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<document id="7EE29AF52CA40D0BBCD3B683FFE95F65" ID-DOI="10.11646/zootaxa.3889.1.1" ID-GBIF-Dataset="3211a768-dcbf-42b4-9892-aa04539b73d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="287509" ID-ZooBank="AAFE8C8C-7A1E-4C5A-BAA8-5D2030E7EC8F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460982723640" checkinUser="plazi" docAuthor="Calderón-Martínez, Nadia R., González-Obando, Ranulfo &amp; García Aldrete, Alfonso N." docDate="2014" docId="763D3C08FF9DFFA7FF3198504DAA43ED" docLanguage="en" docName="zt03889p030.pdf" docOrigin="Zootaxa 3889 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cladiopsocus bidentatus Calderón-Martínez, González-Obando &amp; Aldrete, 2014, n. sp." docType="treatment" docVersion="7" lastPageNumber="6" masterDocId="8A044470FF98FFA2FFA699774836474F" masterDocTitle="Descriptions and records of Cladiopsocidae and Dolabellopsocidae (Insecta: Psocodea: Psocoptera) from Valle del Cauca and National Natural Park Gorgona, Colombia" masterLastPageNumber="30" masterPageNumber="1" pageNumber="6" updateTime="1698560339554" updateUser="plazi">
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<mods:title id="C477D5699FFA5EDC463418E81B84F58F">Descriptions and records of Cladiopsocidae and Dolabellopsocidae (Insecta: Psocodea: Psocoptera) from Valle del Cauca and National Natural Park Gorgona, Colombia</mods:title>
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<mods:namePart id="76FF5A264C7C5555FEF8996A1694B327">Calderón-Martínez, Nadia R.</mods:namePart>
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<mods:namePart id="FE1DB55D28654D43267A1FDA143FA1DA">González-Obando, Ranulfo</mods:namePart>
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<mods:namePart id="3DE89D6B8D87E06AB3B7C37F03AF8685">García Aldrete, Alfonso N.</mods:namePart>
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<treatment id="763D3C08FF9DFFA7FF3198504DAA43ED" ID-DOI="http://doi.org/10.5281/zenodo.5620611" ID-GBIF-Taxon="127670883" ID-Zenodo-Dep="5620611" LSID="urn:lsid:plazi:treatment:763D3C08FF9DFFA7FF3198504DAA43ED" httpUri="http://treatment.plazi.org/id/763D3C08FF9DFFA7FF3198504DAA43ED" lastPageNumber="6" pageId="5" pageNumber="6">
<subSubSection id="B68EDE95FF9DFFA7FF3198504910462C" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="FE2B8D1EFF9DFFA7FF3198504A65460D" blockId="5.[151,595,295,355]" box="[151,595,295,322]" pageId="5" pageNumber="6">
<heading id="A5633A72FF9DFFA7FF3198504A65460D" bold="true" box="[151,595,295,322]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="CCE0510CFF9DFFA7FF3198504A65460D" bold="true" box="[151,595,295,322]" pageId="5" pageNumber="6">
<taxonomicName id="3994F69DFF9DFFA7FF319850498A460E" box="[151,444,295,321]" class="Insecta" family="Cladiopsocidae" genus="Cladiopsocus" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bidentatus" status="sp. nov.">
<emphasis id="CCE0510CFF9DFFA7FF319850498A460E" bold="true" box="[151,444,295,321]" italics="true" pageId="5" pageNumber="6">Cladiopsocus bidentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7D3EC77FF9DFFA7FE62985F4A30460D" box="[452,518,296,322]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
Male.
</emphasis>
</heading>
</paragraph>
<paragraph id="FE2B8D1EFF9DFFA7FF31983D4910462C" blockId="5.[151,595,295,355]" box="[151,294,330,355]" pageId="5" pageNumber="6">
(
<figureCitation id="66AF919BFF9DFFA7FF39983D4929462C" box="[159,287,330,355]" captionStart="FIGURES 13 19" captionStartId="6.[151,264,1899,1921]" captionTargetBox="[151,1436,203,1859]" captionTargetId="figure@6.[151,1436,193,1877]" captionTargetPageId="6" captionText="FIGURES 13 19. Cladiopsocus bidentatus n. sp. Male. 13. Forewing. 14. Hindwing. 15. Front view of head. 16. Hypandrium. 17. Phallosome. 18. Paraproct. 19. Epiproct. Scales in mm." httpUri="https://zenodo.org/record/287512/files/figure.png" pageId="5" pageNumber="6">Figs 1319</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B68EDE95FF9DFFA7FF3198E54B3546BC" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="FE2B8D1EFF9DFFA7FF3198E54B3546BC" blockId="5.[151,1437,402,1186]" pageId="5" pageNumber="6">
<emphasis id="CCE0510CFF9DFFA7FF3198E5492746E4" bold="true" box="[151,273,402,427]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
Similar to
<taxonomicName id="3994F69DFF9DFFA7FE3598E44A9146E4" authority="Eertmoed" authorityName="Eertmoed" box="[403,679,402,427]" class="Insecta" family="Cladiopsocidae" genus="Cladiopsocus" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="interruptus">
<emphasis id="CCE0510CFF9DFFA7FE3598E44A0446E4" box="[403,562,403,427]" italics="true" pageId="5" pageNumber="6">C. interruptus</emphasis>
Eertmoed
</taxonomicName>
and to
<taxonomicName id="3994F69DFF9DFFA7FD5898E44C3F46E4" authority="Eertmoed" authorityName="Eertmoed" box="[766,1033,402,427]" class="Insecta" family="Cladiopsocidae" genus="Cladiopsocus" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mockfordi">
<emphasis id="CCE0510CFF9DFFA7FD5898E44BA746E4" box="[766,913,402,427]" italics="true" pageId="5" pageNumber="6">C. mockfordi</emphasis>
Eertmoed
</taxonomicName>
, but differing from them by having the phallosome densely spiculate (
<figureCitation id="66AF919BFF9DFFA7FDBB98C14A474680" box="[541,625,438,463]" captionStart="FIGURES 13 19" captionStartId="6.[151,264,1899,1921]" captionTargetBox="[151,1436,203,1859]" captionTargetId="figure@6.[151,1436,193,1877]" captionTargetPageId="6" captionText="FIGURES 13 19. Cladiopsocus bidentatus n. sp. Male. 13. Forewing. 14. Hindwing. 15. Front view of head. 16. Hypandrium. 17. Phallosome. 18. Paraproct. 19. Epiproct. Scales in mm." httpUri="https://zenodo.org/record/287512/files/figure.png" pageId="5" pageNumber="6">Fig. 17</figureCitation>
), by having two triangular teeth on the distal edge of the male epiproct and by lacking a median rise on this structure (
<figureCitation id="66AF919BFF9DFFA7FD0598AD4AC046BC" box="[675,758,474,499]" captionStart="FIGURES 13 19" captionStartId="6.[151,264,1899,1921]" captionTargetBox="[151,1436,203,1859]" captionTargetId="figure@6.[151,1436,193,1877]" captionTargetPageId="6" captionText="FIGURES 13 19. Cladiopsocus bidentatus n. sp. Male. 13. Forewing. 14. Hindwing. 15. Front view of head. 16. Hypandrium. 17. Phallosome. 18. Paraproct. 19. Epiproct. Scales in mm." httpUri="https://zenodo.org/record/287512/files/figure.png" pageId="5" pageNumber="6">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B68EDE95FF9DFFA7FF61988948D34379" pageId="5" pageNumber="6" type="description">
<paragraph id="FE2B8D1EFF9DFFA7FF6198894B8E45E8" blockId="5.[151,1437,402,1186]" pageId="5" pageNumber="6">
<emphasis id="CCE0510CFF9DFFA7FF61988949C54558" bold="true" box="[199,499,510,535]" pageId="5" pageNumber="6">Description. Male. Color.</emphasis>
Head (
<figureCitation id="66AF919BFF9DFFA7FDE398894AA04558" box="[581,662,510,535]" captionStart="FIGURES 13 19" captionStartId="6.[151,264,1899,1921]" captionTargetBox="[151,1436,203,1859]" captionTargetId="figure@6.[151,1436,193,1877]" captionTargetPageId="6" captionText="FIGURES 13 19. Cladiopsocus bidentatus n. sp. Male. 13. Forewing. 14. Hindwing. 15. Front view of head. 16. Hypandrium. 17. Phallosome. 18. Paraproct. 19. Epiproct. Scales in mm." httpUri="https://zenodo.org/record/287512/files/figure.png" pageId="5" pageNumber="6">Fig. 15</figureCitation>
) dark brown, postclypeus with V-shaped brown bands, with outward curved ends; compound eyes black, ocelli hyaline. Forewing membrane (
<figureCitation id="66AF919BFF9DFFA7FC779B554C154574" box="[977,1059,546,571]" captionStart="FIGURES 13 19" captionStartId="6.[151,264,1899,1921]" captionTargetBox="[151,1436,203,1859]" captionTargetId="figure@6.[151,1436,193,1877]" captionTargetPageId="6" captionText="FIGURES 13 19. Cladiopsocus bidentatus n. sp. Male. 13. Forewing. 14. Hindwing. 15. Front view of head. 16. Hypandrium. 17. Phallosome. 18. Paraproct. 19. Epiproct. Scales in mm." httpUri="https://zenodo.org/record/287512/files/figure.png" pageId="5" pageNumber="6">Fig. 13</figureCitation>
) slightly fumose, brown spots on vein ends, a brown spot on areola postica at wing margin as illustrated, pterostigma with central brown band; veins irregularly brown pigmented. Hindwing slightly smoked (
<figureCitation id="66AF919BFF9DFFA7FC869B1D4B4545CC" box="[800,883,618,643]" captionStart="FIGURES 13 19" captionStartId="6.[151,264,1899,1921]" captionTargetBox="[151,1436,203,1859]" captionTargetId="figure@6.[151,1436,193,1877]" captionTargetPageId="6" captionText="FIGURES 13 19. Cladiopsocus bidentatus n. sp. Male. 13. Forewing. 14. Hindwing. 15. Front view of head. 16. Hypandrium. 17. Phallosome. 18. Paraproct. 19. Epiproct. Scales in mm." httpUri="https://zenodo.org/record/287512/files/figure.png" pageId="5" pageNumber="6">Fig. 14</figureCitation>
), with brown spots at vein ends. Clunium brown, anteriorly more pigmented, paraprocts brown, hypandrium light brown.
</paragraph>
<paragraph id="FE2B8D1EFF9DFFA7FF619BC54D0444ED" blockId="5.[151,1437,402,1186]" pageId="5" pageNumber="6">
<emphasis id="CCE0510CFF9DFFA7FF619BC549564584" bold="true" box="[199,352,690,715]" pageId="5" pageNumber="6">Morphology.</emphasis>
As in diagnosis, plus the following: hypandrium (
<figureCitation id="66AF919BFF9DFFA7FC109BC54C3A4584" box="[950,1036,690,715]" captionStart="FIGURES 13 19" captionStartId="6.[151,264,1899,1921]" captionTargetBox="[151,1436,203,1859]" captionTargetId="figure@6.[151,1436,193,1877]" captionTargetPageId="6" captionText="FIGURES 13 19. Cladiopsocus bidentatus n. sp. Male. 13. Forewing. 14. Hindwing. 15. Front view of head. 16. Hypandrium. 17. Phallosome. 18. Paraproct. 19. Epiproct. Scales in mm." httpUri="https://zenodo.org/record/287512/files/figure.png" pageId="5" pageNumber="6">Fig. 16</figureCitation>
), setose, with sides converging to round apex. Phallosome (
<figureCitation id="66AF919BFF9DFFA7FE119BA14A3E45A0" box="[439,520,726,751]" captionStart="FIGURES 13 19" captionStartId="6.[151,264,1899,1921]" captionTargetBox="[151,1436,203,1859]" captionTargetId="figure@6.[151,1436,193,1877]" captionTargetPageId="6" captionText="FIGURES 13 19. Cladiopsocus bidentatus n. sp. Male. 13. Forewing. 14. Hindwing. 15. Front view of head. 16. Hypandrium. 17. Phallosome. 18. Paraproct. 19. Epiproct. Scales in mm." httpUri="https://zenodo.org/record/287512/files/figure.png" pageId="5" pageNumber="6">Fig. 17</figureCitation>
) broad, anteriorly obtusely concave, antero-lateral corners projected to form two short, stout columns. External parameres short and broadly bilobed, with a field of small spicules along the margin. Epiproct (
<figureCitation id="66AF919BFF9DFFA7FEAC9A6949684478" box="[266,350,798,823]" captionStart="FIGURES 13 19" captionStartId="6.[151,264,1899,1921]" captionTargetBox="[151,1436,203,1859]" captionTargetId="figure@6.[151,1436,193,1877]" captionTargetPageId="6" captionText="FIGURES 13 19. Cladiopsocus bidentatus n. sp. Male. 13. Forewing. 14. Hindwing. 15. Front view of head. 16. Hypandrium. 17. Phallosome. 18. Paraproct. 19. Epiproct. Scales in mm." httpUri="https://zenodo.org/record/287512/files/figure.png" pageId="5" pageNumber="6">Fig. 19</figureCitation>
) broadly trapeziform, with sides sclerotized, and postero-lateral corners protruding; a field of setae mesally and along posterior border. Paraprocts (
<figureCitation id="66AF919BFF9DFFA7FD4A9A354B0B4414" box="[748,829,834,859]" captionStart="FIGURES 13 19" captionStartId="6.[151,264,1899,1921]" captionTargetBox="[151,1436,203,1859]" captionTargetId="figure@6.[151,1436,193,1877]" captionTargetPageId="6" captionText="FIGURES 13 19. Cladiopsocus bidentatus n. sp. Male. 13. Forewing. 14. Hindwing. 15. Front view of head. 16. Hypandrium. 17. Phallosome. 18. Paraproct. 19. Epiproct. Scales in mm." httpUri="https://zenodo.org/record/287512/files/figure.png" pageId="5" pageNumber="6">Fig. 18</figureCitation>
) anteriorly concave, mesal tubercle distinct, with long setae as illustrated; mesal prong stout, strongly sclerotized, with two apical teeth and a small rounded mesal prominence, sensory fields with 23 trichobothria on basal rosettes. Legs and wings damaged in light trap.
</paragraph>
<paragraph id="FE2B8D1EFF9DFFA7FF619AD94C7F4488" blockId="5.[151,1437,402,1186]" box="[199,1097,942,971]" pageId="5" pageNumber="6">
<emphasis id="CCE0510CFF9DFFA7FF619AD9494B4488" bold="true" box="[199,381,942,967]" pageId="5" pageNumber="6">Measurements.</emphasis>
f1: 90, Mx4: 170, IO: 170, D: 240, d: 330, IO/d: 0.51, PO: 1.37.
</paragraph>
<paragraph id="FE2B8D1EFF9DFFA7FF619AA248D34379" blockId="5.[151,1437,402,1186]" pageId="5" pageNumber="6">
<emphasis id="CCE0510CFF9DFFA7FF619AA2499644A1" bold="true" box="[199,416,981,1006]" pageId="5" pageNumber="6">Specimen studied.</emphasis>
<typeStatus id="212F33BCFF9DFFA7FE0E9AA24A2644A1" box="[424,528,981,1006]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="8683CD8EFF9DFFA7FDF89AA14AC144A1" box="[606,759,982,1006]" name="Colombia" pageId="5" pageNumber="6">COLOMBIA</collectingCountry>
. Cauca. Guapi. NNP Gorgona, Cerro
<collectingCountry id="8683CD8EFF9DFFA7FB059AA24D3444A1" box="[1187,1282,981,1006]" name="Trinidad and Tobago" pageId="5" pageNumber="6">Trinidad</collectingCountry>
trail.
<geoCoordinate id="9BA0EBD9FF9DFFA7FAE19AA148C0435E" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="1" value="2.970389">02° 58 13.4” N</geoCoordinate>
:
<geoCoordinate id="9BA0EBD9FF9DFFA7FEA29A8D49F0435E" box="[260,454,1017,1042]" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="1" value="-78.18272">78° 10 57.8” W</geoCoordinate>
.
<specimenCount id="E8924697FF9DFFA7FE759A8E4A12435E" box="[467,548,1017,1042]" pageId="5" pageNumber="6" type="generic">
<quantity id="396C20FBFF9DFFA7FE759A8E4A16435E" box="[467,544,1017,1042]" metricMagnitude="2" metricUnit="m" metricValue="2.26" pageId="5" pageNumber="6" unit="m" value="226.0">226 m</quantity>
.
</specimenCount>
<date id="8A2AABDEFF9DFFA7FD889A8E4A97435D" box="[558,673,1017,1042]" pageId="5" pageNumber="6" value="2011-02-26">26.ii.2011</date>
. R. González. Collected in blue light trap. MUSENUV slide code 25716.
</paragraph>
</subSubSection>
<subSubSection id="B68EDE95FF9DFFA7FF619D364CBA4315" box="[199,1164,1089,1114]" pageId="5" pageNumber="6" type="etymology">
<paragraph id="FE2B8D1EFF9DFFA7FF619D364CBA4315" blockId="5.[151,1437,402,1186]" box="[199,1164,1089,1114]" pageId="5" pageNumber="6">
<emphasis id="CCE0510CFF9DFFA7FF619D3649784315" bold="true" box="[199,334,1089,1114]" pageId="5" pageNumber="6">Etymology.</emphasis>
The specific name refers to the two distal teeth on the paraproctal prongs.
</paragraph>
</subSubSection>
<subSubSection id="B68EDE95FF9DFFA7FF619D124DAA43ED" pageId="5" pageNumber="6" type="discussion">
<paragraph id="FE2B8D1EFF9DFFA7FF619D124DAA43ED" blockId="5.[151,1437,402,1186]" pageId="5" pageNumber="6">
<emphasis id="CCE0510CFF9DFFA7FF619D12490D4331" bold="true" box="[199,315,1125,1150]" pageId="5" pageNumber="6">Remarks.</emphasis>
This species belongs in Group 1, subgroup 2, of Eertmoeds (1986) classification of
<taxonomicName id="3994F69DFF9DFFA7FB4D9D124DB54331" box="[1259,1411,1125,1150]" class="Insecta" family="Cladiopsocidae" genus="Cladiopsocus" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="CCE0510CFF9DFFA7FB4D9D124DB54331" box="[1259,1411,1125,1150]" italics="true" pageId="5" pageNumber="6">Cladiopsocus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7D3EC77FF9DFFA7FA2C9D11488D43EE" pageId="5" pageNumber="6" sensu="stricto">s. str.</taxonomicNameLabel>
, together with
<taxonomicName id="3994F69DFF9DFFA7FEC69DFD49E143ED" box="[352,471,1161,1186]" class="Insecta" family="Cladiopsocidae" genus="Cladiopsocus" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="roesleri">
<emphasis id="CCE0510CFF9DFFA7FEC69DFD49E143ED" box="[352,471,1161,1186]" italics="true" pageId="5" pageNumber="6">C. roesleri</emphasis>
</taxonomicName>
(Figs 94 and 100),
<taxonomicName id="3994F69DFF9DFFA7FD169DFD4B7743ED" box="[688,833,1161,1186]" class="Insecta" family="Cladiopsocidae" genus="Cladiopsocus" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mockfordi">
<emphasis id="CCE0510CFF9DFFA7FD169DFD4B7743ED" box="[688,833,1161,1186]" italics="true" pageId="5" pageNumber="6">C. mockfordi</emphasis>
</taxonomicName>
(Figs 93 and 99), and
<taxonomicName id="3994F69DFF9DFFA7FB9D9DFD4CEE43EE" box="[1083,1240,1162,1185]" class="Insecta" family="Cladiopsocidae" genus="Cladiopsocus" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="interruptus">
<emphasis id="CCE0510CFF9DFFA7FB9D9DFD4CEE43EE" box="[1083,1240,1162,1185]" italics="true" pageId="5" pageNumber="6">C. interruptus</emphasis>
</taxonomicName>
(Figs 92 and 98).
</paragraph>
</subSubSection>
</treatment>
</document>