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<document id="F63F0CE0CFD4F0D609739722E958DA5D" ID-CLB-Dataset="37318" ID-DOI="10.5281/zenodo.245827" ID-GBIF-Dataset="69cf893c-7b35-48e8-ad78-d9775626f287" ID-ISSN="1175-5326" ID-Zenodo-Dep="245827" ID-ZooBank="9345C596-8656-4B5C-AD8C-2FACF4E9240C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484506177574" checkinUser="GgServerImporter" docAuthor="Blake, James A." docDate="2017" docId="8F2387DD065D0946FF31F94EFEE5F91F" docLanguage="en" docName="Zootaxa.4218.1.001-145" docOrigin="Zootaxa 4218 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Leodamas hyphalos Blake, 2017, new species" docType="treatment" docVersion="14" lastPageNumber="76" masterDocId="731AFFA50615090DFFA6FFEBFFDCFFC0" masterDocTitle="Polychaeta Orbiniidae from Antarctica, the Southern Ocean, the Abyssal Pacific Ocean, and off South America" masterLastPageNumber="145" masterPageNumber="1" pageNumber="73" updateTime="1698435671898" updateUser="ExternalLinkService">
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<mods:title id="6155B28B48183C608AB3687870481B8A">Polychaeta Orbiniidae from Antarctica, the Southern Ocean, the Abyssal Pacific Ocean, and off South America</mods:title>
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<mods:name id="8C5A929B9EB6083A02729AD2753D347D" type="personal">
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<mods:namePart id="74EA9B86DE95FC6B7CF3AF63B33F1732">Blake, James A.</mods:namePart>
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<mods:date id="DD9CDD09880E2E81173A92770D7EF0A5">2017</mods:date>
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<mods:identifier id="A2B3312D6387F339F8D2BDB37CC518A2" type="DOI">10.5281/zenodo.245827</mods:identifier>
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<treatment id="8F2387DD065D0946FF31F94EFEE5F91F" ID-DOI="http://doi.org/10.5281/zenodo.4901785" ID-GBIF-Taxon="127422017" ID-Zenodo-Dep="4901785" LSID="urn:lsid:plazi:treatment:8F2387DD065D0946FF31F94EFEE5F91F" httpUri="http://treatment.plazi.org/id/8F2387DD065D0946FF31F94EFEE5F91F" lastPageId="75" lastPageNumber="76" pageId="72" pageNumber="73">
<subSubSection id="4F906540065D0945FF31F94EFEE4F921" pageId="72" pageNumber="73" type="nomenclature">
<paragraph id="073536CB065D0945FF31F94EFDC2F900" blockId="72.[151,542,1701,1761]" box="[151,542,1701,1728]" pageId="72" pageNumber="73">
<heading id="5C7D81A7065D0945FF31F94EFDC2F900" bold="true" box="[151,542,1701,1728]" fontSize="11" level="1" pageId="72" pageNumber="73" reason="1">
<emphasis id="35FEEAD9065D0945FF31F94EFDC2F900" bold="true" box="[151,542,1701,1728]" pageId="72" pageNumber="73">
<taxonomicName id="C08A4D48065D0945FF31F94EFE5BF97F" ID-CoL="3T6BF" box="[151,391,1701,1727]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="72" pageNumber="73" phylum="Annelida" rank="species" species="hyphalos" status="sp. nov.">
<emphasis id="35FEEAD9065D0945FF31F94EFE5BF97F" bold="true" box="[151,391,1701,1727]" italics="true" pageId="72" pageNumber="73">Leodamas hyphalos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2ECD57A2065D0945FE28F94DFDC2F900" box="[398,542,1702,1728]" pageId="72" pageNumber="73" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="073536CB065D0945FF31F923FEE4F921" blockId="72.[151,542,1701,1761]" box="[151,312,1736,1761]" pageId="72" pageNumber="73">
<figureCitation id="9FB12A4E065D0945FF31F923FEE4F921" box="[151,312,1736,1761]" captionStart-0="FIGURE 33" captionStart-1="FIGURE 34" captionStartId-0="73.[151,249,1872,1894]" captionStartId-1="74.[151,249,1825,1847]" captionTargetBox-0="[158,1431,209,1834]" captionTargetBox-1="[164,1426,202,1792]" captionTargetId-0="figure@73.[151,1436,193,1851]" captionTargetId-1="figure@74.[151,1436,193,1803]" captionTargetPageId-0="73" captionTargetPageId-1="74" captionText-0="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." captionText-1="FIGURE 34. Leodamas hyphalos n. sp. (ANDEEP I, Sta. PS- 61 / 046 - 3, SEMs, JAB) A, anterior end, dorsal view; B, two thoracic neuropodia in anterior view; C, thoracic neuropodial uncini, detail; D, abdominal parapodium, anterior view; E, furcate seta, arrow denotes opening in tip of tyne." httpUri-0="https://zenodo.org/record/245859/files/figure.png" httpUri-1="https://zenodo.org/record/245860/files/figure.png" pageId="72" pageNumber="73">Figures 3334</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4F906540065D0945FF31F8FBFD0FF8E6" box="[151,723,1808,1830]" pageId="72" pageNumber="73" type="description">
