confocal photos from the frontal view and suggest that the absence of any conspicuous accessory granules are artifacts. Remarkably, another Burmese amber fossil,
|
||
<taxonomicName authority="(Lourenco & Velten, 2016 c)" baseAuthorityName="Lourenco & Velten" baseAuthorityYear="2016" class="Arachnida" family="Buthidae" genus="Archaeoananteroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="maderai">
|
||
<emphasis italics="true" pageId="8" pageNumber="72">Archaeoananteroides maderai</emphasis>
|
||
(
|
||
<bibRefCitation author="Lourenco, W. R. & Velten, J." box="[924,1243,446,472]" pageId="8" pageNumber="72" pagination="2 - 9" refId="ref13528" refString="Lourenco, W. R. & Velten, J. (2016 c) A new genus and species of fossil scorpion from Burmese Cretaceous amber (Scorpiones: Buthoidea: Buthidae). Arachnida, Rivista Aracnologica Italiana, 10, 2 - 9." type="journal article" year="2016">Lourenço & Velten, 2016c</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
shows the same state in denticle rows with our studied specimens and this species is likely to be a member of
|
||
<taxonomicName box="[1205,1426,518,544]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1205,1426,518,544]" italics="true" pageId="8" pageNumber="72">Betaburmesebuthus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="8.[810,1452,158,1301]" pageId="8" pageNumber="72">
|
||
<bibRefCitation author="Santiago-Blay, J. A. & Soleglad, M. E. & Fet, V. & Craig, P. R. & Smith, L. B." box="[863,1176,554,581]" pageId="8" pageNumber="72" pagination="4 - 52" refId="ref13726" refString="Santiago-Blay, J. A., Soleglad, M. E., Fet, V., Craig, P. R. & Smith, L. B. (2022) Two new species of Cretaceous scorpions from Burmese amber (Arachnida: Scorpiones: Palaeoburmesebuthidae). The Excitement of Biology, 10 (1), 4 - 52." type="journal article" year="2022">
|
||
Santiago-Blay
|
||
<emphasis box="[1033,1085,554,580]" italics="true" pageId="8" pageNumber="72">et al</emphasis>
|
||
. (2022)
|
||
</bibRefCitation>
|
||
indicated that the tibial spurs III and IV are variable in
|
||
<taxonomicName box="[1230,1451,590,616]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1230,1451,590,616]" italics="true" pageId="8" pageNumber="72">Betaburmesebuthus</emphasis>
|
||
</taxonomicName>
|
||
(present in three species, absent or reduced in other three species). A detailed examination of our material, however, revealed the presence of tibial spurs III and IV is fixed in members of
|
||
<taxonomicName box="[1041,1262,734,760]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1041,1262,734,760]" italics="true" pageId="8" pageNumber="72">Betaburmesebuthus</emphasis>
|
||
</taxonomicName>
|
||
. We suggest that such an absent or reduced condition were probably due to the poor preservation or are artifacts of observation. Furthermore, the tibial spurs of immature individuals are usually undeveloped and not easily to identify.
|
||
</paragraph>
|
||
<paragraph blockId="8.[810,1452,158,1301]" pageId="8" pageNumber="72">
|
||
Another interesting character is telotarsus armature. After the examination of all material, we observed
|
||
<specimenCount pageId="8" pageNumber="72" type="generic" typeStatus="types">two types</specimenCount>
|
||
: few ventrosubmedian setae and a ventromedian spinules row versus numerous ventrosubmedian setae. The feature is rarely mentioned in published papers and should be regarded as a diagnostic character, despite that the telotarsus armature was considered an important taxonomic character in extant scorpion at higher-level taxa (
|
||
<bibRefCitation author="Soleglad, M. E. & Fet, V." box="[817,1064,1202,1229]" pageId="8" pageNumber="72" pagination="1 - 175" refId="ref13888" refString="Soleglad, M. E. & Fet, V. (2003) High-level systematics and phylogeny of the extant scorpions (Scorpiones: Orthosterni). Euscorpius, 11, 1 - 175. https: // doi. org / 10.18590 / euscorpius. 2003. vol 2003. iss 11.1" type="journal article" year="2003">Soleglad & Fet, 2003</bibRefCitation>
|
||
). Lacking more distinct diagnostic characters, we refrain from establishing a new genus to divide the two forms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |