321 lines
31 KiB
XML
321 lines
31 KiB
XML
<document id="E712705594EE82F2573B736442C416D0" ID-DOI="10.5252/cryptogamie-bryologie2023v44a1" ID-ISSN="1776-0992" ID-Zenodo-Dep="7822577" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1681312727720" checkinUser="felipe" docAuthor="Thouvenot, Louis" docDate="2023" docId="CD138796FFC6FFBCFC102496FBD5F8EE" docLanguage="en" docName="CryptogamieBryologie.44.1.1-60.pdf" docOrigin="Cryptogamie, Bryologie 20 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.4878.3.2" docStyle="DocumentStyle:3F4A000F5B047ABA0EAB2943AA9373AE.3:CryptogamieBryologie.2019-.journal_article" docStyleId="3F4A000F5B047ABA0EAB2943AA9373AE" docStyleName="CryptogamieBryologie.2019-.journal_article" docStyleVersion="3" docTitle="Heteroscyphus cornutistipulus Thouvenot 2023, comb. nov." docType="treatment" docVersion="9" lastPageNumber="21" masterDocId="312AFFEEFFD3FFAAFFDE2540FF9AFFE2" masterDocTitle="A taxonomic revision of the Lophocoleaceae Vanden Berghen (Marchantiophyta) of New Caledonia" masterLastPageNumber="60" masterPageNumber="1" pageNumber="20" updateTime="1707318641535" updateUser="guilherme" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="A168677818C6041D4ADD1003E61CE9B0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F3D03ECD4FE2E74B2DF3B2DF91E32F74">
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<mods:title id="40E12B7E71E62E8C577B549D9ABCE361">A taxonomic revision of the Lophocoleaceae Vanden Berghen (Marchantiophyta) of New Caledonia</mods:title>
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</mods:titleInfo>
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<mods:name id="2BA5CE5AEF3263F209CFD78A54DF250C" type="personal">
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<mods:roleTerm id="5491B4654573DB9FB4D8F874871E8981">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2683A9DC068D0B23695C4BE86B23D017">Thouvenot, Louis</mods:namePart>
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<mods:affiliation id="C11830FD1313FA59FFE3DED49BCF5044">Saint Léon, 66000 Perpignan (France) thouvenot. louis @ orange. fr</mods:affiliation>
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<mods:nameIdentifier id="9F7642A1D358485A3949F65498B4C8FF" type="email">louis@orange.fr</mods:nameIdentifier>
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<mods:typeOfResource id="18CE02609C032C48811A4A94B0AB5228">text</mods:typeOfResource>
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<mods:titleInfo id="942164712C3B239135F8AE1039DDBD92">
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<mods:title id="A76A4E9C370D78BF1B4169728246D712">Cryptogamie, Bryologie</mods:title>
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</mods:titleInfo>
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<mods:part id="0469212B0C23DA54F1DA9607B80E2FFE">
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<mods:date id="ED0BA9C778178137D364EB1637A1A633">2023</mods:date>
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<mods:detail id="413DD6F9A5E43DADC20C32C1499575D2" type="pubDate">
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<mods:number id="464EA875442FC8E5502289627C49BE68">2023-01-16</mods:number>
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</mods:detail>
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<mods:detail id="D40DDA17DC9DEED7B054B76D27CC2D89" type="volume">
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<mods:number id="9A73190EE93CC6F4FC76C1E5084AB2E2">20</mods:number>
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<mods:detail id="FAFC55D0D676C75E8D1084AA6187CB37" type="issue">
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<mods:start id="61E2BE111B1099A84DDD0D7BAAD9C9D5">1</mods:start>
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<mods:url id="780C15804AD8CB7DEA857B17EA17BB0F">http://dx.doi.org/10.11646/zootaxa.4878.3.2</mods:url>
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</mods:location>
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<mods:classification id="45973638FCAFCC8CF1DDE2486C37BA33">journal article</mods:classification>
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<mods:identifier id="2A04A09F22B1AA4DE21B669E460DB9A8" type="DOI">10.5252/cryptogamie-bryologie2023v44a1</mods:identifier>
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<mods:identifier id="E9400091AC57A4FD8ED37267BCE5417B" type="ISSN">1776-0992</mods:identifier>
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<mods:identifier id="4C23D6C6F39D4D22D9B19F395E1C3D9E" type="Zenodo-Dep">7822577</mods:identifier>
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<treatment id="CD138796FFC6FFBCFC102496FBD5F8EE" LSID="urn:lsid:plazi:treatment:CD138796FFC6FFBCFC102496FBD5F8EE" httpUri="http://treatment.plazi.org/id/CD138796FFC6FFBCFC102496FBD5F8EE" lastPageId="22" lastPageNumber="21" pageId="21" pageNumber="20">
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<subSubSection id="0DA0650BFFC6FFBFFC102496FA84FDF3" pageId="21" pageNumber="20" type="nomenclature">
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<paragraph id="45053680FFC6FFBFFC102496FA84FDF3" blockId="21.[958,1310,470,529]" pageId="21" pageNumber="20">
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<heading id="1E4D81ECFFC6FFBFFC102496FA84FDF3" centered="true" fontSize="11" level="2" pageId="21" pageNumber="20" reason="4">
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<taxonomicName id="82BA4D03FFC6FFBFFC102496FB02FDF2" ID-CoL="BLRH4" authority="(Steph.) Thouvenot" authorityName="Thouvenot" authorityYear="2023" baseAuthorityName="Thouvenot" baseAuthorityYear="2023" class="Jungermanniopsida" family="Lophocoleaceae" genus="Heteroscyphus" kingdom="Plantae" order="Jungermanniales" pageId="21" pageNumber="20" phylum="Marchantiophyta" rank="species" species="cornutistipulus" status="comb. nov.">
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<emphasis id="77CEEA92FFC6FFBFFC102496FA94FE13" bold="true" box="[974,1294,470,497]" italics="true" pageId="21" pageNumber="20">Heteroscyphus cornutistipulus</emphasis>
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(Steph.) Thouvenot
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</taxonomicName>
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,
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<taxonomicNameLabel id="6CFD57E9FFC6FFBFFB7D24B6FA84FDF3" box="[1187,1310,502,529]" pageId="21" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0DA0650BFFC6FFBFFBE12756FB07FDD3" box="[1087,1181,534,561]" pageId="21" pageNumber="20" type="description">
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<paragraph id="45053680FFC6FFBFFBE12756FB07FDD3" blockId="21.[1087,1181,534,561]" box="[1087,1181,534,561]" pageId="21" pageNumber="20">
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<heading id="1E4D81ECFFC6FFBFFBE12756FB07FDD3" box="[1087,1181,534,561]" centered="true" fontSize="11" level="3" pageId="21" pageNumber="20" reason="3">
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(
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<figureCitation id="DD812A05FFC6FFBFFB992756FB0EFDD2" box="[1095,1172,534,560]" captionStart="FIG" captionStartId="23.[132,143,1564,1582]" captionTargetBox="[132,1455,212,1519]" captionTargetId="figure-292@23.[132,1455,212,1519]" captionTargetPageId="23" captionText="FIG. 10. — Heteroscyphus cornutistipulus (Steph.) Thouvenot,comb.nov.:A, C, shoot portions, ventral view;B, leaf; D-G, underleaves;H, mid-leaf cells.All drawn from the lectotype of Chiloscyphus cornutistipulus Steph. (G00069490). Scale bars: A-E, G, 500 µm; F, 500 µm; H, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.7822606" httpUri="https://zenodo.org/record/7822606/files/figure.png" pageId="21" pageNumber="20">Fig. 10</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0DA0650BFFC6FFBFFC9D270FFC4BFD60" pageId="21" pageNumber="20" type="reference_group">
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<paragraph id="45053680FFC6FFBFFC9D270FFC4BFD60" blockId="21.[812,1455,591,695]" pageId="21" pageNumber="20">
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<treatmentCitation id="C41B1091FFC6FFBFFC9D270FFC57FD60" author="Steph." journal="Species Hepaticarum" page="303" pageId="21" pageNumber="20" score="0.0033280253576670055" volume="6" year="1922">
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<taxonomicName id="82BA4D03FFC6FFBFFC9D270FFB0DFD85" ID-CoL="5XSDR" authority="Steph." authorityName="Steph." box="[835,1175,591,615]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Chiloscyphus" kingdom="Plantae" order="Jungermanniales" pageId="21" pageNumber="20" phylum="Marchantiophyta" rank="species" species="cornutistipulus">
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<emphasis id="77CEEA92FFC6FFBFFC9D270FFBD7FD85" box="[835,1101,591,615]" italics="true" pageId="21" pageNumber="20">Chiloscyphus cornutistipulus</emphasis>
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Steph.
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</taxonomicName>
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,
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<bibCitation id="90903EECFFC6FFBFFB7A270FFC57FD60" journal="Species Hepaticarum" page="303" pageId="21" pageNumber="20" score="0.0033280253576670055" volume="6" year="1922">
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<emphasis id="77CEEA92FFC6FFBFFB7A270FFAF2FD85" box="[1188,1384,591,615]" italics="true" pageId="21" pageNumber="20">Species Hepaticarum</emphasis>
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6: 303 (
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<bibRefCitation id="212B4B71FFC6FFBFFCEB2729FC59FD60" author="STEPHANI F." box="[821,963,617,642]" pageId="21" pageNumber="20" refId="ref42340" refString="STEPHANI F. 1922. - Species Hepaticarum 6. Geneve & Bale, George & Cie, 763 p." type="book" year="1922">Stephani 1922</bibRefCitation>
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)
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</bibCitation>
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</treatmentCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="0DA0650BFFC6FFBFFC06272AFC6BFD63" box="[984,1009,618,641]" pageId="21" pageNumber="20" type="multiple">
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<paragraph id="45053680FFC6FFBFFC06272AFC6BFD63" blockId="21.[812,1455,591,695]" box="[984,1009,618,641]" pageId="21" pageNumber="20">—</paragraph>
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</subSubSection>
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<subSubSection id="0DA0650BFFC6FFBFFC28272AFB35FCF9" pageId="21" pageNumber="20" type="materials_examined">
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<paragraph id="45053680FFC6FFBFFC28272AFC5BFD55" blockId="21.[812,1455,591,695]" pageId="21" pageNumber="20">
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<materialsCitation id="F5D23CDDFFC6FFBFFC28272AFC5BFD55" collectingDate="1909-11-01" collectionCode="G" collectorName="Franc" country="New Caledonia" location="Mts. Koghis" pageId="21" pageNumber="20" specimenCode="G00069490" specimenCount="1" typeStatus="lectotype">
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Type:
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<emphasis id="77CEEA92FFC6FFBFFBEF272AFB51FD61" bold="true" box="[1073,1227,618,643]" pageId="21" pageNumber="20">
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<collectingCountry id="3DAD7610FFC6FFBFFBEF272AFB51FD61" box="[1073,1227,618,643]" name="New Caledonia" pageId="21" pageNumber="20">New Caledonia</collectingCountry>
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</emphasis>
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.
