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<mods:title id="7C919D04DAB225CBA0B86E4C479B67D6">- A-new-species-of-phytotelm-breeding-frog- (Anura: - Rhacophoridae) - from-the-Central-Highlands-of-Vietnam</mods:title>
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<mods:namePart id="BF031B03A28440E836D224C8D776A565">Rowley, Jodi J. L.</mods:namePart>
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<mods:namePart id="D50B0E35A9CF6A9234527650DCEDFBCB">Le, Duong Thi Thuy</mods:namePart>
<mods:affiliation id="FF813CFCC9FAC04DB5415DFDCFF6C796">University of Science, Faculty of Biology and Biotechnology, 227 Nguyen Van Cu, 5 District, Ho Chi Minh City, Vietnam. &amp; Vietnam National University Ho Chi Minh City, Linh Trung Ward, Thu Duc District, Ho Chi Minh City. &amp; lttduong @ hcmus. edu. vn; https: // orcid. org / 0000 - 0002 - 3832 - 7944</mods:affiliation>
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<mods:namePart id="AB30305AFC1B00A932C901B3C03C2EEC">Hoang, Huy Duc</mods:namePart>
<mods:affiliation id="8419536710EE38A8A33EF790C51F46BF">University of Science, Faculty of Biology and Biotechnology, 227 Nguyen Van Cu, 5 District, Ho Chi Minh City, Vietnam. &amp; Vietnam National University Ho Chi Minh City, Linh Trung Ward, Thu Duc District, Ho Chi Minh City. &amp; hdhuy @ hcmus. edu. vn; https: // orcid. org / 0000 - 0001 - 6528 - 193 X</mods:affiliation>
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<mods:namePart id="F7CFBDBB870C0D0A6549BCD730FF3FB4">Cao, Trung Tien</mods:namePart>
<mods:affiliation id="8DC02A79B93F263F1CBA5D7A5E4D5481">Institute of Biology, Chemistry and Environment, Vinh University, 182 Le Duan St, Vinh City, Vietnam. trungct. vinhuni @ gmail. com; https: // orcid. org / 0000 - 0002 - 4256 - 8951</mods:affiliation>
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<mods:namePart id="74A614AC4058D0E94C8BE2B0A48F895D">Dau, Vinh Quang</mods:namePart>
<mods:affiliation id="38001F524C886BEB2AAFBFF6687E6696">Hong Duc University, 565 Quang Trung Street-Dong Ve Ward-Thanh Hoa City, Vietnam.</mods:affiliation>
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<mods:title id="D985394DAC3B0D1B1A57213F2AFC1F66">Zootaxa</mods:title>
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<mods:date id="CFC8C444E33D764368A2948321D97001">2020</mods:date>
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<treatment id="8B1487E31502526865D11812190AB488" ID-DOI="http://doi.org/10.5281/zenodo.3851654" ID-GBIF-Taxon="164243346" ID-Zenodo-Dep="3851654" LSID="urn:lsid:plazi:treatment:8B1487E31502526865D11812190AB488" httpUri="http://treatment.plazi.org/id/8B1487E31502526865D11812190AB488" lastPageId="10" lastPageNumber="351" pageId="2" pageNumber="343">
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<taxonomicName id="C4BD4D761502526065D118121952B75A" box="[151,364,1546,1573]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="343" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71502526065D118121952B75A" bold="true" box="[151,364,1546,1573]" italics="true" pageId="2" pageNumber="343">Gracixalus trieng</emphasis>
</taxonomicName>
­sp.­nov.
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</heading>
</paragraph>
</subSubSection>
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:
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<specimenCode id="531B9E8E150252606414184A1982B712" box="[338,444,1618,1644]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="343">R176206</specimenCode>
, adult male, collected from tree hole (approx.
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diameter entrance,
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off ground) with eggs (
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), in montane evergreen forest in
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,
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,
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,
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(
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,
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,
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; Fig. 1). Collected at 19:45 h on
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by
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,
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.
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. D.,
<collectorName id="AE48532315025260645818A71951B7A6" box="[286,367,1726,1753]" pageId="2" pageNumber="343">Dau Q.</collectorName>
<collectionCode id="65ACAE3015025260643318A719B7B7A7" box="[373,393,1727,1753]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="343" type="Museum">V</collectionCode>
. and
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</paragraph>
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<typeStatus id="DC06885715025260658118FB1901B783" box="[199,319,1763,1789]" pageId="2" pageNumber="343" type="paratype">Paratypes</typeStatus>
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:
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00342
</materialsCitation>
/
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,
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adult male, on dead fallen tree,
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from ground, in montane evergreen forest in
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,
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,
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,
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(
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,
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,
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), collected at 19:30 h on
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<collectingDate id="6747E9DD15025260648F19321A4DB63A" box="[457,627,1834,1860]" pageId="2" pageNumber="343" value="2010-03-29">29 March 2010</collectingDate>
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by
<collectorName id="AE4853231502526067DB19331B7BB63A" box="[669,837,1834,1861]" pageId="2" pageNumber="343">J. J. L. Rowley</collectorName>
,
<collectorName id="AE48532315025260661619331B50B63B" box="[848,878,1835,1861]" pageId="2" pageNumber="343">Le</collectorName>
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.
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. D.,
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Q
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.
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<collectionCode id="65ACAE3015025260616B19331C7FB63B" box="[1069,1089,1835,1861]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="343" type="Museum">V</collectionCode>
. and
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D. H
</materialsCitation>
.
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<collectionCode id="65ACAE3015025260605619321D76B63A" box="[1296,1352,1834,1860]" country="Viet Nam" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="2" pageNumber="343">UNS</collectionCode>
00343
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/
<materialsCitation id="B3D53CA81502526065D11956197AB617" ID-GBIF-Occurrence="2621059305" box="[151,324,1870,1897]" collectionCode="AMS" country="Vietnam" pageId="2" pageNumber="343" specimenCode="R176205" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
<collectionCode id="65ACAE301502526065D1195618EAB616" box="[151,212,1870,1896]" pageId="2" pageNumber="343">AMS</collectionCode>
<specimenCode id="531B9E8E15025260659D1956197AB617" box="[219,324,1870,1897]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="343">R176205</specimenCode>
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,
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adult male, in tree hole filled with water (
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long,
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deep,
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above ground), in montane evergreen forest in
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,
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,
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,
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(
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,
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,
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</quantity>
), collected at 19:55 h on
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<collectingDate id="6747E9DD15025260676A198E1A84B6CE" box="[556,698,1942,1968]" pageId="2" pageNumber="343" value="2010-04-03">3 April 2010</collectingDate>
</date>
by
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,
<collectorName id="AE4853231502526066DC198F1B86B6CF" box="[922,952,1943,1969]" pageId="2" pageNumber="343">Le</collectorName>
</materialsCitation>
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<collectionCode id="65ACAE301502526066F9198F1BEFB6CF" box="[959,977,1943,1969]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="343">T</collectionCode>
.
<collectionCode id="65ACAE3015025260669A198F1BD0B6CF" box="[988,1006,1943,1969]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="343">T</collectionCode>
. D.,
<collectorName id="AE485323150252606165198F1C6CB6CF" box="[1059,1106,1943,1969]" pageId="2" pageNumber="343">Dau</collectorName>
Q
</materialsCitation>
.
<materialsCitation id="B3D53CA815025260613D198F1D6BB6CF" ID-GBIF-Occurrence="2621059313" box="[1147,1365,1942,1969]" collectionCode="V" country="Vietnam" location="Hoang" pageId="2" pageNumber="343" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
<collectionCode id="65ACAE3015025260613D198F1CB1B6CF" box="[1147,1167,1943,1969]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="343" type="Museum">V</collectionCode>
. and
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D. H
</materialsCitation>
.
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<collectionCode id="65ACAE30150252606026198E1DA3B6CE" box="[1376,1437,1942,1968]" pageId="2" pageNumber="343">AMS</collectionCode>
<specimenCode id="531B9E8E1502526065D119A218C0B6AA" box="[151,254,1978,2004]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="343">R176207</specimenCode>
and
</materialsCitation>
<materialsCitation id="B3D53CA815025260647219A219EDB6AA" ID-GBIF-Occurrence="2621059301" box="[308,467,1978,2004]" collectionCode="NCSM" country="Vietnam" pageId="2" pageNumber="343" specimenCode="NCSM 79748" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
<specimenCode id="531B9E8E15025260647219A219EDB6AA" box="[308,467,1978,2004]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="2" pageNumber="343" type="Museum">NCSM 79748</specimenCode>
</materialsCitation>
,
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adult males, in tree hole filled with water (approximately
<quantity id="C4459B1015025260612519A21CB0B6AB" box="[1123,1166,1978,2005]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="343" unit="m" value="1.0">
<elevation id="8890D1C615025260612519A21CB0B6AB" box="[1123,1166,1978,2005]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="343" unit="m" value="1.0">1 m</elevation>
</quantity>
above ground), in montane evergreen forest in
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,
<collectingMunicipality id="E366AC8F15025260679B19C61B9EB686" box="[733,928,2014,2040]" pageId="2" pageNumber="343">Dak Glei District</collectingMunicipality>
,
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,
<collectingCountry id="7BAA76651502526061CA19C61CD5B686" box="[1164,1259,2014,2040]" name="Vietnam" pageId="2" pageNumber="343">Vietnam</collectingCountry>
(approximately
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,
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,
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), collected at approximately 11:00 h on
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by local
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guides (
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,
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,
<collectorName id="AE4853231503526164731EA51941B1A9" box="[309,383,189,215]" pageId="3" pageNumber="344">A Nap</collectorName>
,
<collectorName id="AE4853231503526164CF1EA519EFB1A6" box="[393,465,189,216]" pageId="3" pageNumber="344">A Doi</collectorName>
)
</materialsCitation>
.
<materialsCitation id="B3D53CA81503526164A51EA51A59B1A9" ID-GBIF-Occurrence="2621059331" box="[483,615,188,215]" collectionCode="UNS" country="Vietnam" pageId="3" pageNumber="344" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
<collectionCode id="65ACAE301503526164A51EA51A25B1A9" box="[483,539,189,215]" country="Viet Nam" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="3" pageNumber="344">UNS</collectionCode>
00351
</materialsCitation>
/
<materialsCitation id="B3D53CA81503526167361EA41B22B1A9" ID-GBIF-Occurrence="2621059321" box="[624,796,188,215]" collectionCode="AMS" country="Vietnam" pageId="3" pageNumber="344" specimenCode="R176208" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
<collectionCode id="65ACAE301503526167361EA41A92B1A9" box="[624,684,188,215]" pageId="3" pageNumber="344">AMS</collectionCode>
<specimenCode id="531B9E8E1503526167F51EA51B22B1A9" box="[691,796,189,215]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="344">R176208</specimenCode>
</materialsCitation>
,
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juvenile, on epiphyte leaf,
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above ground in montane evergreen/bamboo forest in
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,
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,
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,
<collectingCountry id="7BAA76651503526161F01EF91D2BB185" box="[1206,1301,225,251]" name="Vietnam" pageId="3" pageNumber="344">Vietnam</collectingCountry>
(
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,
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,
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</quantity>
), collected at 20:10 h on
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<collectingDate id="6747E9DD1503526167D31F1D1B1DB061" box="[661,803,261,287]" pageId="3" pageNumber="344" value="2010-04-03">3 April 2010</collectingDate>
</date>
by
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,
<collectorName id="AE4853231503526161441F1D1C1EB061" box="[1026,1056,261,287]" pageId="3" pageNumber="344">Le</collectorName>
</materialsCitation>
<materialsCitation id="B3D53CA81503526161601F1D1CE8B061" ID-GBIF-Occurrence="2621059323" box="[1062,1238,261,287]" collectionCode="T" collectorName="Dau" country="Vietnam" pageId="3" pageNumber="344" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
<collectionCode id="65ACAE301503526161601F1D1C06B061" box="[1062,1080,261,287]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="344">T</collectionCode>
.
<collectionCode id="65ACAE301503526161051F1D1C6BB061" box="[1091,1109,261,287]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="344">T</collectionCode>
. D.,
<collectorName id="AE4853231503526161CF1F1D1C86B061" box="[1161,1208,261,287]" pageId="3" pageNumber="344">Dau</collectorName>
Q
</materialsCitation>
.
<materialsCitation id="B3D53CA81503526161A71F1D1890B03D" ID-GBIF-Occurrence="2621059318" collectionCode="V" country="Vietnam" location="Hoang" pageId="3" pageNumber="344" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
<collectionCode id="65ACAE301503526161A71F1D1CCBB061" box="[1249,1269,261,287]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="344" type="Museum">V</collectionCode>
. and
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D. H
</materialsCitation>
.
<materialsCitation id="B3D53CA81503526165FF1F311958B03D" ID-GBIF-Occurrence="2621059335" box="[185,358,297,323]" collectionCode="AMS" country="Vietnam" pageId="3" pageNumber="344" specimenCode="R176209" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
<collectionCode id="65ACAE301503526165FF1F3118C8B03D" box="[185,246,297,323]" pageId="3" pageNumber="344">AMS</collectionCode>
<specimenCode id="531B9E8E1503526165BB1F311958B03D" box="[253,358,297,323]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="344">R176209</specimenCode>
</materialsCitation>
,
<materialsCitation id="B3D53CA81503526164341F311B46B0F5" ID-GBIF-Occurrence="2621059317" collectingDate="2010-04-04" collectorName="J. J. L. Rowley &amp; Le" country="Vietnam" elevation="3" latitude="15.063" location="Ngoc Linh Nature Reserve" longLatPrecision="76" longitude="107.859" municipality="Dak Glei District" pageId="3" pageNumber="344" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
juvenile, on plant lead in stream bed in montane evergreen/bamboo forest
<quantity id="C4459B101503526161FC1F311CD8B03D" box="[1210,1254,297,323]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="344" unit="m" value="3.0">
<elevation id="8890D1C61503526161FC1F311CD8B03D" box="[1210,1254,297,323]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="344" unit="m" value="3.0">3 m</elevation>
</quantity>
from
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<elevation id="8890D1C615035261606A1F311D69B03D" box="[1324,1367,297,323]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="344" unit="m" value="3.0">3 m</elevation>
</quantity>
wide, rocky stream in
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,
<collectingMunicipality id="E366AC8F1503526167C41F551B7AB019" box="[642,836,333,359]" pageId="3" pageNumber="344">Dak Glei District</collectingMunicipality>
,
<collectingRegion id="C179F8171503526166081F551C1AB019" box="[846,1060,333,359]" country="Vietnam" name="Kon Tum" pageId="3" pageNumber="344">Kon Tum Province</collectingRegion>
,
<collectingCountry id="7BAA76651503526161691F551CB0B019" box="[1071,1166,333,359]" name="Vietnam" pageId="3" pageNumber="344">Vietnam</collectingCountry>
(
<geoCoordinate id="668950321503526161DD1F541D2EB019" box="[1179,1296,332,359]" degrees="15.063" direction="north" orientation="latitude" pageId="3" pageNumber="344" precision="55" value="15.063">15.063º N</geoCoordinate>
,
<geoCoordinate id="6689503215035261605C1F541DA9B019" box="[1306,1431,332,359]" degrees="107.859" direction="east" orientation="longitude" pageId="3" pageNumber="344" precision="55" value="107.859">107.859º E</geoCoordinate>
,
<quantity id="C4459B101503526165D11F6918D1B0F5" box="[151,239,369,395]" metricMagnitude="3" metricUnit="m" metricValue="1.716" pageId="3" pageNumber="344" unit="m" value="1716.0">
<elevation id="8890D1C61503526165D11F6918D1B0F5" box="[151,239,369,395]" metricMagnitude="3" metricUnit="m" metricValue="1.716" pageId="3" pageNumber="344" unit="m" value="1716.0">1716 m</elevation>
</quantity>
), collected at night on
<date id="770310351503526164A81F691A42B0F5" box="[494,636,369,395]" pageId="3" pageNumber="344" value="2010-04-04">
<collectingDate id="6747E9DD1503526164A81F691A42B0F5" box="[494,636,369,395]" pageId="3" pageNumber="344" value="2010-04-04">4 April 2010</collectingDate>
</date>
by
<collectorName id="AE4853231503526167E01F691B71B0F5" box="[678,847,369,395]" pageId="3" pageNumber="344">J. J. L. Rowley</collectorName>
,
<collectorName id="AE48532315035261661C1F691B46B0F5" box="[858,888,369,395]" pageId="3" pageNumber="344">Le</collectorName>
</materialsCitation>
<materialsCitation id="B3D53CA81503526166381F691C13B0F5" ID-GBIF-Occurrence="2621059306" box="[894,1069,369,395]" collectionCode="T" collectorName="Dau" country="Vietnam" pageId="3" pageNumber="344" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
<collectionCode id="65ACAE301503526166381F691BAEB0F5" box="[894,912,369,395]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="344">T</collectionCode>
.
<collectionCode id="65ACAE301503526166DC1F691B92B0F5" box="[922,940,369,395]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="344">T</collectionCode>
. D.,
<collectorName id="AE4853231503526166A71F691C2EB0F5" box="[993,1040,369,395]" pageId="3" pageNumber="344">Dau</collectorName>
Q
</materialsCitation>
.
<materialsCitation id="B3D53CA815035261617E1F691D2FB0F5" ID-GBIF-Occurrence="2621059312" box="[1080,1297,369,395]" collectionCode="V" country="Vietnam" location="Hoang" pageId="3" pageNumber="344" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
<collectionCode id="65ACAE3015035261617E1F691C72B0F5" box="[1080,1100,369,395]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="344" type="Museum">V</collectionCode>
. and
<location id="0662602E1503526161C01F691CEFB0F5" LSID="urn:lsid:plazi:treatment:8B1487E31502526865D11812190AB488:0662602E1503526161C01F691CEFB0F5" box="[1158,1233,369,395]" country="Vietnam" name="Hoang" pageId="3" pageNumber="344" stateProvince="Kon Tum">Hoang</location>
D. H
</materialsCitation>
.
</paragraph>
<paragraph id="030236F51503526165811F8C1CF0B0D1" blockId="3.[151,1437,152,431]" box="[199,1230,404,431]" pageId="3" pageNumber="344">
<emphasis id="31C9EAE71503526165811F8C1982B0D1" bold="true" box="[199,444,404,431]" pageId="3" pageNumber="344">Referred­Specimen:­</emphasis>
<materialsCitation id="B3D53CA81503526164FA1F8D1CF5B0D1" ID-GBIF-Occurrence="2621059319" box="[444,1227,405,431]" collectingDate="2010-04-03" collectionCode="AMS" collectorName="J. J. L. Rowley &amp; Le &amp; Dau Q. &amp; Hoang D. H." country="Vietnam" elevation="2045" latitude="15.063" location="Ngoc Linh Nature Reserve" longLatPrecision="76" longitude="107.865" municipality="Dak Glei District" pageId="3" pageNumber="344" specimenCode="R176211" specimenCount="1" stateProvince="Kon Tum">
<collectionCode id="65ACAE301503526164FA1F8D19C7B0D1" box="[444,505,405,431]" pageId="3" pageNumber="344">AMS</collectionCode>
<specimenCode id="531B9E8E1503526167461F8D1A57B0D1" box="[512,617,405,431]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="344">R176211</specimenCode>
, eggs, collected at same time and location as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="030236F51503526165D1187D1D01B7B8" blockId="3.[151,1436,1637,1734]" pageId="3" pageNumber="344">
<emphasis id="31C9EAE71503526165D1187D192BB700" bold="true" box="[151,277,1637,1662]" pageId="3" pageNumber="344">FIGURE­1.</emphasis>
Collection sites of
<taxonomicName id="C4BD4D76150352616499187E1AABB703" box="[479,661,1638,1662]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="344" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE7150352616499187E1AABB703" box="[479,661,1638,1662]" italics="true" pageId="3" pageNumber="344">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE71503526167DA187E1ACAB700" bold="true" box="[668,756,1638,1662]" pageId="3" pageNumber="344">sp.­nov.­</emphasis>
Collection locality of
<typeStatus id="DC06885715035261669E187D1C0FB700" box="[984,1073,1637,1662]" pageId="3" pageNumber="344" type="holotype">holotype</typeStatus>
is marked with a yellow star. Note that the high elevation habitat of the new species is geographically isolated and of all known species in ‘Clade II’ (
<bibRefCitation id="672C4B04150352616052189118F4B7B8" author="Rowley, J. J. L. &amp; Dau, Q. V. &amp; Nguyen, T. T. &amp; Cao, T. T. &amp; Nguyen, S. N." pageId="3" pageNumber="344" pagination="22 - 38" refId="ref8737" refString="Rowley, J. J. L., Dau, Q. V., Nguyen, T. T., Cao, T. T. &amp; Nguyen, S. N. (2011) A new species of Gracixalus (Anura: Rhacophoridae) with a hyperextended vocal repertoire from Vietnam. Zootaxa, 3125, 22 - 38." type="journal article" year="2011">
Rowley
<emphasis id="31C9EAE715035261602C18921DA8B7DC" box="[1386,1430,1674,1698]" italics="true" pageId="3" pageNumber="344">et al</emphasis>
. 2011
</bibRefCitation>
) of the genus, only
<taxonomicName id="C4BD4D761503526164D218B61A5AB7B8" box="[404,612,1710,1734]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="344" phylum="Chordata" rank="species" species="lumarius">
<emphasis id="31C9EAE71503526164D218B61A5AB7B8" box="[404,612,1710,1734]" italics="true" pageId="3" pageNumber="344">Gracixalus lumarius</emphasis>
</taxonomicName>
occurs in the mapped area (in syntopy with
<taxonomicName id="C4BD4D7615035261616018B61CE4B7BB" box="[1062,1242,1710,1734]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="344" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE715035261616018B61CE4B7BB" box="[1062,1242,1710,1734]" italics="true" pageId="3" pageNumber="344">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE71503526161A618B61D0EB7B8" bold="true" box="[1248,1328,1710,1734]" pageId="3" pageNumber="344">sp.­nov.</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="4BA7657E15035261658118EC1C92B605" pageId="3" pageNumber="344" type="etymology">
<paragraph id="030236F515035261658118EC19E5B628" blockId="3.[151,1437,1779,2022]" pageId="3" pageNumber="344">
<emphasis id="31C9EAE715035261658118EC1976B670" bold="true" box="[199,328,1780,1806]" pageId="3" pageNumber="344">Etymology</emphasis>
: The specific name is in reference to the Giẻ Triêng people, most of whom live in
<collectingRegion id="C179F8171503526161AA18EC1D69B670" box="[1260,1367,1780,1806]" country="Vietnam" name="Kon Tum" pageId="3" pageNumber="344">Kon Tum</collectingRegion>
Prov- ince in the Central Highlands of
<collectingCountry id="7BAA766515035261675619001A4CB64C" box="[528,626,1816,1842]" name="Vietnam" pageId="3" pageNumber="344">Vietnam</collectingCountry>
, and who assisted us greatly during our surveys. The species epithet is used as a noun in apposition.
</paragraph>
<paragraph id="030236F515035261658119471C92B605" blockId="3.[151,1437,1779,2022]" box="[199,1196,1887,1915]" pageId="3" pageNumber="344">
<emphasis id="31C9EAE715035261658119471A3FB604" bold="true" box="[199,513,1887,1914]" pageId="3" pageNumber="344">Suggested­Common­Name</emphasis>
: Trieng tree frog (English). Ếch cây giẻ
<taxonomicName id="C4BD4D7615035261668119781C96B605" authority="(Vietnamese)" baseAuthorityName="Vietnamese" box="[967,1192,1888,1915]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="344" phylum="Chordata" rank="species" species="trieng">triêng (Vietnamese)</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="4BA7657E150352676581199B1CADB721" lastPageId="5" lastPageNumber="346" pageId="3" pageNumber="344" type="diagnosis">
<paragraph id="030236F5150352666581199B1C6BB089" blockId="3.[151,1437,1779,2022]" lastBlockId="4.[151,1437,152,503]" lastPageId="4" lastPageNumber="345" pageId="3" pageNumber="344">
<emphasis id="31C9EAE7150352616581199B1904B6E0" bold="true" box="[199,314,1923,1950]" pageId="3" pageNumber="344">Diagnosis</emphasis>
: The new species is assigned to the family
<taxonomicName id="C4BD4D76150352616669199C1BE7B6E0" authorityName="Hoffman" authorityYear="1932" box="[815,985,1924,1950]" class="Amphibia" family="Rhacophoridae" kingdom="Animalia" order="Anura" pageId="3" pageNumber="344" phylum="Chordata" rank="family">Rhacophoridae</taxonomicName>
by the presence of intercalary cartilage between the terminal and penultimate phalanges of digits, tips of digits expanded into large discs bearing circummarginal grooves, vomerine teeth absent, pupil horizontal, large, conical tubercles/spines present on the upper eye- lid (
<bibRefCitation id="672C4B041504526665851E81197FB1CD" author="Liem, S. S." box="[195,321,153,179]" pageId="4" pageNumber="345" pagination="1 - 145" refId="ref8278" refString="Liem, S. S. (1970) The morphology, systematics, and evolution of the old world treefrogs (Rhacophoridae and Hyperoliidae). Fieldiana Zoology, 57, 1 - 145. https: // doi. org / 10.5962 / bhl. title. 2939" type="journal article" year="1970">Liem 1970</bibRefCitation>
;
<bibRefCitation id="672C4B0415045266640A1E811A5EB1CD" author="Duellman, W. E. &amp; Trueb, L." box="[332,608,153,179]" pageId="4" pageNumber="345" refId="ref7831" refString="Duellman, W. E. &amp; Trueb, L. (1986) Biology of Amphibians. The John Hopkins University Press, Baltimore and London, 670 pp. https: // doi. org / 10.2307 / 1445022" type="book" year="1986">Duellman &amp; Trueb 1986</bibRefCitation>
;
<bibRefCitation id="672C4B0415045266672A1E811B5AB1CD" author="Brown, W. C. &amp; Alcala, A. C." box="[620,868,153,179]" pageId="4" pageNumber="345" pagination="185 - 220" refId="ref7578" refString="Brown, W. C. &amp; Alcala, A. C. (1994) Philippine frogs of the family Rhacophoridae. Proceedings of the Californian Academy of Sciences, 48, 185 - 220." type="journal article" year="1994">Brown &amp; Alcala 1994</bibRefCitation>
;
<bibRefCitation id="672C4B041504526666291E811C6FB1CD" author="Delorme, M. &amp; Dubois, A. &amp; Grosjean, S. &amp; Ohler, A." box="[879,1105,152,179]" pageId="4" pageNumber="345" pagination="165 - 171" refId="ref7763" refString="Delorme, M., Dubois, A., Grosjean, S. &amp; Ohler, A. (2005) Une nouvelle classification generique et subgenerique de la tribu des Philautini (Amphibia, Anura, Rhacophorinae). Bulletin Mensuel de la Societe Linneenne de Lyon, 74, 165 - 171. https: // doi. org / 10.3406 / linly. 2005.13595" type="journal article" year="2005">
Delorme
<emphasis id="31C9EAE71504526666911E821C31B1CD" box="[983,1039,153,179]" italics="true" pageId="4" pageNumber="345">et al.</emphasis>
2005
</bibRefCitation>
), and to the genus
<taxonomicName id="C4BD4D761504526660661E811DA2B1CD" baseAuthorityName="Frost" baseAuthorityYear="2020" box="[1312,1436,153,179]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="345" phylum="Chordata" rank="genus">
<emphasis id="31C9EAE71504526660661E811DA2B1CD" box="[1312,1436,153,179]" italics="true" pageId="4" pageNumber="345">Gracixalus</emphasis>
</taxonomicName>
by the presence of a dark X, inverted V or inverted Y-shape marking present on the dorsal surface of the trunk (
<bibRefCitation id="672C4B041504526660311EA51928B185" author="Fei, L. &amp; Hu, S. &amp; Ye, C. &amp; Huang, Y." pageId="4" pageNumber="345" refId="ref7872" refString="Fei, L., Hu, S., Ye, C. &amp; Huang, Y. (2009) Fauna Sinica. Amphibia. Vol. 2 Anura. Science Press, Beijing, 957 pp." type="book" year="2009">
Fei
<emphasis id="31C9EAE71504526665D11EFA18F4B185" box="[151,202,225,251]" italics="true" pageId="4" pageNumber="345">et al</emphasis>
. 2009
</bibRefCitation>
).
<taxonomicName id="C4BD4D7615045266646F1EF919CFB185" box="[297,497,225,251]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="345" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE715045266646F1EF919CFB185" box="[297,497,225,251]" italics="true" pageId="4" pageNumber="345">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE71504526664BF1EF91A65B185" bold="true" box="[505,603,225,251]" pageId="4" pageNumber="345">sp.­nov.­</emphasis>
is distinguished from all congeners by a combination of 1) body size medium (
<quantity id="C4459B101504526665A41F1D19B9B061" box="[226,391,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="4.14" metricValueMin="3.72" pageId="4" pageNumber="345" unit="mm" value="39.3" valueMax="41.4" valueMin="37.2">37.2–41.4 mm</quantity>
in five adult males), (2) snout rounded in dorsal and lateral views, (3) dorsal surface brown or yellowish with a darker brown interorbital crossbar and inverse-Y shape on the back, (4) throat and chest yellow or yellowish brown with pinkish mottling and belly and ventral surfaces of limbs including hands and feet pinkish, (5) tympanum and supratympanic fold distinct, (6) iris pale gold with darker gold radiating out from anterior and posterior edges of pupil, (7) majority of dorsal body and limb surfaces smooth in adults, with some individuals having sparsely distributed low, irregular tubercles, (8) nuptial pads on fingers I and II in adult males, and (9) eggs deposited as a tightly spaced array of non-pendent eggs on the wall of a phytotelmon.
</paragraph>
<paragraph id="030236F51504526665D118181A86B743" blockId="4.[151,1437,1536,1597]" pageId="4" pageNumber="345">
<emphasis id="31C9EAE71504526665D11818192CB767" bold="true" box="[151,274,1536,1561]" pageId="4" pageNumber="345">FIGURE­2.</emphasis>
Diurnal dorsolateral view (A), nocturnal dorsolateral view (B), dorsal view (C) and ventral view (D) of the
<typeStatus id="DC06885715045266600218181DA3B767" box="[1348,1437,1536,1561]" pageId="4" pageNumber="345" type="holotype">holotype</typeStatus>
of
<taxonomicName id="C4BD4D761504526665F5183C1959B742" box="[179,359,1572,1596]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="345" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71504526665F5183C1959B742" box="[179,359,1572,1596]" italics="true" pageId="4" pageNumber="345">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE715045266642A183D1983B743" bold="true" box="[364,445,1573,1597]" pageId="4" pageNumber="345">sp.­nov.</emphasis>
(AMS R176206) in life.
</paragraph>
<paragraph id="030236F515045267658118721BBEB089" blockId="4.[151,1437,1642,2030]" lastBlockId="5.[151,1437,153,503]" lastPageId="5" lastPageNumber="346" pageId="4" pageNumber="345">
<emphasis id="31C9EAE7150452666581187219E2B7FB" bold="true" box="[199,476,1642,1669]" pageId="4" pageNumber="345">Description­of­holotype</emphasis>
: A medium-sized rhacophorid (
<quantity id="C4459B1015045266667B18731B98B7FB" box="[829,934,1643,1669]" metricMagnitude="-2" metricUnit="m" metricValue="3.72" pageId="4" pageNumber="345" unit="mm" value="37.2">37.2 mm</quantity>
), body robust (Figs. 2,3), head length 91% of head width, snout broadly rounded in dorsal view, rounded in profile, canthus rostralis rounded; loreal region slightly sloping, interorbital region convex, nostrils rounded, slightly protuberant, without flap of skin laterally, closer to tip of snout than eye; pupil oval, horizontal, tympanic annulus distinct, approximately one-third of eye diameter, pineal ocellus absent, skin not co-ossified to skull, vomerine teeth absent, choanae small, round, separated by about six times that of choanae diameter; tongue attached anteriorly, deeply notched posteriorly, tooth-like projections on lower jaw absent, supratympanic fold distinct, extending from eye to axilla. Vocal sac present, connected to buccal cavity by pair of distinct, oval openings at bases of jaws, only slightly baggy gular region. Forelimbs relatively robust, fingers narrow; relative length of fingers I &lt;II &lt;IV &lt;III. Tips of all fingers with well-developed discs with circummarginal grooves; discs relatively wide compared to finger width (third finger disc 239% of third finger width at penultimate phalange), third finger disc width greater (131%) than tympanum diameter; slight dermal fringes on fingers, basal webbing at base of fingers II–IV. Subarticular tubercles prominent, rounded, formula 1, 1, 2, 2. Accessory palmar tubercles present; supernumerary tubercles absent; palmar tubercle divided, irregular, flat; thenar tubercle indistinct; prepollex elongate, with oval tubercle; distinct (Fig. 4A), nuptial pads present on inner surface of the prepollex and dorsal and inner surfaces of fingers I and II (Fig. 4B). Relative length of toes I &lt;II &lt;V &lt;III &lt;IV, tips of toes with well-developed discs with distinct circummarginal grooves, wide when compared to toe length. Webbing present, formula I 2
<superScript id="F4C89BBD15055267671D1F541A5EB024" attach="left" box="[603,608,332,346]" fontSize="6" pageId="5" pageNumber="346">-</superScript>
– 2
<superScript id="F4C89BBD1505526767CC1F541AADB024" attach="left" box="[650,659,332,346]" fontSize="6" pageId="5" pageNumber="346">+</superScript>
II 1
<superScript id="F4C89BBD1505526767871F541AF4B024" attach="left" box="[705,714,332,346]" fontSize="6" pageId="5" pageNumber="346">+</superScript>
– 2
<superScript id="F4C89BBD1505526767B21F541AC3B024" attach="left" box="[756,765,332,346]" fontSize="6" pageId="5" pageNumber="346">+</superScript>
III 2
<superScript id="F4C89BBD1505526766731F541B04B024" attach="left" box="[821,826,332,346]" fontSize="6" pageId="5" pageNumber="346">-</superScript>
– 3 IV 2
<superScript id="F4C89BBD1505526766DA1F541B9BB024" attach="none" box="[924,933,332,346]" fontSize="6" pageId="5" pageNumber="346">+</superScript>
– 1 V, with slight dermal fringes, subarticular tubercles prominent, rounded, formula 1, 1, 2, 3, 2, inner metatarsal tubercle distinct, outer metatarsal tubercle and supernumerary tubercles absent (Fig. 4C). Dorsal surface smooth, with only sparsely distributed low, irregular, smooth tubercles. Ventral surface of thighs and abdomen coarsely granular, throat finely granular. No dermal fringes on limbs, no tibiotarsal projection, no distinct supracloacal glands.
</paragraph>
<paragraph id="030236F51505526765D11A5618D3B5F4" blockId="5.[151,1436,1102,1163]" pageId="5" pageNumber="346">
<emphasis id="31C9EAE71505526765D11A56192DB519" bold="true" box="[151,275,1102,1127]" pageId="5" pageNumber="346">FIGURE­3.</emphasis>
Dorsal (A) ventral (B) view of preserved male
<typeStatus id="DC0688571505526767BE1A561B6FB519" box="[760,849,1102,1127]" pageId="5" pageNumber="346" type="holotype">holotype</typeStatus>
of
<taxonomicName id="C4BD4D761505526766351A561C16B518" box="[883,1064,1102,1126]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="346" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71505526766351A561C16B518" box="[883,1064,1102,1126]" italics="true" pageId="5" pageNumber="346">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE71505526761681A571C41B519" bold="true" box="[1070,1151,1103,1127]" pageId="5" pageNumber="346">sp.­nov.</emphasis>
(AMS R176206). Scale bar =
<quantity id="C4459B101505526765EA1A6A18D4B5F4" box="[172,234,1138,1163]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="346" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
<paragraph id="030236F51505526765811AA01ADFB48D" blockId="5.[151,1437,1208,2028]" pageId="5" pageNumber="346">
<emphasis id="31C9EAE71505526765811AA01A3EB5AD" bold="true" box="[199,512,1208,1235]" pageId="5" pageNumber="346">Colour­of­holotype­in­life:</emphasis>
Dorsal surface brown (diurnally) or yellowish-tan (nocturnally) with a darker interorbital crossbar and inverted Y-shaped marking on the back starting between the eye and covering most of head; limbs with dark brown barring (Fig. 2). Patterning more distinct diurnally. Fingers brown or yellowish-tan, webbing pinkish brown. Diffuse darker brown line under canthus rostralis from eye to nostril, no obvious tympanic markings. Flanks mottled brown and dark pink (diurnally) or yellow and pink (nocturnally). Throat and chest mostly yellowish brown, with dark pinkish brown mottling (diurnally) or yellow with pink mottling (nocturnally); belly and ventral surfaces of limbs including hands and feet pinkish brown (diurnally) or pink (nocturnally). Iris pale gold with a sparse network of fine dark gold reticulations darker gold radiating out of anterior and posterior edges of pupil; iris periphery black; eye periphery pale blue posteriorly.
</paragraph>
<paragraph id="030236F51505526765811BE41CADB721" blockId="5.[151,1437,1208,2028]" pageId="5" pageNumber="346">
<emphasis id="31C9EAE71505526765811BE41A4EB769" bold="true" box="[199,624,1532,1559]" pageId="5" pageNumber="346">Colour­of­holotype­in­preservative:</emphasis>
Dorsal surface as in life, but paler with less distinct markings (Fig. 3). Ventral surface paler, throat pale brown with pale creamy yellow mottling concentrated on chest. Belly and ventral surface of limbs pinkish grey. Dorsal and ventral surfaces of discs on fingers and toes grey.
</paragraph>
</subSubSection>
<subSubSection id="4BA7657E15055264658118711B01B628" lastPageId="6" lastPageNumber="347" pageId="5" pageNumber="346" type="description">
<paragraph id="030236F515055267658118711B6FB7D9" blockId="5.[151,1437,1208,2028]" pageId="5" pageNumber="346">
<emphasis id="31C9EAE715055267658118711948B7FD" bold="true" box="[199,374,1641,1667]" pageId="5" pageNumber="346">Measurements</emphasis>
:
<typeStatus id="DC0688571505526764C3187119CCB7FD" box="[389,498,1641,1667]" pageId="5" pageNumber="346" type="holotype">Holotype</typeStatus>
: SVL 37.2, HDL 13.0, HDW 14.3, SNT 5.9, EYE 4.4, IOD 4.1, TMP 2.2, TEY 1.2, IN 4.1, NS 2.6, EN 2.9, TIB 18.3. ML 12.7, PL 18.1, IMT 2.0.
</paragraph>
<paragraph id="030236F515055264658118A81DAAB089" blockId="5.[151,1437,1208,2028]" lastBlockId="6.[151,1437,152,503]" lastPageId="6" lastPageNumber="347" pageId="5" pageNumber="346">
<emphasis id="31C9EAE715055267658118A81906B7B5" bold="true" box="[199,312,1712,1739]" pageId="5" pageNumber="346">Variation</emphasis>
:
<materialsCitation id="B3D53CA815055267640E18A91A76B625" ID-GBIF-Occurrence="2621059322" collectionCode="UNS, AMS" pageId="5" pageNumber="346" specimenCode="R176204, R176207, NCSM 79748" specimenCount="1" typeStatus="paratype">
Measurements of the type series are shown in Table 2.
<typeStatus id="DC0688571505526766FD18A91C16B7B5" box="[955,1064,1713,1739]" pageId="5" pageNumber="346" type="paratype">Paratypes</typeStatus>
vary in slightly in their colouration and pattern (Fig. 5). Skin texture also varies from almost completely smooth to sparsely distributed with low, irregular tubercles. Tiny, conical tubercles are present on the eyelids, under the eye and on the tympanum in some individuals (
<collectionCode id="65ACAE3015055267646219051960B649" box="[292,350,1821,1847]" country="Viet Nam" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="5" pageNumber="346">UNS</collectionCode>
00342/
<collectionCode id="65ACAE301505526764F2190519D1B649" box="[436,495,1821,1847]" pageId="5" pageNumber="346">AMS</collectionCode>
<specimenCode id="531B9E8E1505526764BE19051A5FB649" box="[504,609,1821,1847]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="346">R176204</specimenCode>
,
<collectionCode id="65ACAE3015055267672A19051A97B649" box="[620,681,1821,1847]" pageId="5" pageNumber="346">AMS</collectionCode>
<specimenCode id="531B9E8E1505526767F719051B26B649" box="[689,792,1821,1847]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="346">R176207</specimenCode>
and to a lesser extent,
<specimenCode id="531B9E8E15055267615919051CFFB649" box="[1055,1217,1821,1847]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="5" pageNumber="346" type="Museum">NCSM 79748</specimenCode>
). The tympanic annulus is less distinct in
<specimenCode id="531B9E8E1505526764E519591A7AB625" box="[419,580,1857,1883]" collectionCode="NCSM" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="5" pageNumber="346" type="Museum">NCSM 79748</specimenCode>
,
</materialsCitation>
<materialsCitation id="B3D53CA815055267671719591AE9B625" ID-GBIF-Occurrence="2621059310" box="[593,727,1857,1883]" collectionCode="UNS" isTailEnumeration="true" pageId="5" pageNumber="346" specimenCount="1" typeStatus="paratype">
<collectionCode id="65ACAE3015055267671719591AB7B625" box="[593,649,1857,1883]" country="Viet Nam" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="5" pageNumber="346">UNS</collectionCode>
00343
</materialsCitation>
/
<materialsCitation id="B3D53CA81505526767A619591BB5B625" ID-GBIF-Occurrence="2621059307" box="[736,907,1856,1883]" collectionCode="AMS" pageId="5" pageNumber="346" specimenCode="R176205" specimenCount="1" typeStatus="paratype">
<collectionCode id="65ACAE301505526767A619591B22B625" box="[736,796,1857,1883]" pageId="5" pageNumber="346">AMS</collectionCode>
<specimenCode id="531B9E8E15055267666219581BB5B625" box="[804,907,1856,1883]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="346">R176205</specimenCode>
</materialsCitation>
<materialsCitation id="B3D53CA81505526766D519591C4CB625" ID-GBIF-Occurrence="2621059304" box="[915,1138,1857,1883]" collectionCode="AMS" pageId="5" pageNumber="346" specimenCode="R176206" specimenCount="1" typeStatus="paratype">
and
<collectionCode id="65ACAE3015055267668519591C3EB625" box="[963,1024,1857,1883]" pageId="5" pageNumber="346">AMS</collectionCode>
<specimenCode id="531B9E8E15055267614E19591C4CB625" box="[1032,1138,1857,1883]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="346">R176206</specimenCode>
</materialsCitation>
.
<materialsCitation id="B3D53CA815055264613819591B25B061" ID-GBIF-Occurrence="2621059316" collectionCode="AMS, UNS" lastPageId="6" lastPageNumber="347" pageId="5" pageNumber="346" specimenCode="R176206, R176204, R176207, R176208, R176209" specimenCount="1" typeStatus="paratype">
Considerable variation in colouration occurs in individuals over time in life, with several specimens changing from mostly pale yellow dorsally and pale pink ventrally without distinct dorsal markings (typical nocturnal coloration), to dark brown dorsally and dark pinkish brown ventrally, with distinct dark brown dorsal markings (typical diurnal coloration) over the course of 24 h (Fig. 2B;
<collectionCode id="65ACAE301505526764EF19C919D8B695" box="[425,486,2001,2027]" pageId="5" pageNumber="346">AMS</collectionCode>
<specimenCode id="531B9E8E1505526764AA19C91A69B695" box="[492,599,2001,2027]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="346">R176206</specimenCode>
;
<collectionCode id="65ACAE3015055267672419C91AA4B695" box="[610,666,2001,2027]" country="Viet Nam" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="5" pageNumber="346">UNS</collectionCode>
00342/
<collectionCode id="65ACAE301505526767B619C91B15B695" box="[752,811,2001,2027]" pageId="5" pageNumber="346">AMS</collectionCode>
<specimenCode id="531B9E8E15055267667519C91BA2B695" box="[819,924,2001,2027]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="346">R176204</specimenCode>
,
<collectionCode id="65ACAE301505526766E019C91BDDB695" box="[934,995,2001,2027]" pageId="5" pageNumber="346">AMS</collectionCode>
<specimenCode id="531B9E8E1505526766AC19C91C68B692" box="[1002,1110,2001,2028]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="346">R176207</specimenCode>
). In life, juveniles of the new species (
<collectionCode id="65ACAE301506526465B11E81190BB1CD" box="[247,309,153,179]" pageId="6" pageNumber="347">AMS</collectionCode>
<specimenCode id="531B9E8E15065264647A1E81199BB1CD" box="[316,421,153,179]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="347">R176207</specimenCode>
,
<collectionCode id="65ACAE301506526464F61E8119D6B1CD" box="[432,488,153,179]" country="Viet Nam" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="6" pageNumber="347">UNS</collectionCode>
00351/
<collectionCode id="65ACAE3015065264677B1E801A47B1CD" box="[573,633,152,179]" pageId="6" pageNumber="347">AMS</collectionCode>
<specimenCode id="531B9E8E1506526467C61E811AD6B1CA" box="[640,744,153,180]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="347">R176208</specimenCode>
) have a pale tan dorsum with an indistinct underlying pattern, yellowish flanks, and anterior and posterior surfaces of thighs, with dark brown blotches, and transparent creamy yellow venters (
<collectionCode id="65ACAE301506526464011EF919BFB185" box="[327,385,225,251]" country="Viet Nam" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="6" pageNumber="347">UNS</collectionCode>
00351/
<collectionCode id="65ACAE301506526464931EF81A2FB185" box="[469,529,224,251]" pageId="6" pageNumber="347">AMS</collectionCode>
<specimenCode id="531B9E8E1506526467511EF91A41B182" box="[535,639,225,252]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="347">R176208</specimenCode>
) or transparent pale pink venters with white blotches on chest and throat (
<collectionCode id="65ACAE301506526465D91F1D18E3B061" box="[159,221,261,287]" pageId="6" pageNumber="347">AMS</collectionCode>
<specimenCode id="531B9E8E1506526465A01F1D196CB05E" box="[230,338,261,288]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="347">R176209</specimenCode>
). In preservative, all but
<collectionCode id="65ACAE301506526467291F1D1A92B061" box="[623,684,261,287]" pageId="6" pageNumber="347">AMS</collectionCode>
<specimenCode id="531B9E8E1506526467F21F1D1B25B061" box="[692,795,261,287]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="347">R176207</specimenCode>
</materialsCitation>
<materialsCitation id="B3D53CA81506526466621F1D1A0AB019" ID-GBIF-Occurrence="2621059329" collectionCode="UNS" pageId="6" pageNumber="347" specimenCount="1" typeStatus="paratype">
have a distinct cross-mark on the back. In preservative, all adult
<typeStatus id="DC0688571506526465BB1F311956B03D" box="[253,360,297,323]" pageId="6" pageNumber="347" type="paratype">paratypes</typeStatus>
are pinkish-grey ventrally, with variable amounts of indistinct, darker greyish-brown blotches concentrated anteriorly.
<collectionCode id="65ACAE301506526464ED1F5519DDB019" box="[427,483,333,359]" country="Viet Nam" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="6" pageNumber="347">UNS</collectionCode>
00342
</materialsCitation>
/
<materialsCitation id="B3D53CA815065264677A1F551AD6B019" ID-GBIF-Occurrence="2621059330" box="[572,744,333,359]" collectionCode="AMS" isTailEnumeration="true" pageId="6" pageNumber="347" specimenCode="R176204" specimenCount="1" typeStatus="paratype">
<collectionCode id="65ACAE3015065264677A1F551A49B019" box="[572,631,333,359]" pageId="6" pageNumber="347">AMS</collectionCode>
<specimenCode id="531B9E8E1506526467C71F551AD6B019" box="[641,744,333,359]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="347">R176204</specimenCode>
</materialsCitation>
<materialsCitation id="B3D53CA81506526467B71F551C58B019" ID-GBIF-Occurrence="2621059333" box="[753,1126,333,359]" collectionCode="AMS" pageId="6" pageNumber="347" specimenCode="R176206" specimenCount="1" typeStatus="holotype">
and the
<typeStatus id="DC0688571506526466091F551B8FB019" box="[847,945,333,359]" pageId="6" pageNumber="347" type="holotype">holotype</typeStatus>
<collectionCode id="65ACAE301506526466FF1F551BC8B019" box="[953,1014,333,359]" pageId="6" pageNumber="347">AMS</collectionCode>
<specimenCode id="531B9E8E1506526466B91F551C58B019" box="[1023,1126,333,359]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="347">R176206</specimenCode>
</materialsCitation>
<materialsCitation id="B3D53CA81506526461291F5518E2B089" ID-GBIF-Occurrence="2621059311" collectionCode="AMS, UNS" pageId="6" pageNumber="347" specimenCode="R176207, R176208" specimenCount="3" typeStatus="paratype">
have darker brown throats and juveniles of the new species (
<collectionCode id="65ACAE301506526467551F691A6FB0F5" box="[531,593,369,395]" pageId="6" pageNumber="347">AMS</collectionCode>
<specimenCode id="531B9E8E15065264671F1F691AFCB0F5" box="[601,706,369,395]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="347">R176207</specimenCode>
,
<collectionCode id="65ACAE301506526467881F691B38B0F5" box="[718,774,369,395]" country="Viet Nam" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="6" pageNumber="347">UNS</collectionCode>
00351/
<collectionCode id="65ACAE3015065264661D1F681BA9B0F5" box="[859,919,368,395]" pageId="6" pageNumber="347">AMS</collectionCode>
<specimenCode id="531B9E8E1506526466D91F691C36B0F2" box="[927,1032,369,396]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="347">R176208</specimenCode>
) have pale grey venters with darker blotches. All
<typeStatus id="DC06885715065264646C1F8D19ABB0D1" box="[298,405,405,431]" pageId="6" pageNumber="347" type="paratype">paratypes</typeStatus>
have distinct supratympanic folds and dark brown canthal and supratympanic lines. A single adult male
<typeStatus id="DC0688571506526464541FA1194CB0AD" box="[274,370,441,467]" pageId="6" pageNumber="347" type="paratype">paratype</typeStatus>
(
<collectionCode id="65ACAE301506526464C71FA11981B0AD" box="[385,447,441,467]" pageId="6" pageNumber="347">AMS</collectionCode>
<specimenCode id="531B9E8E1506526464831FA11A10B0AA" box="[453,558,441,468]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="347">R176207</specimenCode>
) had fine, small tubercles on the dorsum in life, and the juvenile
<typeStatus id="DC0688571506526461B81FA11D60B0AD" box="[1278,1374,441,467]" pageId="6" pageNumber="347" type="paratype">paratype</typeStatus>
<collectionCode id="65ACAE301506526460221FA11DA2B0AD" box="[1380,1436,441,467]" country="Viet Nam" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="6" pageNumber="347">UNS</collectionCode>
00351
</materialsCitation>
/
<materialsCitation id="B3D53CA81506526465A31FC419C0B089" ID-GBIF-Occurrence="2621059308" box="[229,510,476,503]" collectionCode="AMS" isTailEnumeration="true" pageId="6" pageNumber="347" specimenCode="R176208" specimenCount="1" typeStatus="paratype">
<collectionCode id="65ACAE301506526465A31FC4191FB089" box="[229,289,476,503]" pageId="6" pageNumber="347">AMS</collectionCode>
<specimenCode id="531B9E8E15065264646F1FC519AEB089" box="[297,400,477,503]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="347">R176208</specimenCode>
(Fig. 5D)
</materialsCitation>
was slightly shagreened. All male
<typeStatus id="DC0688571506526466C01FC51BCFB089" box="[902,1009,477,503]" pageId="6" pageNumber="347" type="paratype">paratypes</typeStatus>
have nuptial pads on fingers I and II.
</paragraph>
<paragraph id="030236F51506526465D118EE1B01B628" blockId="6.[151,1437,1782,1879]" pageId="6" pageNumber="347">
<emphasis id="31C9EAE71506526465D118EE192CB671" bold="true" box="[151,274,1782,1807]" pageId="6" pageNumber="347">FIGURE­4.</emphasis>
(A) Plantar surface of left hand showing nuptial pad on prepollex (marked with red arrow), (B) dorsal surface of left hand showing nuptial pads of Fingers I and II (marked with red arrows), and (C) plantar surface of left foot of the male
<typeStatus id="DC06885715065264600219021DA3B64D" box="[1348,1437,1818,1843]" pageId="6" pageNumber="347" type="holotype">holotype</typeStatus>
of
<taxonomicName id="C4BD4D761506526465F519271959B629" box="[179,359,1855,1879]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="347" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71506526465F519271959B629" box="[179,359,1855,1879]" italics="true" pageId="6" pageNumber="347">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE715065264642A19271983B629" bold="true" box="[364,445,1855,1879]" pageId="6" pageNumber="347">sp.­nov.</emphasis>
(AMS R176206). Scale bars =
<quantity id="C4459B101506526467B819271B02B628" box="[766,828,1855,1879]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="347" unit="mm" value="2.0">2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="4BA7657E150652686581199D190AB488" lastPageId="10" lastPageNumber="351" pageId="6" pageNumber="347" type="biology_ecology">
<paragraph id="030236F5150652646581199D1DA2B699" blockId="6.[151,1437,1925,2024]" pageId="6" pageNumber="347">
<emphasis id="31C9EAE7150652646581199D1908B6E1" bold="true" box="[199,310,1925,1951]" pageId="6" pageNumber="347">Ecology:­</emphasis>
<materialsCitation id="B3D53CA815065264647E199D1DA2B699" ID-GBIF-Occurrence="2621059332" collectionCode="AMS" pageId="6" pageNumber="347" specimenCode="R176209" specimenCount="130" specimenCount-egg="130" typeStatus="holotype">
All specimens were found in
<emphasis id="31C9EAE7150652646738199D1ABBB6E1" bold="true" box="[638,645,1925,1951]" pageId="6" pageNumber="347">­</emphasis>
montane evergreen forest (Fig. 6A). The
<typeStatus id="DC06885715065264611E199D1C84B6E1" box="[1112,1210,1925,1951]" pageId="6" pageNumber="347" type="holotype">holotype</typeStatus>
was found in a tree hole, with
<specimenCount id="15BBFD7C15065264644B19B1194EB6BA" box="[269,368,1961,1988]" pageId="6" pageNumber="347" type="egg">130 eggs</specimenCount>
deposited on the trunk of the tree above the water (Fig. 6B). All other adult males in the type series were found in tree holes, and both juveniles were found on arboreal vegetation. All but juvenile
<collectionCode id="65ACAE301506526461B419D51D11B699" box="[1266,1327,1997,2023]" pageId="6" pageNumber="347">AMS</collectionCode>
<specimenCode id="531B9E8E15065264607319D51DA2B699" box="[1333,1436,1997,2023]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="347">R176209</specimenCode>
</materialsCitation>
</paragraph>
<paragraph id="030236F51507526565D11E811B36B1A9" blockId="7.[151,1436,153,215]" pageId="7" pageNumber="348">
were found away from any streams or ponds. The species occurs in syntopy with
<taxonomicName id="C4BD4D7615075265615E1E811CA3B1CD" box="[1048,1181,153,179]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="lumarius">
<emphasis id="31C9EAE715075265615E1E811C12B1CD" box="[1048,1068,153,179]" italics="true" pageId="7" pageNumber="348">G</emphasis>
.
<emphasis id="31C9EAE715075265617F1E811CA3B1CD" box="[1081,1181,153,179]" italics="true" pageId="7" pageNumber="348">lumarius</emphasis>
</taxonomicName>
. The females, tadpoles and advertisement call of the new species are unknown.
</paragraph>
<paragraph id="030236F51507526565D11F1E1CD9B061" blockId="7.[151,1255,262,287]" box="[151,1255,262,287]" pageId="7" pageNumber="348">
<tableNote id="FE5B377B1507526565D11F1E1CD9B061" box="[151,1255,262,287]" pageId="7" pageNumber="348" targetBox="[166,1392,302,1280]" targetPageId="7">
<emphasis id="31C9EAE71507526565D11F1E193DB061" bold="true" box="[151,259,262,287]" pageId="7" pageNumber="348">TABLE­2</emphasis>
. Measurements (mm) of
<taxonomicName id="C4BD4D761507526567541F1F1AF1B060" box="[530,719,263,287]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71507526567541F1F1AF1B060" box="[530,719,263,287]" italics="true" pageId="7" pageNumber="348">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE71507526567931F1F1B15B061" bold="true" box="[725,811,263,287]" pageId="7" pageNumber="348">sp.­nov.</emphasis>
Abbreviations defined in text. *holotype.
</tableNote>
</paragraph>
<paragraph id="030236F51507526564161F371D3CB47E" pageId="7" pageNumber="348">
<table id="71BDC4551507AD9D65E01F361D4EB47E" box="[166,1392,302,1280]" gridcols="8" gridrows="22" pageId="7" pageNumber="348">
<tr id="BD8D34B71507AD9D65E01F361D4EB0CD" box="[166,1392,302,435]" gridrow="0" pageId="7" pageNumber="348" rowspan-0="1">
<th id="FE5C5DCB1507AD9D64161F361984B0CD" box="[336,442,302,435]" gridcol="1" gridrow="0" pageId="7" pageNumber="348">UNS 00342/ AMS R176204</th>
<th id="FE5C5DCB1507AD9D64A51F361A73B0CD" box="[483,589,302,435]" gridcol="2" gridrow="0" pageId="7" pageNumber="348">UNS 00343/ AMS R176205</th>
<th id="FE5C5DCB1507AD9D673F1F361ADAB0CD" box="[633,740,302,435]" gridcol="3" gridrow="0" pageId="7" pageNumber="348">
<emphasis id="31C9EAE715075265673F1F361ADAB015" bold="true" box="[633,740,302,363]" pageId="7" pageNumber="348">AMS­ R176206*</emphasis>
</th>
<th id="FE5C5DCB1507AD9D66491F361B47B0CD" box="[783,889,302,435]" gridcol="4" gridrow="0" pageId="7" pageNumber="348">AMS R176207</th>
<th id="FE5C5DCB1507AD9D66EC1F361C2AB0CD" box="[938,1044,302,435]" gridcol="5" gridrow="0" pageId="7" pageNumber="348">NCSM 79748</th>
<th id="FE5C5DCB1507AD9D61061F361CA3B0CD" box="[1088,1181,302,435]" gridcol="6" gridrow="0" pageId="7" pageNumber="348">UNS 00351/ AMS R176208</th>
<th id="FE5C5DCB1507AD9D61901F361D4EB0CD" box="[1238,1392,302,435]" gridcol="7" gridrow="0" pageId="7" pageNumber="348">AMS R176209</th>
</tr>
<tr id="BD8D34B71507AD9D65E01FDA1D4EB0A5" box="[166,1392,450,475]" gridrow="1" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01FDA192CB0A5" box="[166,274,450,475]" gridcol="0" gridrow="1" pageId="7" pageNumber="348">Sex</th>
<td id="FE5C5DCB1507AD9D64161FDA1984B0A5" box="[336,442,450,475]" gridcol="1" gridrow="1" pageId="7" pageNumber="348">adult male</td>
<td id="FE5C5DCB1507AD9D64A51FDA1A73B0A5" box="[483,589,450,475]" gridcol="2" gridrow="1" pageId="7" pageNumber="348">adult male</td>
<td id="FE5C5DCB1507AD9D673F1FDA1ADAB0A5" box="[633,740,450,475]" gridcol="3" gridrow="1" pageId="7" pageNumber="348">adult male</td>
<td id="FE5C5DCB1507AD9D66491FDA1B47B0A5" box="[783,889,450,475]" gridcol="4" gridrow="1" pageId="7" pageNumber="348">adult male</td>
<td id="FE5C5DCB1507AD9D66EC1FDA1C2AB0A5" box="[938,1044,450,475]" gridcol="5" gridrow="1" pageId="7" pageNumber="348">adult male</td>
<td id="FE5C5DCB1507AD9D61061FDA1CA3B0A5" box="[1088,1181,450,475]" gridcol="6" gridrow="1" pageId="7" pageNumber="348">juvenile</td>
<td id="FE5C5DCB1507AD9D61901FDA1D4EB0A5" box="[1238,1392,450,475]" gridcol="7" gridrow="1" pageId="7" pageNumber="348">juvenile</td>
</tr>
<tr id="BD8D34B71507AD9D65E01FF31D4EB37D" box="[166,1392,491,515]" gridrow="2" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01FF3192CB37D" box="[166,274,491,515]" gridcol="0" gridrow="2" pageId="7" pageNumber="348">SVL</th>
<td id="FE5C5DCB1507AD9D64161FF31984B37D" box="[336,442,491,515]" gridcol="1" gridrow="2" pageId="7" pageNumber="348">40.9</td>
<td id="FE5C5DCB1507AD9D64A51FF31A73B37D" box="[483,589,491,515]" gridcol="2" gridrow="2" pageId="7" pageNumber="348">37.8</td>
<td id="FE5C5DCB1507AD9D673F1FF31ADAB37D" box="[633,740,491,515]" gridcol="3" gridrow="2" pageId="7" pageNumber="348">37.2</td>
<td id="FE5C5DCB1507AD9D66491FF31B47B37D" box="[783,889,491,515]" gridcol="4" gridrow="2" pageId="7" pageNumber="348">38.2</td>
<td id="FE5C5DCB1507AD9D66EC1FF31C2AB37D" box="[938,1044,491,515]" gridcol="5" gridrow="2" pageId="7" pageNumber="348">41.4</td>
<td id="FE5C5DCB1507AD9D61061FF31CA3B37D" box="[1088,1181,491,515]" gridcol="6" gridrow="2" pageId="7" pageNumber="348">24.3</td>
<td id="FE5C5DCB1507AD9D61901FF31D4EB37D" box="[1238,1392,491,515]" gridcol="7" gridrow="2" pageId="7" pageNumber="348">19.9</td>
</tr>
<tr id="BD8D34B71507AD9D65E01C0B1D4EB352" box="[166,1392,531,556]" gridrow="3" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01C0B192CB352" box="[166,274,531,556]" gridcol="0" gridrow="3" pageId="7" pageNumber="348">HDL</th>
<td id="FE5C5DCB1507AD9D64161C0B1984B352" box="[336,442,531,556]" gridcol="1" gridrow="3" pageId="7" pageNumber="348">12.5</td>
<td id="FE5C5DCB1507AD9D64A51C0B1A73B352" box="[483,589,531,556]" gridcol="2" gridrow="3" pageId="7" pageNumber="348">12.2</td>
<td id="FE5C5DCB1507AD9D673F1C0B1ADAB352" box="[633,740,531,556]" gridcol="3" gridrow="3" pageId="7" pageNumber="348">13</td>
<td id="FE5C5DCB1507AD9D66491C0B1B47B352" box="[783,889,531,556]" gridcol="4" gridrow="3" pageId="7" pageNumber="348">13.4</td>
<td id="FE5C5DCB1507AD9D66EC1C0B1C2AB352" box="[938,1044,531,556]" gridcol="5" gridrow="3" pageId="7" pageNumber="348">13.8</td>
<td id="FE5C5DCB1507AD9D61061C0B1CA3B352" box="[1088,1181,531,556]" gridcol="6" gridrow="3" pageId="7" pageNumber="348">9.4</td>
<td id="FE5C5DCB1507AD9D61901C0B1D4EB352" box="[1238,1392,531,556]" gridcol="7" gridrow="3" pageId="7" pageNumber="348">7.5</td>
</tr>
<tr id="BD8D34B71507AD9D65E01C231D4EB32A" box="[166,1392,571,596]" gridrow="4" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01C23192CB32A" box="[166,274,571,596]" gridcol="0" gridrow="4" pageId="7" pageNumber="348">HDW</th>
<td id="FE5C5DCB1507AD9D64161C231984B32A" box="[336,442,571,596]" gridcol="1" gridrow="4" pageId="7" pageNumber="348">14.9</td>
<td id="FE5C5DCB1507AD9D64A51C231A73B32A" box="[483,589,571,596]" gridcol="2" gridrow="4" pageId="7" pageNumber="348">14.2</td>
<td id="FE5C5DCB1507AD9D673F1C231ADAB32A" box="[633,740,571,596]" gridcol="3" gridrow="4" pageId="7" pageNumber="348">14.3</td>
<td id="FE5C5DCB1507AD9D66491C231B47B32A" box="[783,889,571,596]" gridcol="4" gridrow="4" pageId="7" pageNumber="348">14.5</td>
<td id="FE5C5DCB1507AD9D66EC1C231C2AB32A" box="[938,1044,571,596]" gridcol="5" gridrow="4" pageId="7" pageNumber="348">15.1</td>
<td id="FE5C5DCB1507AD9D61061C231CA3B32A" box="[1088,1181,571,596]" gridcol="6" gridrow="4" pageId="7" pageNumber="348">9.4</td>
<td id="FE5C5DCB1507AD9D61901C231D4EB32A" box="[1238,1392,571,596]" gridcol="7" gridrow="4" pageId="7" pageNumber="348">7.5</td>
</tr>
<tr id="BD8D34B71507AD9D65E01C7B1D4EB302" box="[166,1392,611,636]" gridrow="5" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01C7B192CB302" box="[166,274,611,636]" gridcol="0" gridrow="5" pageId="7" pageNumber="348">SNT</th>
<td id="FE5C5DCB1507AD9D64161C7B1984B302" box="[336,442,611,636]" gridcol="1" gridrow="5" pageId="7" pageNumber="348">6.1</td>
<td id="FE5C5DCB1507AD9D64A51C7B1A73B302" box="[483,589,611,636]" gridcol="2" gridrow="5" pageId="7" pageNumber="348">5.3</td>
<td id="FE5C5DCB1507AD9D673F1C7B1ADAB302" box="[633,740,611,636]" gridcol="3" gridrow="5" pageId="7" pageNumber="348">5.9</td>
<td id="FE5C5DCB1507AD9D66491C7B1B47B302" box="[783,889,611,636]" gridcol="4" gridrow="5" pageId="7" pageNumber="348">5.6</td>
<td id="FE5C5DCB1507AD9D66EC1C7B1C2AB302" box="[938,1044,611,636]" gridcol="5" gridrow="5" pageId="7" pageNumber="348">6.0</td>
<td id="FE5C5DCB1507AD9D61061C7B1CA3B302" box="[1088,1181,611,636]" gridcol="6" gridrow="5" pageId="7" pageNumber="348">3.5</td>
<td id="FE5C5DCB1507AD9D61901C7B1D4EB302" box="[1238,1392,611,636]" gridcol="7" gridrow="5" pageId="7" pageNumber="348">2.9</td>
</tr>
<tr id="BD8D34B71507AD9D65E01C941D4EB3DA" box="[166,1392,652,676]" gridrow="6" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01C94192CB3DA" box="[166,274,652,676]" gridcol="0" gridrow="6" pageId="7" pageNumber="348">EYE</th>
<td id="FE5C5DCB1507AD9D64161C941984B3DA" box="[336,442,652,676]" gridcol="1" gridrow="6" pageId="7" pageNumber="348">4.4</td>
<td id="FE5C5DCB1507AD9D64A51C941A73B3DA" box="[483,589,652,676]" gridcol="2" gridrow="6" pageId="7" pageNumber="348">4.0</td>
<td id="FE5C5DCB1507AD9D673F1C941ADAB3DA" box="[633,740,652,676]" gridcol="3" gridrow="6" pageId="7" pageNumber="348">4.4</td>
<td id="FE5C5DCB1507AD9D66491C941B47B3DA" box="[783,889,652,676]" gridcol="4" gridrow="6" pageId="7" pageNumber="348">4.7</td>
<td id="FE5C5DCB1507AD9D66EC1C941C2AB3DA" box="[938,1044,652,676]" gridcol="5" gridrow="6" pageId="7" pageNumber="348">4.8</td>
<td id="FE5C5DCB1507AD9D61061C941CA3B3DA" box="[1088,1181,652,676]" gridcol="6" gridrow="6" pageId="7" pageNumber="348">3.0</td>
<td id="FE5C5DCB1507AD9D61901C941D4EB3DA" box="[1238,1392,652,676]" gridcol="7" gridrow="6" pageId="7" pageNumber="348">2.9</td>
</tr>
<tr id="BD8D34B71507AD9D65E01CAC1D4EB3B3" box="[166,1392,692,717]" gridrow="7" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01CAC192CB3B3" box="[166,274,692,717]" gridcol="0" gridrow="7" pageId="7" pageNumber="348">IOD</th>
<td id="FE5C5DCB1507AD9D64161CAC1984B3B3" box="[336,442,692,717]" gridcol="1" gridrow="7" pageId="7" pageNumber="348">4.6</td>
<td id="FE5C5DCB1507AD9D64A51CAC1A73B3B3" box="[483,589,692,717]" gridcol="2" gridrow="7" pageId="7" pageNumber="348">3.9</td>
<td id="FE5C5DCB1507AD9D673F1CAC1ADAB3B3" box="[633,740,692,717]" gridcol="3" gridrow="7" pageId="7" pageNumber="348">4.1</td>
<td id="FE5C5DCB1507AD9D66491CAC1B47B3B3" box="[783,889,692,717]" gridcol="4" gridrow="7" pageId="7" pageNumber="348">4.0</td>
<td id="FE5C5DCB1507AD9D66EC1CAC1C2AB3B3" box="[938,1044,692,717]" gridcol="5" gridrow="7" pageId="7" pageNumber="348">3.9</td>
<td id="FE5C5DCB1507AD9D61061CAC1CA3B3B3" box="[1088,1181,692,717]" gridcol="6" gridrow="7" pageId="7" pageNumber="348">2.9</td>
<td id="FE5C5DCB1507AD9D61901CAC1D4EB3B3" box="[1238,1392,692,717]" gridcol="7" gridrow="7" pageId="7" pageNumber="348">2.7</td>
</tr>
<tr id="BD8D34B71507AD9D65E01CC51D4EB38B" box="[166,1392,733,757]" gridrow="8" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01CC5192CB38B" box="[166,274,733,757]" gridcol="0" gridrow="8" pageId="7" pageNumber="348">TMP</th>
<td id="FE5C5DCB1507AD9D64161CC51984B38B" box="[336,442,733,757]" gridcol="1" gridrow="8" pageId="7" pageNumber="348">1.8</td>
<td id="FE5C5DCB1507AD9D64A51CC51A73B38B" box="[483,589,733,757]" gridcol="2" gridrow="8" pageId="7" pageNumber="348">1.7</td>
<td id="FE5C5DCB1507AD9D673F1CC51ADAB38B" box="[633,740,733,757]" gridcol="3" gridrow="8" pageId="7" pageNumber="348">2.2</td>
<td id="FE5C5DCB1507AD9D66491CC51B47B38B" box="[783,889,733,757]" gridcol="4" gridrow="8" pageId="7" pageNumber="348">2.1</td>
<td id="FE5C5DCB1507AD9D66EC1CC51C2AB38B" box="[938,1044,733,757]" gridcol="5" gridrow="8" pageId="7" pageNumber="348">2.0</td>
<td id="FE5C5DCB1507AD9D61061CC51CA3B38B" box="[1088,1181,733,757]" gridcol="6" gridrow="8" pageId="7" pageNumber="348">1.3</td>
<td id="FE5C5DCB1507AD9D61901CC51D4EB38B" box="[1238,1392,733,757]" gridcol="7" gridrow="8" pageId="7" pageNumber="348">1.1</td>
</tr>
<tr id="BD8D34B71507AD9D65E01D1C1D4EB263" box="[166,1392,772,797]" gridrow="9" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01D1C192CB263" box="[166,274,772,797]" gridcol="0" gridrow="9" pageId="7" pageNumber="348">TEY</th>
<td id="FE5C5DCB1507AD9D64161D1C1984B263" box="[336,442,772,797]" gridcol="1" gridrow="9" pageId="7" pageNumber="348">1.1</td>
<td id="FE5C5DCB1507AD9D64A51D1C1A73B263" box="[483,589,772,797]" gridcol="2" gridrow="9" pageId="7" pageNumber="348">1.5</td>
<td id="FE5C5DCB1507AD9D673F1D1C1ADAB263" box="[633,740,772,797]" gridcol="3" gridrow="9" pageId="7" pageNumber="348">1.2</td>
<td id="FE5C5DCB1507AD9D66491D1C1B47B263" box="[783,889,772,797]" gridcol="4" gridrow="9" pageId="7" pageNumber="348">1.6</td>
<td id="FE5C5DCB1507AD9D66EC1D1C1C2AB263" box="[938,1044,772,797]" gridcol="5" gridrow="9" pageId="7" pageNumber="348">1.7</td>
<td id="FE5C5DCB1507AD9D61061D1C1CA3B263" box="[1088,1181,772,797]" gridcol="6" gridrow="9" pageId="7" pageNumber="348">0.8</td>
<td id="FE5C5DCB1507AD9D61901D1C1D4EB263" box="[1238,1392,772,797]" gridcol="7" gridrow="9" pageId="7" pageNumber="348">0.7</td>
</tr>
<tr id="BD8D34B71507AD9D65E01D351D4EB238" box="[166,1392,813,838]" gridrow="10" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01D35192CB238" box="[166,274,813,838]" gridcol="0" gridrow="10" pageId="7" pageNumber="348">IN</th>
<td id="FE5C5DCB1507AD9D64161D351984B238" box="[336,442,813,838]" gridcol="1" gridrow="10" pageId="7" pageNumber="348">4.4</td>
<td id="FE5C5DCB1507AD9D64A51D351A73B238" box="[483,589,813,838]" gridcol="2" gridrow="10" pageId="7" pageNumber="348">4.2</td>
<td id="FE5C5DCB1507AD9D673F1D351ADAB238" box="[633,740,813,838]" gridcol="3" gridrow="10" pageId="7" pageNumber="348">4.1</td>
<td id="FE5C5DCB1507AD9D66491D351B47B238" box="[783,889,813,838]" gridcol="4" gridrow="10" pageId="7" pageNumber="348">4.2</td>
<td id="FE5C5DCB1507AD9D66EC1D351C2AB238" box="[938,1044,813,838]" gridcol="5" gridrow="10" pageId="7" pageNumber="348">4.5</td>
<td id="FE5C5DCB1507AD9D61061D351CA3B238" box="[1088,1181,813,838]" gridcol="6" gridrow="10" pageId="7" pageNumber="348">2.9</td>
<td id="FE5C5DCB1507AD9D61901D351D4EB238" box="[1238,1392,813,838]" gridcol="7" gridrow="10" pageId="7" pageNumber="348">2.3</td>
</tr>
<tr id="BD8D34B71507AD9D65E01D4D1D4EB210" box="[166,1392,853,878]" gridrow="11" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01D4D192CB210" box="[166,274,853,878]" gridcol="0" gridrow="11" pageId="7" pageNumber="348">NS</th>
<td id="FE5C5DCB1507AD9D64161D4D1984B210" box="[336,442,853,878]" gridcol="1" gridrow="11" pageId="7" pageNumber="348">2.5</td>
<td id="FE5C5DCB1507AD9D64A51D4D1A73B210" box="[483,589,853,878]" gridcol="2" gridrow="11" pageId="7" pageNumber="348">2.3</td>
<td id="FE5C5DCB1507AD9D673F1D4D1ADAB210" box="[633,740,853,878]" gridcol="3" gridrow="11" pageId="7" pageNumber="348">2.6</td>
<td id="FE5C5DCB1507AD9D66491D4D1B47B210" box="[783,889,853,878]" gridcol="4" gridrow="11" pageId="7" pageNumber="348">2.7</td>
<td id="FE5C5DCB1507AD9D66EC1D4D1C2AB210" box="[938,1044,853,878]" gridcol="5" gridrow="11" pageId="7" pageNumber="348">2.3</td>
<td id="FE5C5DCB1507AD9D61061D4D1CA3B210" box="[1088,1181,853,878]" gridcol="6" gridrow="11" pageId="7" pageNumber="348">1.5</td>
<td id="FE5C5DCB1507AD9D61901D4D1D4EB210" box="[1238,1392,853,878]" gridcol="7" gridrow="11" pageId="7" pageNumber="348">0.9</td>
</tr>
<tr id="BD8D34B71507AD9D65E01D661D4EB2E8" box="[166,1392,894,918]" gridrow="12" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01D66192CB2E8" box="[166,274,894,918]" gridcol="0" gridrow="12" pageId="7" pageNumber="348">EN</th>
<td id="FE5C5DCB1507AD9D64161D661984B2E8" box="[336,442,894,918]" gridcol="1" gridrow="12" pageId="7" pageNumber="348">3.2</td>
<td id="FE5C5DCB1507AD9D64A51D661A73B2E8" box="[483,589,894,918]" gridcol="2" gridrow="12" pageId="7" pageNumber="348">2.8</td>
<td id="FE5C5DCB1507AD9D673F1D661ADAB2E8" box="[633,740,894,918]" gridcol="3" gridrow="12" pageId="7" pageNumber="348">2.9</td>
<td id="FE5C5DCB1507AD9D66491D661B47B2E8" box="[783,889,894,918]" gridcol="4" gridrow="12" pageId="7" pageNumber="348">3.2</td>
<td id="FE5C5DCB1507AD9D66EC1D661C2AB2E8" box="[938,1044,894,918]" gridcol="5" gridrow="12" pageId="7" pageNumber="348">3.7</td>
<td id="FE5C5DCB1507AD9D61061D661CA3B2E8" box="[1088,1181,894,918]" gridcol="6" gridrow="12" pageId="7" pageNumber="348">2.2</td>
<td id="FE5C5DCB1507AD9D61901D661D4EB2E8" box="[1238,1392,894,918]" gridcol="7" gridrow="12" pageId="7" pageNumber="348">1.9</td>
</tr>
<tr id="BD8D34B71507AD9D65E01DBE1D4EB2C0" box="[166,1392,934,958]" gridrow="13" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01DBE192CB2C0" box="[166,274,934,958]" gridcol="0" gridrow="13" pageId="7" pageNumber="348">TIB</th>
<td id="FE5C5DCB1507AD9D64161DBE1984B2C0" box="[336,442,934,958]" gridcol="1" gridrow="13" pageId="7" pageNumber="348">19.2</td>
<td id="FE5C5DCB1507AD9D64A51DBE1A73B2C0" box="[483,589,934,958]" gridcol="2" gridrow="13" pageId="7" pageNumber="348">19.3</td>
<td id="FE5C5DCB1507AD9D673F1DBE1ADAB2C0" box="[633,740,934,958]" gridcol="3" gridrow="13" pageId="7" pageNumber="348">18.3</td>
<td id="FE5C5DCB1507AD9D66491DBE1B47B2C0" box="[783,889,934,958]" gridcol="4" gridrow="13" pageId="7" pageNumber="348">18.7</td>
<td id="FE5C5DCB1507AD9D66EC1DBE1C2AB2C0" box="[938,1044,934,958]" gridcol="5" gridrow="13" pageId="7" pageNumber="348">20.2</td>
<td id="FE5C5DCB1507AD9D61061DBE1CA3B2C0" box="[1088,1181,934,958]" gridcol="6" gridrow="13" pageId="7" pageNumber="348">12.3</td>
<td id="FE5C5DCB1507AD9D61901DBE1D4EB2C0" box="[1238,1392,934,958]" gridcol="7" gridrow="13" pageId="7" pageNumber="348">9.3</td>
</tr>
<tr id="BD8D34B71507AD9D65E01DD51D4EB298" box="[166,1392,973,998]" gridrow="14" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01DD5192CB298" box="[166,274,973,998]" gridcol="0" gridrow="14" pageId="7" pageNumber="348">ML</th>
<td id="FE5C5DCB1507AD9D64161DD51984B298" box="[336,442,973,998]" gridcol="1" gridrow="14" pageId="7" pageNumber="348">14.0</td>
<td id="FE5C5DCB1507AD9D64A51DD51A73B298" box="[483,589,973,998]" gridcol="2" gridrow="14" pageId="7" pageNumber="348">13.3</td>
<td id="FE5C5DCB1507AD9D673F1DD51ADAB298" box="[633,740,973,998]" gridcol="3" gridrow="14" pageId="7" pageNumber="348">12.7</td>
<td id="FE5C5DCB1507AD9D66491DD51B47B298" box="[783,889,973,998]" gridcol="4" gridrow="14" pageId="7" pageNumber="348">14.0</td>
<td id="FE5C5DCB1507AD9D66EC1DD51C2AB298" box="[938,1044,973,998]" gridcol="5" gridrow="14" pageId="7" pageNumber="348">14.0</td>
<td id="FE5C5DCB1507AD9D61061DD51CA3B298" box="[1088,1181,973,998]" gridcol="6" gridrow="14" pageId="7" pageNumber="348">7.6</td>
<td id="FE5C5DCB1507AD9D61901DD51D4EB298" box="[1238,1392,973,998]" gridcol="7" gridrow="14" pageId="7" pageNumber="348">6.4</td>
</tr>
<tr id="BD8D34B71507AD9D65E01DEE1D4EB571" box="[166,1392,1014,1039]" gridrow="15" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01DEE192CB571" box="[166,274,1014,1039]" gridcol="0" gridrow="15" pageId="7" pageNumber="348">PL</th>
<td id="FE5C5DCB1507AD9D64161DEE1984B571" box="[336,442,1014,1039]" gridcol="1" gridrow="15" pageId="7" pageNumber="348">18.6</td>
<td id="FE5C5DCB1507AD9D64A51DEE1A73B571" box="[483,589,1014,1039]" gridcol="2" gridrow="15" pageId="7" pageNumber="348">17.8</td>
<td id="FE5C5DCB1507AD9D673F1DEE1ADAB571" box="[633,740,1014,1039]" gridcol="3" gridrow="15" pageId="7" pageNumber="348">18.1</td>
<td id="FE5C5DCB1507AD9D66491DEE1B47B571" box="[783,889,1014,1039]" gridcol="4" gridrow="15" pageId="7" pageNumber="348">18.7</td>
<td id="FE5C5DCB1507AD9D66EC1DEE1C2AB571" box="[938,1044,1014,1039]" gridcol="5" gridrow="15" pageId="7" pageNumber="348">19.7</td>
<td id="FE5C5DCB1507AD9D61061DEE1CA3B571" box="[1088,1181,1014,1039]" gridcol="6" gridrow="15" pageId="7" pageNumber="348">10.8</td>
<td id="FE5C5DCB1507AD9D61901DEE1D4EB571" box="[1238,1392,1014,1039]" gridcol="7" gridrow="15" pageId="7" pageNumber="348">7.1</td>
</tr>
<tr id="BD8D34B71507AD9D65E01A071D4EB549" box="[166,1392,1055,1079]" gridrow="16" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01A07192CB549" box="[166,274,1055,1079]" gridcol="0" gridrow="16" pageId="7" pageNumber="348">IMT</th>
<td id="FE5C5DCB1507AD9D64161A071984B549" box="[336,442,1055,1079]" gridcol="1" gridrow="16" pageId="7" pageNumber="348">2.1</td>
<td id="FE5C5DCB1507AD9D64A51A071A73B549" box="[483,589,1055,1079]" gridcol="2" gridrow="16" pageId="7" pageNumber="348">1.9</td>
<td id="FE5C5DCB1507AD9D673F1A071ADAB549" box="[633,740,1055,1079]" gridcol="3" gridrow="16" pageId="7" pageNumber="348">2.0</td>
<td id="FE5C5DCB1507AD9D66491A071B47B549" box="[783,889,1055,1079]" gridcol="4" gridrow="16" pageId="7" pageNumber="348">2.2</td>
<td id="FE5C5DCB1507AD9D66EC1A071C2AB549" box="[938,1044,1055,1079]" gridcol="5" gridrow="16" pageId="7" pageNumber="348">2.2</td>
<td id="FE5C5DCB1507AD9D61061A071CA3B549" box="[1088,1181,1055,1079]" gridcol="6" gridrow="16" pageId="7" pageNumber="348">0.9</td>
<td id="FE5C5DCB1507AD9D61901A071D4EB549" box="[1238,1392,1055,1079]" gridcol="7" gridrow="16" pageId="7" pageNumber="348">-</td>
</tr>
<tr id="BD8D34B71507AD9D65E01A5E1D4EB521" box="[166,1392,1094,1119]" gridrow="17" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01A5E192CB521" box="[166,274,1094,1119]" gridcol="0" gridrow="17" pageId="7" pageNumber="348">HW/HL</th>
<td id="FE5C5DCB1507AD9D64161A5E1984B521" box="[336,442,1094,1119]" gridcol="1" gridrow="17" pageId="7" pageNumber="348">1.19</td>
<td id="FE5C5DCB1507AD9D64A51A5E1A73B521" box="[483,589,1094,1119]" gridcol="2" gridrow="17" pageId="7" pageNumber="348">1.16</td>
<td id="FE5C5DCB1507AD9D673F1A5E1ADAB521" box="[633,740,1094,1119]" gridcol="3" gridrow="17" pageId="7" pageNumber="348">1.10</td>
<td id="FE5C5DCB1507AD9D66491A5E1B47B521" box="[783,889,1094,1119]" gridcol="4" gridrow="17" pageId="7" pageNumber="348">1.08</td>
<td id="FE5C5DCB1507AD9D66EC1A5E1C2AB521" box="[938,1044,1094,1119]" gridcol="5" gridrow="17" pageId="7" pageNumber="348">1.09</td>
<td id="FE5C5DCB1507AD9D61061A5E1CA3B521" box="[1088,1181,1094,1119]" gridcol="6" gridrow="17" pageId="7" pageNumber="348">1.00</td>
<td id="FE5C5DCB1507AD9D61901A5E1D4EB521" box="[1238,1392,1094,1119]" gridcol="7" gridrow="17" pageId="7" pageNumber="348">1.00</td>
</tr>
<tr id="BD8D34B71507AD9D65E01A761D4EB5F9" box="[166,1392,1134,1159]" gridrow="18" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01A76192CB5F9" box="[166,274,1134,1159]" gridcol="0" gridrow="18" pageId="7" pageNumber="348">HL/HW</th>
<td id="FE5C5DCB1507AD9D64161A761984B5F9" box="[336,442,1134,1159]" gridcol="1" gridrow="18" pageId="7" pageNumber="348">0.84</td>
<td id="FE5C5DCB1507AD9D64A51A761A73B5F9" box="[483,589,1134,1159]" gridcol="2" gridrow="18" pageId="7" pageNumber="348">0.86</td>
<td id="FE5C5DCB1507AD9D673F1A761ADAB5F9" box="[633,740,1134,1159]" gridcol="3" gridrow="18" pageId="7" pageNumber="348">0.91</td>
<td id="FE5C5DCB1507AD9D66491A761B47B5F9" box="[783,889,1134,1159]" gridcol="4" gridrow="18" pageId="7" pageNumber="348">0.92</td>
<td id="FE5C5DCB1507AD9D66EC1A761C2AB5F9" box="[938,1044,1134,1159]" gridcol="5" gridrow="18" pageId="7" pageNumber="348">0.91</td>
<td id="FE5C5DCB1507AD9D61061A761CA3B5F9" box="[1088,1181,1134,1159]" gridcol="6" gridrow="18" pageId="7" pageNumber="348">1.00</td>
<td id="FE5C5DCB1507AD9D61901A761D4EB5F9" box="[1238,1392,1134,1159]" gridcol="7" gridrow="18" pageId="7" pageNumber="348">1.00</td>
</tr>
<tr id="BD8D34B71507AD9D65E01A8F1D4EB5CE" box="[166,1392,1175,1200]" gridrow="19" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01A8F192CB5CE" box="[166,274,1175,1200]" gridcol="0" gridrow="19" pageId="7" pageNumber="348">TIB/SVL</th>
<td id="FE5C5DCB1507AD9D64161A8F1984B5CE" box="[336,442,1175,1200]" gridcol="1" gridrow="19" pageId="7" pageNumber="348">0.47</td>
<td id="FE5C5DCB1507AD9D64A51A8F1A73B5CE" box="[483,589,1175,1200]" gridcol="2" gridrow="19" pageId="7" pageNumber="348">0.51</td>
<td id="FE5C5DCB1507AD9D673F1A8F1ADAB5CE" box="[633,740,1175,1200]" gridcol="3" gridrow="19" pageId="7" pageNumber="348">0.49</td>
<td id="FE5C5DCB1507AD9D66491A8F1B47B5CE" box="[783,889,1175,1200]" gridcol="4" gridrow="19" pageId="7" pageNumber="348">0.49</td>
<td id="FE5C5DCB1507AD9D66EC1A8F1C2AB5CE" box="[938,1044,1175,1200]" gridcol="5" gridrow="19" pageId="7" pageNumber="348">0.49</td>
<td id="FE5C5DCB1507AD9D61061A8F1CA3B5CE" box="[1088,1181,1175,1200]" gridcol="6" gridrow="19" pageId="7" pageNumber="348">0.51</td>
<td id="FE5C5DCB1507AD9D61901A8F1D4EB5CE" box="[1238,1392,1175,1200]" gridcol="7" gridrow="19" pageId="7" pageNumber="348">0.47</td>
</tr>
<tr id="BD8D34B71507AD9D65E01AA71D4EB5A6" box="[166,1392,1215,1240]" gridrow="20" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01AA7192CB5A6" box="[166,274,1215,1240]" gridcol="0" gridrow="20" pageId="7" pageNumber="348">HDL/SVL</th>
<td id="FE5C5DCB1507AD9D64161AA71984B5A6" box="[336,442,1215,1240]" gridcol="1" gridrow="20" pageId="7" pageNumber="348">0.31</td>
<td id="FE5C5DCB1507AD9D64A51AA71A73B5A6" box="[483,589,1215,1240]" gridcol="2" gridrow="20" pageId="7" pageNumber="348">0.32</td>
<td id="FE5C5DCB1507AD9D673F1AA71ADAB5A6" box="[633,740,1215,1240]" gridcol="3" gridrow="20" pageId="7" pageNumber="348">0.35</td>
<td id="FE5C5DCB1507AD9D66491AA71B47B5A6" box="[783,889,1215,1240]" gridcol="4" gridrow="20" pageId="7" pageNumber="348">0.35</td>
<td id="FE5C5DCB1507AD9D66EC1AA71C2AB5A6" box="[938,1044,1215,1240]" gridcol="5" gridrow="20" pageId="7" pageNumber="348">0.33</td>
<td id="FE5C5DCB1507AD9D61061AA71CA3B5A6" box="[1088,1181,1215,1240]" gridcol="6" gridrow="20" pageId="7" pageNumber="348">0.39</td>
<td id="FE5C5DCB1507AD9D61901AA71D4EB5A6" box="[1238,1392,1215,1240]" gridcol="7" gridrow="20" pageId="7" pageNumber="348">0.38</td>
</tr>
<tr id="BD8D34B71507AD9D65E01AFF1D4EB47E" box="[166,1392,1255,1280]" gridrow="21" pageId="7" pageNumber="348">
<th id="FE5C5DCB1507AD9D65E01AFF192CB47E" box="[166,274,1255,1280]" gridcol="0" gridrow="21" pageId="7" pageNumber="348">TMP/EYE</th>
<td id="FE5C5DCB1507AD9D64161AFF1984B47E" box="[336,442,1255,1280]" gridcol="1" gridrow="21" pageId="7" pageNumber="348">0.41</td>
<td id="FE5C5DCB1507AD9D64A51AFF1A73B47E" box="[483,589,1255,1280]" gridcol="2" gridrow="21" pageId="7" pageNumber="348">0.43</td>
<td id="FE5C5DCB1507AD9D673F1AFF1ADAB47E" box="[633,740,1255,1280]" gridcol="3" gridrow="21" pageId="7" pageNumber="348">0.50</td>
<td id="FE5C5DCB1507AD9D66491AFF1B47B47E" box="[783,889,1255,1280]" gridcol="4" gridrow="21" pageId="7" pageNumber="348">0.45</td>
<td id="FE5C5DCB1507AD9D66EC1AFF1C2AB47E" box="[938,1044,1255,1280]" gridcol="5" gridrow="21" pageId="7" pageNumber="348">0.42</td>
<td id="FE5C5DCB1507AD9D61061AFF1CA3B47E" box="[1088,1181,1255,1280]" gridcol="6" gridrow="21" pageId="7" pageNumber="348">0.43</td>
<td id="FE5C5DCB1507AD9D61901AFF1D4EB47E" box="[1238,1392,1255,1280]" gridcol="7" gridrow="21" pageId="7" pageNumber="348">0.38</td>
</tr>
</table>
</paragraph>
<paragraph id="030236F51507526565811B371C6EB7CD" blockId="7.[151,1437,1327,2002]" pageId="7" pageNumber="348">
<emphasis id="31C9EAE71507526565811B371A47B434" bold="true" box="[199,633,1327,1354]" pageId="7" pageNumber="348">Distribution­and­conservation­status</emphasis>
: The new species is known only from between
<quantity id="C4459B101507526561CB1B371D16B434" box="[1165,1320,1327,1354]" metricMagnitude="3" metricUnit="m" metricValue="1.8855" metricValueMax="2.055" metricValueMin="1.716" pageId="7" pageNumber="348" unit="m" value="1885.5" valueMax="2055.0" valueMin="1716.0">1716–2055 m</quantity>
asl within Ngoc Linh Nature Reserve,
<collectingRegion id="C179F8171507526564881B4C1A9DB410" box="[462,675,1364,1390]" country="Vietnam" name="Kon Tum" pageId="7" pageNumber="348">Kon Tum Province</collectingRegion>
,
<collectingCountry id="7BAA76651507526567EB1B4C1B2EB410" box="[685,784,1364,1390]" name="Vietnam" pageId="7" pageNumber="348">Vietnam</collectingCountry>
. Mount Ngoc Linh and adjacent high-elevation peaks form an isolated area of high elevation and many faunal elements at this location are thought to be endemic (
<bibRefCitation id="672C4B041507526560501B6018CBB4CB" author="Abramov, A. V. &amp; Rozhnov, V. V. &amp; Morozov, P. N." pageId="7" pageNumber="348" pagination="85 - 92" refId="ref7423" refString="Abramov, A. V., Rozhnov, V. V. &amp; Morozov, P. N. (2006) Notes on mammals of the Ngoc Linh Nature Reserve (Vietnam, Kon Tum Province). Russian Journal of Therology, 5, 85 - 92. https: // doi. org / 10.15298 / rusjtheriol. 05.2.06" type="journal article" year="2006">
Abramov
<emphasis id="31C9EAE71507526560CE1B601893B4C8" italics="true" pageId="7" pageNumber="348">et al</emphasis>
. 2006
</bibRefCitation>
,
<bibRefCitation id="672C4B041507526565B91B8419EDB4C8" author="Jenkins, P. D. &amp; Abramov, A. V. &amp; Rozhnov, V. V. &amp; Makarova, O. V." box="[255,467,1436,1462]" pageId="7" pageNumber="348" pagination="57 - 68" refId="ref8088" refString="Jenkins, P. D., Abramov, A. V., Rozhnov, V. V. &amp; Makarova, O. V. (2007) Description of two new species of white-toothed shrews belonging to the genus Crocidura (Soricomorpha: Soricidae) from Ngoc Linh Mountain, Vietnam. Zootaxa, 1589 (1), 57 - 68. https: // doi. org / 10.11646 / zootaxa. 1589.1.5" type="journal article" year="2007">
Jenkins
<emphasis id="31C9EAE715075265641F1B8419AFB4C8" box="[345,401,1436,1462]" italics="true" pageId="7" pageNumber="348">et al.</emphasis>
2007
</bibRefCitation>
). The species was not located at lower elevations, despite surveys over 19 nights during March/
<date id="770310351507526565AE1BD8195FB4A4" box="[232,353,1472,1498]" pageId="7" pageNumber="348" value="2009-04">April 2009</date>
and
<date id="770310351507526564D01BD81A3DB4A4" box="[406,515,1472,1498]" pageId="7" pageNumber="348" value="2010-07">July 2010</date>
at elevations from ~
<quantity id="C4459B101507526567A01BD81B13B4A4" box="[742,813,1472,1498]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="7" pageNumber="348" unit="m" value="930.0">930 m</quantity>
and above (Rowley
<emphasis id="31C9EAE71507526561571BD81C7CB4A4" box="[1041,1090,1472,1498]" italics="true" pageId="7" pageNumber="348">et al</emphasis>
. unpub. data). The range of the new species is not expected to extend outside of Mount Ngoc Linh and closely surrounding peaks but may occur in adjacent
<collectingRegion id="C179F8171507526565BA181019CCB75C" box="[252,498,1544,1570]" country="Vietnam" name="Quang Nam" pageId="7" pageNumber="348">Quang Nam Province</collectingRegion>
. Thus, we assume that the new species is restricted geographically, likely having an extent of occurrence (EOO) of &lt;
<quantity id="C4459B1015075265674318341A57B738" box="[517,617,1580,1606]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="348" unit="km" value="1000.0">1000 km</quantity>
<superScript id="F4C89BBD15075265672F18321A4FB746" attach="left" box="[617,625,1578,1592]" fontSize="6" pageId="7" pageNumber="348">2</superScript>
. The new species also most likely occurs in one threat-defined location, which has been characterized as an area with continuing decline in the quality of its habitat due to deforestation (
<bibRefCitation id="672C4B041507526565D8186C19A8B7F0" author="Meyfroidt, P. &amp; Vu, T. P. &amp; Hoang, V. A." box="[158,406,1652,1678]" pageId="7" pageNumber="348" pagination="1187 - 1198" refId="ref8434" refString="Meyfroidt, P., Vu, T. P. &amp; Hoang, V. A. (2013) Trajectories of deforestation, coffee expansion and displacement of shifting cultivation in the Central Highlands of Vietnam. Global Environmental Change, 23, 1187 - 1198. https: // doi. org / 10.1016 / j. gloenvcha. 2013.04.005" type="journal article" year="2013">
Meyfroidt
<emphasis id="31C9EAE715075265645F186C1975B7F0" box="[281,331,1652,1678]" italics="true" pageId="7" pageNumber="348">et al</emphasis>
. 2013
</bibRefCitation>
).
<taxonomicName id="C4BD4D761507526564EF186C1A51B7F3" box="[425,623,1652,1678]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71507526564EF186C1A51B7F3" box="[425,623,1652,1678]" italics="true" pageId="7" pageNumber="348">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE7150752656729186C1AEEB7F0" bold="true" box="[623,720,1652,1678]" pageId="7" pageNumber="348">­sp.­nov.</emphasis>
therefore likely qualifies as Endangered in accordance with the IUCN Red List of Threatened Species categories and criteria (
<bibRefCitation id="672C4B0415075265660818801BE8B7CC" author="IUCN" box="[846,982,1688,1714]" pageId="7" pageNumber="348" refId="ref8059" refString="IUCN (2012) IUCN Red List Categories and Criteria: Version 3.1. 2 nd Edition. IUCN, Gland and Cambridge, iv + 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
) B1ab(iii).
