269 lines
30 KiB
XML
269 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.81737" ID-GBIF-Dataset="d148b8fa-4f4c-4069-80e5-f04f98561bfe" ID-Pensoft-Pub="2747-8106-96-51" ID-Pensoft-UUID="FCEF2A912BC258BBB022681E86486B06" ID-ZooBank="CD8C70BA025345408EFCD430A12F6B35" ModsDocID="2747-8106-96-51" checkinTime="1649795137967" checkinUser="pensoft" docAuthor="Schwarzhans, Werner & Keupp, Helmut" docDate="2022" docId="6CEBFA1EA2EA5A989BADEB52261BA48A" docLanguage="en" docName="Zitteliana 96: 51-67" docOrigin="Zitteliana 96" docPubDate="2022-04-12" docSource="http://dx.doi.org/10.3897/zitteliana.96.81737" docTitle="Leptolepis curvisulcatus" docType="treatment" docVersion="2" id="FCEF2A912BC258BBB022681E86486B06" lastPageNumber="51" masterDocId="FCEF2A912BC258BBB022681E86486B06" masterDocTitle="Early teleost otolith morphogenesis observed in the Jurassic of Franconia, Bavaria, southern Germany" masterLastPageNumber="67" masterPageNumber="51" pageId="0" pageNumber="51" updateTime="1649795760618" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Early teleost otolith morphogenesis observed in the Jurassic of Franconia, Bavaria, southern Germany</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schwarzhans, Werner</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4842-7989</mods:nameIdentifier>
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<mods:affiliation>Zoological Museum, Natural History Museum of Denmark, Universitetsparken 15, 2100 Kobenhavn, Denmark & Ahrensburger Weg 103, 22359 Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">wwschwarz@aol.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Keupp, Helmut</mods:namePart>
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<mods:affiliation>Freie Universitaet Berlin, Institut fuer Geologische Wissenschaften, Fachrichtung Palaeontologie, Malteser Strasse 74 - 100, Haus D, 12249 Berlin, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zitteliana</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-04-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>96</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>51</mods:start>
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<mods:end>67</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zitteliana.96.81737</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zitteliana.96.81737</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2747-8106-96-51</mods:identifier>
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<mods:identifier type="ZooBank">CD8C70BA025345408EFCD430A12F6B35</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FCEF2A912BC258BBB022681E86486B06</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:6CEBFA1EA2EA5A989BADEB52261BA48A" httpUri="http://treatment.plazi.org/id/6CEBFA1EA2EA5A989BADEB52261BA48A" lastPageNumber="51" pageId="0" pageNumber="51">
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<subSubSection pageId="0" pageNumber="51" type="nomenclature">
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName LSID="6CEBFA1E-A2EA-5A98-9BAD-EB52261BA48A" authority="(Schroeder, 1956)" baseAuthorityName="Schroeder" baseAuthorityYear="1956" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Leptolepis curvisulcatus" order="Leptolepidiformes" pageId="0" pageNumber="51" rank="species" species="curvisulcatus">
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Leptolepis curvisulcatus (
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<normalizedToken originalValue="Schröder">Schroeder</normalizedToken>
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, 1956)
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="description">
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<paragraph pageId="0" pageNumber="51">
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<figureCitation captionStart="Plate 1" captionStartId="F3" captionText="Plate 1. Otoliths of Leptolepis normandica and L. curvisulcatus. figs 1 - 15: Leptolepis normandica Nybelin, 1962; 1 - 11: From late Pliensbachian of Buttenheim, 1 - 8, 11: EPH horizon (SNSB-BSPG 2022 IV 3); 9: " Quellhorizon " (SNSB-BSPG 2022 IV 2); 10: Apyrenum subzone (SNSB-BSPG 2022 IV 1); 12: paratype of Ot. cristatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 5); 13: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 20); 14: paratype of Ot. vastus Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 34); 15: holotype of Ot. vastus from Peulendorf (SNSB-BSPG 2022 III 30). figs 15 - 19 Leptolepis curvisulcatus (Schroeder, 1956); 15: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 17); 16: holotype of Ot. curvisulcatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 42); 17: holotype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 2); 18: paratype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 47); 19: holotype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 16)." figureDoi="10.3897/zitteliana.96.81737.plate1" httpUri="https://binary.pensoft.net/fig/671631" pageId="0" pageNumber="51">Plate 1, figs 15-19</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="reference_group">
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName authority="Koken, 1884" authorityName="Koken" authorityYear="1884" class="Actinopterygii" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otolithus curvisulcatus" order="Leptolepidiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="curvisulcatus">Otolithus curvisulcatus</taxonomicName>
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1956
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<taxonomicName authority="Koken, 1884" authorityName="Koken" authorityYear="1884" class="Actinopterygii" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otolithus curvisulcatus" order="Leptolepidiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="curvisulcatus">Otolithus curvisulcatus</taxonomicName>
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-
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<normalizedToken originalValue="Schröder">Schroeder</normalizedToken>
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: pl. 7, figs 25-27.