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<emphasis id="77CEEA92FFC6FFBFFB07272AFAAFFD63" box="[1241,1333,618,641]" italics="true" pageId="21" pageNumber="20">
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<collectorName id="E84F5356FFC6FFBFFB07272AFA94FD63" box="[1241,1294,618,641]" pageId="21" pageNumber="20">Franc</collectorName>
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s.n.
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</emphasis>
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(
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<typeStatus id="9A018822FFC6FFBFFA9F2729FAE6FD63" box="[1345,1404,617,641]" pageId="21" pageNumber="20">lecto-</typeStatus>
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, here designated,
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<collectionCode id="23ABAE45FFC6FFBFFC4527C4FC37FD7E" box="[923,941,644,668]" pageId="21" pageNumber="20">G</collectionCode>
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[“Nlle Calédonie,
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<location id="4065605BFFC6FFBFFB8727C4FB54FD7E" LSID="urn:lsid:plazi:treatment:CD138796FFC6FFBCFC102496FBD5F8EE:4065605BFFC6FFBFFB8727C4FB54FD7E" box="[1113,1230,644,668]" country="New Caledonia" name="Mts. Koghis" pageId="21" pageNumber="20">Mts. Koghis</location>
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”,
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<date id="31041040FFC6FFBFFB0127C4FAD9FD7E" box="[1247,1347,644,668]" pageId="21" pageNumber="20" value="1909-11-01">
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<collectingDate id="2140E9A8FFC6FFBFFB0127C4FAD9FD7E" box="[1247,1347,644,668]" pageId="21" pageNumber="20" value="1909-11-01">1.XI.1909</collectingDate>
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</date>
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,
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<emphasis id="77CEEA92FFC6FFBFFA9427C5FA3DFD7E" box="[1354,1447,644,668]" italics="true" pageId="21" pageNumber="20">Franc 166</emphasis>
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,
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<specimenCode id="151C9EFBFFC6FFBFFCF327DEFC3FFD54" box="[813,933,670,694]" pageId="21" pageNumber="20">G00069490</specimenCode>
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]!).
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</materialsCitation>
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</paragraph>
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<paragraph id="45053680FFC6FFBFFCF3278FFB35FCF9" blockId="21.[811,1456,718,795]" pageId="21" pageNumber="20">
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FURTHER SPECIMEN EXAMINED. —
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<materialsCitation id="F5D23CDDFFC6FFBFFBAB278EFB35FCF9" collectedFrom="on dead wood in wet forest" collectingDate="2016-09-26" collectionCode="PC" collectorName="Thouvenot" country="New Caledonia" elevation="855" location="Dogny" municipality="Sarramea" pageId="21" pageNumber="20" specimenCode="PC0779841" specimenCount="1" stateProvince="South Province">
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<emphasis id="77CEEA92FFC6FFBFFBAB278EFA96FD05" bold="true" box="[1141,1292,718,743]" pageId="21" pageNumber="20">
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<collectingCountry id="3DAD7610FFC6FFBFFBAB278EFA96FD05" box="[1141,1292,718,743]" name="New Caledonia" pageId="21" pageNumber="20">New Caledonia</collectingCountry>
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</emphasis>
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.
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<collectingRegion id="877EF862FFC6FFBFFAC6278EFA36FD04" box="[1304,1452,718,742]" country="New Caledonia" name="Province Sud" pageId="21" pageNumber="20">South Province</collectingRegion>
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,
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<collectingMunicipality id="A561ACFAFFC6FFBFFCF527A9FC1DFCE3" box="[811,903,745,769]" pageId="21" pageNumber="20">Sarraméa</collectingMunicipality>
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,
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<location id="4065605BFFC6FFBFFC4E27A9FC4EFCE3" LSID="urn:lsid:plazi:treatment:CD138796FFC6FFBCFC102496FBD5F8EE:4065605BFFC6FFBFFC4E27A9FC4EFCE3" box="[912,980,745,769]" country="New Caledonia" municipality="Sarramea" name="Dogny" pageId="21" pageNumber="20" stateProvince="South Province">Dogny</location>
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,
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<quantity id="82429B65FFC6FFBFFC0327A8FB84FCE2" box="[989,1054,744,768]" metricMagnitude="2" metricUnit="m" metricValue="8.55" pageId="21" pageNumber="20" unit="m" value="855.0">
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<elevation id="CE97D1B3FFC6FFBFFC0327A8FB84FCE2" box="[989,1054,744,768]" metricMagnitude="2" metricUnit="m" metricValue="8.55" pageId="21" pageNumber="20" unit="m" value="855.0">855 m</elevation>
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</quantity>
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,
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<collectedFrom id="F6A6757DFFC6FFBFFBF927A9FAB7FCE2" box="[1063,1325,744,768]" pageId="21" pageNumber="20">on dead wood in wet forest</collectedFrom>
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,
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<date id="31041040FFC6FFBFFAE827A8FA30FCE2" box="[1334,1450,744,768]" pageId="21" pageNumber="20" value="2016-09-26">
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<collectingDate id="2140E9A8FFC6FFBFFAE827A8FA30FCE2" box="[1334,1450,744,768]" pageId="21" pageNumber="20" value="2016-09-26">26.IX.2016</collectingDate>
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</date>
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,
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<emphasis id="77CEEA92FFC6FFBFFCF52643FC73FCF9" box="[811,1001,771,795]" italics="true" pageId="21" pageNumber="20">
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<collectorName id="E84F5356FFC6FFBFFCF52643FC17FCF9" box="[811,909,771,795]" pageId="21" pageNumber="20">Thouvenot</collectorName>
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NC3283
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</emphasis>
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(
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<collectionCode id="23ABAE45FFC6FFBFFC262643FB80FCF9" box="[1016,1050,771,795]" pageId="21" pageNumber="20">PC</collectionCode>
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[
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<specimenCode id="151C9EFBFFC6FFBFFBFE2643FB03FCF9" box="[1056,1177,771,795]" pageId="21" pageNumber="20">PC0779841</specimenCode>
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]).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="0DA0650BFFC6FFBFFCF32673FBDBFC4D" pageId="21" pageNumber="20" type="distribution">
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<paragraph id="45053680FFC6FFBFFCF32673FC7CFC9D" blockId="21.[813,1455,819,895]" pageId="21" pageNumber="20">
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DISTRIBUTION IN
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<collectingCountry id="3DAD7610FFC6FFBFFC3C2673FB0AFCAB" box="[994,1168,819,843]" name="New Caledonia" pageId="21" pageNumber="20">NEW CALEDONIA</collectingCountry>
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. — Only known from scattered localities in
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<collectingRegion id="877EF862FFC6FFBFFC00260DFBEDFC87" box="[990,1143,845,869]" country="New Caledonia" name="Province Sud" pageId="21" pageNumber="20">South Province</collectingRegion>
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, on dead wood in wet forest at medium elevation.
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</paragraph>
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<paragraph id="45053680FFC6FFBFFCF326D8FBDBFC4D" blockId="21.[813,1089,919,943]" box="[813,1089,919,943]" pageId="21" pageNumber="20">TOTAL RANGE. — Endemic.</paragraph>
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</subSubSection>
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<subSubSection id="0DA0650BFFC6FFBFFCF32695FAAEF809" pageId="21" pageNumber="20" type="description">
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<paragraph id="45053680FFC6FFBFFCF32695FC27FC0F" blockId="21.[813,957,981,1007]" box="[813,957,981,1007]" pageId="21" pageNumber="20">
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<heading id="1E4D81ECFFC6FFBFFCF32695FC27FC0F" box="[813,957,981,1007]" fontSize="9" level="3" pageId="21" pageNumber="20" reason="5">DESCRIPTION</heading>
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</paragraph>
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<paragraph id="45053680FFC6FFBFFCF326B4FC3BFBEC" blockId="21.[813,929,1012,1038]" box="[813,929,1012,1038]" pageId="21" pageNumber="20">Dioecious.</paragraph>
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<paragraph id="45053680FFC6FFBFFCF32174FCF1FBAC" blockId="21.[813,875,1076,1102]" box="[813,875,1076,1102]" pageId="21" pageNumber="20">
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<heading id="1E4D81ECFFC6FFBFFCF32174FCF1FBAC" box="[813,875,1076,1102]" fontSize="11" level="3" pageId="21" pageNumber="20" reason="9">
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<emphasis id="77CEEA92FFC6FFBFFCF32174FCF1FBAC" box="[813,875,1076,1102]" italics="true" pageId="21" pageNumber="20">Habit</emphasis>
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</heading>
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</paragraph>
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<paragraph id="45053680FFC6FFBFFCF32114FA2BFB2C" blockId="21.[810,1457,1108,1230]" pageId="21" pageNumber="20">Plants medium sized, branching lateral-intercalary; wet shoots 3.50-4.00 mm wide, somewhat complanate,with leaves alternate, horizontally spreading, slightly convex, antical margins free, separated by 2 cortical cell files, shortly decurrent on the stem.</paragraph>
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<paragraph id="45053680FFC6FFBFFCF321B4FCF5FAEF" blockId="21.[813,879,1268,1293]" box="[813,879,1268,1293]" pageId="21" pageNumber="20">
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<heading id="1E4D81ECFFC6FFBFFCF321B4FCF5FAEF" box="[813,879,1268,1293]" fontSize="11" level="3" pageId="21" pageNumber="20" reason="9">
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<emphasis id="77CEEA92FFC6FFBFFCF321B4FCF5FAEF" box="[813,879,1268,1293]" italics="true" pageId="21" pageNumber="20">Leaves</emphasis>
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</heading>
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</paragraph>
|
|
<paragraph id="45053680FFC6FFBFFCF32053FAD9FA8F" blockId="21.[813,1456,1299,1389]" pageId="21" pageNumber="20">
|
|
<quantity id="82429B65FFC6FFBFFCF32053FC54FACC" box="[813,974,1299,1326]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="21" pageNumber="20" unit="mm" value="1.9" valueMax="2.2" valueMin="1.6">1.60-2.20 mm</quantity>
|
|
long, 1.20-2.00 mm wide, oval to oblong, slightly longer than wide, apices entire to slightly retuse, widely rounded, margins entire, hardly crenulate.
|
|
</paragraph>
|
|
<paragraph id="45053680FFC6FFBFFCF320D3FCC4FA4F" blockId="21.[813,862,1427,1453]" box="[813,862,1427,1453]" pageId="21" pageNumber="20">
|
|
<heading id="1E4D81ECFFC6FFBFFCF320D3FCC4FA4F" box="[813,862,1427,1453]" fontSize="11" level="3" pageId="21" pageNumber="20" reason="9">
|
|
<emphasis id="77CEEA92FFC6FFBFFCF320D3FCC4FA4F" box="[813,862,1427,1453]" italics="true" pageId="21" pageNumber="20">Cells</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="45053680FFC6FFBFFCF320F3FA36FA0F" blockId="21.[810,1455,1459,1517]" pageId="21" pageNumber="20">Median leaf cells rounded-hexagonal 40-80 µm wide, with thin walls and small to medium acute trigones up to 9 µm wide.</paragraph>
|
|
<paragraph id="45053680FFC6FFBFFCF32352FC33F9CE" blockId="21.[813,937,1554,1580]" box="[813,937,1554,1580]" pageId="21" pageNumber="20">
|
|
<heading id="1E4D81ECFFC6FFBFFCF32352FC33F9CE" box="[813,937,1554,1580]" fontSize="11" level="3" pageId="21" pageNumber="20" reason="9">
|
|
<emphasis id="77CEEA92FFC6FFBFFCF32352FC33F9CE" box="[813,937,1554,1580]" italics="true" pageId="21" pageNumber="20">Underleaves</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="45053680FFC6FFBFFCF32372FBF1F92E" blockId="21.[811,1456,1586,1741]" pageId="21" pageNumber="20">Narrowly to conspicuously connate on single or both sides to the closest leaves, connation 1-6 cells wide, underleaves 3-5 times as wide as the stem, bifid, lobes narrowly triangular, set wide apart, sinus lunate, disc transversely elongate, 1-2 short teeth on both lateral margins.</paragraph>
|
|
<paragraph id="45053680FFC6FFBFFCF323B2FC36F8EE" blockId="21.[813,940,1778,1804]" box="[813,940,1778,1804]" pageId="21" pageNumber="20">
|
|
<heading id="1E4D81ECFFC6FFBFFCF323B2FC36F8EE" box="[813,940,1778,1804]" fontSize="11" level="3" pageId="21" pageNumber="20" reason="9">
|
|
<emphasis id="77CEEA92FFC6FFBFFCF323B2FC36F8EE" box="[813,940,1778,1804]" italics="true" pageId="21" pageNumber="20">Gametangia</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="45053680FFC6FFBFFCF32252FAAEF809" blockId="21.[811,1457,1810,2027]" pageId="21" pageNumber="20">
|
|
Androecia and gynoecia on short leafless lateral branches; androecia in spikes made of 6 pairs of bracts very smaller than the normal leaves and inflated; female bracts not seen, female bracteoles 1/3-1/2 bifid, with 1 tooth on both sides; perianths cyathiform,
|
|
<quantity id="82429B65FFC6FFBFFC6F22D1FB8FF84E" box="[945,1045,1937,1964]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="20" unit="mm" value="2.2">2.20 mm</quantity>
|
|
long,
|
|
<quantity id="82429B65FFC6FFBFFB8422D1FB24F84E" box="[1114,1214,1937,1964]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="20" unit="mm" value="1.2">1.20 mm</quantity>
|
|
wide, 3-plicate, plicae rounded, mouth wide, lobate-laciniate, margins roughly toothed; calyptrae 2/3 shorter than the perianths.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<caption id="11C56608FFC5FFBCFF5A20EDFDFAFA0E" ID-DOI="http://doi.org/10.5281/zenodo.7822604" ID-Zenodo-Dep="7822604" httpUri="https://zenodo.org/record/7822604/files/figure.png" pageId="22" pageNumber="21" startId="22.[132,143,1453,1471]" targetBox="[137,1455,215,1407]" targetPageId="22" targetType="figure">
|
|
<paragraph id="45053680FFC5FFBCFF5A20EDFDFAFA0E" blockId="22.[132,1457,1453,1516]" pageId="22" pageNumber="21">
|
|
FIG. 9. —
|
|
<taxonomicName id="82BA4D03FFC5FFBCFF0920EDFDCFFA5D" authority="(Steph.) Thouvenot" authorityName="Thouvenot" baseAuthorityName="Steph." box="[215,597,1453,1471]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Heteroscyphus" kingdom="Plantae" order="Jungermanniales" pageId="22" pageNumber="21" phylum="Marchantiophyta" rank="species" species="confertus">
|
|
<emphasis id="77CEEA92FFC5FFBCFF0920EDFE31FA5D" box="[215,427,1453,1471]" italics="true" pageId="22" pageNumber="21">Heteroscyphus confertus</emphasis>
|
|
(Steph.) Thouvenot
|
|
</taxonomicName>
|
|
:
|
|
<emphasis id="77CEEA92FFC5FFBCFD8020EDFDF1FA5D" bold="true" box="[606,619,1453,1471]" pageId="22" pageNumber="21">A</emphasis>
|
|
, shoot portion, ventral view;
|
|
<emphasis id="77CEEA92FFC5FFBCFCB820EDFCE9FA5D" bold="true" box="[870,883,1453,1471]" pageId="22" pageNumber="21">B</emphasis>
|
|
,
|
|
<emphasis id="77CEEA92FFC5FFBCFCA120EDFC11FA5D" bold="true" box="[895,907,1453,1471]" pageId="22" pageNumber="21">E</emphasis>
|
|
,
|
|
<emphasis id="77CEEA92FFC5FFBCFC4920EDFC3FFA5D" bold="true" box="[919,933,1453,1471]" pageId="22" pageNumber="21">G</emphasis>
|
|
, underleaves;
|
|
<emphasis id="77CEEA92FFC5FFBCFBFD20EDFBABFA5D" bold="true" box="[1059,1073,1453,1471]" pageId="22" pageNumber="21">C</emphasis>
|
|
,
|
|
<emphasis id="77CEEA92FFC5FFBCFBE220EDFBD0FA5D" bold="true" box="[1084,1098,1453,1471]" pageId="22" pageNumber="21">D</emphasis>
|
|
,
|
|
<emphasis id="77CEEA92FFC5FFBCFB8820EDFBFBFA5D" bold="true" box="[1110,1121,1453,1471]" pageId="22" pageNumber="21">F</emphasis>
|
|
, leaves from different shoot segments.
|
|
<emphasis id="77CEEA92FFC5FFBCFF5A2084FF0BFA34" bold="true" box="[132,145,1476,1494]" pageId="22" pageNumber="21">A</emphasis>
|
|
,
|
|
<emphasis id="77CEEA92FFC5FFBCFF422084FF30FA34" bold="true" box="[156,170,1476,1494]" pageId="22" pageNumber="21">H</emphasis>
|
|
,
|
|
<emphasis id="77CEEA92FFC5FFBCFF6B2084FF21FA34" bold="true" box="[181,187,1476,1494]" pageId="22" pageNumber="21">I</emphasis>
|
|
, drawn from a syntype of
|
|
<taxonomicName id="82BA4D03FFC5FFBCFE462083FDC1FA34" box="[408,603,1475,1494]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Chiloscyphus" kingdom="Plantae" order="Jungermanniales" pageId="22" pageNumber="21" phylum="Marchantiophyta" rank="species" species="confertus">
|
|
<emphasis id="77CEEA92FFC5FFBCFE462083FDC1FA34" box="[408,603,1475,1494]" italics="true" pageId="22" pageNumber="21">Chiloscyphus confertus</emphasis>
|
|
</taxonomicName>
|
|
(PC0101951);
|
|
<emphasis id="77CEEA92FFC5FFBCFD062084FD7FFA34" bold="true" box="[728,741,1476,1494]" pageId="22" pageNumber="21">B</emphasis>
|
|
,
|
|
<emphasis id="77CEEA92FFC5FFBCFD2E2083FD64FA37" bold="true" box="[752,766,1475,1493]" pageId="22" pageNumber="21">C</emphasis>
|
|
, from
|
|
<emphasis id="77CEEA92FFC5FFBCFCEB2084FC42FA37" box="[821,984,1475,1494]" italics="true" pageId="22" pageNumber="21">Thouvenot NC2477</emphasis>
|
|
;
|
|
<emphasis id="77CEEA92FFC5FFBCFC3C2084FB9FFA37" bold="true" box="[994,1029,1475,1494]" pageId="22" pageNumber="21">D -G</emphasis>
|
|
, from the lectotype of
|
|
<taxonomicName id="82BA4D03FFC5FFBCFB1B2083FF27FA0E" authority="Steph." authorityName="Steph." class="Jungermanniopsida" family="Lophocoleaceae" genus="Chiloscyphus" kingdom="Plantae" order="Jungermanniales" pageId="22" pageNumber="21" phylum="Marchantiophyta" rank="species" species="rotundiphyllus">
|
|
<emphasis id="77CEEA92FFC5FFBCFB1B2083FA2AFA34" box="[1221,1456,1475,1494]" italics="true" pageId="22" pageNumber="21">Chiloscyphus rotundiphyllus</emphasis>
|
|
Steph.
|
|
</taxonomicName>
|
|
(G00069432). Scale bars: A-G, 1 mm; H, I, 50 µm.
|
|
</paragraph>
|
|
</caption>
|
|
<subSubSection id="0DA0650BFFC5FFBCFF5A2373FBD5F8EE" pageId="22" pageNumber="21" type="discussion">
|
|
<paragraph id="45053680FFC5FFBCFF5A2373FE9EF9A9" blockId="22.[132,260,1587,1613]" box="[132,260,1587,1613]" pageId="22" pageNumber="21">
|
|
<heading id="1E4D81ECFFC5FFBCFF5A2373FE9EF9A9" box="[132,260,1587,1613]" fontSize="9" level="3" pageId="22" pageNumber="21" reason="5">COMMENTS</heading>
|
|
</paragraph>
|
|
<paragraph id="45053680FFC5FFBCFF5A2312FBD5F8EE" blockId="22.[130,778,1618,2028]" lastBlockId="22.[811,1457,1586,1804]" pageId="22" pageNumber="21">
|
|
The
|
|
<typeStatus id="9A018822FFC5FFBCFF6F2313FF44F98F" box="[177,222,1619,1645]" pageId="22" pageNumber="21">type</typeStatus>
|
|
specimen has no gametangium, although the packet is annotated “
|
|
<emphasis id="77CEEA92FFC5FFBCFEDE2333FEA1F96E" box="[256,315,1651,1676]" italics="true" pageId="22" pageNumber="21">c. per.</emphasis>
|
|
”. A recently collected specimen has androecia and gynoecia (
|
|
<emphasis id="77CEEA92FFC5FFBCFE9C23D2FD89F94E" box="[322,531,1682,1708]" italics="true" pageId="22" pageNumber="21">Thouvenot NC3283</emphasis>
|
|
) and indicates that the species belongs in
|
|
<taxonomicName id="82BA4D03FFC5FFBCFE9B23F2FE4EF92E" box="[325,468,1714,1740]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Heteroscyphus" kingdom="Plantae" order="Jungermanniales" pageId="22" pageNumber="21" phylum="Marchantiophyta" rank="genus">
|
|
<emphasis id="77CEEA92FFC5FFBCFE9B23F2FE4EF92E" box="[325,468,1714,1740]" italics="true" pageId="22" pageNumber="21">Heteroscyphus</emphasis>
|
|
</taxonomicName>
|
|
. The fertile plant has smaller trigones but is otherwise vegetatively identical to the
|
|
<typeStatus id="9A018822FFC5FFBCFD652392FD73F90E" box="[699,745,1746,1772]" pageId="22" pageNumber="21">type</typeStatus>
|
|
of
|
|
<taxonomicName id="82BA4D03FFC5FFBCFF5A23B2FE39F8EE" box="[132,419,1778,1804]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Chiloscyphus" kingdom="Plantae" order="Jungermanniales" pageId="22" pageNumber="21" phylum="Marchantiophyta" rank="species" species="cornutistipulus">
|
|
<emphasis id="77CEEA92FFC5FFBCFF5A23B2FE39F8EE" box="[132,419,1778,1804]" italics="true" pageId="22" pageNumber="21">Chiloscyphus cornutistipulus</emphasis>
|
|
</taxonomicName>
|
|
. Like in
|
|
<taxonomicName id="82BA4D03FFC5FFBCFE2023B3FD09F8EE" box="[510,659,1778,1804]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Chiloscyphus" kingdom="Plantae" order="Jungermanniales" pageId="22" pageNumber="21" phylum="Marchantiophyta" rank="species" species="caledonicus">
|
|
<emphasis id="77CEEA92FFC5FFBCFE2023B3FD09F8EE" box="[510,659,1778,1804]" italics="true" pageId="22" pageNumber="21">H. caledonicus</emphasis>
|
|
</taxonomicName>
|
|
(see under that species), some vegetative characters of
|
|
<taxonomicName id="82BA4D03FFC5FFBCFD922252FECEF8A9" authority="(Steph.) Thouvenot" authorityName="Thouvenot" authorityYear="2023" baseAuthorityName="Steph." class="Jungermanniopsida" family="Lophocoleaceae" genus="Chiloscyphus" kingdom="Plantae" order="Jungermanniales" pageId="22" pageNumber="21" phylum="Marchantiophyta" rank="species" species="cornutistipulus" status="comb. nov.">
|
|
<emphasis id="77CEEA92FFC5FFBCFD922252FC9CF8CE" box="[588,774,1810,1836]" italics="true" pageId="22" pageNumber="21">H. cornutistipulus</emphasis>
|
|
(Steph.) Thouvenot
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel id="6CFD57E9FFC5FFBCFE822271FE4AF8AE" box="[348,464,1841,1868]" pageId="22" pageNumber="21" rank="species">comb. nov.</taxonomicNameLabel>
|
|
are chiloscyphoid, such as the alternate leaves and the narrow connation of the underleaves.
|
|
<taxonomicName id="82BA4D03FFC5FFBCFF452231FD17F869" authority="(Steph.) Thouvenot" authorityName="Thouvenot" authorityYear="2023" baseAuthorityName="Thouvenot" baseAuthorityYear="2023" box="[155,653,1905,1932]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Heteroscyphus" kingdom="Plantae" order="Jungermanniales" pageId="22" pageNumber="21" phylum="Marchantiophyta" rank="species" species="cornutistipulus" status="comb. nov.">
|
|
<emphasis id="77CEEA92FFC5FFBCFF452231FE21F869" box="[155,443,1905,1931]" italics="true" pageId="22" pageNumber="21">Heteroscyphus cornutistipulus</emphasis>
|
|
(Steph.) Thouvenot
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel id="6CFD57E9FFC5FFBCFD4B2231FC92F86E" box="[661,776,1905,1932]" pageId="22" pageNumber="21" rank="species">comb. nov.</taxonomicNameLabel>
|
|
is close to
|
|
<taxonomicName id="82BA4D03FFC5FFBCFF2522D1FD99F849" box="[251,515,1937,1963]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Chiloscyphus" kingdom="Plantae" order="Jungermanniales" pageId="22" pageNumber="21" phylum="Marchantiophyta" rank="species" species="trigonifolius">
|
|
<emphasis id="77CEEA92FFC5FFBCFF2522D1FD99F849" box="[251,515,1937,1963]" italics="true" pageId="22" pageNumber="21">Chiloscyphus trigonifolius</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="82BA4D03FFC5FFBCFDE322D2FD48F849" box="[573,722,1937,1963]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Heteroscyphus" kingdom="Plantae" order="Jungermanniales" pageId="22" pageNumber="21" phylum="Marchantiophyta" rank="species" species="succulentus">
|
|
<emphasis id="77CEEA92FFC5FFBCFDE322D2FD48F849" box="[573,722,1937,1963]" italics="true" pageId="22" pageNumber="21">H. succulentus</emphasis>
|
|
</taxonomicName>
|
|
, but
|
|
<taxonomicName id="82BA4D03FFC5FFBCFF5822F2FEB0F829" box="[134,298,1969,1996]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Chiloscyphus" kingdom="Plantae" order="Jungermanniales" pageId="22" pageNumber="21" phylum="Marchantiophyta" rank="species" species="trigonifolius">
|
|
<emphasis id="77CEEA92FFC5FFBCFF5822F2FEB0F829" box="[134,298,1969,1996]" italics="true" pageId="22" pageNumber="21">C. trigonifolius</emphasis>
|
|
</taxonomicName>
|
|
is easily separated from
|
|
<taxonomicName id="82BA4D03FFC5FFBCFD9B22F2FECDF809" authority="(Steph.) Thouvenot" authorityName="Thouvenot" authorityYear="2023" baseAuthorityName="Thouvenot" baseAuthorityYear="2023" class="Jungermanniopsida" family="Lophocoleaceae" genus="Heteroscyphus" kingdom="Plantae" order="Jungermanniales" pageId="22" pageNumber="21" phylum="Marchantiophyta" rank="species" species="cornutistipulus" status="comb. nov.">
|
|
<emphasis id="77CEEA92FFC5FFBCFD9B22F2FC93F829" box="[581,777,1969,1995]" italics="true" pageId="22" pageNumber="21">H. cornutistipulus</emphasis>
|
|
(Steph.) Thouvenot
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel id="6CFD57E9FFC5FFBCFEBF2291FE4FF80E" box="[353,469,2001,2028]" pageId="22" pageNumber="21" rank="species">comb. nov.</taxonomicNameLabel>
|
|
by its smaller size with shoots
|
|
<quantity id="82429B65FFC5FFBCFCF52372FC01F9AF" box="[811,923,1586,1613]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="22" pageNumber="21" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5-2 mm</quantity>
|
|
, strongly involute leaves when dry and the ovatetriangular leaf shape. Furthermore, the androecia are in spikes on normal-leaved shoots, male bracts being quite similar to the normal leaves.
|
|
<taxonomicName id="82BA4D03FFC5FFBCFC3523D2FB70F94E" box="[1003,1258,1682,1708]" class="Jungermanniopsida" family="Lophocoleaceae" genus="Heteroscyphus" kingdom="Plantae" order="Jungermanniales" pageId="22" pageNumber="21" phylum="Marchantiophyta" rank="species" species="succulentus">
|
|
<emphasis id="77CEEA92FFC5FFBCFC3523D2FB70F94E" box="[1003,1258,1682,1708]" italics="true" pageId="22" pageNumber="21">Heteroscyphus succulentus</emphasis>
|
|
</taxonomicName>
|
|
differs by its larger size, subopposite leaves, underleaves margins long laciniate instead of short toothed, bulging leaf cells without trigones, and crenulate leaf margins.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |