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<document id="9DCBCFF18528A63DB97721AE61AF73F1" ID-CLB-Dataset="45703" ID-DOI="10.5281/zenodo.211332" ID-GBIF-Dataset="7ee29426-108f-4393-ab96-28df5a11092b" ID-ISSN="1175-5326" ID-Zenodo-Dep="211332" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460424169765" checkinUser="plazi" docAuthor="Purchart, Luboš" docDate="2012" docId="732487F5BD4E6E7EEBEFA673FF0CFA39" docLanguage="en" docName="zt03153p068.pdf" docOrigin="Zootaxa 3153" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Deretus denticollis Gahan 1900" docType="treatment" docVersion="11" lastPageNumber="61" masterDocId="8F1DFF8DBD4D6E7AEB78A553FFF4FFF9" masterDocTitle="Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae)" masterLastPageNumber="68" masterPageNumber="57" pageNumber="60" updateTime="1698330379695" updateUser="ExternalLinkService">
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<mods:title id="FA69BB1EFC85FCA854B4D191BF5382EF">Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae)</mods:title>
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<mods:date id="9EF9CB9FB513787DB486A151E3C64813">2012</mods:date>
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<treatment id="732487F5BD4E6E7EEBEFA673FF0CFA39" ID-DOI="http://doi.org/10.5281/zenodo.6180202" ID-GBIF-Taxon="119428104" ID-Zenodo-Dep="6180202" LSID="urn:lsid:plazi:treatment:732487F5BD4E6E7EEBEFA673FF0CFA39" httpUri="http://treatment.plazi.org/id/732487F5BD4E6E7EEBEFA673FF0CFA39" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60">
<subSubSection id="B3976568BD4E6E79EBEFA673FE71FCA5" pageId="3" pageNumber="60" type="nomenclature">
<paragraph id="FB3236E3BD4E6E79EBEFA673FDF8FCC1" blockId="3.[151,524,799,860]" box="[151,524,799,824]" pageId="3" pageNumber="60">
<heading id="A07A818FBD4E6E79EBEFA673FDF8FCC1" bold="true" box="[151,524,799,824]" fontSize="10" level="2" pageId="3" pageNumber="60" reason="2">
<taxonomicName id="3C8D4D60BD4E6E79EBEFA673FDF8FCC1" authority="Gahan, 1900" authorityName="Gahan" authorityYear="1900" box="[151,524,799,824]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="C9F9EAF1BD4E6E79EBEFA673FDF8FCC1" bold="true" box="[151,524,799,824]" pageId="3" pageNumber="60">
<emphasis id="C9F9EAF1BD4E6E79EBEFA673FE9AFCC1" bold="true" box="[151,366,799,824]" italics="true" pageId="3" pageNumber="60">Deretus denticollis</emphasis>
<bibRefCitation id="9F1C4B12BD4E6E79EA0DA64CFDF8FCC1" author="Gahan" box="[373,524,799,824]" pageId="3" pageNumber="60" refString="Gahan, C. J. (1900) Expedition to Sokotra. XI. Descriptions of the new genera and species of Coleoptera. Bulletin of the Liverpool Museums, 3, 8 - 13." type="journal article" year="1900">Gahan, 1900</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="FB3236E3BD4E6E79EBEFA617FE71FCA5" blockId="3.[151,524,799,860]" box="[151,389,836,860]" pageId="3" pageNumber="60">
(
<figureCitation id="63B62A66BD4E6E79EBE7A617FF0BFCA5" box="[159,255,836,860]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1945,1968]" captionTargetBox="[161,1425,339,1921]" captionTargetId="figure@2.[161,1426,339,1921]" captionTargetPageId="2" captionText="FIGURES 1 7. Habitus, dorsal view. 1. Deretus bezdeki sp. nov. (paratype), male; 2. D. bezdeki sp. nov. (paratype), female; 3. D. denticollis Gahan, 1900, male; 4. D. denticollis Gahan, 1900, female; 5. D. hulai sp. nov. (paratype), female; 6. D. maderai sp. nov. (holotype), male; 7. D. maderai sp. nov. (paratype), female." httpUri="https://zenodo.org/record/211333/files/figure.png" pageId="3" pageNumber="60">Fig. 3, 4</figureCitation>
,
<figureCitation id="63B62A66BD4E6E79EA74A617FEDEFCA5" box="[268,298,836,860]" captionStart="FIGURES 13 18" captionStartId="9.[151,264,1753,1776]" captionText="FIGURES 13 18. Pronotum, dorsal view. 13. Deretus bezdeki sp. nov. (paratype), male; 14. D. denticollis Gahan, 1900, male; 15. D. hulai sp. nov. (paratype), female; 16. D. maderai sp. nov. (holotype), male; 17. Deretus necopinatus sp. nov. (paratype), male; 18. D. spinicollis Schawaller, 2004, male." pageId="3" pageNumber="60">14</figureCitation>
,
<figureCitation id="63B62A66BD4E6E79EA4DA617FEA7FCA5" box="[309,339,836,860]" captionStart="FIGURES 19 23" captionStartId="9.[151,264,1845,1868]" captionTargetBox="[220,1367,193,1738]" captionTargetId="figure@9.[220,1367,193,1739]" captionTargetPageId="9" captionText="FIGURES 19 23. Aedeagus, dorsal view. 19. Deretus bezdeki sp. nov. (paratype), male; 20. D. denticollis Gahan, 1900, male; 21. D. maderai sp. nov. (holotype), male; 22. Deretus necopinatus sp. nov. (paratype), male; 23. D. spinicollis Schawaller, 2004, male." httpUri="https://zenodo.org/record/211335/files/figure.png" pageId="3" pageNumber="60">20</figureCitation>
,
<figureCitation id="63B62A66BD4E6E79EA27A617FE88FCA5" box="[351,380,836,860]" captionStart="FIGURES 24 28" captionStartId="9.[151,264,1937,1960]" captionText="FIGURES 24 28. Aedeagus, lateral view. 24. Deretus bezdeki sp. nov. (paratype), male; 25. D. denticollis Gahan, 1900, male; 26. D. maderai sp. nov. (holotype), male; 27. Deretus necopinatus sp. nov. (paratype), male; 28. D. spinicollis Schawaller, 2004, male." pageId="3" pageNumber="60">25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B3976568BD4E6E79EBEFA6DEFCD4FC07" pageId="3" pageNumber="60" type="reference_group">
<paragraph id="FB3236E3BD4E6E79EBEFA6DEFC3AFC39" blockId="3.[151,1355,907,1022]" pageId="3" pageNumber="60">
<treatmentCitationGroup id="DB9D11CDBD4E6E79EBEFA6DEFC3AFC39" pageId="3" pageNumber="60">
<taxonomicName id="3C8D4D60BD4E6E79EBEFA6DEFDF6FC58" authority="Gahan, 1900: 10" authorityName="Gahan" authorityPageNumber="10" authorityYear="1900" box="[151,514,907,930]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="C9F9EAF1BD4E6E79EBEFA6DEFEA2FC5B" box="[151,342,908,930]" italics="true" pageId="3" pageNumber="60">Deretus denticollis</emphasis>
<treatmentCitation id="7A2C10F2BD4E6E79EA24A6D8FDF6FC58" author="Gahan" box="[348,514,907,929]" page="10" pageId="3" pageNumber="60" year="1900">
<bibRefCitation id="9F1C4B12BD4E6E79EA24A6D8FE15FC58" author="Gahan" box="[348,481,907,929]" pageId="3" pageNumber="60" refString="Gahan, C. J. (1900) Expedition to Sokotra. XI. Descriptions of the new genera and species of Coleoptera. Bulletin of the Liverpool Museums, 3, 8 - 13." type="journal article" year="1900">Gahan, 1900</bibRefCitation>
: 10
</treatmentCitation>
</taxonomicName>
.—
<treatmentCitation id="7A2C10F2BD4E6E79E958A6D8FD2DFC58" author="Gebien" box="[544,729,907,929]" page="902" pageId="3" pageNumber="60" year="1942">
<bibRefCitation id="9F1C4B12BD4E6E79E958A6D8FD5DFC58" author="Gebien" box="[544,681,907,929]" pageId="3" pageNumber="60" refString="Gebien, H. (1942 - 1944) Katalog der Tenebrioniden. Teil III. Mitteilungen der Munchener Entomologischen Gesellschaft, 32 (1942), 729 - 760 [746 - 777]; 33 (1943), 399 - 430 [778 - 809], 895 - 926 [810 - 841]; 34 (1944), 497 - 555 [842 - 899]." type="journal article" year="1942">Gebien 1942</bibRefCitation>
: 902
</treatmentCitation>
[817];
<treatmentCitation id="7A2C10F2BD4E6E79E85CA6D8FC33FC58" author="Koch" box="[804,967,907,929]" page="124" pageId="3" pageNumber="60" year="1970">
<bibRefCitation id="9F1C4B12BD4E6E79E85CA6D8FC6CFC58" author="Koch" box="[804,920,907,929]" pageId="3" pageNumber="60" refString="Koch, C. (1970) Die Tenebrioniden (Coleoptera) des Archipels von Socotra. Italian Journal of Zoology, 4, 69 - 132." type="journal article" year="1970">Koch 1970</bibRefCitation>
: 124
</treatmentCitation>
127 (with figure of habitus) (partim).
<taxonomicName id="3C8D4D60BD4E6E79EBEFA6F8FDE4FC39" authority="Gahan 1903: 281" authorityName="Gahan" authorityPageNumber="281" authorityYear="1903" box="[151,528,938,960]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="C9F9EAF1BD4E6E79EBEFA6F8FEA2FC39" box="[151,342,938,960]" italics="true" pageId="3" pageNumber="60">Deretus denticollis</emphasis>
:
<treatmentCitation id="7A2C10F2BD4E6E79EA1BA6F9FDE4FC39" author="Gahan" box="[355,528,938,960]" page="281" pageId="3" pageNumber="60" year="1903">
<bibRefCitation id="9F1C4B12BD4E6E79EA1BA6F9FE16FC39" author="Gahan" box="[355,482,938,960]" pageId="3" pageNumber="60" refString="Gahan, C. J. (1903) Insecta: Coleoptera. In: Forbes, H. O. (Ed.), The natural history of Socotra and Abd-El-Kuri. Henry Young &amp; Sons, London, pp. 262 - 292." type="book chapter" year="1903">Gahan 1903</bibRefCitation>
: 281
</treatmentCitation>
</taxonomicName>
282 (redescription, with figure of habitus).
</treatmentCitationGroup>
</paragraph>
<paragraph id="FB3236E3BD4E6E79EBEFA699FC83FC26" blockId="3.[151,1355,907,1022]" box="[151,887,969,991]" pageId="3" pageNumber="60">
<treatmentCitationGroup id="DB9D11CDBD4E6E79EBEFA699FC83FC26" box="[151,887,969,991]" pageId="3" pageNumber="60">
<taxonomicName id="3C8D4D60BD4E6E79EBEFA699FDD6FC26" authority="Schawaller, 2004: 457" authorityName="Schawaller" authorityPageNumber="457" authorityYear="2004" box="[151,546,969,991]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="wraniki">
<emphasis id="C9F9EAF1BD4E6E79EBEFA699FECDFC26" box="[151,313,969,991]" italics="true" pageId="3" pageNumber="60">Deretus wraniki</emphasis>
<treatmentCitation id="7A2C10F2BD4E6E79EA38A69AFDD6FC26" author="Schawaller" box="[320,546,969,991]" page="457" pageId="3" pageNumber="60" year="2004">
<bibRefCitation id="9F1C4B12BD4E6E79EA38A69AFE07FC26" author="Schawaller" box="[320,499,969,991]" pageId="3" pageNumber="60" refString="Schawaller, W. (2004) New species and records of Tenebrionidae (Coleoptera) from the Socotra Archipelago. Fauna of Arabia, 20, 439 - 458." type="journal article" year="2004">Schawaller, 2004</bibRefCitation>
: 457
</treatmentCitation>
</taxonomicName>
(with figure of habitus)
<emphasis id="C9F9EAF1BD4E6E79E862A699FC83FC26" bold="true" box="[794,887,970,991]" pageId="3" pageNumber="60">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="FB3236E3BD4E6E79EBEFA6BBFCD4FC07" blockId="3.[151,1355,907,1022]" box="[151,800,1000,1022]" pageId="3" pageNumber="60">
(see also
<taxonomicName id="3C8D4D60BD4E6E79EB8DA6BAFE30FC07" box="[245,452,1001,1022]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="necopinatus" status="sp. nov.">
<emphasis id="C9F9EAF1BD4E6E79EB8DA6BAFE30FC07" box="[245,452,1001,1022]" italics="true" pageId="3" pageNumber="60">Deretus necopinatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2CA578ABD4E6E79EAB3A6BBFDE8FC04" box="[459,540,1000,1021]" pageId="3" pageNumber="60" rank="species">
<emphasis id="C9F9EAF1BD4E6E79EAB3A6BBFDE8FC04" bold="true" box="[459,540,1000,1021]" pageId="3" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="3C8D4D60BD4E6E79E935A6BAFD35FC07" box="[589,705,1000,1022]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="maderai" status="sp. nov.">
<emphasis id="C9F9EAF1BD4E6E79E935A6BAFD35FC07" box="[589,705,1000,1022]" italics="true" pageId="3" pageNumber="60">D. maderai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2CA578ABD4E6E79E9BFA6BBFCE3FC04" box="[711,791,1000,1021]" pageId="3" pageNumber="60" rank="species">
<emphasis id="C9F9EAF1BD4E6E79E9BFA6BBFCE3FC04" bold="true" box="[711,791,1000,1021]" pageId="3" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
)
</paragraph>
</subSubSection>
<subSubSection id="B3976568BD4E6E7EEBEFA178FECDFF2D" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60" type="materials_examined">
<paragraph id="FB3236E3BD4E6E79EBEFA178FD2DFBBB" blockId="3.[151,1437,1066,2027]" box="[151,729,1066,1091]" pageId="3" pageNumber="60">
<emphasis id="C9F9EAF1BD4E6E79EBEFA178FEC1FBBA" bold="true" box="[151,309,1066,1091]" pageId="3" pageNumber="60">
<typeStatus id="24368841BD4E6E79EBEFA178FF26FBBA" box="[151,210,1067,1091]" pageId="3" pageNumber="60">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="839A7673BD4E6E79EA44A178FE79FBBB" box="[316,397,1067,1090]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
,
<collectingCountry id="839A7673BD4E6E79EAE0A179FE19FBBB" box="[408,493,1066,1090]" name="Yemen" pageId="3" pageNumber="60">Socotra</collectingCountry>
Island, Jena-agahan.
</paragraph>
<paragraph id="FB3236E3BD4E6E79EBBDA11CFD80FB57" blockId="3.[151,1437,1066,2027]" pageId="3" pageNumber="60">
<emphasis id="C9F9EAF1BD4E6E79EBBDA11CFE1FFB9E" bold="true" box="[197,491,1102,1127]" pageId="3" pageNumber="60">
<typeStatus id="24368841BD4E6E79EBBDA11CFEF4FB9E" box="[197,256,1103,1127]" pageId="3" pageNumber="60">Type</typeStatus>
material examined.
</emphasis>
<taxonomicName id="3C8D4D60BD4E6E79EA8AA103FD30FB9F" box="[498,708,1102,1127]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="C9F9EAF1BD4E6E79EA8AA103FD30FB9F" box="[498,708,1102,1127]" italics="true" pageId="3" pageNumber="60">Deretus denticollis</emphasis>
</taxonomicName>
:
<typeStatus id="24368841BD4E6E79E9AAA103FCCBFB9E" box="[722,831,1104,1127]" pageId="3" pageNumber="60" type="lectotype">
<emphasis id="C9F9EAF1BD4E6E79E9AAA103FCCBFB9E" box="[722,831,1104,1127]" italics="true" pageId="3" pageNumber="60">Lectotype</emphasis>
</typeStatus>
(designated here)—(Ƥ
<collectionCode id="9D9CAE26BD4E6E79EF3FA11DFB55FB9F" box="[1095,1185,1102,1126]" pageId="3" pageNumber="60">BMNH</collectionCode>
):
<typeStatus id="24368841BD4E6E79EFCBA11CFB1EFB9E" box="[1203,1258,1103,1127]" pageId="3" pageNumber="60">Type</typeStatus>
{white and red, rounded, printed} // Jena-agahan,
<collectingCountry id="839A7673BD4E6E79E96AA121FD99FB73" box="[530,621,1138,1162]" name="Yemen" pageId="3" pageNumber="60">Sokotra</collectingCountry>
.
<quantity id="3C759B06BD4E6E79E90FA121FD16FB73" box="[631,738,1138,1162]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="3" pageNumber="60" unit="ft" value="1200.0">1200 feet</quantity>
,
<date id="8F331023BD4E6E79E995A120FCCAFB73" box="[749,830,1138,1162]" pageId="3" pageNumber="60" value="1999-01">Jan. 99</date>
, W.R.O. Grant, 99-85 {white, printed} //
<taxonomicName id="3C8D4D60BD4E6E79EE74A120FEC2FB57" authority="Gahan" authorityName="Gahan" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="denticollis">Deretus denticollis Gahan</taxonomicName>
,
<typeStatus id="24368841BD4E6E79EA39A1C4FE8DFB56" box="[321,377,1175,1199]" pageId="3" pageNumber="60">Type</typeStatus>
{white, handwritten}.
</paragraph>
<paragraph id="FB3236E3BD4E6E79EBBDA1EFFDCEFA9B" blockId="3.[151,1437,1066,2027]" pageId="3" pageNumber="60">
<taxonomicName id="3C8D4D60BD4E6E79EBBDA1EFFE88FB2A" box="[197,380,1211,1235]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="wraniki">
<emphasis id="C9F9EAF1BD4E6E79EBBDA1EFFE88FB2A" box="[197,380,1211,1235]" italics="true" pageId="3" pageNumber="60">Deretus wraniki</emphasis>
</taxonomicName>
:
<typeStatus id="24368841BD4E6E79EAF6A1E8FE01FB2A" box="[398,501,1211,1235]" pageId="3" pageNumber="60" type="holotype">
<emphasis id="C9F9EAF1BD4E6E79EAF6A1E8FE01FB2A" box="[398,501,1211,1235]" italics="true" pageId="3" pageNumber="60">Holotype</emphasis>
</typeStatus>
—(13
<collectionCode id="9D9CAE26BD4E6E79E93FA1E9FD6DFB2B" LSID="urn:lsid:biocol.org:col:34840" box="[583,665,1210,1234]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="3" pageNumber="60">SMNS</collectionCode>
): ZOOLOGISCHE EXKURSION,
<collectingCountry id="839A7673BD4E6E79EF57A1E9FB73FB2B" box="[1071,1159,1210,1234]" name="Yemen" pageId="3" pageNumber="60">Socotra</collectingCountry>
, Diksam, 11. 1997, leg. Wranik {white label, handwritten}//
<typeStatus id="24368841BD4E6E79E94EA18DFD14FB0F" box="[566,736,1246,1270]" pageId="3" pageNumber="60" type="holotype">HOLOTYPUS</typeStatus>
,
<taxonomicName id="3C8D4D60BD4E6E79E993A18CFC54FB0F" box="[747,928,1246,1270]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="wraniki" status="sp. nov.">Deretus wraniki</taxonomicName>
<taxonomicNameLabel id="D2CA578ABD4E6E79E8D0A18CFC13FB0F" box="[936,999,1247,1270]" pageId="3" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
, Schawaller 2000 {red label, printed};
<typeStatus id="24368841BD4E6E79EBEFA057FF0AFAE2" box="[151,254,1284,1307]" pageId="3" pageNumber="60" type="paratype">
<emphasis id="C9F9EAF1BD4E6E79EBEFA057FF0AFAE2" box="[151,254,1284,1307]" italics="true" pageId="3" pageNumber="60">Paratype</emphasis>
</typeStatus>
[actually
<taxonomicName id="3C8D4D60BD4E6E79EA16A057FE19FAE2" box="[366,493,1283,1307]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="maderai" status="sp. nov.">
<emphasis id="C9F9EAF1BD4E6E79EA16A057FE19FAE2" box="[366,493,1283,1307]" italics="true" pageId="3" pageNumber="60">D. maderai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2CA578ABD4E6E79EA8DA050FDC4FAE3" box="[501,560,1283,1306]" pageId="3" pageNumber="60" rank="species">sp. n.</taxonomicNameLabel>
] (1Ƥ
<collectionCode id="9D9CAE26BD4E6E79E9FBA051FD39FAE3" box="[643,717,1282,1306]" pageId="3" pageNumber="60">SBSC</collectionCode>
):
<collectingCountry id="839A7673BD4E6E79E998A050FCB7FAE3" box="[736,835,1283,1306]" name="Yemen" pageId="3" pageNumber="60">YEMEN</collectingCountry>
E,
<collectingCountry id="839A7673BD4E6E79E811A051FC4AFAE3" box="[873,958,1282,1306]" name="Yemen" pageId="3" pageNumber="60">Socotra</collectingCountry>
Isl.,
<date id="8F331023BD4E6E79E880A051FB61FAE3" box="[1016,1173,1282,1306]" pageId="3" pageNumber="60" value="1999-09-06" valueMax="1999-09-24" valueMin="1999-09-06">6.24.ix.1999</date>
, V.Bejček &amp; K.Šťastný lgt. {printed}// COLLECTION STANISLAV BEČVÁŘ {yellow, printed} //
<typeStatus id="24368841BD4E6E79E88FA075FB6CFAC7" box="[1015,1176,1318,1342]" pageId="3" pageNumber="60" type="paratype">PARATYPUS</typeStatus>
,
<taxonomicName id="3C8D4D60BD4E6E79EFDCA074FAACFAC7" box="[1188,1368,1318,1342]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="wraniki" status="sp. nov.">Deretus wraniki</taxonomicName>
<taxonomicNameLabel id="D2CA578ABD4E6E79EE18A074FA63FAC7" box="[1376,1431,1319,1342]" pageId="3" pageNumber="60" rank="species">n.sp.</taxonomicNameLabel>
, SCHAWALLER 2000 {red, printed}.
</paragraph>
<paragraph id="FB3236E3BD4E6E7EEBBDA03DFECDFF2D" blockId="3.[151,1437,1066,2027]" lastBlockId="4.[151,1437,151,1472]" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60">
<emphasis id="C9F9EAF1BD4E6E79EBBDA03DFE43FA7F" bold="true" box="[197,439,1390,1415]" pageId="3" pageNumber="60">Additional material.</emphasis>
(43+ 4Ƥ
<collectionCode id="9D9CAE26BD4E6E79E952A03DFD82FA7F" box="[554,630,1390,1414]" pageId="3" pageNumber="60">LPCB</collectionCode>
):
<collectingCountry id="839A7673BD4E6E79E9F0A03CFD2DFA7F" box="[648,729,1391,1414]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
,
<collectingCountry id="839A7673BD4E6E79E99CA03DFCCDFA7F" box="[740,825,1390,1414]" name="Yemen" pageId="3" pageNumber="60">Socotra</collectingCountry>
Island, wadi Ayhaft, 12°36.5N, 53°58.9E,
<quantity id="3C759B06BD4E6E79EE54A03DFA80FA7F" box="[1324,1396,1390,1414]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="60" unit="m" value="200.0">200 m</quantity>
,
<date id="8F331023BD4E6E79EEF8A03CFEF7FA53" pageId="3" pageNumber="60" value="2010-11-07" valueMax="2010-11-08" valueMin="2010-11-07">7 8.xi.2010</date>
, L. Purchart lgt.; (43+6Ƥ
<collectionCode id="9D9CAE26BD4E6E79E94FA0C1FD77FA53" box="[567,643,1426,1450]" pageId="3" pageNumber="60">LPCB</collectionCode>
): same data as previous, but J. Bezdĕk lgt.; (1Ƥ
<collectionCode id="9D9CAE26BD4E6E79EFC9A0C1FB03FA53" box="[1201,1271,1426,1450]" pageId="3" pageNumber="60">JBCP</collectionCode>
): same data as previous, but Jan Batelka leg.; (53+4Ƥ
<collectionCode id="9D9CAE26BD4E6E79E920A0E5FD59FA37" box="[600,685,1462,1486]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="60">NMPC</collectionCode>
): same data as previous, but Jiří Hájek leg.; (13
<collectionCode id="9D9CAE26BD4E6E79EFB6A0E5FAEEFA37" box="[1230,1306,1462,1486]" pageId="3" pageNumber="60">LPCB</collectionCode>
): same data as previous, but J. Bezdĕk lgt., Alcohol 70 % + acetic acid; (1Ƥ
<collectionCode id="9D9CAE26BD4E6E79E8F9A089FC3AFA0B" box="[897,974,1498,1522]" pageId="3" pageNumber="60">SBCB</collectionCode>
):
<collectingCountry id="839A7673BD4E6E79E89BA088FBC0FA0B" box="[995,1076,1499,1522]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
,
<collectingCountry id="839A7673BD4E6E79EF38A089FB62FA0B" box="[1088,1174,1498,1522]" name="Yemen" pageId="3" pageNumber="60">Socotra</collectingCountry>
Isl., Hadiboh env., 6. 24.1999, V.Bejček &amp; K.Šťastný lgt.; (13
<collectionCode id="9D9CAE26BD4E6E79E910A0ADFD41F9EF" box="[616,693,1534,1558]" pageId="3" pageNumber="60">SBCB</collectionCode>
):
<collectingCountry id="839A7673BD4E6E79E9B0A0ADFCEAF9EF" box="[712,798,1534,1558]" name="Yemen" pageId="3" pageNumber="60">Socotra</collectingCountry>
Isl. (
<collectingCountry id="839A7673BD4E6E79E82DA0ADFC5DF9EF" box="[853,937,1534,1558]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
), Hamadero,
<date id="8F331023BD4E6E79EF47A0ADFB0DF9EF" box="[1087,1273,1534,1558]" pageId="3" pageNumber="60" value="2000-11-20" valueMax="2000-11-21" valueMin="2000-11-20">20.21. xi. 2000</date>
, lgt. V. Bejček + K. Šťastný; (13
<collectionCode id="9D9CAE26BD4E6E79EA0CA371FE35F9C3" box="[372,449,1570,1594]" pageId="3" pageNumber="60">SBCB</collectionCode>
):
<collectingCountry id="839A7673BD4E6E79EAAEA371FDDFF9C3" box="[470,555,1570,1594]" name="Yemen" pageId="3" pageNumber="60">Socotra</collectingCountry>
Isl. (
<collectingCountry id="839A7673BD4E6E79E912A371FD4AF9C3" box="[618,702,1570,1594]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
), Ayhaft,
<date id="8F331023BD4E6E79E848A371FC45F9C3" box="[816,945,1570,1594]" pageId="3" pageNumber="60" value="2000-11-03">3. xi. 2000</date>
, lgt. V. Bejček + K. Šťastný; (23
<collectionCode id="9D9CAE26BD4E6E79EE3BA371FA64F9C3" box="[1347,1424,1570,1594]" pageId="3" pageNumber="60">SBCB</collectionCode>
):
<collectingCountry id="839A7673BD4E6E79EBEFA315FF18F9A7" box="[151,236,1606,1630]" name="Yemen" pageId="3" pageNumber="60">Socotra</collectingCountry>
Isl. (
<collectingCountry id="839A7673BD4E6E79EA5EA315FE8EF9A7" box="[294,378,1606,1630]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
), Lahas,
<date id="8F331023BD4E6E79EA98A315FD6FF9A7" box="[480,667,1606,1630]" pageId="3" pageNumber="60" value="2000-11-17" valueMax="2000-11-18" valueMin="2000-11-17">17.18. xi. 2000</date>
, lgt. V. Bejček + K. Šťastný; (23+ 2Ƥ
<collectionCode id="9D9CAE26BD4E6E79EF2EA315FB5FF9A7" box="[1110,1195,1606,1630]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="60">NMPC</collectionCode>
):
<collectingCountry id="839A7673BD4E6E79EFC6A314FAFBF9A7" box="[1214,1295,1607,1630]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
,
<collectingCountry id="839A7673BD4E6E79EE63A315FA86F9A7" box="[1307,1394,1606,1630]" name="Yemen" pageId="3" pageNumber="60">Soqotra</collectingCountry>
Is.,
<date id="8F331023BD4E6E79EBEFA339FE93F97B" box="[151,359,1642,1666]" pageId="3" pageNumber="60" value="2003-11-21" valueMax="2003-12-12" valueMin="2003-11-21">21.xi.12.xii.2003</date>
, Hadiboh env., ca
<quantity id="3C759B06BD4E6E79E931A339FD35F97B" box="[585,705,1642,1666]" metricMagnitude="1" metricUnit="m" metricValue="5.5" metricValueMax="10.0" metricValueMin="1.0" pageId="3" pageNumber="60" unit="m" value="55.0" valueMax="100.0" valueMin="10.0">10100 m</quantity>
,
<geoCoordinate id="9EB95024BD4E6E79E9AAA338FC9FF97B" box="[722,875,1642,1666]" direction="north" orientation="latitude" pageId="3" pageNumber="60" precision="15" value="13.083889">N 12°6502</geoCoordinate>
<geoCoordinate id="9EB95024BD4E6E79E80FA338FBF9F97B" box="[887,1037,1642,1666]" direction="east" orientation="longitude" pageId="3" pageNumber="60" precision="15" value="54.034443">E 54°0204</geoCoordinate>
[GPS], David Král lgt.//
<collectingCountry id="839A7673BD4E6E79EE38A338FA7AF97B" box="[1344,1422,1643,1666]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
<collectingCountry id="839A7673BD4E6E79EBEFA3DDFF05F95F" box="[151,241,1678,1702]" name="Yemen" pageId="3" pageNumber="60">Soqotra</collectingCountry>
, 2003, Expedition; Jan Farkač, Petr Kabátek &amp; David Král; (23+ 4Ƥ
<collectionCode id="9D9CAE26BD4E6E79EF65A3DDFB86F95F" box="[1053,1138,1678,1702]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="60">NMPC</collectionCode>
):
<collectingCountry id="839A7673BD4E6E79EFFFA3DCFB2CF95F" box="[1159,1240,1679,1702]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
,
<collectingCountry id="839A7673BD4E6E79EF9EA3DDFAC9F95F" box="[1254,1341,1678,1702]" name="Yemen" pageId="3" pageNumber="60">Soqotra</collectingCountry>
Is.,
<date id="8F331023BD4E6E79EE0AA3DDFEE5F933" pageId="3" pageNumber="60" value="2003-11-24" valueMax="2003-11-26" valueMin="2003-11-24">24 26.xi.2003</date>
, wadi Ayhaft,
<quantity id="3C759B06BD4E6E79EACBA3E1FE0FF933" box="[435,507,1714,1738]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="60" unit="m" value="190.0">190 m</quantity>
,
<geoCoordinate id="9EB95024BD4E6E79E97FA3E1FD64F933" box="[519,656,1714,1738]" direction="north" orientation="latitude" pageId="3" pageNumber="60" precision="15" value="12.610556">N12°3638”</geoCoordinate>
<geoCoordinate id="9EB95024BD4E6E79E9EFA3E1FCE9F933" box="[663,797,1714,1738]" direction="east" orientation="longitude" pageId="3" pageNumber="60" precision="15" value="53.980278">E53°5849”</geoCoordinate>
[GPS], David Král lgt. //
<collectingCountry id="839A7673BD4E6E79EF45A3E1FB7AF933" box="[1085,1166,1714,1738]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
<collectingCountry id="839A7673BD4E6E79EFE4A3E1FB07F933" box="[1180,1267,1714,1738]" name="Yemen" pageId="3" pageNumber="60">Soqotra</collectingCountry>
, 2003, Expedition; Jan Farkač, Petr Kabátek &amp; David Král; (13
<collectionCode id="9D9CAE26BD4E6E79E992A385FCD8F917" box="[746,812,1750,1774]" pageId="3" pageNumber="60">JFCP</collectionCode>
): same as previous, but Jan Farkač lgt.; (13
<collectionCode id="9D9CAE26BD4E6E79EE44A385FA65F917" box="[1340,1425,1750,1774]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="60">NMPC</collectionCode>
):
<collectingCountry id="839A7673BD4E6E79EBEFA3A8FF1CF8EB" box="[151,232,1787,1810]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
,
<collectingCountry id="839A7673BD4E6E79EB8BA3A9FEBEF8EB" box="[243,330,1786,1810]" name="Yemen" pageId="3" pageNumber="60">Soqotra</collectingCountry>
Is., Homhil protected area,
<date id="8F331023BD4E6E79E9F8A3A9FCD1F8EB" box="[640,805,1786,1810]" pageId="3" pageNumber="60" value="2003-11-28" valueMax="2003-11-29" valueMin="2003-11-28">
2829.xi.
<quantity id="3C759B06BD4E6E79E991A3A9FCD1F8EB" box="[745,805,1786,1810]" metricMagnitude="6" metricUnit="m" metricValue="2.003364" pageId="3" pageNumber="60" unit="m" value="2003364.0">2003</quantity>
</date>
, 364 m,
<geoCoordinate id="9EB95024BD4E6E79E8FCA3A9FBF9F8EB" box="[900,1037,1786,1810]" direction="north" orientation="latitude" pageId="3" pageNumber="60" precision="15" value="12.574166">N12°3427”</geoCoordinate>
<geoCoordinate id="9EB95024BD4E6E79EF6CA3A9FB6DF8EB" box="[1044,1177,1786,1810]" direction="east" orientation="longitude" pageId="3" pageNumber="60" precision="15" value="54.308887">E54°1832”</geoCoordinate>
[GPS], David Král lgt. //
<collectingCountry id="839A7673BD4E6E79EBD7A24DFEF4F8CF" box="[175,256,1822,1846]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
<collectingCountry id="839A7673BD4E6E79EA77A24DFE92F8CF" box="[271,358,1822,1846]" name="Yemen" pageId="3" pageNumber="60">Soqotra</collectingCountry>
, 2003, Expedition; Jan Farkač, Petr Kabátek &amp; David Král; (33
<collectionCode id="9D9CAE26BD4E6E79EF37A24DFB66F8CF" box="[1103,1170,1822,1846]" pageId="3" pageNumber="60">JFCP</collectionCode>
): same as previous, but Jan Farkač lgt.; (1Ƥ
<collectionCode id="9D9CAE26BD4E6E79EA04A211FE4AF8A3" box="[380,446,1858,1882]" pageId="3" pageNumber="60">JFCP</collectionCode>
):
<collectingCountry id="839A7673BD4E6E79EAA9A210FDD6F8A3" box="[465,546,1859,1882]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
,
<collectingCountry id="839A7673BD4E6E79E955A211FD70F8A3" box="[557,644,1858,1882]" name="Yemen" pageId="3" pageNumber="60">Soqotra</collectingCountry>
Is.,
<date id="8F331023BD4E6E79E9CCA211FCBEF8A3" box="[692,842,1858,1882]" pageId="3" pageNumber="60" value="2003-12-06" valueMax="2003-12-07" valueMin="2003-12-06">6.7.xii.2003</date>
, Noged plain: wadi Ireeh,
<geoCoordinate id="9EB95024BD4E6E79EF0AA210FAF6F8A3" box="[1138,1282,1858,1882]" direction="north" orientation="latitude" pageId="3" pageNumber="60" precision="15" value="12.386389">N 12°2311”</geoCoordinate>
<geoCoordinate id="9EB95024BD4E6E79EE70A210FA63F8A3" box="[1288,1431,1858,1882]" direction="east" orientation="longitude" pageId="3" pageNumber="60" precision="15" value="53.996387">E 53°5947”</geoCoordinate>
,
<quantity id="3C759B06BD4E6E79EBEFA235FF26F887" box="[151,210,1894,1918]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="3" pageNumber="60" unit="m" value="95.0">95 m</quantity>
[GPS]; Jan Farkač lgt. //
<collectingCountry id="839A7673BD4E6E79EA86A235FDBBF887" box="[510,591,1894,1918]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
<collectingCountry id="839A7673BD4E6E79E926A235FD41F887" box="[606,693,1894,1918]" name="Yemen" pageId="3" pageNumber="60">Soqotra</collectingCountry>
, 2003, Expedition; Jan Farkač, Petr Kabátek &amp; David Král; (23
<collectionCode id="9D9CAE26BD4E6E79EBEFA2D9FF18F85B" box="[151,236,1930,1954]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="60">NMPC</collectionCode>
):
<collectingCountry id="839A7673BD4E6E79EB87A2D8FEA4F85B" box="[255,336,1931,1954]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
,
<collectingCountry id="839A7673BD4E6E79EA24A2D9FE45F85B" box="[348,433,1930,1954]" name="Yemen" pageId="3" pageNumber="60">Socotra</collectingCountry>
Island, Aloove area, Hassan vill. env., 12°31.2N, 54°07.4E,
<quantity id="3C759B06BD4E6E79EF0CA2D9FB4AF85B" box="[1140,1214,1930,1954]" metricMagnitude="2" metricUnit="m" metricValue="2.21" pageId="3" pageNumber="60" unit="m" value="221.0">221 m</quantity>
, Jiří Hájek leg.,
<date id="8F331023BD4E6E79EE01A2D9FEE6F83F" pageId="3" pageNumber="60" value="2010-11-09" valueMax="2010-11-10" valueMin="2010-11-09">9. 10.xi.2010</date>
; (1
<collectionCode id="9D9CAE26BD4E6E79EA4FA2FCFE6AF83E" box="[311,414,1967,1991]" pageId="3" pageNumber="60">ƤSMNS</collectionCode>
):
<collectingCountry id="839A7673BD4E6E79EACAA2FDFDF3F83F" box="[434,519,1966,1990]" name="Yemen" pageId="3" pageNumber="60">Socotra</collectingCountry>
Archipelago,
<collectingCountry id="839A7673BD4E6E79E9D3A2FDFCF7F83F" box="[683,771,1966,1990]" name="Yemen" pageId="3" pageNumber="60">Socotra</collectingCountry>
, Hadibo,
<date id="8F331023BD4E6E79E80CA2FDFC14F83F" box="[884,992,1966,1990]" pageId="3" pageNumber="60" value="2008-11-01">1.xi.2008</date>
, leg. T. van Harten; (33+ 1Ƥ
<collectionCode id="9D9CAE26BD4E6E79EE46A2FDFA64F83F" LSID="urn:lsid:biocol.org:col:34840" box="[1342,1424,1966,1990]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="3" pageNumber="60">SMNS</collectionCode>
):
<collectingCountry id="839A7673BD4E6E79EBEFA281FF18F813" box="[151,236,2002,2026]" name="Yemen" pageId="3" pageNumber="60">Socotra</collectingCountry>
Archipelago,
<collectingCountry id="839A7673BD4E6E79EAF6A281FE12F813" box="[398,486,2002,2026]" name="Yemen" pageId="3" pageNumber="60">Socotra</collectingCountry>
, Wadi Danegan,
<quantity id="3C759B06BD4E6E79E9D0A281FD10F813" box="[680,740,2002,2026]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="60" unit="m" value="90.0">90 m</quantity>
,
<date id="8F331023BD4E6E79E98AA281FC91F813" box="[754,869,2002,2026]" pageId="3" pageNumber="60" value="2000-10-30">30.x.2000</date>
, leg. T. van Harten; (1Ƥ
<collectionCode id="9D9CAE26BD4E6E79EFFAA281FB20F813" LSID="urn:lsid:biocol.org:col:34840" box="[1154,1236,2002,2026]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="3" pageNumber="60">SMNS</collectionCode>
):
<collectingCountry id="839A7673BD4E6E79EF9FA280FACCF813" box="[1255,1336,2003,2026]" name="Yemen" pageId="3" pageNumber="60">Yemen</collectingCountry>
,
<collectingCountry id="839A7673BD4E6E79EE3DA281FA68F813" box="[1349,1436,2002,2026]" name="Yemen" pageId="3" pageNumber="60">Soqotra</collectingCountry>
Archipelago,
<collectingCountry id="839A7673BD496E7EEA49A5C4FE7FFF56" box="[305,395,151,175]" name="Yemen" pageId="4" pageNumber="61">Soqotra</collectingCountry>
, Noged, Farnihin, Steroh wadi, 12°2426 N,54°0810 E,
<date id="8F331023BD496E7EEF4EA5C4FB4FFF56" box="[1078,1211,151,175]" pageId="4" pageNumber="61" value="2010-10-24">24. x. 2010</date>
, leg. T. van Harten, SOQ 2000/06.