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</paragraph>
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName authority="Koken, 1884" authorityName="Koken" authorityYear="1884" class="Actinopterygii" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otolithus opalini" order="Leptolepidiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="opalini">Otolithus opalini</taxonomicName>
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1956
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<taxonomicName authority="Koken, 1884" authorityName="Koken" authorityYear="1884" class="Actinopterygii" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otolithus opalini" order="Leptolepidiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="opalini">Otolithus opalini</taxonomicName>
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-
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<normalizedToken originalValue="Schröder">Schroeder</normalizedToken>
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: pl. 6, figs 7-8 (non fig. 9).
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</paragraph>
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName authority="Koken, 1884" authorityName="Koken" authorityYear="1884" class="Actinopterygii" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otolithus schattenbergi" order="Leptolepidiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="schattenbergi">Otolithus schattenbergi</taxonomicName>
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1956
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<taxonomicName authority="Koken, 1884" authorityName="Koken" authorityYear="1884" class="Actinopterygii" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otolithus schattenbergi" order="Leptolepidiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="schattenbergi">Otolithus schattenbergi</taxonomicName>
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-
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<normalizedToken originalValue="Schröder">Schroeder</normalizedToken>
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: pl. 6, figs 10-12.
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</paragraph>
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<paragraph pageId="0" pageNumber="51">
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<taxonomicName authorityName="Schwarzhans & Keupp" authorityYear="2022" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Leptolepis ornatus" order="Leptolepidiformes" pageId="0" pageNumber="51" rank="species" species="ornatus">Leptolepis ornatus</taxonomicName>
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2018
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<taxonomicName authorityName="Schwarzhans & Keupp" authorityYear="2022" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Leptolepis ornatus" order="Leptolepidiformes" pageId="0" pageNumber="51" rank="species" species="ornatus">Leptolepis ornatus</taxonomicName>
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(Weiler, 1953) - Schwarzhans: fig. 1I-J,?K.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="materials_examined">
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<paragraph pageId="0" pageNumber="51">Material.</paragraph>
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<paragraph pageId="0" pageNumber="51">
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<specimenCount type="generic">15 specimens</specimenCount>
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from
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<normalizedToken originalValue="Schröder’s">Schroeder's</normalizedToken>
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collection from the late Toarcian to early Aalenian:
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<typeStatus>holotype</typeStatus>
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of
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<taxonomicName genus="Ot." lsidName="Ot. curvisulcatus" pageId="0" pageNumber="51" rank="species" species="curvisulcatus">
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<emphasis italics="true" pageId="0" pageNumber="51">Ot. curvisulcatus</emphasis>
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</taxonomicName>
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from Kremmeldorf (SNSB-BSPG
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<date value="2022-03">2022 III</date>
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42, Plate
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<figureCitation captionStart="Plate 1" captionStartId="F3" captionText="Plate 1. Otoliths of Leptolepis normandica and L. curvisulcatus. figs 1 - 15: Leptolepis normandica Nybelin, 1962; 1 - 11: From late Pliensbachian of Buttenheim, 1 - 8, 11: EPH horizon (SNSB-BSPG 2022 IV 3); 9: " Quellhorizon " (SNSB-BSPG 2022 IV 2); 10: Apyrenum subzone (SNSB-BSPG 2022 IV 1); 12: paratype of Ot. cristatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 5); 13: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 20); 14: paratype of Ot. vastus Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 34); 15: holotype of Ot. vastus from Peulendorf (SNSB-BSPG 2022 III 30). figs 15 - 19 Leptolepis curvisulcatus (Schroeder, 1956); 15: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 17); 16: holotype of Ot. curvisulcatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 42); 17: holotype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 2); 18: paratype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 47); 19: holotype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 16)." figureDoi="10.3897/zitteliana.96.81737.plate1" httpUri="https://binary.pensoft.net/fig/671631" pageId="0" pageNumber="51">1</figureCitation>
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, fig. 16) and
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<specimenCount type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
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of
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<taxonomicName genus="Ot." lsidName="Ot. curvisulcatus" pageId="0" pageNumber="51" rank="species" species="curvisulcatus">
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<emphasis italics="true" pageId="0" pageNumber="51">Ot. curvisulcatus</emphasis>
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</taxonomicName>
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, thereof 2 from Kremmeldorf and 1 from Freiahorn (SNSB-BSPG
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<date value="2022-03">2022 III</date>
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43-45),
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<typeStatus>holotype</typeStatus>
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of
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<taxonomicName genus="Ot." lsidName="Ot. opalini" pageId="0" pageNumber="51" rank="species" species="opalini">
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<emphasis italics="true" pageId="0" pageNumber="51">Ot. opalini</emphasis>
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</taxonomicName>
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from Kremmeldorf (SNSB-BSPG
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<date value="2022-03-16">2022 III 16</date>
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, Plate
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<figureCitation captionStart="Plate 1" captionStartId="F3" captionText="Plate 1. Otoliths of Leptolepis normandica and L. curvisulcatus. figs 1 - 15: Leptolepis normandica Nybelin, 1962; 1 - 11: From late Pliensbachian of Buttenheim, 1 - 8, 11: EPH horizon (SNSB-BSPG 2022 IV 3); 9: " Quellhorizon " (SNSB-BSPG 2022 IV 2); 10: Apyrenum subzone (SNSB-BSPG 2022 IV 1); 12: paratype of Ot. cristatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 5); 13: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 20); 14: paratype of Ot. vastus Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 34); 15: holotype of Ot. vastus from Peulendorf (SNSB-BSPG 2022 III 30). figs 15 - 19 Leptolepis curvisulcatus (Schroeder, 1956); 15: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 17); 16: holotype of Ot. curvisulcatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 42); 17: holotype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 2); 18: paratype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 47); 19: holotype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 16)." figureDoi="10.3897/zitteliana.96.81737.plate1" httpUri="https://binary.pensoft.net/fig/671631" pageId="0" pageNumber="51">1</figureCitation>
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, fig. 19) and
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<specimenCount type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
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of
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<taxonomicName genus="Ot." lsidName="Ot. opalini" pageId="0" pageNumber="51" rank="species" species="opalini">
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<emphasis italics="true" pageId="0" pageNumber="51">Ot. opalini</emphasis>
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</taxonomicName>
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from Kremmeldorf (SNSB-BSPG
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<date value="2022-03-17" valueMax="2022-03-19" valueMin="2022-03-17">2022 III 17-19</date>
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, Plate
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<figureCitation captionStart="Plate 1" captionStartId="F3" captionText="Plate 1. Otoliths of Leptolepis normandica and L. curvisulcatus. figs 1 - 15: Leptolepis normandica Nybelin, 1962; 1 - 11: From late Pliensbachian of Buttenheim, 1 - 8, 11: EPH horizon (SNSB-BSPG 2022 IV 3); 9: " Quellhorizon " (SNSB-BSPG 2022 IV 2); 10: Apyrenum subzone (SNSB-BSPG 2022 IV 1); 12: paratype of Ot. cristatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 5); 13: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 20); 14: paratype of Ot. vastus Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 34); 15: holotype of Ot. vastus from Peulendorf (SNSB-BSPG 2022 III 30). figs 15 - 19 Leptolepis curvisulcatus (Schroeder, 1956); 15: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 17); 16: holotype of Ot. curvisulcatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 42); 17: holotype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 2); 18: paratype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 47); 19: holotype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 16)." figureDoi="10.3897/zitteliana.96.81737.plate1" httpUri="https://binary.pensoft.net/fig/671631" pageId="0" pageNumber="51">1</figureCitation>
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, fig. 15),
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<typeStatus>holotype</typeStatus>
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of
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<taxonomicName genus="Ot." lsidName="Ot. schattenbergi" pageId="0" pageNumber="51" rank="species" species="schattenbergi">
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<emphasis italics="true" pageId="0" pageNumber="51">Ot. schattenbergi</emphasis>
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</taxonomicName>
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from Kremmelsdorf (SNSB-BSPG
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<date value="2022-03-02">2022 III 2</date>
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, Plate
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<figureCitation captionStart="Plate 1" captionStartId="F3" captionText="Plate 1. Otoliths of Leptolepis normandica and L. curvisulcatus. figs 1 - 15: Leptolepis normandica Nybelin, 1962; 1 - 11: From late Pliensbachian of Buttenheim, 1 - 8, 11: EPH horizon (SNSB-BSPG 2022 IV 3); 9: " Quellhorizon " (SNSB-BSPG 2022 IV 2); 10: Apyrenum subzone (SNSB-BSPG 2022 IV 1); 12: paratype of Ot. cristatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 5); 13: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 20); 14: paratype of Ot. vastus Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 34); 15: holotype of Ot. vastus from Peulendorf (SNSB-BSPG 2022 III 30). figs 15 - 19 Leptolepis curvisulcatus (Schroeder, 1956); 15: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 17); 16: holotype of Ot. curvisulcatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 42); 17: holotype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 2); 18: paratype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 47); 19: holotype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 16)." figureDoi="10.3897/zitteliana.96.81737.plate1" httpUri="https://binary.pensoft.net/fig/671631" pageId="0" pageNumber="51">1</figureCitation>
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, fig. 17),
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<specimenCount type="generic" typeStatus="paratypes">6 paratypes</specimenCount>
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of
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<taxonomicName genus="Ot." lsidName="Ot. schattenbergi" pageId="0" pageNumber="51" rank="species" species="schattenbergi">
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<emphasis italics="true" pageId="0" pageNumber="51">Ot. schattenbergi</emphasis>
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</taxonomicName>
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with 5 from Kremmelsdorf (SNSB-BSPG
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<date value="2022-03-03" valueMax="2022-03-05" valueMin="2022-03-03">2022 III 3-5</date>
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, 47-48, Plate
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<figureCitation captionStart="Plate 1" captionStartId="F3" captionText="Plate 1. Otoliths of Leptolepis normandica and L. curvisulcatus. figs 1 - 15: Leptolepis normandica Nybelin, 1962; 1 - 11: From late Pliensbachian of Buttenheim, 1 - 8, 11: EPH horizon (SNSB-BSPG 2022 IV 3); 9: " Quellhorizon " (SNSB-BSPG 2022 IV 2); 10: Apyrenum subzone (SNSB-BSPG 2022 IV 1); 12: paratype of Ot. cristatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 5); 13: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 20); 14: paratype of Ot. vastus Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 34); 15: holotype of Ot. vastus from Peulendorf (SNSB-BSPG 2022 III 30). figs 15 - 19 Leptolepis curvisulcatus (Schroeder, 1956); 15: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 17); 16: holotype of Ot. curvisulcatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 42); 17: holotype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 2); 18: paratype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 47); 19: holotype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 16)." figureDoi="10.3897/zitteliana.96.81737.plate1" httpUri="https://binary.pensoft.net/fig/671631" pageId="0" pageNumber="51">1</figureCitation>
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, fig. 18) and 1 from Peulendorf (SNSB-BSPG
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<date value="2022-03">2022 III</date>
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46).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="51" type="differential diagnosis">
|
||
<paragraph pageId="0" pageNumber="51">Differential diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="51">
|
||
<taxonomicName baseAuthorityName="Schroeder" baseAuthorityYear="1956" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" kingdom="Animalia" lsidName="Leptolepis curvisulcatus" order="Elopiformes" pageId="0" pageNumber="51" rank="species" species="curvisulcatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">Leptolepis curvisulcatus</emphasis>
|
||
</taxonomicName>
|
||
is slightly more elongate than
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. normandica" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="normandica">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. normandica</emphasis>
|
||
</taxonomicName>
|
||
with an OL:OH ratio of 1.4-1.6 (vs. 1.25-1.45), which is mainly caused by a relatively low postdorsal rim, which is also more rounded. Other proportions and characteristics are shared with
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. normandica" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="normandica">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. normandica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName baseAuthorityName="Schroeder" baseAuthorityYear="1956" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" kingdom="Animalia" lsidName="Leptolepis curvisulcatus" order="Elopiformes" pageId="0" pageNumber="51" rank="species" species="curvisulcatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">Leptolepis curvisulcatus</emphasis>
|
||
</taxonomicName>
|
||
also differs from
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. normandica" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="normandica">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. normandica</emphasis>
|
||
</taxonomicName>
|
||
in its longer ostium as compared to the length of its cauda, but because of the gradual transition from ostium to cauda, this feature cannot be measured unequivocally. The rounded and short postdorsal rim distinguishes
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. curvisulcatus" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="curvisulcatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. curvisulcatus</emphasis>
|
||
</taxonomicName>
|
||
from the coeval
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. kremmeldorfensis" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="kremmeldorfensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. kremmeldorfensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="51" type="discussion">
|
||
<paragraph pageId="0" pageNumber="51">Discussion.</paragraph>
|
||
<paragraph pageId="0" pageNumber="51">
|
||
The distinctions among the three nominal late Toarcian to Aalenian species
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. normandica" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="normandica">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. normandica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. curvisulcatus" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="curvisulcatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. curvisulcatus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. kremmeldorfensis" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="kremmeldorfensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. kremmeldorfensis</emphasis>
|
||
</taxonomicName>
|
||
are all very subtle and it is possible that with more material becoming available from this time interval their nature and validity may have to be revised again. This is also because most specimens so far known from
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. curvisulcatus" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="curvisulcatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. curvisulcatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. kremmeldorfensis" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="kremmeldorfensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. kremmeldorfensis</emphasis>
|
||
</taxonomicName>
|
||
are smaller than 1.5 mm in length and, therefore, cannot be considered morphologically mature.
|
||
<bibRefCitation author="Schroeder, G" journalOrPublisher="Geologische Blaetter fuer Nordost-Bayern und angrenzende Gebiete" pageId="0" pageNumber="51" pagination="128 - 153" refId="B29" refString="Schroeder, G, 1956. Otolithen aus dem Lias und Dogger des Fraenkischen Jura. Geologische Blaetter fuer Nordost-Bayern und angrenzende Gebiete 6: 128 - 153" title="Otolithen aus dem Lias und Dogger des Fraenkischen Jura." volume="6" year="1956">
|
||
<normalizedToken originalValue="Schröder">Schroeder</normalizedToken>
|
||
(1956)
|
||
</bibRefCitation>
|
||
mentioned that the slightly curved shape of the sulcus (cauda) is a distinctive character of the species (hence the name curvisulcatus). However, this character does not seem to be reliable for its distinction from otoliths of
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. normandica" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="normandica">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. normandica</emphasis>
|
||
</taxonomicName>
|
||
, and we therefore consider it to reflect a degree of variability.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="51">
|
||
The best-preserved and largest specimen that we attribute to
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. curvisulcatus" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="curvisulcatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. curvisulcatus</emphasis>
|
||
</taxonomicName>
|
||
is a paratype of
|
||
<normalizedToken originalValue="Schröder’s">Schroeder's</normalizedToken>
|
||
<taxonomicName authority="Koken, 1884" authorityName="Koken" authorityYear="1884" class="Actinopterygii" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otolithus opalini" order="Elopiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="opalini">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">Otolithus opalini</emphasis>
|
||
</taxonomicName>
|
||
of 2.55 mm in length (Plate
|
||
<figureCitation captionStart="Plate 1" captionStartId="F3" captionText="Plate 1. Otoliths of Leptolepis normandica and L. curvisulcatus. figs 1 - 15: Leptolepis normandica Nybelin, 1962; 1 - 11: From late Pliensbachian of Buttenheim, 1 - 8, 11: EPH horizon (SNSB-BSPG 2022 IV 3); 9: " Quellhorizon " (SNSB-BSPG 2022 IV 2); 10: Apyrenum subzone (SNSB-BSPG 2022 IV 1); 12: paratype of Ot. cristatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 5); 13: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 20); 14: paratype of Ot. vastus Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 34); 15: holotype of Ot. vastus from Peulendorf (SNSB-BSPG 2022 III 30). figs 15 - 19 Leptolepis curvisulcatus (Schroeder, 1956); 15: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 17); 16: holotype of Ot. curvisulcatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 42); 17: holotype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 2); 18: paratype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 47); 19: holotype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 16)." figureDoi="10.3897/zitteliana.96.81737.plate1" httpUri="https://binary.pensoft.net/fig/671631" pageId="0" pageNumber="51">1</figureCitation>
|
||
, fig. 16), while the specimen selected as holotype represents a small and poorly preserved otolith of 1.25 mm in length (Plate
|
||
<figureCitation captionStart="Plate 1" captionStartId="F3" captionText="Plate 1. Otoliths of Leptolepis normandica and L. curvisulcatus. figs 1 - 15: Leptolepis normandica Nybelin, 1962; 1 - 11: From late Pliensbachian of Buttenheim, 1 - 8, 11: EPH horizon (SNSB-BSPG 2022 IV 3); 9: " Quellhorizon " (SNSB-BSPG 2022 IV 2); 10: Apyrenum subzone (SNSB-BSPG 2022 IV 1); 12: paratype of Ot. cristatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 5); 13: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 20); 14: paratype of Ot. vastus Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 34); 15: holotype of Ot. vastus from Peulendorf (SNSB-BSPG 2022 III 30). figs 15 - 19 Leptolepis curvisulcatus (Schroeder, 1956); 15: paratype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 17); 16: holotype of Ot. curvisulcatus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 42); 17: holotype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 2); 18: paratype of Ot. schattenbergi Schroeder, 1956 from Kremmelsdorf (SNSB-BSPG 2022 III 47); 19: holotype of Ot. opalini Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 16)." figureDoi="10.3897/zitteliana.96.81737.plate1" httpUri="https://binary.pensoft.net/fig/671631" pageId="0" pageNumber="51">1</figureCitation>
|
||
, fig. 19). The holotype of
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. curvisulcatus" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="curvisulcatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. curvisulcatus</emphasis>
|
||
</taxonomicName>
|
||
is the next best-preserved specimen of 2.25 mm in length, and we have thus chosen this name to represent this nominal species.
|
||
<bibRefCitation DOI="https://doi.org/10.1127/njgpa/2018/0707" author="Schwarzhans, W" journalOrPublisher="Neues Jahrbuch Geologie und Palaeontologie Abhandlungen" pageId="0" pageNumber="51" pagination="75 - 121" refId="B32" refString="Schwarzhans, W, 2018. A review of Jurassic and Early Cretaceous otoliths and the development of early morphological diversity in otoliths. Neues Jahrbuch Geologie und Palaeontologie Abhandlungen 287: 75 - 121, DOI: https://doi.org/10.1127/njgpa/2018/0707" title="A review of Jurassic and Early Cretaceous otoliths and the development of early morphological diversity in otoliths." url="https://doi.org/10.1127/njgpa/2018/0707" volume="287" year="2018">Schwarzhans (2018)</bibRefCitation>
|
||
could not review
|
||
<normalizedToken originalValue="Schröder’s">Schroeder's</normalizedToken>
|
||
specimens and attributed similar specimens from the same stratigraphic interval to
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. ornatus" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="ornatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. ornatus</emphasis>
|
||
</taxonomicName>
|
||
(Weiler, 1953). Unfortunately,
|
||
<normalizedToken originalValue="Weiler’s">Weiler's</normalizedToken>
|
||
type-material has not been located and is not available for revision. His documentation is rather indistinct, so we therefore propose no longer using this name and instead refer to
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. curvisulcatus" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="curvisulcatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. curvisulcatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |