620 lines
89 KiB
XML
620 lines
89 KiB
XML
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<mods:title id="599189078F05C5224E3277744163C405">The arboreal snail genus Amphidromus Albers, 1850 (Eupulmonata, Camaenidae) of Southeast Asia: 1. Molecular systematics of some Vietnamese species and related species from Cambodia, Indonesia, and Laos</mods:title>
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<mods:namePart id="04C71F3D0C0B2CF3861E4C05484CABEE">Jirapatrasilp, Parin</mods:namePart>
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<mods:affiliation id="B05EE5FAF9D4BE732DE26AE45CD621D1">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand & Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, Hamburg, Germany</mods:affiliation>
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<mods:namePart id="1FE1821EFABBDA537874E92FD56C6848">Huang, Chih-Wei</mods:namePart>
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<mods:affiliation id="B517C6CE70D0759B472F2C01D0D23B7C">School of Life Science, National Taiwan Normal University, Taipei, Taiwan</mods:affiliation>
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<mods:namePart id="A5ECB5739097F92A45115910314DF97A">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="DC325D74A9645C96FF9ED68D99EBE0A6">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand</mods:affiliation>
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<mods:affiliation id="D6ABA0BB5B86E6F26DB6E5D42362069C">Department of Life Sciences, National Chung Hsing University, Taichung, Taiwan</mods:affiliation>
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<treatment id="6083D4A531595FA79F2C8B8BD37A108B" LSID="urn:lsid:plazi:treatment:6083D4A531595FA79F2C8B8BD37A108B" httpUri="http://treatment.plazi.org/id/6083D4A531595FA79F2C8B8BD37A108B" lastPageNumber="15" pageId="0" pageNumber="15">
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<taxonomicName id="910079C40D2F9132D9073CE543615F92" ID-CoL="87KP2" LSID="6083D4A5-3159-5FA7-9F2C-8B8BD37A108B" authority="Fruhstorfer, 1905" authorityName="Fruhstorfer" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus buelowi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">Amphidromus buelowi Fruhstorfer, 1905</taxonomicName>
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<subSubSection id="20EEF1FE9C55A2DFAF2EDE74949D2F13" pageId="0" pageNumber="15" type="description">
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<figureCitation id="E02FDDC75BEDBFE81F39158738ADEE47" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Living Amphidromus spp A-C Amphidromus ingens Moellendorff, 1900 from Dak Lak, Vietnam D Amphidromus placostylus Moellendorff, 1900 from Hoai An, An Lao, Binh Dinh, Vietnam E, F Amphidromus ingensoides sp. nov. from Hon Ba, Khanh Son, Khanh Hoa, Vietnam G Amphidromus asperoides sp. nov. from Ea Tu, Buon Ma Thuat city, Dak Lak, Vietnam H, I Amphidromus buelowi Fruhstorfer, 1905 from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam J Amphidromus thachi Huber, 2015 from Krong Bong, Dak Lak, Vietnam." figureDoi="10.3897/zookeys.1196.112146.figure6" httpUri="https://binary.pensoft.net/fig/1007824" pageId="0" pageNumber="15">Figs 6H, I</figureCitation>
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<figureCitation id="F478F51F0EAC58D37BE1B26047F10394" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. SEM images of the radula A Amphidromus placostylus Moellendorff, 1900 from Dak Po, Gia Lai, Vietnam (NMNS- 8764 - 217) B Amphidromus buelowi Fruhstorfer, 1905 from Mount Singgalang, Sepuluh Koto, Tanah Datar Regency, West Sumatra, Indonesia (NMNS- 8764 - 024) C Amphidromus thachi Huber, 2015 from Buon Don, Dak Lak, Vietnam (NMNS- 8764 - 271). Central teeth are marked in yellow. The left and right images show the outer and inner sections of each radula, respectively." figureDoi="10.3897/zookeys.1196.112146.figure12" httpUri="https://binary.pensoft.net/fig/1007830" pageId="0" pageNumber="15">, 12B</figureCitation>
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<figureCitation id="AB1FD1D9BA50FA3B6CBD9E9C99A0B512" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Genitalia of Amphidromus spp A-C Amphidromus ingensoides sp. nov. from Hon Ba, Khanh Son, Khanh Hoa, Vietnam (NMNS- 8764 - 107), showing A general view of genitalia B interior structures of penis C interior structures of vagina chamber D-F Amphidromus buelowi Fruhstorfer, 1905 from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 031), showing D general view of genitalia E interior structures of penis F interior structures of vagina chamber. Red dots indicate the shape of the missing gametolytic sac. Green arrows indicate the genital openings. Abbreviations: ap, appendix; e, epiphallus; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; ov, oviduct; p, penis; pp, penial pilaster; pr, penial retractor muscle; pv, penial verge; v, vagina; vd, vas deferens; vp, vaginal pilaster" figureDoi="10.3897/zookeys.1196.112146.figure14" httpUri="https://binary.pensoft.net/fig/1007832" pageId="0" pageNumber="15">, 14D-F</figureCitation>
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<figureCitation id="F8FC6887BB32BBC06E402A642AFE79BA" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Shells of Amphidromus buelowi Fruhstorfer, 1905 A lectotype of " Amphidromus buelowi " (NHMUK 1910.12.30.98) B holotype of " Amphidromus asper " (SMF 7762) C holotype of " Amphidromus franzhuberi " (MNHN-IM- 2000 - 31892) D specimen from Padang Soekeli, Singalang, Indonesia (RBINS I. G. 10591) E specimen from Mount Singgalang, Sepuluh Koto, Tanah Datar Regency, West Sumatra, Indonesia (NMNS- 8764 - 025) F specimen from Lang-Biang, Annam, Vietnam (RBINS I. G. 10591) G specimen from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam (NMNS- 8764 - 027) H, I specimen from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 030, NMNS- 8764 - 031). Credit: H. Taylor, NHM (A), M. Caballer, MNHN (C), RBINS (D, F)." figureDoi="10.3897/zookeys.1196.112146.figure15" httpUri="https://binary.pensoft.net/fig/1007833" pageId="0" pageNumber="15">, 15</figureCitation>
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<figureCitation id="B50B4CC3E8BEF3FEB15EFF63089DE720" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Mitochondrial haplotype minimum spanning networks of Amphidromus buelowi Fruhstorfer, 1905 A COI and B 16 S rRNA. The size of each circle corresponds to the frequency of that haplotype, also shown as the number in that circle. The bars on the branches indicate the number of mutational steps between haplotypes. Specimen codes correspond to those in Table 1." figureDoi="10.3897/zookeys.1196.112146.figure16" httpUri="https://binary.pensoft.net/fig/1007834" pageId="0" pageNumber="15">, 16</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="A5F0B3AB02DFF5EA4118AF54B5E7BC3B" pageId="0" pageNumber="15" type="reference_group">
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<paragraph id="D771393720B6653EE216437CAF90789F" pageId="0" pageNumber="15">
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<taxonomicName id="7D32985BC347ED06B4C9D19BF399C0C1" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="genus">Amphidromus</taxonomicName>
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<taxonomicName id="357CF2E539369B76A45262869722A35E" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="genus">Amphidromus</taxonomicName>
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(
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<taxonomicName id="A69A12B304B2791DB45CB94264BDF6C5" class="Gastropoda" family="Camaenidae" genus="Goniodromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Goniodromus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="genus">Goniodromus</taxonomicName>
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)
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<normalizedToken id="6E0ADEF8945EE9061189F82D7E96C220" originalValue="bülowi">buelowi</normalizedToken>
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Fruhstorfer, 1905: 83-84, pl. 1, fig. 2. Type locality: West-Sumatra.