</paragraph>
</subSubSection>
<subSubSection id="B3976568BD496E7EEBBDA5B3FF0CFA39" pageId="4" pageNumber="61" type="discussion">
<paragraph id="FB3236E3BD496E7EEBBDA5B3FC30FEC6" blockId="4.[151,1437,151,1472]" pageId="4" pageNumber="61">
<emphasis id="C9F9EAF1BD496E7EEBBDA5B3FECDFF0E" bold="true" box="[197,313,224,248]" pageId="4" pageNumber="61">Remarks.</emphasis>
<taxonomicName id="3C8D4D60BD496E7EEA47A5B2FE22FF01" box="[319,470,224,248]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="C9F9EAF1BD496E7EEA47A5B2FE22FF01" box="[319,470,224,248]" italics="true" pageId="4" pageNumber="61">D. denticollis</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="9F1C4B12BD496E7EE9DCA58CFCCBFF0E" author="Gahan" box="[676,831,223,247]" pageId="4" pageNumber="61" refString="Gahan, C. J. (1900) Expedition to Sokotra. XI. Descriptions of the new genera and species of Coleoptera. Bulletin of the Liverpool Museums, 3, 8 - 13." type="journal article" year="1900">Gahan (1900)</bibRefCitation>
probably based on a single female, as the only known
<typeStatus id="24368841BD496E7EEBEFA456FF33FEE4" box="[151,199,261,285]" pageId="4" pageNumber="61">type</typeStatus>
is a female deposited in the BMNH. However, Gahan stated neither the number of described specimens nor their deposition. For that reason I designate this specimen as a
<typeStatus id="24368841BD496E7EE82EA474FC34FEC6" box="[854,960,295,319]" pageId="4" pageNumber="61" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="FB3236E3BD496E7EEBBDA41FFD86FDC5" blockId="4.[151,1437,151,1472]" pageId="4" pageNumber="61">
<bibRefCitation id="9F1C4B12BD496E7EEBBDA41FFEA0FE9D" author="Koch" box="[197,340,332,356]" pageId="4" pageNumber="61" refString="Koch, C. (1970) Die Tenebrioniden (Coleoptera) des Archipels von Socotra. Italian Journal of Zoology, 4, 69 - 132." type="journal article" year="1970">Koch (1970)</bibRefCitation>
made a detailed description of a single male specimen deposited in the Frey collection (Natural History Museum, Basel) and designated it as the allotype of
<taxonomicName id="3C8D4D60BD496E7EE845A422FC20FE71" box="[829,980,368,392]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="C9F9EAF1BD496E7EE845A422FC20FE71" box="[829,980,368,392]" italics="true" pageId="4" pageNumber="61">D. denticollis</emphasis>
</taxonomicName>
. However, this is not a valid name-bearing
<typeStatus id="24368841BD496E7EEBB9A4C6FF05FE54" box="[193,241,405,429]" pageId="4" pageNumber="61">type</typeStatus>
(
<bibRefCitation id="9F1C4B12BD496E7EEB87A4C7FE34FE55" author="ICZN" box="[255,448,404,428]" pageId="4" pageNumber="61" refString="ICZN (1999) International Code of Zoological Nomenclature. The International Trust for Zoological Nomenclature, London. 306 pp." type="book" year="1999">ICZN 1999: 72A</bibRefCitation>
) and therefore it cannot be used as a
<typeStatus id="24368841BD496E7EE823A4C6FC7FFE54" box="[859,907,405,429]" pageId="4" pageNumber="61">type</typeStatus>
specimen. Koch undoubtedly believed that this male belonged to the same species as the above mentioned female
<typeStatus id="24368841BD496E7EE8F0A4EBFC4CFE29" box="[904,952,440,464]" pageId="4" pageNumber="61">type</typeStatus>
specimen in the BMNH. Much later
<bibRefCitation id="9F1C4B12BD496E7EEE24A4E4FEDAFE0D" author="Schawaller" pageId="4" pageNumber="61" refString="Schawaller, W. (2004) New species and records of Tenebrionidae (Coleoptera) from the Socotra Archipelago. Fauna of Arabia, 20, 439 - 458." type="journal article" year="2004">Schawaller (2004)</bibRefCitation>
listed an additional
<specimenCount id="ED8BFD6ABD496E7EE968A48FFD88FE0D" box="[528,636,476,500]" pageId="4" pageNumber="61" type="female">4 females</specimenCount>
as
<taxonomicName id="3C8D4D60BD496E7EE9D9A48EFCCCFE0D" box="[673,824,476,500]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="C9F9EAF1BD496E7EE9D9A48EFCCCFE0D" box="[673,824,476,500]" italics="true" pageId="4" pageNumber="61">D. denticollis</emphasis>
</taxonomicName>
that were compared with this non-type male described by Koch. The same author also described
<taxonomicName id="3C8D4D60BD496E7EE914A752FC1CFDEE" authority="Schawaller, 2004" authorityName="Schawaller" authorityYear="2004" box="[620,1000,511,536]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="wraniki">
<emphasis id="C9F9EAF1BD496E7EE914A752FCD5FDE1" box="[620,801,512,536]" italics="true" pageId="4" pageNumber="61">Deretus wraniki</emphasis>
<bibRefCitation id="9F1C4B12BD496E7EE85FA4ACFC1CFDEE" author="Schawaller" box="[807,1000,511,535]" pageId="4" pageNumber="61" refString="Schawaller, W. (2004) New species and records of Tenebrionidae (Coleoptera) from the Socotra Archipelago. Fauna of Arabia, 20, 439 - 458." type="journal article" year="2004">Schawaller, 2004</bibRefCitation>
</taxonomicName>
which was differentiated from
<taxonomicName id="3C8D4D60BD496E7EEE30A752FF17FDC5" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="C9F9EAF1BD496E7EEE30A752FF17FDC5" italics="true" pageId="4" pageNumber="61">D. denticollis</emphasis>
</taxonomicName>
mainly by its very small body size.
</paragraph>
<paragraph id="FB3236E3BD496E7EEBBDA714FEC0FC31" blockId="4.[151,1437,151,1472]" pageId="4" pageNumber="61">
During expeditions to
<collectingCountry id="839A7673BD496E7EEAB8A714FDE1FDA6" box="[448,533,583,607]" name="Yemen" pageId="4" pageNumber="61">Socotra</collectingCountry>
Island the author collected, with his colleagues, a larger series of
<taxonomicName id="3C8D4D60BD496E7EEF82A71AFAA6FD99" box="[1274,1362,585,608]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="C9F9EAF1BD496E7EEF82A71AFAA6FD99" box="[1274,1362,585,608]" italics="true" pageId="4" pageNumber="61">Deretus</emphasis>
</taxonomicName>
specimens comprising males and females of several species. These specimens were compared with the
<typeStatus id="24368841BD496E7EEF91A73EFAEDFD7C" box="[1257,1305,621,645]" pageId="4" pageNumber="61">type</typeStatus>
material of
<taxonomicName id="3C8D4D60BD496E7EEBEFA7C2FEDAFD51" box="[151,302,656,680]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="C9F9EAF1BD496E7EEBEFA7C2FEDAFD51" box="[151,302,656,680]" italics="true" pageId="4" pageNumber="61">D. denticollis</emphasis>
</taxonomicName>
and
<taxonomicName id="3C8D4D60BD496E7EEA1EA7C2FE2BFD51" box="[358,479,656,680]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="wraniki">
<emphasis id="C9F9EAF1BD496E7EEA1EA7C2FE2BFD51" box="[358,479,656,680]" italics="true" pageId="4" pageNumber="61">D. wraniki</emphasis>
</taxonomicName>
and the non-type male of
<taxonomicName id="3C8D4D60BD496E7EE870A7C2FC54FD51" box="[776,928,656,680]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="C9F9EAF1BD496E7EE870A7C2FC54FD51" box="[776,928,656,680]" italics="true" pageId="4" pageNumber="61">D. denticollis</emphasis>
</taxonomicName>
from the Frey collection. The author discovered that the male
<typeStatus id="24368841BD496E7EEA17A7E7FE25FD35" box="[367,465,692,716]" pageId="4" pageNumber="61" type="holotype">holotype</typeStatus>
of
<taxonomicName id="3C8D4D60BD496E7EEA83A7E6FD81FD35" box="[507,629,692,716]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="wraniki">
<emphasis id="C9F9EAF1BD496E7EEA83A7E6FD81FD35" box="[507,629,692,716]" italics="true" pageId="4" pageNumber="61">D. wraniki</emphasis>
</taxonomicName>
is identical to the
<typeStatus id="24368841BD496E7EE837A7E7FC41FD35" box="[847,949,692,716]" pageId="4" pageNumber="61" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="3C8D4D60BD496E7EE8A7A7E6FB8DFD35" box="[991,1145,692,716]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="C9F9EAF1BD496E7EE8A7A7E6FB8DFD35" box="[991,1145,692,716]" italics="true" pageId="4" pageNumber="61">D. denticollis</emphasis>
</taxonomicName>
and therefore
<taxonomicName id="3C8D4D60BD496E7EEE5AA7E6FA68FD35" box="[1314,1436,692,716]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="wraniki">
<emphasis id="C9F9EAF1BD496E7EEE5AA7E6FA68FD35" box="[1314,1436,692,716]" italics="true" pageId="4" pageNumber="61">D. wraniki</emphasis>
</taxonomicName>
should be synonymised. He also observed, based on a large series of this species, that the body size of this species is strongly variable and cannot be used for differential diagnoses. The
<typeStatus id="24368841BD496E7EE8DDA7AFFBF3FCED" box="[933,1031,764,788]" pageId="4" pageNumber="61" type="holotype">holotype</typeStatus>
of
<taxonomicName id="3C8D4D60BD496E7EEF53A7AEFB57FCED" box="[1067,1187,764,788]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="wraniki">
<emphasis id="C9F9EAF1BD496E7EEF53A7AEFB57FCED" box="[1067,1187,764,788]" italics="true" pageId="4" pageNumber="61">D. wraniki</emphasis>
</taxonomicName>
simply represents one extreme of the body-size range. It was also found that the female
<typeStatus id="24368841BD496E7EE802A672FC2EFCC0" box="[890,986,801,825]" pageId="4" pageNumber="61" type="paratype">paratype</typeStatus>
of
<taxonomicName id="3C8D4D60BD496E7EEF79A672FB8EFCC1" box="[1025,1146,800,824]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="wraniki">
<emphasis id="C9F9EAF1BD496E7EEF79A672FB8EFCC1" box="[1025,1146,800,824]" italics="true" pageId="4" pageNumber="61">D. wraniki</emphasis>
</taxonomicName>
is a different species (see remarks in
<taxonomicName id="3C8D4D60BD496E7EEA6CA616FE67FCA5" box="[276,403,836,860]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="maderai" status="sp. nov.">
<emphasis id="C9F9EAF1BD496E7EEA6CA616FE67FCA5" box="[276,403,836,860]" italics="true" pageId="4" pageNumber="61">D. maderai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2CA578ABD496E7EEAE3A617FE00FCA5" box="[411,500,836,860]" pageId="4" pageNumber="61" rank="species">
<emphasis id="C9F9EAF1BD496E7EEAE3A617FE00FCA5" bold="true" box="[411,500,836,860]" pageId="4" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
). The most surprising discovery is that all specimens previously identified by Koch and Schawaller as
<taxonomicName id="3C8D4D60BD496E7EEA10A63AFE0BFC79" box="[360,511,872,896]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="C9F9EAF1BD496E7EEA10A63AFE0BFC79" box="[360,511,872,896]" italics="true" pageId="4" pageNumber="61">D. denticollis</emphasis>
</taxonomicName>
differ from the
<typeStatus id="24368841BD496E7EE9CAA634FCEDFC86" box="[690,793,871,895]" pageId="4" pageNumber="61" type="lectotype">lectotype</typeStatus>
(designated in this paper) and from new specimens of
<taxonomicName id="3C8D4D60BD496E7EEEF9A63AFEF8FC5D" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="C9F9EAF1BD496E7EEEF9A63AFEF8FC5D" italics="true" pageId="4" pageNumber="61">D. denticollis</emphasis>
</taxonomicName>
collected in the field. These specimens represent a new species, which has been named
<taxonomicName id="3C8D4D60BD496E7EEF91A6DEFF20FC31" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="necopinatus" status="sp. nov.">
<emphasis id="C9F9EAF1BD496E7EEF91A6DEFF20FC31" italics="true" pageId="4" pageNumber="61">Deretus necopinatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2CA578ABD496E7EEBA3A6E3FEC0FC31" box="[219,308,944,968]" pageId="4" pageNumber="61" rank="species">
<emphasis id="C9F9EAF1BD496E7EEBA3A6E3FEC0FC31" bold="true" box="[219,308,944,968]" pageId="4" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="FB3236E3BD496E7EEBBDA680FAE6FBA1" blockId="4.[151,1437,151,1472]" pageId="4" pageNumber="61">
<emphasis id="C9F9EAF1BD496E7EEBBDA680FE3FFC15" bold="true" box="[197,459,979,1004]" pageId="4" pageNumber="61">Differential diagnosis.</emphasis>
The
<typeStatus id="24368841BD496E7EE97DA686FDC1FC14" box="[517,565,981,1005]" pageId="4" pageNumber="61">type</typeStatus>
specimen was redescribed by
<bibRefCitation id="9F1C4B12BD496E7EE8F4A687FBD2FC15" author="Gahan" box="[908,1062,980,1004]" pageId="4" pageNumber="61" refString="Gahan, C. J. (1903) Insecta: Coleoptera. In: Forbes, H. O. (Ed.), The natural history of Socotra and Abd-El-Kuri. Henry Young &amp; Sons, London, pp. 262 - 292." type="book chapter" year="1903">Gahan (1903)</bibRefCitation>
and figured by
<bibRefCitation id="9F1C4B12BD496E7EEFA0A687FA9EFC15" author="Koch" box="[1240,1386,980,1004]" pageId="4" pageNumber="61" refString="Koch, C. (1970) Die Tenebrioniden (Coleoptera) des Archipels von Socotra. Italian Journal of Zoology, 4, 69 - 132." type="journal article" year="1970">Koch (1970)</bibRefCitation>
. For this reason, and as the male and female do not differ significantly from each other in the shape of habitus (in males the apical antennomere is somewhat longer than in females), the description is not repeated here. Size of
<typeStatus id="24368841BD496E7EEE4DA14FFA68FBCD" box="[1333,1436,1052,1076]" pageId="4" pageNumber="61" type="lectotype">lectotype</typeStatus>
11.0 x 4.0 mm. Variability of size in females—5.711.0 x
<quantity id="3C759B06BD496E7EE859A113FC5FFBA1" box="[801,939,1088,1112]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="4.1" metricValueMin="2.1" pageId="4" pageNumber="61" unit="mm" value="3.0999999999999996" valueMax="4.1" valueMin="2.1">2.14.1 mm</quantity>
; males—4.79.5 x
<quantity id="3C759B06BD496E7EEFFCA113FAFAFBA1" box="[1156,1294,1088,1112]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.8" metricValueMin="1.8" pageId="4" pageNumber="61" unit="mm" value="2.8" valueMax="3.8" valueMin="1.8">1.83.8 mm</quantity>
.
</paragraph>
<paragraph id="FB3236E3BD496E7EEBBDA137FF0CFA39" blockId="4.[151,1437,151,1472]" pageId="4" pageNumber="61">
This species, in its general appearance, most resembles
<taxonomicName id="3C8D4D60BD496E7EE840A136FC16FB85" box="[824,994,1125,1148]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="necopinatus" status="sp. nov.">
<emphasis id="C9F9EAF1BD496E7EE840A136FC16FB85" box="[824,994,1125,1148]" italics="true" pageId="4" pageNumber="61">D. necopinatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2CA578ABD496E7EE891A137FBB7FB85" box="[1001,1091,1124,1148]" pageId="4" pageNumber="61" rank="species">
<emphasis id="C9F9EAF1BD496E7EE891A137FBB7FB85" bold="true" box="[1001,1091,1124,1148]" pageId="4" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
from which it is differentiated by the shiny integument, the broadly rounded pronotal anterior corners (
<figureCitation id="63B62A66BD496E7EE8B7A1DBFBD6FB59" box="[975,1058,1160,1184]" captionStart="FIGURES 13 18" captionStartId="9.[151,264,1753,1776]" captionText="FIGURES 13 18. Pronotum, dorsal view. 13. Deretus bezdeki sp. nov. (paratype), male; 14. D. denticollis Gahan, 1900, male; 15. D. hulai sp. nov. (paratype), female; 16. D. maderai sp. nov. (holotype), male; 17. Deretus necopinatus sp. nov. (paratype), male; 18. D. spinicollis Schawaller, 2004, male." pageId="4" pageNumber="61">Fig. 14</figureCitation>
) and by the absence of patchs of sensillae on the abdominal ventrites together with the unmodified metatibia in males. From
<taxonomicName id="3C8D4D60BD496E7EEFC8A1FEFAB3FB3D" box="[1200,1351,1196,1220]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="spinicollis">
<emphasis id="C9F9EAF1BD496E7EEFC8A1FEFAB3FB3D" box="[1200,1351,1196,1220]" italics="true" pageId="4" pageNumber="61">D. spinicollis</emphasis>
</taxonomicName>
and
<taxonomicName id="3C8D4D60BD496E7EEEF9A1FEFF1FFB11" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="bezdeki" status="sp. nov.">
<emphasis id="C9F9EAF1BD496E7EEEF9A1FEFF1FFB11" italics="true" pageId="4" pageNumber="61">D. bezdeki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2CA578ABD496E7EEB8CA183FEA4FB11" box="[244,336,1232,1256]" pageId="4" pageNumber="61" rank="species">
<emphasis id="C9F9EAF1BD496E7EEB8CA183FEA4FB11" bold="true" box="[244,336,1232,1256]" pageId="4" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
it can be distinguished mainly by the shape of the pronotum, which in the latter two species is characterized by extremely elongated (extended) anterior corners (
<figureCitation id="63B62A66BD496E7EE8FAA1A7FBF4FAF5" box="[898,1024,1268,1292]" captionStart="FIGURES 13 18" captionStartId="9.[151,264,1753,1776]" captionText="FIGURES 13 18. Pronotum, dorsal view. 13. Deretus bezdeki sp. nov. (paratype), male; 14. D. denticollis Gahan, 1900, male; 15. D. hulai sp. nov. (paratype), female; 16. D. maderai sp. nov. (holotype), male; 17. Deretus necopinatus sp. nov. (paratype), male; 18. D. spinicollis Schawaller, 2004, male." pageId="4" pageNumber="61">Fig. 13, 18</figureCitation>
). It differs from
<taxonomicName id="3C8D4D60BD496E7EEFC3A1A6FAE3FAF5" box="[1211,1303,1268,1292]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="hulai" status="sp. nov.">
<emphasis id="C9F9EAF1BD496E7EEFC3A1A6FAE3FAF5" box="[1211,1303,1268,1292]" italics="true" pageId="4" pageNumber="61">D. hulai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2CA578ABD496E7EEE67A1A7FA8DFAF5" box="[1311,1401,1268,1292]" pageId="4" pageNumber="61" rank="species">
<emphasis id="C9F9EAF1BD496E7EEE67A1A7FA8DFAF5" bold="true" box="[1311,1401,1268,1292]" pageId="4" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
by its smooth, almost glabrous (only ultra-microscopically and very sparsely pubescent), elytra (conspicuously pubescent and sculptured with small tubercles in the latter species). From
<taxonomicName id="3C8D4D60BD496E7EE8E2A06EFBEEFAAD" box="[922,1050,1340,1364]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="maderai" status="sp. nov.">
<emphasis id="C9F9EAF1BD496E7EE8E2A06EFBEEFAAD" box="[922,1050,1340,1364]" italics="true" pageId="4" pageNumber="61">D. maderai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2CA578ABD496E7EEF5AA06FFB89FAAD" box="[1058,1149,1340,1364]" pageId="4" pageNumber="61" rank="species">
<emphasis id="C9F9EAF1BD496E7EEF5AA06FFB89FAAD" bold="true" box="[1058,1149,1340,1364]" pageId="4" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
it can be separated by its almost flat pronotum (lateral view) and by its elytra length / pronotum width ratio, that is larger than 2.0. In
<taxonomicName id="3C8D4D60BD496E7EEEF9A032FF00FA65" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="maderai" status="sp. nov.">
<emphasis id="C9F9EAF1BD496E7EEEF9A032FF00FA65" italics="true" pageId="4" pageNumber="61">D. maderai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2CA578ABD496E7EEB84A0D7FEA2FA65" box="[252,342,1412,1436]" pageId="4" pageNumber="61" rank="species">
<emphasis id="C9F9EAF1BD496E7EEB84A0D7FEA2FA65" bold="true" box="[252,342,1412,1436]" pageId="4" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
the pronotum is distinctly convex in lateral view and elytra length / pronotum width ratio is less than 1.8.
</paragraph>
</subSubSection>
</treatment>
</document>