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<document id="418608016E3E62875246A711D946D59E" ID-CLB-Dataset="37318" ID-DOI="10.5281/zenodo.245827" ID-GBIF-Dataset="69cf893c-7b35-48e8-ad78-d9775626f287" ID-ISSN="1175-5326" ID-Zenodo-Dep="245827" ID-ZooBank="9345C596-8656-4B5C-AD8C-2FACF4E9240C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484506177574" checkinUser="GgServerImporter" docAuthor="Blake, James A." docDate="2017" docId="8F2387DD061D0904FF31F92DFB83FD30" docLanguage="en" docName="Zootaxa.4218.1.001-145" docOrigin="Zootaxa 4218 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Berkeleyia Hartman, 1971 Emended" docType="treatment" docVersion="14" lastPageNumber="10" masterDocId="731AFFA50615090DFFA6FFEBFFDCFFC0" masterDocTitle="Polychaeta Orbiniidae from Antarctica, the Southern Ocean, the Abyssal Pacific Ocean, and off South America" masterLastPageNumber="145" masterPageNumber="1" pageNumber="9" updateTime="1698435671898" updateUser="ExternalLinkService">
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<mods:title id="0A440389781627C42850ED3574E03221">Polychaeta Orbiniidae from Antarctica, the Southern Ocean, the Abyssal Pacific Ocean, and off South America</mods:title>
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<mods:namePart id="1C638D5428742F2052BCCC53049C285F">Blake, James A.</mods:namePart>
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<mods:date id="3E9D50C566C8C79ADD4F6ED675931277">2017</mods:date>
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<treatment id="8F2387DD061D0904FF31F92DFB83FD30" ID-DOI="http://doi.org/10.5281/zenodo.4901729" ID-GBIF-Taxon="127422030" ID-Zenodo-Dep="4901729" LSID="urn:lsid:plazi:treatment:8F2387DD061D0904FF31F92DFB83FD30" httpUri="http://treatment.plazi.org/id/8F2387DD061D0904FF31F92DFB83FD30" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<subSubSection id="4F906540061D0905FF31F92DFD6DF920" box="[151,689,1734,1760]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="073536CB061D0905FF31F92DFD6DF920" blockId="8.[151,689,1734,1760]" box="[151,689,1734,1760]" pageId="8" pageNumber="9">
<heading id="5C7D81A7061D0905FF31F92DFD6DF920" bold="true" box="[151,689,1734,1760]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="35FEEAD9061D0905FF31F92DFD6DF920" bold="true" box="[151,689,1734,1760]" pageId="8" pageNumber="9">
Genus
<taxonomicName id="C08A4D48061D0905FF48F92DFD6DF920" ID-CoL="84MF8" authority="Hartman, 1971 Emended" authorityName="Hartman, 1971 Emended" box="[238,689,1734,1760]" class="Polychaeta" family="Orbiniidae" genus="Berkeleyia" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Annelida" rank="genus">
<emphasis id="35FEEAD9061D0905FF48F92DFEB3F920" bold="true" box="[238,367,1734,1760]" italics="true" pageId="8" pageNumber="9">Berkeleyia</emphasis>
<bibRefCitation id="631B4B3A061D0905FED0F92DFDEFF920" author="Hartman" box="[374,563,1734,1760]" pageId="8" pageNumber="9" refString="Hartman, O. (1971) Abyssal polychaetous annelids from the Mozambique Basin off Southeast Africa, with a compendium of abyssal polychaetous annelids from world-wide areas. Journal of the Fisheries Research Board, Canada, 28, 1407 - 1428." type="journal article" year="1971">Hartman, 1971</bibRefCitation>
Emended
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4F906540061D0905FF31F8E6FBD6F8E3" box="[151,1034,1805,1827]" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="073536CB061D0905FF31F8E6FBD6F8E3" blockId="8.[151,1034,1805,1827]" box="[151,1034,1805,1827]" pageId="8" pageNumber="9">
<materialsCitation id="B7E23C96061D0905FF31F8E6FBD6F8E3" ID-GBIF-Occurrence="1424727812" box="[151,1034,1805,1827]" country="Mozambique" elevation="4978" location="Mozambique" pageId="8" pageNumber="9" specimenCount="1" typeStatus="holotype">
<emphasis id="35FEEAD9061D0905FF31F8E6FEF9F8E3" bold="true" box="[151,293,1805,1827]" pageId="8" pageNumber="9">
<typeStatus id="D8318869061D0905FF31F8E6FF10F8E3" box="[151,204,1805,1827]" pageId="8" pageNumber="9">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="C08A4D48061D0905FE8DF8E6FD46F8E3" authority="Hartman, 1971" authorityName="Hartman" authorityYear="1971" box="[299,666,1805,1827]" class="Polychaeta" family="Orbiniidae" genus="Berkeleyia" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="profunda">
<emphasis id="35FEEAD9061D0905FE8DF8E6FE26F8E3" box="[299,506,1805,1827]" italics="true" pageId="8" pageNumber="9">Berkeleyia profunda</emphasis>
<bibRefCitation id="631B4B3A061D0905FDA6F8E5FD46F8E3" author="Hartman" box="[512,666,1805,1827]" pageId="8" pageNumber="9" refString="Hartman, O. (1971) Abyssal polychaetous annelids from the Mozambique Basin off Southeast Africa, with a compendium of abyssal polychaetous annelids from world-wide areas. Journal of the Fisheries Research Board, Canada, 28, 1407 - 1428." type="journal article" year="1971">Hartman, 1971</bibRefCitation>
</taxonomicName>
,
<collectingCountry id="7F9D765B061D0905FD02F8E6FCF6F8E3" box="[676,810,1805,1827]" name="Mozambique" pageId="8" pageNumber="9">Mozambique</collectingCountry>
Basin,
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<elevation id="8CA7D1F8061D0905FCD0F8E6FBDAF8E3" box="[886,1030,1805,1827]" metricMagnitude="3" metricUnit="m" metricValue="4.9775" metricValueMax="5.069" metricValueMin="4.886" pageId="8" pageNumber="9" unit="m" value="4977.5" valueMax="5069.0" valueMin="4886.0">48865069 m</elevation>
</quantity>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4F906540061D0904FF31F8A4FCCBFF15" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="073536CB061D0904FF31F8A4FCCBFF15" blockId="8.[151,1436,1871,2005]" lastBlockId="9.[151,1437,151,752]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="35FEEAD9061D0905FF31F8A4FECDF8A8" bold="true" box="[151,273,1871,1896]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
Prostomium pointed, conical; peristomium with one achaetous ring. Branchiae from posterior thoracic or abdominal segments. Thoracic noto- and neuropodia with one postsetal lobe, sometimes absent or inconspicuous on anteriormost setigers; subpodial lobes absent; abdominal setigers with simple noto- and neuropodia; neuropodia with or without ventral cirrus; interramal cirri absent. Thoracic noto- and neurosetae all capillaries. Abdominal notosetae include capillaries; pointed spines present or absent; furcate setae present or absent; neuropodia with capillaries and protruding acicular spines; or only spines.
</paragraph>
</subSubSection>
<subSubSection id="4F906540061C0904FF61FF34FC0CFE6D" pageId="9" pageNumber="10" type="discussion">
<paragraph id="073536CB061C0904FF61FF34FC0CFE6D" blockId="9.[151,1437,151,752]" pageId="9" pageNumber="10">
<emphasis id="35FEEAD9061C0904FF61FF34FEE7FF38" bold="true" box="[199,315,223,248]" pageId="9" pageNumber="10">Remarks.</emphasis>
The genus
<taxonomicName id="C08A4D48061C0904FE62FF0BFDE0FF38" box="[452,572,224,248]" class="Polychaeta" family="Orbiniidae" genus="Berkeleyia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="genus">
<emphasis id="35FEEAD9061C0904FE62FF0BFDE0FF38" box="[452,572,224,248]" italics="true" pageId="9" pageNumber="10">Berkeleyia</emphasis>
</taxonomicName>
was established by
<bibRefCitation id="631B4B3A061C0904FC80FF0BFC00FF38" author="Hartman" box="[806,988,224,248]" pageId="9" pageNumber="10" refString="Hartman, O. (1971) Abyssal polychaetous annelids from the Mozambique Basin off Southeast Africa, with a compendium of abyssal polychaetous annelids from world-wide areas. Journal of the Fisheries Research Board, Canada, 28, 1407 - 1428." type="journal article" year="1971">Hartman (1971)</bibRefCitation>
for a single species,
<taxonomicName id="C08A4D48061C0904FB72FF0AFA80FF38" box="[1236,1372,224,248]" class="Polychaeta" family="Orbiniidae" genus="Berkeleyia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="profunda">
<emphasis id="35FEEAD9061C0904FB72FF0AFA80FF38" box="[1236,1372,224,248]" italics="true" pageId="9" pageNumber="10">B. profunda</emphasis>
</taxonomicName>
from abyssal depths in the Mozambique Basin.
<taxonomicName id="C08A4D48061C0904FDD4FEEFFD36FEDC" box="[626,746,260,284]" class="Polychaeta" family="Orbiniidae" genus="Berkeleyia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="genus">
<emphasis id="35FEEAD9061C0904FDD4FEEFFD36FEDC" box="[626,746,260,284]" italics="true" pageId="9" pageNumber="10">Berkeleyia</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="C08A4D48061C0904FC72FEEFFBABFEDC" box="[980,1143,260,284]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="genus">
<emphasis id="35FEEAD9061C0904FC72FEEFFBABFEDC" box="[980,1143,260,284]" italics="true" pageId="9" pageNumber="10">Leitoscoloplos</emphasis>
</taxonomicName>
in having only camerated capillaries in thoracic neuropodia. The two genera differ markedly, however, in that species of
<emphasis id="35FEEAD9061C0904FB41FEC3FA8DFE80" box="[1255,1361,296,320]" italics="true" pageId="9" pageNumber="10">Berkeleyi</emphasis>
a have well-developed acicular spines protruding from abdominal neuropodia and sometimes in abdominal notopodia. Five species of
<taxonomicName id="C08A4D48061C0904FEE8FE9BFE1AFE48" box="[334,454,368,392]" class="Polychaeta" family="Orbiniidae" genus="Berkeleyia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="genus">
<emphasis id="35FEEAD9061C0904FEE8FE9BFE1AFE48" box="[334,454,368,392]" italics="true" pageId="9" pageNumber="10">Berkeleyia</emphasis>
</taxonomicName>
are now known, all from the Southern Hemisphere and all but one are from abyssal depths; four new species have been encountered as part the present study:
</paragraph>
</subSubSection>
<subSubSection id="4F906540061C0904FF61FE52FA41FE35" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="073536CB061C0904FF61FE52FC4FFE0F" blockId="9.[151,1437,151,752]" box="[199,915,439,464]" pageId="9" pageNumber="10">
<materialsCitation id="B7E23C96061C0904FF61FE52FC4FFE0F" ID-GBIF-Occurrence="1424727827" box="[199,915,439,464]" country="Mozambique" elevation="4968" location="Mozambique" pageId="9" pageNumber="10" specimenCount="1">
<taxonomicName id="C08A4D48061C0904FF61FE52FE22FE10" ID-CoL="LL62" authority="Hartman, 1971" authorityName="Hartman" authorityYear="1971" box="[199,510,440,464]" class="Polychaeta" family="Orbiniidae" genus="Berkeleyia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="profunda">
<emphasis id="35FEEAD9061C0904FF61FE52FE90FE10" box="[199,332,440,464]" italics="true" pageId="9" pageNumber="10">B. profunda</emphasis>
<bibRefCitation id="631B4B3A061C0904FEF5FE53FE22FE10" author="Hartman" box="[339,510,440,464]" pageId="9" pageNumber="10" refString="Hartman, O. (1971) Abyssal polychaetous annelids from the Mozambique Basin off Southeast Africa, with a compendium of abyssal polychaetous annelids from world-wide areas. Journal of the Fisheries Research Board, Canada, 28, 1407 - 1428." type="journal article" year="1971">Hartman, 1971</bibRefCitation>
</taxonomicName>
.
<collectingCountry id="7F9D765B061C0904FDAEFE5CFD40FE10" box="[520,668,439,464]" name="Mozambique" pageId="9" pageNumber="10">Mozambique</collectingCountry>
Basin,
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<elevation id="8CA7D1F8061C0904FD57FE5CFC53FE0F" box="[753,911,439,464]" metricMagnitude="3" metricUnit="m" metricValue="4.967499999999999" metricValueMax="5.069" metricValueMin="4.866" pageId="9" pageNumber="10" unit="m" value="4967.5" valueMax="5069.0" valueMin="4866.0">48665069 m</elevation>
</quantity>
.
</materialsCitation>
</paragraph>
<paragraph id="073536CB061C0904FF61FE36FA41FE35" blockId="9.[151,1437,151,752]" box="[199,1437,476,501]" pageId="9" pageNumber="10">
<materialsCitation id="B7E23C96061C0904FF61FE36FA41FE35" ID-GBIF-Occurrence="1424727814" box="[199,1437,476,501]" country="Iceland" location="South America" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Tierra del Fuego">
<taxonomicName id="C08A4D48061C0904FF61FE36FEEDFE34" ID-CoL="LL5Z" box="[199,305,476,500]" class="Polychaeta" family="Orbiniidae" genus="Berkeleyia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="heroae" status="sp. nov.">
<emphasis id="35FEEAD9061C0904FF61FE36FEEDFE34" box="[199,305,476,500]" italics="true" pageId="9" pageNumber="10">B. heroae</emphasis>
</taxonomicName>
,
<emphasis id="35FEEAD9061C0904FE99FE37FEA1FE35" bold="true" box="[319,381,476,501]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="2ECD57A2061C0904FE99FE37FEA1FE35" box="[319,381,476,501]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
<location id="02556010061C0904FE22FE37FDF3FE35" LSID="urn:lsid:plazi:treatment:8F2387DD061D0904FF31F92DFB83FD30:02556010061C0904FE22FE37FDF3FE35" box="[388,559,476,501]" country="Iceland" name="South America" pageId="9" pageNumber="10" stateProvince="Tierra del Fuego">South America</location>
, Staten
<collectingCountry id="7F9D765B061C0904FD20FE37FD11FE35" box="[646,717,476,501]" name="Iceland" pageId="9" pageNumber="10">Island</collectingCountry>
, off
<collectingRegion id="C54EF829061C0904FD58FE37FC65FE35" box="[766,953,476,501]" country="Argentina" name="Tierra del Fuego - Antartida e Islas del Atlantico Sur" pageId="9" pageNumber="10">Tierra del Fuego</collectingRegion>
, intertidal to shallow subtidal. (See below).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4F906540061C0904FF61FDEAFB83FD30" pageId="9" pageNumber="10" type="description">
<paragraph id="073536CB061C0904FF61FDEAFB36FDD8" blockId="9.[151,1437,151,752]" box="[199,1258,511,536]" pageId="9" pageNumber="10">
<taxonomicName id="C08A4D48061C0904FF61FDEAFE94FDD8" box="[199,328,512,536]" class="Polychaeta" family="Orbiniidae" genus="Berkeleyia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="abyssala" status="sp. nov.">
<emphasis id="35FEEAD9061C0904FF61FDEAFE94FDD8" box="[199,328,512,536]" italics="true" pageId="9" pageNumber="10">B. abyssala</emphasis>
</taxonomicName>
<emphasis id="35FEEAD9061C0904FEE9FE14FE51FDD8" bold="true" box="[335,397,511,536]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="2ECD57A2061C0904FEE9FE14FE51FDD8" box="[335,397,511,536]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Antarctic seas, Drake Passage and Weddell Sea,
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. (See below).
</paragraph>
<paragraph id="073536CB061C0904FF61FDCEFC10FDFD" blockId="9.[151,1437,151,752]" box="[199,972,548,573]" pageId="9" pageNumber="10">
<taxonomicName id="C08A4D48061C0904FF61FDCEFE8FFDFC" box="[199,339,548,572]" class="Polychaeta" family="Orbiniidae" genus="Berkeleyia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="weddellia" status="sp. nov.">
<emphasis id="35FEEAD9061C0904FF61FDCEFE8FFDFC" box="[199,339,548,572]" italics="true" pageId="9" pageNumber="10">B. weddellia</emphasis>
</taxonomicName>
<emphasis id="35FEEAD9061C0904FEFCFDCFFE44FDFD" bold="true" box="[346,408,548,573]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="2ECD57A2061C0904FEFCFDCFFE44FDFD" box="[346,408,548,573]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Antarctic sea, Weddell Sea,
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. (See below).
</paragraph>
<paragraph id="073536CB061C0904FF61FDA2FC51FDA0" blockId="9.[151,1437,151,752]" box="[199,909,583,608]" pageId="9" pageNumber="10">
<taxonomicName id="C08A4D48061C0904FF61FDA2FEE8FDA0" box="[199,308,584,608]" class="Polychaeta" family="Orbiniidae" genus="Berkeleyia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="hadala" status="sp. nov.">
<emphasis id="35FEEAD9061C0904FF61FDA2FF04FDA0" box="[199,216,585,608]" italics="true" pageId="9" pageNumber="10">B</emphasis>
.
<emphasis id="35FEEAD9061C0904FF40FDA3FEE8FDA0" box="[230,308,584,608]" italics="true" pageId="9" pageNumber="10">hadala</emphasis>
</taxonomicName>
<emphasis id="35FEEAD9061C0904FE9DFDACFEA5FDA0" bold="true" box="[315,377,583,608]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="2ECD57A2061C0904FE9DFDACFEA5FDA0" box="[315,377,583,608]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Peru-Chile Trench,
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(See below).
</paragraph>
<paragraph id="073536CB061C0904FF61FD87FB83FD30" blockId="9.[151,1437,151,752]" pageId="9" pageNumber="10">
<taxonomicName id="C08A4D48061C0904FF61FD87FE4FFD44" box="[199,403,620,644]" class="Polychaeta" family="Orbiniidae" genus="Berkeleyia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="heroae" status="sp. nov.">
<emphasis id="35FEEAD9061C0904FF61FD87FE4FFD44" box="[199,403,620,644]" italics="true" pageId="9" pageNumber="10">Berkeleyia heroae</emphasis>
</taxonomicName>
<emphasis id="35FEEAD9061C0904FE3DFD87FE07FD45" bold="true" box="[411,475,620,645]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="2ECD57A2061C0904FE3DFD87FE07FD45" box="[411,475,620,645]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from the four deep-water species in having the abdominal neuropodial spines with a distinctly bifurcated or notched tip, but otherwise all four of the new species and the type-species from the
<collectingCountry id="7F9D765B061C0904FF31FD5FFEF7FD0D" box="[151,299,692,717]" name="Mozambique" pageId="9" pageNumber="10">Mozambique</collectingCountry>
Basin form a well-defined group within the
<taxonomicName id="C08A4D48061C0904FC97FD5FFC70FD0D" box="[817,940,692,717]" class="Polychaeta" family="Orbiniidae" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Annelida" rank="family">Orbiniidae</taxonomicName>
. An additional new deep-water species has been discovered on a seamount in the Atlantic Ocean and will be described separately.
</paragraph>
</subSubSection>
</treatment>
</document>