</paragraph>
<paragraph id="030236F515075265658118A31A60B6AF" blockId="7.[151,1437,1327,2002]" pageId="7" pageNumber="348">
<emphasis id="31C9EAE715075265658118A31951B7A8" bold="true" box="[199,367,1723,1750]" pageId="7" pageNumber="348">Comparisons:</emphasis>
Within the genus
<taxonomicName id="C4BD4D7615075265677718A41A93B7A8" baseAuthorityName="Frost" baseAuthorityYear="2020" box="[561,685,1724,1750]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="genus">
<emphasis id="31C9EAE715075265677718A41A93B7A8" box="[561,685,1724,1750]" italics="true" pageId="7" pageNumber="348">Gracixalus</emphasis>
</taxonomicName>
,
<taxonomicName id="C4BD4D761507526567FC18A41B41B7AB" box="[698,895,1724,1750]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71507526567FC18A41B41B7AB" box="[698,895,1724,1750]" italics="true" pageId="7" pageNumber="348">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE71507526566C218A41BE3B7A8" bold="true" box="[900,989,1724,1750]" pageId="7" pageNumber="348">sp.­nov.</emphasis>
is distinguished from Clade I of
<bibRefCitation id="672C4B0415075265600318A4191FB784" author="Rowley, J. J. L. &amp; Dau, Q. V. &amp; Nguyen, T. T. &amp; Cao, T. T. &amp; Nguyen, S. N." pageId="7" pageNumber="348" pagination="22 - 38" refId="ref8737" refString="Rowley, J. J. L., Dau, Q. V., Nguyen, T. T., Cao, T. T. &amp; Nguyen, S. N. (2011) A new species of Gracixalus (Anura: Rhacophoridae) with a hyperextended vocal repertoire from Vietnam. Zootaxa, 3125, 22 - 38." type="journal article" year="2011">
Rowley
<emphasis id="31C9EAE71507526565D118F818F9B784" box="[151,199,1760,1786]" italics="true" pageId="7" pageNumber="348">et al</emphasis>
. (2011)
</bibRefCitation>
;
<taxonomicName id="C4BD4D7615075265646A18F81A12B784" authority="(Bourret)" baseAuthorityName="Bourret" baseAuthorityYear="1937" box="[300,556,1760,1786]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="gracilipes">
<emphasis id="31C9EAE715075265646A18F81983B784" box="[300,445,1760,1786]" italics="true" pageId="7" pageNumber="348">G. gracilipes</emphasis>
(Bourret)
</taxonomicName>
<taxonomicName id="C4BD4D7615075265677718F81BA1B784" authority="Rowley, Dau, Nguyen" authorityName="Rowley, Dau, Nguyen" box="[561,927,1760,1786]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="quangi">
<emphasis id="31C9EAE715075265677718F81A9EB787" box="[561,672,1760,1786]" italics="true" pageId="7" pageNumber="348">G. quangi</emphasis>
Rowley, Dau, Nguyen
</taxonomicName>
, Cao &amp; Nguyen,
<taxonomicName id="C4BD4D7615075265611918F819A9B661" authority="(Nguyen, Hendrix, Bohme, Vu &amp; Ziegler)" baseAuthorityName="Nguyen, Hendrix, Bohme, Vu &amp; Ziegler" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="quyeti">
<emphasis id="31C9EAE715075265611918F81CFAB787" box="[1119,1220,1760,1786]" italics="true" pageId="7" pageNumber="348">G. quyeti</emphasis>
(Nguyen, Hendrix, Böhme, Vu &amp; Ziegler)
</taxonomicName>
and
<taxonomicName id="C4BD4D76150752656496191C1B9BB660" authority="(Orlov, Ho, &amp; Nguyen)" baseAuthorityName="Orlov, Ho, &amp; Nguyen" box="[464,933,1796,1822]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="supercornutus">
<emphasis id="31C9EAE7150752656496191C1AADB663" box="[464,659,1796,1822]" italics="true" pageId="7" pageNumber="348">G. supercornutus</emphasis>
(Orlov, Ho, &amp; Nguyen)
</taxonomicName>
by having a medium body size (adult males
<quantity id="C4459B101507526565D119301904B63C" box="[151,314,1832,1858]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="4.14" metricValueMin="3.72" pageId="7" pageNumber="348" unit="mm" value="39.3" valueMax="41.4" valueMin="37.2">37.2–41.4 mm</quantity>
SVL in five adult males) versus small body size (males &lt;
<quantity id="C4459B10150752656680193F1C2BB63C" box="[966,1045,1831,1858]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="348" unit="mm" value="25.0">25 mm</quantity>
SVL), brown or yellowish dorsum and a pink or pinkish brown venter (versus greenish dorsum and white or yellow venter), and eggs deposited on wall of a phytotelmon (versus on vegetation above pools for
<emphasis id="31C9EAE715075265665719681C14B6F7" box="[785,1066,1904,1930]" italics="true" pageId="7" pageNumber="348">
<taxonomicName id="C4BD4D7615075265665719681B99B6F4" baseAuthorityName="Bourret" baseAuthorityYear="1937" box="[785,935,1904,1930]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="gracilipes">G. gracilipes</taxonomicName>
,
<taxonomicName id="C4BD4D761507526566F519681C19B6F7" box="[947,1063,1904,1930]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="quangi">G. quangi</taxonomicName>
,
</emphasis>
and
<taxonomicName id="C4BD4D7615075265612519681D1BB6F7" box="[1123,1317,1904,1930]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="supercornutus">
<emphasis id="31C9EAE715075265612519681D1BB6F7" box="[1123,1317,1904,1930]" italics="true" pageId="7" pageNumber="348">G. supercornutus</emphasis>
</taxonomicName>
, unknown in
<taxonomicName id="C4BD4D761507526565F2198C1924B6D3" box="[180,282,1940,1966]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="quyeti">
<emphasis id="31C9EAE71507526565F2198C1924B6D3" box="[180,282,1940,1966]" italics="true" pageId="7" pageNumber="348">G. quyeti</emphasis>
</taxonomicName>
). The round snout (versus pointed snout) further differentiates
<taxonomicName id="C4BD4D76150752656698198C1C9BB6D3" box="[990,1189,1940,1966]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE7150752656698198C1C9BB6D3" box="[990,1189,1940,1966]" italics="true" pageId="7" pageNumber="348">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE71507526561EA198C1D3BB6D0" bold="true" box="[1196,1285,1940,1966]" pageId="7" pageNumber="348">sp.­nov.</emphasis>
from
<taxonomicName id="C4BD4D7615075265600C198C18DEB6AC" baseAuthorityName="Bourret" baseAuthorityYear="1937" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="gracilipes">
<emphasis id="31C9EAE715075265600C198C18DEB6AC" italics="true" pageId="7" pageNumber="348">G. gracilipes</emphasis>
</taxonomicName>
,
<taxonomicName id="C4BD4D761507526565A819A01960B6AF" box="[238,350,1976,2002]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="quangi">
<emphasis id="31C9EAE71507526565A819A01960B6AF" box="[238,350,1976,2002]" italics="true" pageId="7" pageNumber="348">G. quangi</emphasis>
</taxonomicName>
and
<emphasis id="31C9EAE71507526564D319A01A60B6AF" box="[405,606,1976,2002]" italics="true" pageId="7" pageNumber="348">
<taxonomicName id="C4BD4D761507526564D319A01A64B6AF" box="[405,602,1976,2002]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="supercornutus">G. supercornutus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="030236F51508526A65D11DFD1A44B538" blockId="8.[151,1436,997,1094]" pageId="8" pageNumber="349">
<emphasis id="31C9EAE71508526A65D11DFD192DB283" bold="true" box="[151,275,997,1022]" pageId="8" pageNumber="349">FIGURE­5.</emphasis>
<taxonomicName id="C4BD4D761508526A645F1DFD19F3B283" box="[281,461,997,1021]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="349" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71508526A645F1DFD19F3B283" box="[281,461,997,1021]" italics="true" pageId="8" pageNumber="349">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE71508526A64941DFD1A17B283" bold="true" box="[466,553,997,1021]" pageId="8" pageNumber="349">sp.­nov.­</emphasis>
(A)
<emphasis id="31C9EAE71508526A670A1DFD1A6CB283" bold="true" box="[588,594,997,1021]" pageId="8" pageNumber="349">­</emphasis>
male
<typeStatus id="DC0688571508526A67CC1DFE1ADFB280" box="[650,737,998,1022]" pageId="8" pageNumber="349" type="paratype">paratype</typeStatus>
UNS 00342/AMS R176204, nocturnal coloration, (B)
<emphasis id="31C9EAE71508526A604E1DFD1D30B283" bold="true" box="[1288,1294,997,1021]" pageId="8" pageNumber="349">­</emphasis>
male
<typeStatus id="DC0688571508526A60031DFE1DA2B280" box="[1349,1436,998,1022]" pageId="8" pageNumber="349" type="paratype">paratype</typeStatus>
UNS 00343/AMS R176205, diurnal coloration, (C) male
<typeStatus id="DC0688571508526A67AB1A111B78B55C" box="[749,838,1033,1058]" pageId="8" pageNumber="349" type="holotype">holotype</typeStatus>
<emphasis id="31C9EAE71508526A66001A111B72B55F" bold="true" box="[838,844,1033,1057]" pageId="8" pageNumber="349">­</emphasis>
AMS R176206, diurnal coloration, (D) juvenile
<typeStatus id="DC0688571508526A60031A121DA2B55C" box="[1349,1436,1034,1058]" pageId="8" pageNumber="349" type="paratype">paratype</typeStatus>
UNS 00351/AMS R176208, diurnal coloration.
</paragraph>
<paragraph id="030236F51508526A65D1198B1939B68A" blockId="8.[151,1436,1939,2036]" pageId="8" pageNumber="349">
<emphasis id="31C9EAE71508526A65D1198B192CB6D2" bold="true" box="[151,274,1939,1964]" pageId="8" pageNumber="349">FIGURE­6.</emphasis>
(A) Montane evergreen and bamboo forest at ~
<quantity id="C4459B101508526A67B4198C1B7EB6D2" box="[754,832,1940,1964]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="349" unit="m" value="2000.0">2000 m</quantity>
elevation, habitat of
<taxonomicName id="C4BD4D761508526A6152198C1CF6B6D2" box="[1044,1224,1940,1964]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="349" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71508526A6152198C1CF6B6D2" box="[1044,1224,1940,1964]" italics="true" pageId="8" pageNumber="349">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE71508526A618B198C1D20B6D2" bold="true" box="[1229,1310,1940,1964]" pageId="8" pageNumber="349">sp.­nov.</emphasis>
and (B) tree hole with
<taxonomicName id="C4BD4D761508526A65BC19A01993B6AE" box="[250,429,1976,2000]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="349" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71508526A65BC19A01993B6AE" box="[250,429,1976,2000]" italics="true" pageId="8" pageNumber="349">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE71508526A64F719A01A38B6AE" bold="true" box="[433,518,1976,2000]" pageId="8" pageNumber="349">sp.­nov.­</emphasis>
eggs (AMS R176211) deposited, and where the male
<typeStatus id="DC0688571508526A615D19AF1C4AB6AE" box="[1051,1140,1975,2000]" pageId="8" pageNumber="349" type="holotype">holotype</typeStatus>
of
<taxonomicName id="C4BD4D761508526A61D519A11CD0B6AE" box="[1171,1262,1977,2000]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="349" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71508526A61D519A11CD0B6AE" box="[1171,1262,1977,2000]" italics="true" pageId="8" pageNumber="349">G. trieng</emphasis>
</taxonomicName>
(AMS R176206) was found.
</paragraph>
<paragraph id="030236F51509526B65811E811C90B05E" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="350">
The new species differs from
<taxonomicName id="C4BD4D761509526B67581E811BD7B1CA" authority="(Matsui &amp; Orlov)" baseAuthorityName="Matsui &amp; Orlov" baseAuthorityYear="2004" box="[542,1001,153,180]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="ananjevae">
<emphasis id="31C9EAE71509526B67581E811B28B1CD" box="[542,790,153,179]" italics="true" pageId="9" pageNumber="350">Gracixalus ananjevae</emphasis>
(Matsui &amp; Orlov)
</taxonomicName>
by having a medium body size (adult males
<quantity id="C4459B101509526B65A41EA519B6B1A9" box="[226,392,189,215]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="4.14" metricValueMin="3.72" pageId="9" pageNumber="350" unit="mm" value="39.3" valueMax="41.4" valueMin="37.2">37.2–41.4 mm</quantity>
SVL in five adult males), compared to
<quantity id="C4459B101509526B661E1EA51B80B1A9" box="[856,958,189,215]" metricMagnitude="-2" metricUnit="m" metricValue="3.2399999999999998" pageId="9" pageNumber="350" unit="mm" value="32.4">32.4 mm</quantity>
in one adult male; a yellow or yellowish brown throat and chest with pinkish mottling and pinkish belly (versus greyish white venter [likely in preservative] with no mottling), and adult males with nuptial pads on fingers I and II (versus finger I only).
</paragraph>
<paragraph id="030236F51509526B65811F31193AB0AD" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="350">
<taxonomicName id="C4BD4D761509526B65811F3119B0B03D" box="[199,398,297,323]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71509526B65811F3119B0B03D" box="[199,398,297,323]" italics="true" pageId="9" pageNumber="350">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE71509526B64D31F3119D0B03D" bold="true" box="[405,494,297,323]" pageId="9" pageNumber="350">sp.­nov.</emphasis>
is distinguished from
<taxonomicName id="C4BD4D761509526B67AF1F311C50B03A" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1893" box="[745,1134,297,324]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="carinensis">
<emphasis id="31C9EAE71509526B67AF1F311BDEB03D" box="[745,992,297,323]" italics="true" pageId="9" pageNumber="350">Gracixalus carinensis</emphasis>
(Boulenger)
</taxonomicName>
by having a yellow or yellowish-brown throat and chest with pinkish mottling and pinkish belly (versus white venter with no mottling), adult males with nuptial pads on fingers I and II (versus finger I only;
<bibRefCitation id="672C4B041509526B66361F691CBDB0F5" author="Bossuyt, F. &amp; Dubois, A." box="[880,1155,369,395]" pageId="9" pageNumber="350" pagination="1 - 112" refId="ref7615" refString="Bossuyt, F. &amp; Dubois, A. (2001) A review of the frog genus Philautus Gistel, 1848 (Amphibia, Anura, Ranidae, Rhacophorinae). Zeylanica, 6, 1 - 112." type="journal article" year="2001">Bossuyt &amp; Dubois 2001</bibRefCitation>
). In addition,
<taxonomicName id="C4BD4D761509526B60661F6918E4B0D1" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71509526B60661F6918E4B0D1" italics="true" pageId="9" pageNumber="350">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE71509526B65A41F8D197DB0D1" bold="true" box="[226,323,405,431]" pageId="9" pageNumber="350">sp.­nov.­</emphasis>
has less developed webbing (Fig. 4C) compared to
<taxonomicName id="C4BD4D761509526B66C01F8D1C43B0D1" baseAuthorityName="Boulenger" baseAuthorityYear="1893" box="[902,1149,405,431]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="carinensis">
<emphasis id="31C9EAE71509526B66C01F8D1C43B0D1" box="[902,1149,405,431]" italics="true" pageId="9" pageNumber="350">Gracixalus carinensis</emphasis>
</taxonomicName>
(see Fig. 4B in
<bibRefCitation id="672C4B041509526B60751F8D18C6B0AD" author="Matsui, M. &amp; Ohler, A. &amp; Eto, K. &amp; Nguyen, T. T." pageId="9" pageNumber="350" pagination="25 - 37" refId="ref8386" refString="Matsui, M., Ohler, A., Eto, K. &amp; Nguyen, T. T. (2017) Distinction of Gracixalus carinensis from Vietnam and Myanmar, with description of a new species. Alytes, 33, 25 - 37." type="journal article" year="2017">
Matsui
<emphasis id="31C9EAE71509526B60CE1F8E188AB0AD" italics="true" pageId="9" pageNumber="350">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
<paragraph id="030236F51509526B65811FC51C8EB31A" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="350">
From
<taxonomicName id="C4BD4D761509526B644A1FC51BB7B089" authority="(Wang, Zeng, Liu &amp; Wang)" baseAuthorityName="Wang, Zeng, Liu &amp; Wang" box="[268,905,477,503]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="guangdongensis">
<emphasis id="31C9EAE71509526B644A1FC51A78B089" box="[268,582,477,503]" italics="true" pageId="9" pageNumber="350">Gracixalus guangdongensis</emphasis>
(Wang, Zeng, Liu &amp; Wang)
</taxonomicName>
,
<taxonomicName id="C4BD4D761509526B66D31FC51C62B089" box="[917,1116,477,503]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71509526B66D31FC51C62B089" box="[917,1116,477,503]" italics="true" pageId="9" pageNumber="350">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE71509526B61221FC51CF8B089" bold="true" box="[1124,1222,477,503]" pageId="9" pageNumber="350">sp.­nov.­</emphasis>
differs by having a greater body size (adult males
<quantity id="C4459B101509526B64AB1C191AB1B365" box="[493,655,513,539]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="4.14" metricValueMin="3.72" pageId="9" pageNumber="350" unit="mm" value="39.3" valueMax="41.4" valueMin="37.2">37.2–41.4 mm</quantity>
SVL in five adult males, compared to SVL
<quantity id="C4459B101509526B613F1C191D21B365" box="[1145,1311,513,539]" metricMagnitude="-2" metricUnit="m" metricValue="3.04" metricValueMax="3.47" metricValueMin="2.61" pageId="9" pageNumber="350" unit="mm" value="30.4" valueMax="34.7" valueMin="26.1">26.1–34.7 mm</quantity>
), yellow or yellowish brown throat and chest with pinkish mottling and pinkish belly (versus ventral surface of throat, chest and forelimbs dirty white with small dark specks; belly white anteriorly with large dark blotches).
</paragraph>
<paragraph id="030236F51509526B65811C751CE1B38A" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="350">
From
<taxonomicName id="C4BD4D761509526B644C1C751C33B3F9" authority="(Zeng, Zhao, Chen, Chen, Zhang &amp; Wang)" baseAuthorityName="Zeng, Zhao, Chen, Chen, Zhang &amp; Wang" box="[266,1037,621,647]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="jinggangensis">
<emphasis id="31C9EAE71509526B644C1C751A17B3F9" box="[266,553,621,647]" italics="true" pageId="9" pageNumber="350">Gracixalus jinggangensis</emphasis>
(Zeng, Zhao, Chen, Chen, Zhang &amp; Wang)
</taxonomicName>
, the new species differs by having a larger body size (adult males
<quantity id="C4459B101509526B64991C891ABFB3D5" box="[479,641,657,683]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="4.14" metricValueMin="3.72" pageId="9" pageNumber="350" unit="mm" value="39.3" valueMax="41.4" valueMin="37.2">37.2–41.4 mm</quantity>
SVL in five adult males, compared to SVL
<quantity id="C4459B101509526B612E1C891D31B3D5" box="[1128,1295,657,683]" metricMagnitude="-2" metricUnit="m" metricValue="3.085" metricValueMax="3.38" metricValueMin="2.79" pageId="9" pageNumber="350" unit="mm" value="30.849999999999998" valueMax="33.8" valueMin="27.9">27.9–33.8 mm</quantity>
), and yellow or yellowish brown throat and chest with pinkish mottling and pinkish belly (versus ventral surface of throat, chest and forelimbs dirty white with small dark specks; belly white anteriorly with large dark blotches).
</paragraph>
<paragraph id="030236F51509526B65811CE51AA9B2FA" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="350">
From
<taxonomicName id="C4BD4D761509526B644C1CE51A7FB269" authority="(Hu)" baseAuthorityName="Hu" baseAuthorityYear="1978" box="[266,577,765,791]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="jinxiuensis">
<emphasis id="31C9EAE71509526B644C1CE51A3AB269" box="[266,516,765,791]" italics="true" pageId="9" pageNumber="350">Gracixalus jinxiuensis</emphasis>
(Hu)
</taxonomicName>
, the new species is distinguished by having a larger body size (
<quantity id="C4459B101509526B61BF1CE51DA3B269" box="[1273,1437,765,791]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="4.14" metricValueMin="3.72" pageId="9" pageNumber="350" unit="mm" value="39.3" valueMax="41.4" valueMin="37.2">37.2–41.4 mm</quantity>
SVL in five adult males compared to
<quantity id="C4459B101509526B677C1D381AA0B245" box="[570,670,800,827]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="9" pageNumber="350" unit="mm" value="23.5">23.5 mm</quantity>
in adult male
<typeStatus id="DC0688571509526B66781D391B9AB242" box="[830,932,801,828]" pageId="9" pageNumber="350" type="holotype">holotype</typeStatus>
), yellow or yellowish brown throat and chest with pinkish mottling and pinkish belly (versus pale grey-brown with dark marbling), and adult males with nuptial pads on fingers I and II (versus finger I only).
</paragraph>
<paragraph id="030236F51509526B65811D951B92B522" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="350">
From
<taxonomicName id="C4BD4D761509526B644B1D951B1EB2D9" authority="Rowley, Le, Dau, Hoang" authorityName="Rowley, Le, Dau, Hoang" box="[269,800,909,935]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="lumarius">
<emphasis id="31C9EAE71509526B644B1D9519C9B2D9" box="[269,503,909,935]" italics="true" pageId="9" pageNumber="350">Gracixalus lumarius</emphasis>
Rowley, Le, Dau, Hoang
</taxonomicName>
, &amp; Cao, the new species differs by having large, white conical tubercles on the dorsum absent (versus at most sparsely distributed low, irregular tubercles present), iris pale gold with a sparse network of fine dark gold reticulations darker gold radiating out of anterior and posterior edges of pupil (versus a dark gold iris with a dense, relatively uniformly distributed network of black reticulations), yellow or yellowish brown throat and chest with pinkish mottling and pinkish belly (versus uniformly pinkish venter), and adult males with nuptial pads on fingers I and II (versus finger I only).
</paragraph>
<paragraph id="030236F51509526B65811A7D1C10B595" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="350">
From
<taxonomicName id="C4BD4D761509526B644D1A7D1AADB501" authority="(Ye &amp; Hu)" baseAuthorityName="Ye &amp; Hu" baseAuthorityYear="1984" box="[267,659,1125,1151]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="medogensis">
<emphasis id="31C9EAE71509526B644D1A7D1A2CB501" box="[267,530,1125,1151]" italics="true" pageId="9" pageNumber="350">Gracixalus medogensis</emphasis>
(Ye &amp; Hu)
</taxonomicName>
, the new species is distinguished by having a larger body size (
<quantity id="C4459B101509526B601D1A7D18C2B5DD" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="4.14" metricValueMin="3.72" pageId="9" pageNumber="350" unit="mm" value="39.3" valueMax="41.4" valueMin="37.2">37.2– 41.4 mm</quantity>
SVL in five adult males compared to
<quantity id="C4459B101509526B67EA1A901B2FB5DD" box="[684,785,1160,1187]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" pageId="9" pageNumber="350" unit="mm" value="26.5">26.5 mm</quantity>
in one adult male), yellow or yellowish brown throat and chest with pinkish mottling and pinkish belly (versus light grey or whitish venter), adult males with nuptial pads on fingers I and II (versus finger I only), and linea masculina absent (versus present).
</paragraph>
<paragraph id="030236F51509526B65811AED199FB4DE" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="350">
From
<taxonomicName id="C4BD4D761509526B64481AED1B88B471" authority="Mo, Zhang, Luo, Zhou &amp; Chen" authorityName="Mo, Zhang, Luo, Zhou &amp; Chen" authorityYear="2013" box="[270,950,1269,1295]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="nonggangensis">
<emphasis id="31C9EAE71509526B64481AED1A03B471" box="[270,573,1269,1295]" italics="true" pageId="9" pageNumber="350">Gracixalus nonggangensis</emphasis>
Mo, Zhang, Luo, Zhou &amp; Chen
</taxonomicName>
and
<taxonomicName id="C4BD4D761509526B66B51AED19C7B44D" authority="Nguyen, Le, Pham, Nguyen, Bonkowski &amp; Ziegler" authorityName="Nguyen, Le, Pham, Nguyen, Bonkowski &amp; Ziegler" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="waza">
<emphasis id="31C9EAE71509526B66B51AED1C8DB471" box="[1011,1203,1269,1295]" italics="true" pageId="9" pageNumber="350">Gracixalus waza</emphasis>
Nguyen, Le, Pham, Nguyen, Bonkowski &amp; Ziegler
</taxonomicName>
, the new species differs by having a larger body size (
<quantity id="C4459B101509526B61211B011D30B44D" box="[1127,1294,1305,1331]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="4.14" metricValueMin="3.72" pageId="9" pageNumber="350" unit="mm" value="39.3" valueMax="41.4" valueMin="37.2">37.2–41.4 mm</quantity>
SVL in five adult males versus &lt;
<quantity id="C4459B101509526B643D1B2519F4B429" box="[379,458,1341,1367]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="9" pageNumber="350" unit="mm" value="36.0">36 mm</quantity>
in adult males), and a yellow or yellowish brown throat and chest with pinkish mottling and pinkish belly (versus white venter with brown spots), adult males with nuptial pads on fingers I and II (versus absent or finger I only).
</paragraph>
<paragraph id="030236F51509526B65811BB11CDFB772" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="350">
From
<taxonomicName id="C4BD4D761509526B644A1BB11B66B4BD" authority="Matsui, Ohler, Eto &amp; Nguyen" authorityName="Matsui, Ohler, Eto &amp; Nguyen" authorityYear="2017" box="[268,856,1449,1475]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="sapaensis">
<emphasis id="31C9EAE71509526B644A1BB119C0B4BD" box="[268,510,1449,1475]" italics="true" pageId="9" pageNumber="350">Gracixalus sapaensis</emphasis>
Matsui, Ohler, Eto &amp; Nguyen
</taxonomicName>
, the new species differs by having a yellow or yellowish brown throat and chest with pinkish mottling and pinkish belly (versus ventral surface of throat, chest, belly, and forelimb light yellow), adult males with nuptial pads on fingers I and II (versus finger I only).
</paragraph>
<paragraph id="030236F51509526B6581180D1B09B7E2" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="350">
From
<taxonomicName id="C4BD4D761509526B644D180D1B74B751" authority="Matsui, Khonsue, Panha &amp; Eto" authorityName="Matsui, Khonsue, Panha &amp; Eto" authorityYear="2015" box="[267,842,1557,1583]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="seesom">
<emphasis id="31C9EAE71509526B644D180D19DEB751" box="[267,480,1557,1583]" italics="true" pageId="9" pageNumber="350">Gracixalus seesom</emphasis>
Matsui, Khonsue, Panha &amp; Eto
</taxonomicName>
, the new species differs by having a larger body size (
<quantity id="C4459B101509526B65D91821197AB72D" box="[159,324,1593,1619]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="4.14" metricValueMin="3.72" pageId="9" pageNumber="350" unit="mm" value="39.3" valueMax="41.4" valueMin="37.2">37.2–41.4 mm</quantity>
SVL in five adult males versus 21.6–23.0 mm), round (versus pointed) snout, yellow or yellowish brown throat and chest with pinkish mottling and pinkish belly (versus opaque white venter), and adult males with nuptial pads on fingers I and II (versus nuptial pads absent).
</paragraph>
<paragraph id="030236F51509526B658118BD197EB652" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="350">
From
<taxonomicName id="C4BD4D761509526B644C18BD1B66B7C1" authority="Chen, Bei, Liao, Zhou &amp; Mo" authorityName="Chen, Bei, Liao, Zhou &amp; Mo" authorityYear="2018" box="[266,856,1701,1727]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="tianlinensis">
<emphasis id="31C9EAE71509526B644C18BD1A30B7C1" box="[266,526,1701,1727]" italics="true" pageId="9" pageNumber="350">Gracixalus tianlinensis</emphasis>
Chen, Bei, Liao, Zhou &amp; Mo
</taxonomicName>
, the new species differs by having a larger body size (
<quantity id="C4459B101509526B65D918D1197BB79D" box="[159,325,1737,1763]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="4.14" metricValueMin="3.72" pageId="9" pageNumber="350" unit="mm" value="39.3" valueMax="41.4" valueMin="37.2">37.2–41.4 mm</quantity>
SVL in five adult males versus
<quantity id="C4459B101509526B67F018D01B61B79D" box="[694,863,1736,1763]" metricMagnitude="-2" metricUnit="m" metricValue="3.31" metricValueMax="3.59" metricValueMin="3.03" pageId="9" pageNumber="350" unit="mm" value="33.1" valueMax="35.9" valueMin="30.3">30.3–35.9 mm</quantity>
), yellow or yellowish brown throat and chest with pinkish mottling and pinkish belly (versus ventral surface of the throat, chest grey with small dark specks and belly creamy white).
</paragraph>
<paragraph id="030236F51509526B6581192D1D08B6C2" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="350">
From
<taxonomicName id="C4BD4D761509526B644A192D1BB4B631" authority="Yu, Li, Wang, Rao, Wu &amp; Yang" authorityName="Yu, Li, Wang, Rao, Wu &amp; Yang" authorityYear="2019" box="[268,906,1845,1871]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="31C9EAE71509526B644A192D1A24B631" box="[268,538,1845,1871]" italics="true" pageId="9" pageNumber="350">Gracixalus yunnanensis</emphasis>
Yu, Li, Wang, Rao, Wu &amp; Yang
</taxonomicName>
,
<taxonomicName id="C4BD4D761509526B66D0192D1C63B631" box="[918,1117,1845,1871]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE71509526B66D0192D1C63B631" box="[918,1117,1845,1871]" italics="true" pageId="9" pageNumber="350">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE71509526B6123192D1CF8B631" bold="true" box="[1125,1222,1845,1871]" pageId="9" pageNumber="350">sp.­nov.­</emphasis>
differs by having a larger body size (
<quantity id="C4459B101509526B641C194119C1B60D" box="[346,511,1881,1907]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="4.14" metricValueMin="3.72" pageId="9" pageNumber="350" unit="mm" value="39.3" valueMax="41.4" valueMin="37.2">37.2–41.4 mm</quantity>
SVL in five adult males versus 26.0–
<quantity id="C4459B101509526B66EF19411C2FB60D" box="[937,1041,1881,1907]" metricMagnitude="-2" metricUnit="m" metricValue="3.42" pageId="9" pageNumber="350" unit="mm" value="34.2">34.2 mm</quantity>
), yellow or yellowish brown throat and chest with pinkish mottling and pinkish belly (versus semi-transparent orangish with yellow spots), linea masculina absent (versus present), and adult males with nuptial pads on fingers I and II (versus finger I only).
</paragraph>
<paragraph id="030236F515095268658119DC1A66B03D" blockId="9.[151,1437,153,2015]" lastBlockId="10.[151,1437,152,323]" lastPageId="10" lastPageNumber="351" pageId="9" pageNumber="350">
<emphasis id="31C9EAE71509526B658119DC1A4FB6A1" bold="true" box="[199,625,1988,2015]" pageId="9" pageNumber="350">Mitochondrial­sequence­divergence</emphasis>
: The phylogenetic placement of the new species was not well-resolved (Fig. 7). Uncorrected sequence divergences between
<taxonomicName id="C4BD4D76150A526867B01E801B80B1CC" box="[758,958,152,178]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE7150A526867B01E801B80B1CC" box="[758,958,152,178]" italics="true" pageId="10" pageNumber="351">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE7150A526866811E811C14B1CD" bold="true" box="[967,1066,153,179]" pageId="10" pageNumber="351">sp.­nov.­</emphasis>
and all other
<taxonomicName id="C4BD4D76150A526861801E801D7CB1CC" baseAuthorityName="Frost" baseAuthorityYear="2020" box="[1222,1346,152,178]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="351" phylum="Chordata" rank="genus">
<emphasis id="31C9EAE7150A526861801E801D7CB1CC" box="[1222,1346,152,178]" italics="true" pageId="10" pageNumber="351">Gracixalus</emphasis>
</taxonomicName>
species for which comparable molecular data are available is&gt; 3.6% at the 16S rRNA gene fragment examined, with the least divergence between the new species and
<taxonomicName id="C4BD4D76150A526867E41EF91B44B184" box="[674,890,224,251]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="guangdongensis">
<emphasis id="31C9EAE7150A526867E41EF91B44B184" box="[674,890,224,251]" italics="true" pageId="10" pageNumber="351">G. guangdongensis</emphasis>
</taxonomicName>
(3.6%) and
<taxonomicName id="C4BD4D76150A526861411EF91CA9B184" authorityName="Matsui, Ohler, Eto &amp; Nguyen" authorityYear="2017" box="[1031,1175,225,251]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="sapaensis">
<emphasis id="31C9EAE7150A526861411EF91CA9B184" box="[1031,1175,225,251]" italics="true" pageId="10" pageNumber="351">G. sapaensis</emphasis>
</taxonomicName>
(3.9%). This degree of pairwise divergence in the 16S rRNA gene in frogs has been interpreted previously as indicative of differentiation at the species level (
<bibRefCitation id="672C4B04150A5268643E1F311A72B03D" author="Vences, M. &amp; Thomas, M. &amp; Bonett, R. M. &amp; Vieites, D. R." box="[376,588,296,323]" pageId="10" pageNumber="351" pagination="1859 - 1868" refId="ref9070" refString="Vences, M., Thomas, M., Bonett, R. M. &amp; Vieites, D. R. (2005) Deciphering amphibian diversity through DNA barcoding: chances and challenges. Philosophical Transactions of the Royal Society London, Series B, 360, 1859 - 1868. https: // doi. org / 10.1098 / rstb. 2005.1717" type="journal article" year="2005">
Vences
<emphasis id="31C9EAE7150A526864961F311A3CB03C" box="[464,514,296,322]" italics="true" pageId="10" pageNumber="351">et al</emphasis>
. 2005
</bibRefCitation>
).
</paragraph>
<paragraph id="030236F5150A526865D11B69190AB488" blockId="10.[151,1437,1393,1526]" pageId="10" pageNumber="351">
<emphasis id="31C9EAE7150A526865D11B69192DB4F4" bold="true" box="[151,275,1393,1418]" pageId="10" pageNumber="351">FIGURE­7.</emphasis>
Maximum likelihood tree on ~550 bp section of 16S (mtDNA) gene for
<taxonomicName id="C4BD4D76150A526866B51B691C99B4F7" box="[1011,1191,1393,1417]" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="trieng">
<emphasis id="31C9EAE7150A526866B51B691C99B4F7" box="[1011,1191,1393,1417]" italics="true" pageId="10" pageNumber="351">Gracixalus trieng</emphasis>
</taxonomicName>
<emphasis id="31C9EAE7150A526861EA1B6A1D3DB4F4" bold="true" box="[1196,1283,1394,1418]" pageId="10" pageNumber="351">sp.­nov.­</emphasis>
and
<emphasis id="31C9EAE7150A5268606E1B6A1D10B4F4" bold="true" box="[1320,1326,1394,1418]" pageId="10" pageNumber="351">­</emphasis>
all
<taxonomicName id="C4BD4D76150A5268600B1B6A18FFB4D3" baseAuthorityName="Frost" baseAuthorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="351" phylum="Chordata" rank="genus">
<emphasis id="31C9EAE7150A5268600B1B6A18FFB4D3" italics="true" pageId="10" pageNumber="351">Gracixalus</emphasis>
</taxonomicName>
species for which comparative sequences are available and for outgroups (
<taxonomicName id="C4BD4D76150A526866FF1B8D1C47B4D0" box="[953,1145,1429,1454]" class="Amphibia" family="Rhacophoridae" genus="Kurixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="effingeri">
<emphasis id="31C9EAE7150A526866FF1B8D1C47B4D0" box="[953,1145,1429,1454]" italics="true" pageId="10" pageNumber="351">Kurixalus effingeri</emphasis>
</taxonomicName>
,
<taxonomicName id="C4BD4D76150A526861C01B8D1D4FB4D0" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[1158,1393,1429,1454]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="aurifasciatus">
<emphasis id="31C9EAE7150A526861C01B8D1D4FB4D0" box="[1158,1393,1429,1454]" italics="true" pageId="10" pageNumber="351">Philautus aurifasciatus</emphasis>
</taxonomicName>
and
<taxonomicName id="C4BD4D76150A526865D11BA119AAB4AF" baseAuthorityName="Schlegel" baseAuthorityYear="1840" box="[151,404,1465,1489]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="reinwardtii">
<emphasis id="31C9EAE7150A526865D11BA119AAB4AF" box="[151,404,1465,1489]" italics="true" pageId="10" pageNumber="351">Rhacophorus reinwardtii</emphasis>
</taxonomicName>
). Node support is indicated on the branches as Maximum Likelihood bootstrap support (&lt;70% = grey. 100% = black).
</paragraph>
</subSubSection>
</treatment>
</document>
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