<paragraph id="073536CB065D0945FF31F8FBFD0FF8E6" blockId="72.[151,723,1808,1830]" box="[151,723,1808,1830]" pageId="72" pageNumber="73">
<treatmentCitationGroup id="279A11E5065D0945FF31F8FBFD0FF8E6" box="[151,723,1808,1830]" pageId="72" pageNumber="73">
<taxonomicName id="C08A4D48065D0945FF31F8FBFEA4F8E6" ID-CoL="9CM26" authority="(Leodamas)" baseAuthorityName="Leodamas" box="[151,376,1808,1830]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="72" pageNumber="73" phylum="Annelida" rank="genus">
<emphasis id="35FEEAD9065D0945FF31F8FBFF26F8E6" box="[151,250,1808,1830]" italics="true" pageId="72" pageNumber="73">Scoloplos</emphasis>
(
<emphasis id="35FEEAD9065D0945FEAEF8FBFEACF8E6" box="[264,368,1808,1830]" italics="true" pageId="72" pageNumber="73">Leodamas</emphasis>
)
</taxonomicName>
spp.
<treatmentCitation id="862B10DA065D0945FE09F8FAFDAFF8E6" author="Hartman" box="[431,627,1808,1830]" page="108" pageId="72" pageNumber="73" year="1967">
<bibRefCitation id="631B4B3A065D0945FE09F8FAFD9FF8E6" author="Hartman" box="[431,579,1808,1830]" pageId="72" pageNumber="73" refString="Hartman, O. (1967) Polychaetous annelids collected by the USNS Eltanin and Staten Island cruises, chiefly from Antarctic seas. Allan Hancock Monographs in Marine Biology, 2, 1 - 387, 51 plates." type="book chapter" year="1967">Hartman 1967</bibRefCitation>
: 108
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="4F906540065D0945FF31F8B8FAECF817" pageId="72" pageNumber="73" type="materials_examined">
<paragraph id="073536CB065D0945FF31F8B8FAECF817" blockId="72.[151,1436,1875,2008]" pageId="72" pageNumber="73">
<emphasis id="35FEEAD9065D0945FF31F8B8FE5EF8AC" bold="true" box="[151,386,1875,1900]" pageId="72" pageNumber="73">Material examined.</emphasis>
Drake Passage,
<emphasis id="35FEEAD9065D0945FDE7F8B8FD48F8AB" box="[577,660,1875,1899]" italics="true" pageId="72" pageNumber="73">Eltanin</emphasis>
Sta. 4-112,
<date id="7334100B065D0945FCB9F8BFFC75F8AC" box="[799,937,1875,1900]" pageId="72" pageNumber="73" value="1962-07-20">20 Jul 1962</date>
, 56.03°S, 61.93° W,
<quantity id="C0729B2E065D0945FB3CF8BFFB2FF8AB" box="[1178,1267,1876,1900]" metricMagnitude="3" metricUnit="m" metricValue="4.008" pageId="72" pageNumber="73" unit="m" value="4008.0">4008 m</quantity>
, 13
<emphasis id="35FEEAD9065D0945FA80F8B8FF7CF850" bold="true" pageId="72" pageNumber="73">paratypes (</emphasis>
USNM 56456); Sta. 4-126,
<date id="7334100B065D0945FE79F89CFDB5F850" box="[479,617,1911,1936]" pageId="72" pageNumber="73" value="1962-07-29">29 Jul 1962</date>
, 57.20°S, 62.75°W,
<quantity id="C0729B2E065D0945FCF4F89CFC2EF84F" box="[850,1010,1911,1936]" metricMagnitude="3" metricUnit="m" metricValue="3.7695" metricValueMax="3.806" metricValueMin="3.7329999999999997" pageId="72" pageNumber="73" unit="m" value="3769.5" valueMax="3806.0" valueMin="3733.0">37333806 m</quantity>
,
<emphasis id="35FEEAD9065D0945FC59F89CFBBAF850" bold="true" box="[1023,1126,1911,1936]" pageId="72" pageNumber="73">holotype</emphasis>
(USNM 1013904); and 13
<emphasis id="35FEEAD9065D0945FF31F870FED2F874" bold="true" box="[151,270,1947,1972]" pageId="72" pageNumber="73">paratypes</emphasis>
(USNM 56457); Sta. 4-155,
<date id="7334100B065D0945FDFEF877FD0FF874" box="[600,723,1947,1972]" pageId="72" pageNumber="73" value="1962-08-17">17 Aug 62</date>
, 56.52°S, 63.25°W
<quantity id="C0729B2E065D0945FC12F877FBD0F873" box="[948,1036,1948,1972]" metricMagnitude="3" metricUnit="m" metricValue="3.9269999999999996" pageId="72" pageNumber="73" unit="m" value="3927.0">3927 m</quantity>
, 7
<emphasis id="35FEEAD9065D0945FB88F870FB79F874" bold="true" box="[1070,1189,1947,1972]" pageId="72" pageNumber="73">paratypes</emphasis>
(USNM 56458); R/V
<emphasis id="35FEEAD9065D0945FF31F854FEC9F817" box="[151,277,1983,2007]" italics="true" pageId="72" pageNumber="73">Polarstern,</emphasis>
ANDEEP I, Sta. PS-61/043-2, 3958 m (1, SEM, JAB); PS-61/046-3, 2888 m (1, SEM, JAB).
</paragraph>
</subSubSection>
<caption id="53F56643065C0944FF31F8BBFBB0F802" httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="73" pageNumber="74" targetBox="[158,1431,209,1834]" targetPageId="73">
<paragraph id="073536CB065C0944FF31F8BBFBB0F802" blockId="73.[151,1436,1872,1986]" pageId="73" pageNumber="74">
<emphasis id="35FEEAD9065C0944FF31F8BBFEC3F8A6" bold="true" box="[151,287,1872,1894]" pageId="73" pageNumber="74">FIGURE 33.</emphasis>
<taxonomicName id="C08A4D48065C0944FE80F8BBFE2DF8A6" box="[294,497,1872,1894]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="73" pageNumber="74" phylum="Annelida" rank="species" species="hyphalos" status="sp. nov.">
<emphasis id="35FEEAD9065C0944FE80F8BBFE2DF8A6" box="[294,497,1872,1894]" italics="true" pageId="73" pageNumber="74">Leodamas hyphalos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2ECD57A2065C0944FE51F8BBFDEDF8A6" box="[503,561,1872,1894]" pageId="73" pageNumber="74" rank="species">
<emphasis id="35FEEAD9065C0944FE51F8BBFDEDF8A6" bold="true" box="[503,561,1872,1894]" pageId="73" pageNumber="74">n. sp.</emphasis>
</taxonomicNameLabel>
(Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); EF, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view.
</paragraph>
</caption>
<caption id="53F56643065F0947FF31F8CAFD52F8B4" httpUri="https://zenodo.org/record/245860/files/figure.png" pageId="74" pageNumber="75" targetBox="[164,1426,202,1792]" targetPageId="74">
<paragraph id="073536CB065F0947FF31F8CAFD52F8B4" blockId="74.[151,1435,1825,1908]" pageId="74" pageNumber="75">
<emphasis id="35FEEAD9065F0947FF31F8CAFEFEF8F7" bold="true" box="[151,290,1825,1847]" pageId="74" pageNumber="75">FIGURE 34.</emphasis>
<taxonomicName id="C08A4D48065F0947FE8DF8CAFE24F8F7" box="[299,504,1825,1847]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="74" pageNumber="75" phylum="Annelida" rank="species" species="hyphalos" status="sp. nov.">
<emphasis id="35FEEAD9065F0947FE8DF8CAFE24F8F7" box="[299,504,1825,1847]" italics="true" pageId="74" pageNumber="75">Leodamas hyphalos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2ECD57A2065F0947FDA7F8CAFDE1F8F7" box="[513,573,1825,1847]" pageId="74" pageNumber="75" rank="species">
<emphasis id="35FEEAD9065F0947FDA7F8CAFDE1F8F7" bold="true" box="[513,573,1825,1847]" pageId="74" pageNumber="75">n. sp.</emphasis>
</taxonomicNameLabel>
(ANDEEP I, Sta. PS-61/046-3, SEMs, JAB) A, anterior end, dorsal view; B, two thoracic neuropodia in anterior view; C, thoracic neuropodial uncini, detail; D, abdominal parapodium, anterior view; E, furcate seta, arrow denotes opening in tip of tyne.
</paragraph>
</caption>
<subSubSection id="4F906540065F0946FF61F870FAC4FB04" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75" type="description">
<paragraph id="073536CB065F0946FF61F870FD54FF15" blockId="74.[151,1437,1947,2008]" lastBlockId="75.[151,1437,151,1760]" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75">
<emphasis id="35FEEAD9065F0947FF61F870FE85F874" bold="true" box="[199,345,1947,1972]" pageId="74" pageNumber="75">Description.</emphasis>
Majority of specimens small, incomplete;
<typeStatus id="D8318869065F0947FCFAF870FC62F874" box="[860,958,1947,1972]" pageId="74" pageNumber="75" type="holotype">holotype</typeStatus>
<quantity id="C0729B2E065F0947FC6CF870FBFAF873" box="[970,1062,1947,1972]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="74" pageNumber="75" unit="mm" value="5.5">5.5 mm</quantity>
long and
<quantity id="C0729B2E065F0947FB01F877FADFF873" box="[1191,1283,1948,1972]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="74" pageNumber="75" unit="mm" value="0.7">0.7 mm</quantity>
wide for 29 setigers; largest
<typeStatus id="D8318869065F0947FEEDF82BFE70F818" box="[331,428,1984,2008]" pageId="74" pageNumber="75" type="paratype">paratype</typeStatus>
9.0 mm long and
<quantity id="C0729B2E065F0947FDD2F854FD16F817" box="[628,714,1983,2008]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="74" pageNumber="75" unit="mm" value="0.6">0.6 mm</quantity>
wide for 34 setigers. A single, much larger incomplete specimen from ANDEEP I Sta. PS-61 046-3 broken into two parts with 82 setigers, measuring
<quantity id="C0729B2E065E0946FBFDFF7CFB77FF6F" box="[1115,1195,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="75" pageNumber="76" unit="mm" value="36.0">36 mm</quantity>
long and
<quantity id="C0729B2E065E0946FABAFF73FA82FF6F" box="[1308,1374,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="75" pageNumber="76" unit="mm" value="3.0">3 mm</quantity>
wide across thorax. Color in alcohol: light brown.
</paragraph>
<paragraph id="073536CB065E0946FF61FF34FAF3FE48" blockId="75.[151,1437,151,1760]" pageId="75" pageNumber="76">
Thoracic region with 1618 setigers, inflated in first 34 setigers (
<figureCitation id="9FB12A4E065E0946FC70FF34FBF1FF38" box="[982,1069,223,248]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
B), subsequent thoracic setigers depressed; transition to abdominal region gradual, with last three thoracic setigers having fewer setae (
<figureCitation id="9FB12A4E065E0946FA81FEEFFAA7FEDC" box="[1319,1403,260,285]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
C). Intersegmental annulations first present from setigers 89; these weakly developed, never prominent (
<figureCitation id="9FB12A4E065E0946FA83FECCFAA4FE80" box="[1317,1400,295,320]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
A). Branchiae from setiger 6 (
<figureCitation id="9FB12A4E065E0946FE18FEA7FDC6FEA4" box="[446,538,332,357]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Figs. 33</figureCitation>
A, 34A); thoracic and anterior abdominal branchiae basally inflated (
<figureCitation id="9FB12A4E065E0946FA83FEA7FA5DFEA4" box="[1317,1409,332,357]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Figs. 33</figureCitation>
A, 34A), subsequent abdominal branchiae expanded (
<figureCitation id="9FB12A4E065E0946FD68FE84FCFCFE48" box="[718,800,367,392]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
J); branchiae absent from far posterior setigers.
</paragraph>
<paragraph id="073536CB065E0946FF61FE7FFC15FE34" blockId="75.[151,1437,151,1760]" pageId="75" pageNumber="76">
Prostomium triangular in outline, pointed on anterior margin (
<figureCitation id="9FB12A4E065E0946FCDBFE7FFC04FE6C" box="[893,984,404,429]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Figs. 33</figureCitation>
A, 34A); eyespots absent; nuchal organs in SEM as transverse slits at border of prostomium and peristomium (
<figureCitation id="9FB12A4E065E0946FC14FE5CFC23FE10" box="[946,1023,439,464]" captionStart="FIGURE 34" captionStartId="74.[151,249,1825,1847]" captionTargetBox="[164,1426,202,1792]" captionTargetId="figure@74.[151,1436,193,1803]" captionTargetPageId="74" captionText="FIGURE 34. Leodamas hyphalos n. sp. (ANDEEP I, Sta. PS- 61 / 046 - 3, SEMs, JAB) A, anterior end, dorsal view; B, two thoracic neuropodia in anterior view; C, thoracic neuropodial uncini, detail; D, abdominal parapodium, anterior view; E, furcate seta, arrow denotes opening in tip of tyne." httpUri="https://zenodo.org/record/245860/files/figure.png" pageId="75" pageNumber="76">Fig 34</figureCitation>
A); proboscis divided into 34 lobes (
<figureCitation id="9FB12A4E065E0946FF39FE37FF2EFE34" box="[159,242,476,501]" captionStart="FIGURE 34" captionStartId="74.[151,249,1825,1847]" captionTargetBox="[164,1426,202,1792]" captionTargetId="figure@74.[151,1436,193,1803]" captionTargetPageId="74" captionText="FIGURE 34. Leodamas hyphalos n. sp. (ANDEEP I, Sta. PS- 61 / 046 - 3, SEMs, JAB) A, anterior end, dorsal view; B, two thoracic neuropodia in anterior view; C, thoracic neuropodial uncini, detail; D, abdominal parapodium, anterior view; E, furcate seta, arrow denotes opening in tip of tyne." httpUri="https://zenodo.org/record/245860/files/figure.png" pageId="75" pageNumber="76">Fig. 34</figureCitation>
A). Peristomium with one short achaetous ring (
<figureCitation id="9FB12A4E065E0946FCACFE37FCBAFE34" box="[778,870,476,501]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Figs. 33</figureCitation>
A, 34A).
</paragraph>
<paragraph id="073536CB065E0946FF61FE14FCC2FCD4" blockId="75.[151,1437,151,1760]" pageId="75" pageNumber="76">
Thoracic notopodia with minute postsetal lamellae on setigers 12, barely seen in SEM (
<figureCitation id="9FB12A4E065E0946FB17FE14FADEFDD8" box="[1201,1282,511,536]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
A), becoming larger more visible from setiger 3 (
<figureCitation id="9FB12A4E065E0946FD8EFDCFFDA7FDFC" box="[552,635,548,573]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
A), short at first, then elongating, becoming unusually long, fingerlike structures (
<figureCitation id="9FB12A4E065E0946FEB5FDACFEB3FDA0" box="[275,367,583,608]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Figs. 33</figureCitation>
AC, 34A), continuing through abdominal region. Thoracic neuropodia with setae arising from elongated thickened lobes (
<figureCitation id="9FB12A4E065E0946FE68FD87FDF0FD44" box="[462,556,620,645]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Figs. 33</figureCitation>
C, 34B); postsetal lamellae from setiger 511, attached to posterior border of upper one-third of setal lobe (
<figureCitation id="9FB12A4E065E0946FE56FD64FD93FD68" box="[496,591,655,680]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Figs. 33</figureCitation>
C, 34B); each lobe short triangular at first, then elongating in transitional region to form fingerlike lamella; a single subpodial lobe from setiger 1415, continuing through anterior abdominal segments (
<figureCitation id="9FB12A4E065E0946FE32FD3CFE36FD30" box="[404,490,727,752]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
C); absent from middle and posterior abdominal segments (
<figureCitation id="9FB12A4E065E0946FB05FD3CFB26FD30" box="[1187,1274,727,752]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
J). Abdominal neuropodia elongated, with short ventral cirrus (
<figureCitation id="9FB12A4E065E0946FD14FD17FCD9FCD4" box="[690,773,764,789]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
J).
</paragraph>
<paragraph id="073536CB065E0946FF61FCF4FAC4FB04" blockId="75.[151,1437,151,1760]" pageId="75" pageNumber="76">
Thoracic notopodia with fascicles of numerous crenulated capillaries and a single furcate seta; abdominal notopodia with three thin crenulated capillaries, two flail setae (
<figureCitation id="9FB12A4E065E0946FCD6FCAFFC1FFC9C" box="[880,963,836,861]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">
Fig.
<date id="7334100B065E0946FC00FCAFFC1FFC9C" box="[934,963,836,860]" pageId="75" pageNumber="76" value="1933-01">33</date>
</figureCitation>
I) and 23 furcate setae (
<figureCitation id="9FB12A4E065E0946FB47FCAFFAE1FC9C" box="[1249,1341,836,861]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Figs. 33</figureCitation>
G, 34E); furcate setae thin, delicate, easily broken, with unequal tynes connected by thin webbing composed of fine needles and shaft with faint annulations (
<figureCitation id="9FB12A4E065E0946FDB6FC67FDB8FC64" box="[528,612,908,933]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
G); with SEM, furcate setae with 910 needles on each side with lateral ones merging with tynes; each tyne with minute apical opening (
<figureCitation id="9FB12A4E065E0946FCDCFC44FC12FC08" box="[890,974,943,968]" captionStart="FIGURE 34" captionStartId="74.[151,249,1825,1847]" captionTargetBox="[164,1426,202,1792]" captionTargetId="figure@74.[151,1436,193,1803]" captionTargetPageId="74" captionText="FIGURE 34. Leodamas hyphalos n. sp. (ANDEEP I, Sta. PS- 61 / 046 - 3, SEMs, JAB) A, anterior end, dorsal view; B, two thoracic neuropodia in anterior view; C, thoracic neuropodial uncini, detail; D, abdominal parapodium, anterior view; E, furcate seta, arrow denotes opening in tip of tyne." httpUri="https://zenodo.org/record/245860/files/figure.png" pageId="75" pageNumber="76">Fig. 34</figureCitation>
E). Flail setae with thick, non-crenulated shaft bearing thin, crenulated tips (
<figureCitation id="9FB12A4E065E0946FD93FC3FFD51FC2C" box="[565,653,980,1005]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">
Fig.
<date id="7334100B065E0946FDC8FC3FFD51FC2C" box="[622,653,980,1004]" pageId="75" pageNumber="76" value="1933-01">33</date>
</figureCitation>
I). Thoracic neuropodia with dense fascicles of blunt-tipped uncini arranged in four rows (
<figureCitation id="9FB12A4E065E0946FE3EFC1CFE36FBD0" box="[408,490,1015,1040]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
BC); uncini of anterior row typically continuing ventrally below rows 2 and 3, then merging with row 4 forming a U-shape (
<figureCitation id="9FB12A4E065E0946FDCBFBF7FD1EFBF4" box="[621,706,1052,1077]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
D); in some neuropodia, rows 2 and three also joined ventrally; uncini of anterior row sharply curved, with 35 transverse ridges (
<figureCitation id="9FB12A4E065E0946FC26FBD4FC0EFB98" box="[896,978,1087,1112]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
E), with SEM ridges appearing irregular, angular (
<figureCitation id="9FB12A4E065E0946FF5AFB8FFE8DFBBC" box="[252,337,1124,1149]" captionStart="FIGURE 34" captionStartId="74.[151,249,1825,1847]" captionTargetBox="[164,1426,202,1792]" captionTargetId="figure@74.[151,1436,193,1803]" captionTargetPageId="74" captionText="FIGURE 34. Leodamas hyphalos n. sp. (ANDEEP I, Sta. PS- 61 / 046 - 3, SEMs, JAB) A, anterior end, dorsal view; B, two thoracic neuropodia in anterior view; C, thoracic neuropodial uncini, detail; D, abdominal parapodium, anterior view; E, furcate seta, arrow denotes opening in tip of tyne." httpUri="https://zenodo.org/record/245860/files/figure.png" pageId="75" pageNumber="76">Fig. 34</figureCitation>
C); uncini of posterior row not as sharply bent, with 910 transverse ridges (
<figureCitation id="9FB12A4E065E0946FB1AFB8FFACCFBBC" box="[1212,1296,1124,1149]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
F); 23 long crenulated capillaries present in superior position of last row of uncini (
<figureCitation id="9FB12A4E065E0946FC65FB6CFBC9FB60" box="[963,1045,1159,1184]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
BC, 34B). Abdominal neurosetae including 12 simple, blunt-tipped acicula (
<figureCitation id="9FB12A4E065E0946FD26FB47FD00FB04" box="[640,732,1196,1221]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Figs. 33</figureCitation>
H, 34D) and 23 thin, simple capillaries (
<figureCitation id="9FB12A4E065E0946FB0AFB47FB22FB04" box="[1196,1278,1196,1221]" captionStart="FIGURE 33" captionStartId="73.[151,249,1872,1894]" captionTargetBox="[158,1431,209,1834]" captionTargetId="figure@73.[151,1436,193,1851]" captionTargetPageId="73" captionText="FIGURE 33. Leodamas hyphalos n. sp. (Holotype USNM 1013904). A, anterior end, dorsal view; B, anterior end, left lateral view; C, posterior thoracic region in left lateral view showing transition to abdominal segments; D,. diagram of middle thoracic neuropodium showing arrangement of uncini (not to scale); E F, thoracic neuropodial uncini; G, furcate seta; H, abdominal neuropodial acicula; I, abdominal notopodial flail seta; J, abdominal parapodium, posterior view." httpUri="https://zenodo.org/record/245859/files/figure.png" pageId="75" pageNumber="76">Fig. 33</figureCitation>
J).
</paragraph>
</subSubSection>
<subSubSection id="4F906540065E0946FF61FB24FC9AFB28" box="[199,838,1231,1256]" pageId="75" pageNumber="76" type="etymology">
<paragraph id="073536CB065E0946FF61FB24FC9AFB28" blockId="75.[151,1437,151,1760]" box="[199,838,1231,1256]" pageId="75" pageNumber="76">
<emphasis id="35FEEAD9065E0946FF61FB24FE92FB28" bold="true" box="[199,334,1231,1256]" pageId="75" pageNumber="76">Etymology.</emphasis>
<taxonomicName id="C08A4D48065E0946FEF2FB3BFDD0FB28" authority="Greek" authorityName="Greek" box="[340,524,1231,1256]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="75" pageNumber="76" phylum="Annelida" rank="species" species="hyphalos">
<emphasis id="35FEEAD9065E0946FEF2FB3BFE65FB28" box="[340,441,1232,1256]" italics="true" pageId="75" pageNumber="76">hyphalos</emphasis>
: Greek
</taxonomicName>
for submerged, in the deep.
</paragraph>
</subSubSection>
<subSubSection id="4F906540065E0946FF61FB1FFEEDF958" pageId="75" pageNumber="76" type="discussion">
<paragraph id="073536CB065E0946FF61FB1FFEEDF958" blockId="75.[151,1437,151,1760]" pageId="75" pageNumber="76">
<emphasis id="35FEEAD9065E0946FF61FB1FFEE7FACD" bold="true" box="[199,315,1268,1293]" pageId="75" pageNumber="76">Remarks.</emphasis>
Of the six species of
<taxonomicName id="C08A4D48065E0946FD96FB1FFD7FFACC" box="[560,675,1268,1292]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="75" pageNumber="76" phylum="Annelida" rank="genus">
<emphasis id="35FEEAD9065E0946FD96FB1FFD7FFACC" box="[560,675,1268,1292]" italics="true" pageId="75" pageNumber="76">Leodamas</emphasis>
</taxonomicName>
encountered in this study,
<taxonomicName id="C08A4D48065E0946FC77FB1EFB8FFACC" box="[977,1107,1268,1292]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="75" pageNumber="76" phylum="Annelida" rank="species" species="hyphalos" status="sp. nov.">
<emphasis id="35FEEAD9065E0946FC77FB1EFB8FFACC" box="[977,1107,1268,1292]" italics="true" pageId="75" pageNumber="76">L. hyphalos</emphasis>
</taxonomicName>
<emphasis id="35FEEAD9065E0946FBFDFB1FFB46FACD" bold="true" box="[1115,1178,1268,1293]" pageId="75" pageNumber="76">
<taxonomicNameLabel id="2ECD57A2065E0946FBFDFB1FFB46FACD" box="[1115,1178,1268,1293]" pageId="75" pageNumber="76" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is the only one to bear flail setae in the abdominal notopodia.
<taxonomicName id="C08A4D48065E0946FDE9FAF3FCF3FAF0" box="[591,815,1304,1328]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="75" pageNumber="76" phylum="Annelida" rank="species" species="hyphalos" status="sp. nov.">
<emphasis id="35FEEAD9065E0946FDE9FAF3FCF3FAF0" box="[591,815,1304,1328]" italics="true" pageId="75" pageNumber="76">Leodamas hyphalos</emphasis>
</taxonomicName>
<emphasis id="35FEEAD9065E0946FC91FAFCFCAAFAF0" bold="true" box="[823,886,1303,1328]" pageId="75" pageNumber="76">
<taxonomicNameLabel id="2ECD57A2065E0946FC91FAFCFCAAFAF0" box="[823,886,1303,1328]" pageId="75" pageNumber="76" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is closest in morphology to
<taxonomicName id="C08A4D48065E0946FB1EFAF2FA88FAEF" box="[1208,1364,1304,1328]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="75" pageNumber="76" phylum="Annelida" rank="species" species="marginatus">
<emphasis id="35FEEAD9065E0946FB1EFAF2FA88FAEF" box="[1208,1364,1304,1328]" italics="true" pageId="75" pageNumber="76">L. marginatus</emphasis>
</taxonomicName>
in the nature and arrangement of the neuropodial uncini; both species have a few superior capillary setae dorsal to the posterior row of uncini. The thoracic neuropodial uncini of
<taxonomicName id="C08A4D48065E0946FC9CFA8AFC0AFAB7" box="[826,982,1376,1400]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="75" pageNumber="76" phylum="Annelida" rank="species" species="marginatus">
<emphasis id="35FEEAD9065E0946FC9CFA8AFC0AFAB7" box="[826,982,1376,1400]" italics="true" pageId="75" pageNumber="76">L. marginatus</emphasis>
</taxonomicName>
are arranged in three long vertical rows and with a short fourth row that continues ventral to the first three; in contrast, there are four long rows of uncini in
<taxonomicName id="C08A4D48065E0946FF31FA42FEC6FA00" box="[151,282,1448,1472]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="75" pageNumber="76" phylum="Annelida" rank="species" species="hyphalos" status="sp. nov.">
<emphasis id="35FEEAD9065E0946FF31FA42FEC6FA00" box="[151,282,1448,1472]" italics="true" pageId="75" pageNumber="76">L. hyphalos</emphasis>
</taxonomicName>
<emphasis id="35FEEAD9065E0946FE85FA4CFEBEFA00" bold="true" box="[291,354,1447,1472]" pageId="75" pageNumber="76">
<taxonomicNameLabel id="2ECD57A2065E0946FE85FA4CFEBEFA00" box="[291,354,1447,1472]" pageId="75" pageNumber="76" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
with the first and last often merged ventrally and forming a U-shape. Although the majority of specimens were small, the much larger specimen from Sta. PS-61 046-3 from lower slope depths of
<quantity id="C0729B2E065E0946FAB4FA27FAB4FA24" box="[1298,1384,1484,1508]" metricMagnitude="3" metricUnit="m" metricValue="2.888" pageId="75" pageNumber="76" unit="m" value="2888.0">2888 m</quantity>
was similar to
<taxonomicName id="C08A4D48065E0946FEAFFA1AFE79F9C7" box="[265,421,1520,1544]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="75" pageNumber="76" phylum="Annelida" rank="species" species="marginatus">
<emphasis id="35FEEAD9065E0946FEAFFA1AFE79F9C7" box="[265,421,1520,1544]" italics="true" pageId="75" pageNumber="76">L. marginatus</emphasis>
</taxonomicName>
in size. The smaller specimens of
<taxonomicName id="C08A4D48065E0946FC8DFA1AFC72F9C8" box="[811,942,1520,1544]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="75" pageNumber="76" phylum="Annelida" rank="species" species="hyphalos" status="sp. nov.">
<emphasis id="35FEEAD9065E0946FC8DFA1AFC72F9C8" box="[811,942,1520,1544]" italics="true" pageId="75" pageNumber="76">L. hyphalos</emphasis>
</taxonomicName>
<emphasis id="35FEEAD9065E0946FC13FA04FC28F9C8" bold="true" box="[949,1012,1519,1544]" pageId="75" pageNumber="76">
<taxonomicNameLabel id="2ECD57A2065E0946FC13FA04FC28F9C8" box="[949,1012,1519,1544]" pageId="75" pageNumber="76" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
all occurred at abyssal depths greater than
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, suggesting that the species may be size limited in the deeper parts of its habitat, possibly due to limited or patchy organic input to the abyssal sediments. The close similarity of
<taxonomicName id="C08A4D48065E0946FBB3F9D2FB44F990" box="[1045,1176,1592,1616]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="75" pageNumber="76" phylum="Annelida" rank="species" species="hyphalos" status="sp. nov.">
<emphasis id="35FEEAD9065E0946FBB3F9D2FB44F990" box="[1045,1176,1592,1616]" italics="true" pageId="75" pageNumber="76">L. hyphalos</emphasis>
</taxonomicName>
<emphasis id="35FEEAD9065E0946FB38F9DCFB00F990" bold="true" box="[1182,1244,1591,1616]" pageId="75" pageNumber="76">
<taxonomicNameLabel id="2ECD57A2065E0946FB38F9DCFB00F990" box="[1182,1244,1591,1616]" pageId="75" pageNumber="76" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
to
<taxonomicName id="C08A4D48065E0946FAA6F9D2FA47F98F" box="[1280,1435,1592,1616]" class="Polychaeta" family="Orbiniidae" genus="Leodamas" kingdom="Animalia" pageId="75" pageNumber="76" phylum="Annelida" rank="species" species="marginatus">
<emphasis id="35FEEAD9065E0946FAA6F9D2FA47F98F" box="[1280,1435,1592,1616]" italics="true" pageId="75" pageNumber="76">L. marginatus</emphasis>
</taxonomicName>
suggests that it may be a lower slope and abyssal sibling of the widely distributed and common shelf and upper slope species.
</paragraph>
</subSubSection>
<subSubSection id="4F906540065E0946FF61F94FFEE5F91F" pageId="75" pageNumber="76" type="distribution">
<paragraph id="073536CB065E0946FF61F94FFEE5F91F" blockId="75.[151,1437,151,1760]" pageId="75" pageNumber="76">
<emphasis id="35FEEAD9065E0946FF61F94FFEBCF97D" bold="true" box="[199,352,1700,1725]" pageId="75" pageNumber="76">Distribution.</emphasis>
Known only from the Drake Passage between South America and the Antarctic Peninsula;
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.
</paragraph>
</subSubSection>
</treatment>
</document>