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<bibRefCitation id="7067B05A6F5E00C8255278961F6DE661" author="Rolle, H" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="15" pagination="63 - 70" publicationUrl="https://biostor.org/reference/190496" refId="B52" refString="Rolle, H, 1908. Zur Fauna von West-Sumatra. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 40: 63 - 70, https://biostor.org/reference/190496" title="Zur Fauna von West-Sumatra." url="https://biostor.org/reference/190496" volume="40" year="1908">Rolle 1908</bibRefCitation>
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: 67.
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<bibRefCitation id="297AE031D07E31FCDD43EE5B8AACA1E3" DOI="https://doi.org/10.5962/bhl.title.2832" author="Laidlaw, FF" journalOrPublisher="Zoology" pageId="0" pageNumber="15" pagination="507 - 677" refId="B34" refString="Laidlaw, FF, Solem, A, 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana. Zoology 41: 507 - 677, DOI: https://doi.org/10.5962/bhl.title.2832" title="The land snail genus Amphidromus: A synoptic catalogue. Fieldiana." url="https://doi.org/10.5962/bhl.title.2832" volume="41" year="1961">Laidlaw and Solem 1961</bibRefCitation>
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: 587, 606, fig. 37.
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</paragraph>
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<paragraph id="9588C61B56459F2C91786A456C504742" pageId="0" pageNumber="15">
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<taxonomicName id="4B1558862645E032B71C3E6A7E5BE827" authorityName="Fruhstorfer" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus bülowi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="bülowi">
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Amphidromus
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<normalizedToken id="8639EBC28E3055AA3251635201D9ABA8" originalValue="bülowi">buelowi</normalizedToken>
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</taxonomicName>
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<taxonomicName id="3D2D379D286FB157F63C73178ED31672" authorityName="Fruhstorfer" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus bülowi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="bülowi">
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Amphidromus
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<normalizedToken id="3E4F24F78CBF639FCAFE1F0BA1A6C67A" originalValue="bülowi">buelowi</normalizedToken>
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</taxonomicName>
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.
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<bibRefCitation id="1BB6EE4CC330543740187303646A2D9A" author="Dautzenberg, P" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="15" pagination="343 - 471" publicationUrl="https://biostor.org/reference/145632" refId="B9" refString="Dautzenberg, P, Fischer, H, 1906. Liste des mollusques recoltes par M.H. Mansuy en Indo-Chine et au Yunnan et description ďespeces nouvelles. Journal de Conchyliologie 53 (1905): 343 - 471, https://biostor.org/reference/145632" title="Liste des mollusques recoltes par M. H. Mansuy en Indo-Chine et au Yunnan et description ďespeces nouvelles." url="https://biostor.org/reference/145632" volume="53" year="1906">Dautzenberg and Fischer 1906</bibRefCitation>
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: 365-366, pl. 8, figs 10-12.
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<bibRefCitation id="402898E5B118CFF3E61C7684AF5DF621" DOI="https://doi.org/10.14203/treubia.v10i2-3.1808" author="Degner, E" journalOrPublisher="Treubia" pageId="0" pageNumber="15" pagination="319 - 354" refId="B11" refString="Degner, E, 1928. Spolia mentaweisiana: Binnen-Mollusken von den Mentawei-Inseln. Treubia 10: 319 - 354, DOI: https://doi.org/10.14203/treubia.v10i2-3.1808" title="Spolia mentaweisiana: Binnen-Mollusken von den Mentawei-Inseln." url="https://doi.org/10.14203/treubia.v10i2-3.1808" volume="10" year="1928">Degner 1928</bibRefCitation>
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: 360.
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<bibRefCitation id="F8B6B40FF93B46D92F52FC21303036E6" author="Benthem Jutting, WSS van" journalOrPublisher="Beaufortia" pageId="0" pageNumber="15" pagination="41 - 191" publicationUrl="https://repository.naturalis.nl/pub/504682" refId="B6" refString="Benthem Jutting, WSS van, 1959. Catalogue of the non-marine Mollusca of Sumatra and of its satellite islands. Beaufortia 7: 41 - 191, https://repository.naturalis.nl/pub/504682" title="Catalogue of the non-marine Mollusca of Sumatra and of its satellite islands." url="https://repository.naturalis.nl/pub/504682" volume="7" year="1959">Benthem Jutting 1959</bibRefCitation>
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: 165.
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</paragraph>
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<paragraph id="64156ECF8A374A841309F25EAC23B045" pageId="0" pageNumber="15">
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<taxonomicName id="76E45D4BC69E658E15679D6F473B1533" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="genus">Amphidromus</taxonomicName>
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<taxonomicName id="601D4F75C83BC3A4DB5E0056EA6426D2" authorityName="Jirapatrasilp & Huang & Sutcharit & Lee" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus (Goniodromus) asper" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asper" subGenus="Goniodromus">Amphidromus (Goniodromus) asper</taxonomicName>
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Haas, 1934: 96, figs 11, 12. Type locality:
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<normalizedToken id="CF669A9CE26DBDDFF48C85481ACC3B12" originalValue="Süd-Annam">Sued-Annam</normalizedToken>
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, 120 km von der
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<normalizedToken id="4DF01F73ADDDBC8DE0BD788E08173D8B" originalValue="Küste">Kueste</normalizedToken>
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, auf dem Wege zum Plateau von Lang-Bian, zw. 600-1000 m [South Annam, 120 km from the coast, on the way to the plateau of Lang-Bian, between 600-1000 m].
|
||
<bibRefCitation id="1D032734253600B84D02B5EAB9FE0A16" DOI="https://doi.org/10.5962/bhl.title.2832" author="Laidlaw, FF" journalOrPublisher="Zoology" pageId="0" pageNumber="15" pagination="507 - 677" refId="B34" refString="Laidlaw, FF, Solem, A, 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana. Zoology 41: 507 - 677, DOI: https://doi.org/10.5962/bhl.title.2832" title="The land snail genus Amphidromus: A synoptic catalogue. Fieldiana." url="https://doi.org/10.5962/bhl.title.2832" volume="41" year="1961">Laidlaw and Solem 1961</bibRefCitation>
|
||
: 588, 601.
|
||
<bibRefCitation id="6998C42883A8EF76E23BB3CD81A1F41E" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="15" pagination="131 - 140" refId="B83" refString="Zilch, A, 1953. Die Typen und Typoide des Natur-Museums Senckenberg, 10: Mollusca, Pleurodontidae (1). Archiv fuer Molluskenkunde 82: 131 - 140" title="Die Typen und Typoide des Natur-Museums Senckenberg, 10: Mollusca, Pleurodontidae (1)." volume="82" year="1953">Zilch 1953</bibRefCitation>
|
||
: 138, pl. 25, fig. 44. syn. nov.
|
||
</paragraph>
|
||
<paragraph id="3BEB4E247DB424CEB9039D43BDACE0EF" pageId="0" pageNumber="15">
|
||
<taxonomicName id="82C6676A950C3689D0530FF0535B37C3" authorityName="Jirapatrasilp & Huang & Sutcharit & Lee" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus asper" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asper">Amphidromus asper</taxonomicName>
|
||
.
|
||
<bibRefCitation id="5FF39CF0AE2DDE1A2864DAD2D7007678" author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="0" pageNumber="15" pagination="1 - 68" publicationUrl="https://www.biotaxa.org/Ruthenica/article/view/3603" refId="B56" refString="Schileyko, AA, 2011. Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). Ruthenica 21: 1 - 68, https://www.biotaxa.org/Ruthenica/article/view/3603" title="Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." url="https://www.biotaxa.org/Ruthenica/article/view/3603" volume="21" year="2011">Schileyko 2011</bibRefCitation>
|
||
: 49.
|
||
<bibRefCitation id="E1E8555AF873973232EDCE02FF55ACDA" DOI="https://doi.org/10.12657/folmal.028.002" author="Pall-Gergely, B" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="15" pagination="35 - 76" refId="B45" refString="Pall-Gergely, B, Hunyadi, A, Auffenberg, K, 2020. Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014-2019). Folia Malacologica 28 (1): 35 - 76, DOI: https://doi.org/10.12657/folmal.028.002" title="Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014 - 2019)." url="https://doi.org/10.12657/folmal.028.002" volume="28" year="2020">
|
||
<normalizedToken id="E06D135E2D31543FFAD7D7BB70F21281" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
||
et al. 2020
|
||
</bibRefCitation>
|
||
: 49, 51, fig. 15.
|
||
<bibRefCitation id="31C06E44A791D5FCA27480AED0BC12E2" author="Thach, NN" journalOrPublisher="48 HRBooks Company, Akron, Ohio, USA" pageId="0" pageNumber="15" refId="B75" refString="Thach, NN, 2020a. New shells of South Asia. Vol. 2. Seashells*Freshwater*Land snails. With one New Genus and 140 New Species & Subspecies, Reply to comments made in error. 48HRBooks Company, Akron, Ohio, USA" title="New shells of South Asia. Vol. 2. Seashells * Freshwater * Land snails. With one New Genus and 140 New Species & Subspecies, Reply to comments made in error." year="2020 a">Thach 2020a</bibRefCitation>
|
||
: pl. 76, fig. 893 right.
|
||
</paragraph>
|
||
<paragraph id="FFE7DB514954AF52E9689E1100AC201F" pageId="0" pageNumber="15">
|
||
<taxonomicName id="B50D52420949527A14741366F24961FC" authorityName="Fruhstorfer" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus bulowi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="bulowi">Amphidromus bulowi</taxonomicName>
|
||
[sic].
|
||
<bibRefCitation id="E97F996440BE53435D81355532989320" author="Huber, F" journalOrPublisher="Gloria Maris" pageId="0" pageNumber="15" pagination="29 - 31" refId="B23" refString="Huber, F, 2015. Amphidromus thachi, a new species (Gastropoda: Camaenidae) from Vietnam. Gloria Maris 54: 29 - 31" title="Amphidromus thachi, a new species (Gastropoda: Camaenidae) from Vietnam." volume="54" year="2015">Huber 2015</bibRefCitation>
|
||
: figs 9, 10.
|
||
<bibRefCitation id="CD3A25EF469836A0C5CB97A058A0F801" DOI="https://doi.org/10.3897/zookeys.492.8641" author="Sutcharit, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="15" pagination="49 - 105" refId="B68" refString="Sutcharit, C, Ablett, J, Tongkerd, P, Naggs, F, Panha, S, 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105, DOI: https://doi.org/10.3897/zookeys.492.8641" title="Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species." url="https://doi.org/10.3897/zookeys.492.8641" volume="492" year="2015">Sutcharit et al. 2015</bibRefCitation>
|
||
: 61, fig. 4e.
|
||
</paragraph>
|
||
<paragraph id="49010B402C592FF757BE7D1966AF0F3B" pageId="0" pageNumber="15">
|
||
<taxonomicName id="60E3A399A3F191EF5CE6DCD8420D1225" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="genus">Amphidromus</taxonomicName>
|
||
<taxonomicName id="C965FF939E413A05C46EAE51112FF0A5" authorityName="Fruhstorfer" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus (Goniodromus) bulowi subsp. bulowi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="subSpecies" species="bulowi" subGenus="Goniodromus" subSpecies="bulowi">Amphidromus (Goniodromus) bulowi bulowi</taxonomicName>
|
||
[sic].
|
||
<bibRefCitation id="66D22925CF54BA775A4AB9F76DBE2588" DOI="https://doi.org/10.54173/F484235" author="Parsons, J" journalOrPublisher="The Festivus" pageId="0" pageNumber="15" pagination="235 - 247" refId="B46" refString="Parsons, J, Abbas, J, 2016. A new subspecies of Amphidromus (Goniodromus) bulowi Fruhstorfer, 1905 (Gastropoda: Pulmonata: Camaenidae) from Sumatra, Indonesia. The Festivus 48 (4): 235 - 247, DOI: https://doi.org/10.54173/F484235" title="A new subspecies of Amphidromus (Goniodromus) bulowi Fruhstorfer, 1905 (Gastropoda: Pulmonata: Camaenidae) from Sumatra, Indonesia." url="https://doi.org/10.54173/F484235" volume="48" year="2016">Parsons and Abbas 2016</bibRefCitation>
|
||
: 240-242, figs 4 bottom, 5, 6a, b, d, 7.
|
||
</paragraph>
|
||
<paragraph id="CED15CD858868238F2CA6791EC840BAB" pageId="0" pageNumber="15">
|
||
<taxonomicName id="88B2CEEC0C205ECFE7638D809357D9EA" authorityName="Thach" authorityYear="2016" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus franzhuberi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="franzhuberi">Amphidromus franzhuberi</taxonomicName>
|
||
Thach, 2016: 64-65, fig. 42; pl. 23, figs 315-319. Type locality: along the border of Nha Trang outskirts and Khanh Vinh District, Khanh Hoa Province (Central Vietnam).
|
||
<bibRefCitation id="DDE4574DB0A1B768B960D6AA40657693" DOI="https://doi.org/10.12657/folmal.028.002" author="Pall-Gergely, B" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="15" pagination="35 - 76" refId="B45" refString="Pall-Gergely, B, Hunyadi, A, Auffenberg, K, 2020. Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014-2019). Folia Malacologica 28 (1): 35 - 76, DOI: https://doi.org/10.12657/folmal.028.002" title="Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014 - 2019)." url="https://doi.org/10.12657/folmal.028.002" volume="28" year="2020">
|
||
<normalizedToken id="1F5358A18DD02F8CC10E1C8CBA777862" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
||
et al. 2020
|
||
</bibRefCitation>
|
||
: 50, fig. 14.
|
||
<bibRefCitation id="A812B7519B69DF8A33C50F4EDD446C0D" author="Thach, NN" journalOrPublisher="48 HRBooks Company, Akron, Ohio, USA" pageId="0" pageNumber="15" refId="B75" refString="Thach, NN, 2020a. New shells of South Asia. Vol. 2. Seashells*Freshwater*Land snails. With one New Genus and 140 New Species & Subspecies, Reply to comments made in error. 48HRBooks Company, Akron, Ohio, USA" title="New shells of South Asia. Vol. 2. Seashells * Freshwater * Land snails. With one New Genus and 140 New Species & Subspecies, Reply to comments made in error." year="2020 a">Thach 2020a</bibRefCitation>
|
||
: 58, pl. 76, fig. 893 left.
|
||
<bibRefCitation id="98FB596E8F48E43F12E8F38FAE9E5056" author="Thach, NN" journalOrPublisher="48 HRBooks Company, Akron, Ohio, USA" pageId="0" pageNumber="15" refId="B77" refString="Thach, NN, 2021. New shells of South Asia and Taiwan, China, Tanzania. Seashells*Freshwater*Land snails. With 116 new species and subspecies and rejected synonyms, accepted species. 48HRBooks Company, Akron, Ohio, USA" title="New shells of South Asia and Taiwan, China, Tanzania. Seashells * Freshwater * Land snails. With 116 new species and subspecies and rejected synonyms, accepted species." year="2021">Thach 2021</bibRefCitation>
|
||
: 60 syn. nov.
|
||
</paragraph>
|
||
<paragraph id="A29BDD1F16776DDDEBD111E21FD89A3C" pageId="0" pageNumber="15">
|
||
<taxonomicName id="24A00832524A361E1205CBFFF7E2B974" authorityName="Fruhstorfer" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphidromus buelowi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">Amphidromus buelowi</taxonomicName>
|
||
.
|
||
<bibRefCitation id="BEA855DAD757658CC201BC9DEA7E11A0" DOI="https://doi.org/10.12657/folmal.028.002" author="Pall-Gergely, B" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="15" pagination="35 - 76" refId="B45" refString="Pall-Gergely, B, Hunyadi, A, Auffenberg, K, 2020. Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014-2019). Folia Malacologica 28 (1): 35 - 76, DOI: https://doi.org/10.12657/folmal.028.002" title="Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014 - 2019)." url="https://doi.org/10.12657/folmal.028.002" volume="28" year="2020">
|
||
<normalizedToken id="942265AAD9FAB5A55BC64740105E69A3" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
||
et al. 2020
|
||
</bibRefCitation>
|
||
: fig. 16.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CC1114F1BE83386296B7DD39FD1E136F" pageId="0" pageNumber="15" type="materials_examined">
|
||
<paragraph id="4618505CB64C5451B7C6FDBC2F18B549" pageId="0" pageNumber="15">Material examined.</paragraph>
|
||
<paragraph id="8CBD3809F94A934483281EB491B9A60A" pageId="0" pageNumber="15">
|
||
<materialsCitation id="F15C2067A738E27386D054B56590DCEE" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Indonesia" location="Sinistral" specimenCount="1" typeStatus="lectotype">
|
||
<collectingCountry id="EBC508BEB19F745DF62A5380BBB7C2A4" name="Indonesia">Indonesia</collectingCountry>
|
||
:
|
||
<location id="C42CB0B9E447AAB68150C393558C6164" LSID="urn:lsid:plazi:treatment:6083D4A531595FA79F2C8B8BD37A108B:C42CB0B9E447AAB68150C393558C6164" country="Indonesia" name="Sinistral">Sinistral</location>
|
||
,
|
||
<emphasis id="2DC691E6B3DAC0CB1A00ADB9AB3BD15B" bold="true" italics="true" pageId="0" pageNumber="15">
|
||
<typeStatus id="F9DF4166A4DAC0794B962B2DFE7F4831" type="lectotype">lectotype</typeStatus>
|
||
</emphasis>
|
||
of "
|
||
<taxonomicName id="87488EA3C7D14891119DD9FFFCF26EFA" authorityName="Fruhstorfer" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus buelowi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="81B6F756014E7A3E04B1BEFFEA234239" italics="true" pageId="0" pageNumber="15">Amphidromus buelowi</emphasis>
|
||
</taxonomicName>
|
||
", NHMUK 1910.12.30.98 (Fig.
|
||
<figureCitation id="8AACC2CB9C01F7302971C91FD1A96472" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Shells of Amphidromus buelowi Fruhstorfer, 1905 A lectotype of " Amphidromus buelowi " (NHMUK 1910.12.30.98) B holotype of " Amphidromus asper " (SMF 7762) C holotype of " Amphidromus franzhuberi " (MNHN-IM- 2000 - 31892) D specimen from Padang Soekeli, Singalang, Indonesia (RBINS I. G. 10591) E specimen from Mount Singgalang, Sepuluh Koto, Tanah Datar Regency, West Sumatra, Indonesia (NMNS- 8764 - 025) F specimen from Lang-Biang, Annam, Vietnam (RBINS I. G. 10591) G specimen from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam (NMNS- 8764 - 027) H, I specimen from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 030, NMNS- 8764 - 031). Credit: H. Taylor, NHM (A), M. Caballer, MNHN (C), RBINS (D, F)." figureDoi="10.3897/zookeys.1196.112146.figure15" httpUri="https://binary.pensoft.net/fig/1007833" pageId="0" pageNumber="15">15A</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="0140B94380CFFFAF30621DF307AD3FBD" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Vietnam" location="Dextral" specimenCount="1" typeStatus="holotype">
|
||
<collectingCountry id="CE96083C202C25E433B29E2D6135EF0F" name="Vietnam">Vietnam</collectingCountry>
|
||
:
|
||
<location id="E95E24CE1EB1F09255E40C82CB7ADD8C" LSID="urn:lsid:plazi:treatment:6083D4A531595FA79F2C8B8BD37A108B:E95E24CE1EB1F09255E40C82CB7ADD8C" country="Vietnam" name="Dextral">Dextral</location>
|
||
,
|
||
<emphasis id="D77C1556E1C9A2B624B6841D97EB8D03" bold="true" italics="true" pageId="0" pageNumber="15">
|
||
<typeStatus id="66920B14F12CCABF1F6D504DFD0B4786" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
of "
|
||
<taxonomicName id="4E0295531DBB5C4C1EF9CF7A57F3F62D" authorityName="Jirapatrasilp & Huang & Sutcharit & Lee" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus asper" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asper">
|
||
<emphasis id="C12B1832CCBAA95F0EE17C194FBC1C3C" italics="true" pageId="0" pageNumber="15">Amphidromus asper</emphasis>
|
||
</taxonomicName>
|
||
", SMF 7762 (Fig.
|
||
<figureCitation id="14BA5A6EE55AF88B96CB67BB5E3D5442" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Shells of Amphidromus buelowi Fruhstorfer, 1905 A lectotype of " Amphidromus buelowi " (NHMUK 1910.12.30.98) B holotype of " Amphidromus asper " (SMF 7762) C holotype of " Amphidromus franzhuberi " (MNHN-IM- 2000 - 31892) D specimen from Padang Soekeli, Singalang, Indonesia (RBINS I. G. 10591) E specimen from Mount Singgalang, Sepuluh Koto, Tanah Datar Regency, West Sumatra, Indonesia (NMNS- 8764 - 025) F specimen from Lang-Biang, Annam, Vietnam (RBINS I. G. 10591) G specimen from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam (NMNS- 8764 - 027) H, I specimen from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 030, NMNS- 8764 - 031). Credit: H. Taylor, NHM (A), M. Caballer, MNHN (C), RBINS (D, F)." figureDoi="10.3897/zookeys.1196.112146.figure15" httpUri="https://binary.pensoft.net/fig/1007833" pageId="0" pageNumber="15">15B</figureCitation>
|
||
). Dextral,
|
||
<emphasis id="14E5F217E05D424A98BA0160ACE28BCC" bold="true" italics="true" pageId="0" pageNumber="15">
|
||
<typeStatus id="3DE3ECC586AB96225D71473DC8D3AC5D" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
of "
|
||
<taxonomicName id="4DC45B30478E08983DD25F51D3E1616E" authorityName="Thach" authorityYear="2016" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus franzhuberi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="franzhuberi">
|
||
<emphasis id="382BD772F44C2889F6A3D61396D3BA17" italics="true" pageId="0" pageNumber="15">Amphidromus franzhuberi</emphasis>
|
||
</taxonomicName>
|
||
", MNHN-IM-2000-31892 (Fig.
|
||
<figureCitation id="6A753939906C67D40A279E8EB9207FE9" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Shells of Amphidromus buelowi Fruhstorfer, 1905 A lectotype of " Amphidromus buelowi " (NHMUK 1910.12.30.98) B holotype of " Amphidromus asper " (SMF 7762) C holotype of " Amphidromus franzhuberi " (MNHN-IM- 2000 - 31892) D specimen from Padang Soekeli, Singalang, Indonesia (RBINS I. G. 10591) E specimen from Mount Singgalang, Sepuluh Koto, Tanah Datar Regency, West Sumatra, Indonesia (NMNS- 8764 - 025) F specimen from Lang-Biang, Annam, Vietnam (RBINS I. G. 10591) G specimen from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam (NMNS- 8764 - 027) H, I specimen from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 030, NMNS- 8764 - 031). Credit: H. Taylor, NHM (A), M. Caballer, MNHN (C), RBINS (D, F)." figureDoi="10.3897/zookeys.1196.112146.figure15" httpUri="https://binary.pensoft.net/fig/1007833" pageId="0" pageNumber="15">15C</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7B15483B311086C3489C21493A4AF52B" pageId="0" pageNumber="15" type="materials_examined">
|
||
<paragraph id="CE35B2CB07E95121C5315D6ECA438818" pageId="0" pageNumber="15">Other material examined.</paragraph>
|
||
<paragraph id="9C60E04471329D0AA020DECBB9DCCE41" pageId="0" pageNumber="15">
|
||
<collectingCountry id="E459D00A3CC590B219950E72F2F51C89" name="Indonesia">Indonesia</collectingCountry>
|
||
: 2D specimens, Padang
|
||
<normalizedToken id="6542F99D5243E6F4448B04791FBD794D" originalValue="Sökeli">Soekeli</normalizedToken>
|
||
, Singalang, RBINS I.G. 10591/1-2 (Fig.
|
||
<figureCitation id="6140CC187AD3F926E6CDD186B44238A9" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Shells of Amphidromus buelowi Fruhstorfer, 1905 A lectotype of " Amphidromus buelowi " (NHMUK 1910.12.30.98) B holotype of " Amphidromus asper " (SMF 7762) C holotype of " Amphidromus franzhuberi " (MNHN-IM- 2000 - 31892) D specimen from Padang Soekeli, Singalang, Indonesia (RBINS I. G. 10591) E specimen from Mount Singgalang, Sepuluh Koto, Tanah Datar Regency, West Sumatra, Indonesia (NMNS- 8764 - 025) F specimen from Lang-Biang, Annam, Vietnam (RBINS I. G. 10591) G specimen from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam (NMNS- 8764 - 027) H, I specimen from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 030, NMNS- 8764 - 031). Credit: H. Taylor, NHM (A), M. Caballer, MNHN (C), RBINS (D, F)." figureDoi="10.3897/zookeys.1196.112146.figure15" httpUri="https://binary.pensoft.net/fig/1007833" pageId="0" pageNumber="15">15D</figureCitation>
|
||
); 4D specimens, Mount Singgalang, Sepuluh Koto, Tanah Datar Regency,
|
||
<collectingRegion id="C81ED204924E981279977AD72E2E0BFD" country="Indonesia" name="Sumatera Barat">West Sumatra</collectingRegion>
|
||
, NMNS-8764-022- NMNS-8764-025 (Fig.
|
||
<figureCitation id="2278C77D245EA9A4353DEDD658BE21C8" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Shells of Amphidromus buelowi Fruhstorfer, 1905 A lectotype of " Amphidromus buelowi " (NHMUK 1910.12.30.98) B holotype of " Amphidromus asper " (SMF 7762) C holotype of " Amphidromus franzhuberi " (MNHN-IM- 2000 - 31892) D specimen from Padang Soekeli, Singalang, Indonesia (RBINS I. G. 10591) E specimen from Mount Singgalang, Sepuluh Koto, Tanah Datar Regency, West Sumatra, Indonesia (NMNS- 8764 - 025) F specimen from Lang-Biang, Annam, Vietnam (RBINS I. G. 10591) G specimen from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam (NMNS- 8764 - 027) H, I specimen from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 030, NMNS- 8764 - 031). Credit: H. Taylor, NHM (A), M. Caballer, MNHN (C), RBINS (D, F)." figureDoi="10.3897/zookeys.1196.112146.figure15" httpUri="https://binary.pensoft.net/fig/1007833" pageId="0" pageNumber="15">15E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="0A8686CB33F157B5A31B8E4A64B0ED25" doi="10.3897/zookeys.1196.112146.figure15" httpUri="https://binary.pensoft.net/fig/1007833" pageId="0" pageNumber="15" start="Figure 15" startId="F15">
|
||
<paragraph id="325BBADEE5BF05E9DDE9427F740A6D76" pageId="0" pageNumber="15">
|
||
<emphasis id="8ABBAF40855FF9B28149D8096443EBF8" bold="true" pageId="0" pageNumber="15">Figure 15.</emphasis>
|
||
Shells of
|
||
<taxonomicName id="9CA62FC6241DE7E2B443348075243BCE" authorityName="Fruhstorfer" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus buelowi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="608C2F7C29D578ABC6249BFA25705F95" italics="true" pageId="0" pageNumber="15">Amphidromus buelowi</emphasis>
|
||
</taxonomicName>
|
||
Fruhstorfer, 1905
|
||
<emphasis id="598F3CBF49D9589A6E0EFB8E7FBB26FE" bold="true" pageId="0" pageNumber="15">A</emphasis>
|
||
<typeStatus id="BEB7F61C697A7A72CAFC4072EB13EA69" type="lectotype">lectotype</typeStatus>
|
||
of "
|
||
<taxonomicName id="1BEE0310B16A355C02797EB04147300F" authorityName="Fruhstorfer" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus buelowi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="930D017A6B5AA79E42FB01C6D110948C" italics="true" pageId="0" pageNumber="15">Amphidromus buelowi</emphasis>
|
||
</taxonomicName>
|
||
" (NHMUK 1910.12.30.98)
|
||
<emphasis id="55198F7CAFA6BA9D877737559AF18A60" bold="true" pageId="0" pageNumber="15">B</emphasis>
|
||
<typeStatus id="822DDEBA6A543E406074716E8843D3BB" type="holotype">holotype</typeStatus>
|
||
of "
|
||
<taxonomicName id="B01E0742A3E3BBE9A66462DE3C58BAC4" authorityName="Jirapatrasilp & Huang & Sutcharit & Lee" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus asper" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asper">
|
||
<emphasis id="4D4FC48B3C410B141858240F1BADBA5C" italics="true" pageId="0" pageNumber="15">Amphidromus asper</emphasis>
|
||
</taxonomicName>
|
||
" (SMF 7762)
|
||
<emphasis id="DA7156D43E067FDE5C9D7EDD61B59312" bold="true" pageId="0" pageNumber="15">C</emphasis>
|
||
<typeStatus id="CAD5BE8A4B718F94DA21635D7AA4E134" type="holotype">holotype</typeStatus>
|
||
of "
|
||
<taxonomicName id="A07D2920DE1ED73B36DC6CD886B75CCE" authorityName="Thach" authorityYear="2016" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus franzhuberi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="franzhuberi">
|
||
<emphasis id="907EB38473AE294AD8419743888CDE8E" italics="true" pageId="0" pageNumber="15">Amphidromus franzhuberi</emphasis>
|
||
</taxonomicName>
|
||
" (MNHN-IM-2000-31892)
|
||
<emphasis id="F973FB3C02B25A5AAAD006FF0C5FD43A" bold="true" pageId="0" pageNumber="15">D</emphasis>
|
||
specimen from Padang
|
||
<normalizedToken id="B57AFDF183E616FF26D160F5AFEEB221" originalValue="Sökeli">Soekeli</normalizedToken>
|
||
, Singalang,
|
||
<collectingCountry id="B0F5A0808EE1CDA80A29B11F62687417" name="Indonesia">Indonesia</collectingCountry>
|
||
(RBINS I.G. 10591)
|
||
<emphasis id="AD314642E7368173769460E0775DAA87" bold="true" pageId="0" pageNumber="15">E</emphasis>
|
||
specimen from Mount Singgalang, Sepuluh Koto, Tanah Datar Regency,
|
||
<collectingRegion id="C203D07184F8041083699F37C3F2E67D" country="Indonesia" name="Sumatera Barat">West Sumatra</collectingRegion>
|
||
,
|
||
<collectingCountry id="865D63B2302C342ED421ACBF3A3B9296" name="Indonesia">Indonesia</collectingCountry>
|
||
(NMNS-8764-025)
|
||
<emphasis id="4B789AA40355E1B87239A7FE5A216399" bold="true" pageId="0" pageNumber="15">F</emphasis>
|
||
specimen from Lang-Biang,
|
||
<collectingCountry id="42C46A420C504DE3E255B162164897C8" name="Vietnam">Annam</collectingCountry>
|
||
,
|
||
<collectingCountry id="75DEC3EE4F3C65E52553ECC580630B5F" name="Vietnam">Vietnam</collectingCountry>
|
||
(RBINS I.G. 10591)
|
||
<emphasis id="0D2D7E7B06131D4404238DD68C03D342" bold="true" pageId="0" pageNumber="15">G</emphasis>
|
||
specimen from Lang-Biang plateau, Lac Duong,
|
||
<collectingRegion id="F32804B9249F086A77D6BB4137353245" country="Vietnam" name="Lam Dong">Lam Dong</collectingRegion>
|
||
,
|
||
<collectingCountry id="1BB82A0D4AB6B5B949400E7BA994F294" name="Vietnam">Vietnam</collectingCountry>
|
||
(NMNS-8764-027)
|
||
<emphasis id="11C1A8A8AAE40415C89240A547894ADA" bold="true" pageId="0" pageNumber="15">H, I</emphasis>
|
||
specimen from Nha Trang,
|
||
<collectingRegion id="9352E318AB06CA381777A441566DAB3D" country="Vietnam" name="Khanh Hoa">Khanh Hoa</collectingRegion>
|
||
,
|
||
<collectingCountry id="CFF752F1718F15AAF102D6B3CB304850" name="Vietnam">Vietnam</collectingCountry>
|
||
(NMNS-8764-030, NMNS-8764-031). Credit: H. Taylor, NHM (
|
||
<emphasis id="2F58ACD27DB6BED7A7C51F9968F0AD1B" bold="true" pageId="0" pageNumber="15">A</emphasis>
|
||
), M. Caballer, MNHN (
|
||
<emphasis id="EF471A3DF6E1FF36B2765A6238F82E3C" bold="true" pageId="0" pageNumber="15">C</emphasis>
|
||
), RBINS (
|
||
<emphasis id="E9F5F83395551981D6D327522E7BE5B6" bold="true" pageId="0" pageNumber="15">D, F</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="456D75ED47A5E88CD4709AA51F8EE839" pageId="0" pageNumber="15">
|
||
<collectingCountry id="E03F2181C89BAB1EB8922FBB301D3FF6" name="Vietnam">Vietnam</collectingCountry>
|
||
: 2D specimens, Lang-Biang,
|
||
<collectingCountry id="ECE767C74FC0248F4CB70971ECC0E429" name="Vietnam">Annam</collectingCountry>
|
||
, RBINS I.G. 10591/3-4 (Fig.
|
||
<figureCitation id="723DB70D26E6C21671E430BFF13819B9" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Shells of Amphidromus buelowi Fruhstorfer, 1905 A lectotype of " Amphidromus buelowi " (NHMUK 1910.12.30.98) B holotype of " Amphidromus asper " (SMF 7762) C holotype of " Amphidromus franzhuberi " (MNHN-IM- 2000 - 31892) D specimen from Padang Soekeli, Singalang, Indonesia (RBINS I. G. 10591) E specimen from Mount Singgalang, Sepuluh Koto, Tanah Datar Regency, West Sumatra, Indonesia (NMNS- 8764 - 025) F specimen from Lang-Biang, Annam, Vietnam (RBINS I. G. 10591) G specimen from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam (NMNS- 8764 - 027) H, I specimen from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 030, NMNS- 8764 - 031). Credit: H. Taylor, NHM (A), M. Caballer, MNHN (C), RBINS (D, F)." figureDoi="10.3897/zookeys.1196.112146.figure15" httpUri="https://binary.pensoft.net/fig/1007833" pageId="0" pageNumber="15">15F</figureCitation>
|
||
); 2D specimens, Lang-Biang plateau, Lac Duong District,
|
||
<collectingRegion id="5A395E3DA6293B2FA5878F9A5AD8F31B" country="Vietnam" name="Lam Dong">Lam Dong Province</collectingRegion>
|
||
, NMNS-8764-026, NMNS-8764-027 (Fig.
|
||
<figureCitation id="6265E13440668F22DAF8CC57F2F148A9" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Shells of Amphidromus buelowi Fruhstorfer, 1905 A lectotype of " Amphidromus buelowi " (NHMUK 1910.12.30.98) B holotype of " Amphidromus asper " (SMF 7762) C holotype of " Amphidromus franzhuberi " (MNHN-IM- 2000 - 31892) D specimen from Padang Soekeli, Singalang, Indonesia (RBINS I. G. 10591) E specimen from Mount Singgalang, Sepuluh Koto, Tanah Datar Regency, West Sumatra, Indonesia (NMNS- 8764 - 025) F specimen from Lang-Biang, Annam, Vietnam (RBINS I. G. 10591) G specimen from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam (NMNS- 8764 - 027) H, I specimen from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 030, NMNS- 8764 - 031). Credit: H. Taylor, NHM (A), M. Caballer, MNHN (C), RBINS (D, F)." figureDoi="10.3897/zookeys.1196.112146.figure15" httpUri="https://binary.pensoft.net/fig/1007833" pageId="0" pageNumber="15">15G</figureCitation>
|
||
); 6D + 1S specimens, Nha Trang,
|
||
<collectingRegion id="ED0E5507B8A9ED06AD4296EA0C2780A2" country="Vietnam" name="Khanh Hoa">Khanh Hoa Province</collectingRegion>
|
||
, NMNS-8764-028- NMNS-8764-034 (Fig.
|
||
<figureCitation id="5E7FF243EDBA51063AAA40EFB97A6CB7" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Shells of Amphidromus buelowi Fruhstorfer, 1905 A lectotype of " Amphidromus buelowi " (NHMUK 1910.12.30.98) B holotype of " Amphidromus asper " (SMF 7762) C holotype of " Amphidromus franzhuberi " (MNHN-IM- 2000 - 31892) D specimen from Padang Soekeli, Singalang, Indonesia (RBINS I. G. 10591) E specimen from Mount Singgalang, Sepuluh Koto, Tanah Datar Regency, West Sumatra, Indonesia (NMNS- 8764 - 025) F specimen from Lang-Biang, Annam, Vietnam (RBINS I. G. 10591) G specimen from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam (NMNS- 8764 - 027) H, I specimen from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 030, NMNS- 8764 - 031). Credit: H. Taylor, NHM (A), M. Caballer, MNHN (C), RBINS (D, F)." figureDoi="10.3897/zookeys.1196.112146.figure15" httpUri="https://binary.pensoft.net/fig/1007833" pageId="0" pageNumber="15">15H, I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4D54724504ADF923186D81117AC414B1" pageId="0" pageNumber="15" type="diagnosis">
|
||
<paragraph id="BA8DC716FB027E88BC532F29A2F52864" pageId="0" pageNumber="15">Diagnosis.</paragraph>
|
||
<paragraph id="293FCB4DF42889EFE21F9EFC98BB2488" pageId="0" pageNumber="15">Shell large and chirally dimorphic. Shell colour with irregularly zigzag of dark radial streaks, and dark triangular blotches. Aperture elliptical ovate with more or less prominent anterior notch and umbilical hump; twisted columella plait. Genitalia with appendix.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="47674CF99135E5DC4F6C4D0A6D86E029" pageId="0" pageNumber="15" type="differential diagnosis">
|
||
<paragraph id="67B7442745E51DC9755470698454BC0E" pageId="0" pageNumber="15">Differential diagnosis.</paragraph>
|
||
<paragraph id="3F2AE461E75B5CA4225292D91021D606" pageId="0" pageNumber="15">
|
||
<taxonomicName id="36461388953D0B4A6F15C50B1EE09DC8" authorityName="Fruhstorfer" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus buelowi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="444E278023EEA8166D510A22A9387B3A" italics="true" pageId="0" pageNumber="15">Amphidromus buelowi</emphasis>
|
||
</taxonomicName>
|
||
differs from the similar species
|
||
<taxonomicName id="540184F6AA71D1D1A34996F0D5473AC1" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. asperoides" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asperoides">
|
||
<emphasis id="A23275A15C6AD0D512288A98C2FFAE34" italics="true" pageId="0" pageNumber="15">A. asperoides</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in having a distinct twisted columella plait, a prominent umbilical hump encircling columellar area, and an apertural notch projecting anteriorly. In contrast,
|
||
<taxonomicName id="D41FDA009D3A7823BBCB1FDB74BBAA4F" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. asperoides" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asperoides">
|
||
<emphasis id="02BA410B7B936DE418A57D743EDF5C57" italics="true" pageId="0" pageNumber="15">A. asperoides</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. possesses a straight columella, and without apertural notch and umbilical hump. In addition, on the soft body of living snail, the entire body of
|
||
<taxonomicName id="79194808CF6BC12185EB3396EF3BB950" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. buelowi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="CAFD984B600F24323E64285B3360AC4F" italics="true" pageId="0" pageNumber="15">A. buelowi</emphasis>
|
||
</taxonomicName>
|
||
is reddish orange, while
|
||
<taxonomicName id="A9ABC0FC2AE7CED1A4F9B1227F8C5EE5" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. asperoides" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asperoides">
|
||
<emphasis id="4A68A1C9F18E719FFACEAECD6146CBE6" italics="true" pageId="0" pageNumber="15">A. asperoides</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. exhibits a uniform brownish yellow to pale brown body.
|
||
<taxonomicName id="9727F0430D3F025A8BB8243D6CFA6208" authorityName="Fruhstorfer" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus buelowi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="F0D2CD720230AFB5C448746D89122463" italics="true" pageId="0" pageNumber="15">Amphidromus buelowi</emphasis>
|
||
</taxonomicName>
|
||
is also recognised by a distinct clade in the molecular phylogeny (Fig.
|
||
<figureCitation id="F58F236188E8FB62D6666C50A983C3E3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bayesian phylogeny of Amphidromus spp. based on mitochondrial COI and 16 S genes. Nodal support values are given as SH-aLRT / aBayes / ultra-fast bootstrap (IQ-TREE, ML) / posterior probability (MrBayes, BI). An asterisk on each branch indicates a clade with all well-supported values (SH-aLRT ≥ 80 %, aBayes ≥ 0.95, BS ≥ 95 %, PP ≥ 0.95). Species numbers correspond to those in Fig. 1 and Tables 1, 2." figureDoi="10.3897/zookeys.1196.112146.figure2" httpUri="https://binary.pensoft.net/fig/1007820" pageId="0" pageNumber="15">2</figureCitation>
|
||
), with the closest
|
||
<emphasis id="F4C7F4A7AC5660FF16CFF3EFD816D427" italics="true" pageId="0" pageNumber="15">p</emphasis>
|
||
-distance to
|
||
<taxonomicName id="8E107462C3063BABC947B53D605895EC" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ingens" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="F50D20CF1D0BBF495DED4DD22E5D42F1" italics="true" pageId="0" pageNumber="15">A. ingens</emphasis>
|
||
</taxonomicName>
|
||
in COI (12.23%) and
|
||
<taxonomicName id="8B2928954A0CDD3ABF8776CA94FD4840" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. asperoides" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asperoides">
|
||
<emphasis id="70C9EF3D266BFC10E1663A2E818C159B" italics="true" pageId="0" pageNumber="15">A. asperoides</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="C3C0E4A20CAC2147AED951191FC850F1" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ingens" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="576E7B706409BE2B93591A613CC24ED2" italics="true" pageId="0" pageNumber="15">A. ingens</emphasis>
|
||
</taxonomicName>
|
||
in 16S (4.61%) (Table
|
||
<tableCitation id="E09EF7543966A87113BA1C9993CE7774" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Percentage of uncorrected pairwise interspecific distances for the partial COI (above the diagonal) and 16 S rRNA (below the diagonal) gene fragments among the Amphidromus species in this study. Intraspecific distances for COI / 16 S rRNA are shown on the diagonal (bold). Species numbers correspond to those in Figs 1, 2, and Table 1. The numbers of sequences used to calculate the distances of each respective gene fragment are given as n = COI, 16 S in the first column." httpUri="http://table.plazi.org/id/A1B4F960F999761C3840BD02D5BE2608" pageId="0" pageNumber="15" tableUuid="A1B4F960F999761C3840BD02D5BE2608">2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="75D1E2ECFC5295D3FE3573E1432F1DAE" pageId="0" pageNumber="15" type="description">
|
||
<paragraph id="EFFF4BBC8D858003416EF8316BE71B1F" pageId="0" pageNumber="15">Description.</paragraph>
|
||
<paragraph id="3DEF368708831E3802C64A6E6C0F32A1" pageId="0" pageNumber="15">
|
||
<emphasis id="D9A5FDE3AAEC6DC0A7F1E01D7D650F27" bold="true" italics="true" pageId="0" pageNumber="15">Shell</emphasis>
|
||
large (height 45.3-51.1 mm, width 26.2-26.6 mm), chirally dimorphic, solid, and ovate conical. Spire conical with white or pale colour; apex acute without black spot on tip. Whorls 6-7 little convex to smooth; suture wide and shallow; last whorl well rounded to slightly elongated and with more or less prominent umbilical hump. Periostracum thin corneous; varices generally present. Shell ground colour pale yellowish, decorated with irregular zigzag of dark radial streaks, and dark triangular blotches connected with dark streaks. Parietal callus thickened, white and much thinner in central area. Aperture elliptical ovate; with more or less anterior notch; inner side of outer wall whitish colour; peristome thickened, expanded, and reflexed but not attached to last whorl; lip whitish. Columella white, straight and with distinct twisted plait. Umbilicus imperforate.
|
||
</paragraph>
|
||
<paragraph id="9CFE85A6A8328B2DE1B519CE9D135F2A" pageId="0" pageNumber="15">
|
||
<emphasis id="8DD7D490251B2554A4BC0D44540A5F18" bold="true" pageId="0" pageNumber="15">
|
||
<emphasis id="2A10314B71D5543C36878C19E328C938" italics="true" pageId="0" pageNumber="15">Radula</emphasis>
|
||
.
|
||
</emphasis>
|
||
Teeth arranged in anteriorly pointed V-shaped rows. Central tooth monocuspid and slightly elongate-spatulate teeth with truncated cusp. Lateral teeth bicuspid; endocone curved with wide notch and blunt cusp; ectocone large with truncated cusp. Lateral teeth gradually transformed to asymmetric tricuspid marginal teeth. Outermost teeth with small and curved cusp on ectocone, and endocone and mesocone with curved cusps (Fig.
|
||
<figureCitation id="9D40377CEF393AC5CF32B3AA5E411659" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. SEM images of the radula A Amphidromus placostylus Moellendorff, 1900 from Dak Po, Gia Lai, Vietnam (NMNS- 8764 - 217) B Amphidromus buelowi Fruhstorfer, 1905 from Mount Singgalang, Sepuluh Koto, Tanah Datar Regency, West Sumatra, Indonesia (NMNS- 8764 - 024) C Amphidromus thachi Huber, 2015 from Buon Don, Dak Lak, Vietnam (NMNS- 8764 - 271). Central teeth are marked in yellow. The left and right images show the outer and inner sections of each radula, respectively." figureDoi="10.3897/zookeys.1196.112146.figure12" httpUri="https://binary.pensoft.net/fig/1007830" pageId="0" pageNumber="15">12B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7B51118F9B37BF6E35FBE25E3509E619" pageId="0" pageNumber="15">
|
||
<emphasis id="A13A289D91E879FECD9E280D88782656" bold="true" pageId="0" pageNumber="15">
|
||
<emphasis id="093A361BA904DD8503DB136BFAA53E71" italics="true" pageId="0" pageNumber="15">Genital organs</emphasis>
|
||
.
|
||
</emphasis>
|
||
Atrium relatively short. Penis slender, conical, and nearly as long as vagina. Penial retractor muscle inserting on epiphallus close to penis. Epiphallus long, slender tube, and almost same diameter as penis. Flagellum short, extending from epiphallus and terminating in slightly enlarged folded coil. Appendix short, thin tube, 3
|
||
<normalizedToken id="35919F5D125C7F323778D2FB87A47DA8" originalValue="×">x</normalizedToken>
|
||
longer than flagellum, and approximately as long as epiphallus. Vas deferens slender tube passing from free oviduct and terminating at epiphallus-flagellum junction (Fig.
|
||
<figureCitation id="0CFC13861F910FF475E6912D6F33493F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Genitalia of Amphidromus spp A-C Amphidromus ingensoides sp. nov. from Hon Ba, Khanh Son, Khanh Hoa, Vietnam (NMNS- 8764 - 107), showing A general view of genitalia B interior structures of penis C interior structures of vagina chamber D-F Amphidromus buelowi Fruhstorfer, 1905 from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 031), showing D general view of genitalia E interior structures of penis F interior structures of vagina chamber. Red dots indicate the shape of the missing gametolytic sac. Green arrows indicate the genital openings. Abbreviations: ap, appendix; e, epiphallus; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; ov, oviduct; p, penis; pp, penial pilaster; pr, penial retractor muscle; pv, penial verge; v, vagina; vd, vas deferens; vp, vaginal pilaster" figureDoi="10.3897/zookeys.1196.112146.figure14" httpUri="https://binary.pensoft.net/fig/1007832" pageId="0" pageNumber="15">14D</figureCitation>
|
||
). Internal wall of penis corrugated, exhibiting series of thickened and swollen longitudinal penial pilasters forming fringe around penial wall, and with weaker folds around base of penial verge. Penial verge short conical with nearly smooth surface (Fig.
|
||
<figureCitation id="90A33F76F5F40DCCDF9BA110307F29B7" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Genitalia of Amphidromus spp A-C Amphidromus ingensoides sp. nov. from Hon Ba, Khanh Son, Khanh Hoa, Vietnam (NMNS- 8764 - 107), showing A general view of genitalia B interior structures of penis C interior structures of vagina chamber D-F Amphidromus buelowi Fruhstorfer, 1905 from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 031), showing D general view of genitalia E interior structures of penis F interior structures of vagina chamber. Red dots indicate the shape of the missing gametolytic sac. Green arrows indicate the genital openings. Abbreviations: ap, appendix; e, epiphallus; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; ov, oviduct; p, penis; pp, penial pilaster; pr, penial retractor muscle; pv, penial verge; v, vagina; vd, vas deferens; vp, vaginal pilaster" figureDoi="10.3897/zookeys.1196.112146.figure14" httpUri="https://binary.pensoft.net/fig/1007832" pageId="0" pageNumber="15">14E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="59A3D2BA24C04D8CF7C65BE9741BEFCC" pageId="0" pageNumber="15">
|
||
Vagina slender, long cylindrical, and ~ 2
|
||
<normalizedToken id="77BDD984F9339B7391D103F521E8306B" originalValue="×">x</normalizedToken>
|
||
longer than penis. Gametolytic duct enlarged cylindrical tube then abruptly tapering to long, slender tube terminally, connected to elongate gametolytic sac. Free oviduct short; oviduct compact, enlarged to form lobule alveoli (Fig.
|
||
<figureCitation id="64E28B6B92F113F2AFD22568475793F7" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Genitalia of Amphidromus spp A-C Amphidromus ingensoides sp. nov. from Hon Ba, Khanh Son, Khanh Hoa, Vietnam (NMNS- 8764 - 107), showing A general view of genitalia B interior structures of penis C interior structures of vagina chamber D-F Amphidromus buelowi Fruhstorfer, 1905 from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 031), showing D general view of genitalia E interior structures of penis F interior structures of vagina chamber. Red dots indicate the shape of the missing gametolytic sac. Green arrows indicate the genital openings. Abbreviations: ap, appendix; e, epiphallus; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; ov, oviduct; p, penis; pp, penial pilaster; pr, penial retractor muscle; pv, penial verge; v, vagina; vd, vas deferens; vp, vaginal pilaster" figureDoi="10.3897/zookeys.1196.112146.figure14" httpUri="https://binary.pensoft.net/fig/1007832" pageId="0" pageNumber="15">14D</figureCitation>
|
||
). Internal wall of vagina possessing corrugated ridges near genital orifice; ridges becoming thinner and smooth longitudinal vaginal pilasters in middle, swollen with irregularly shaped deep crenelations close to free oviduct opening (Fig.
|
||
<figureCitation id="6791A7295E600F29E9AAEB4182D68954" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Genitalia of Amphidromus spp A-C Amphidromus ingensoides sp. nov. from Hon Ba, Khanh Son, Khanh Hoa, Vietnam (NMNS- 8764 - 107), showing A general view of genitalia B interior structures of penis C interior structures of vagina chamber D-F Amphidromus buelowi Fruhstorfer, 1905 from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 031), showing D general view of genitalia E interior structures of penis F interior structures of vagina chamber. Red dots indicate the shape of the missing gametolytic sac. Green arrows indicate the genital openings. Abbreviations: ap, appendix; e, epiphallus; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; ov, oviduct; p, penis; pp, penial pilaster; pr, penial retractor muscle; pv, penial verge; v, vagina; vd, vas deferens; vp, vaginal pilaster" figureDoi="10.3897/zookeys.1196.112146.figure14" httpUri="https://binary.pensoft.net/fig/1007832" pageId="0" pageNumber="15">14F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C6AC80DAC3AB9C7E9AB102D382780775" pageId="0" pageNumber="15">
|
||
<emphasis id="391449A5B4DC513F7F078D8E050D3631" bold="true" italics="true" pageId="0" pageNumber="15">Living specimens</emphasis>
|
||
with soft body morphology generally similar to
|
||
<taxonomicName id="0B3F3C6166E33BAC0D4A369E03C886C5" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. ingens" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="ingens">
|
||
<emphasis id="DF4D7B437D1C5F0751446BED1688217C" italics="true" pageId="0" pageNumber="15">A. ingens</emphasis>
|
||
</taxonomicName>
|
||
. Animals with reddish orange body covered with reticulated skin. Lateral of body vary from yellowish (in younger specimen) to dark reddish orange colour (older specimens). Foot broad and long with reddish orange colour near foot sole margin. Head and dorsal of anterior body with reddish orange to dark colour. Upper tentacles pale reddish orange to brownish; lower tentacles short and paler in colour (Fig.
|
||
<figureCitation id="2DCEFCA48C708AF54E7A230093036B23" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Living Amphidromus spp A-C Amphidromus ingens Moellendorff, 1900 from Dak Lak, Vietnam D Amphidromus placostylus Moellendorff, 1900 from Hoai An, An Lao, Binh Dinh, Vietnam E, F Amphidromus ingensoides sp. nov. from Hon Ba, Khanh Son, Khanh Hoa, Vietnam G Amphidromus asperoides sp. nov. from Ea Tu, Buon Ma Thuat city, Dak Lak, Vietnam H, I Amphidromus buelowi Fruhstorfer, 1905 from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam J Amphidromus thachi Huber, 2015 from Krong Bong, Dak Lak, Vietnam." figureDoi="10.3897/zookeys.1196.112146.figure6" httpUri="https://binary.pensoft.net/fig/1007824" pageId="0" pageNumber="15">6H, I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E161BE85CDC6BF261796FEBA4B98D471" pageId="0" pageNumber="15" type="haplotype network">
|
||
<paragraph id="0A5EA0D38B721E8AB109914E1B862795" pageId="0" pageNumber="15">Haplotype network.</paragraph>
|
||
<paragraph id="22041AE562BD91FD72FA2C6EC69FFEBF" pageId="0" pageNumber="15">
|
||
There was a total of four COI haplotypes (Fig.
|
||
<figureCitation id="6DDCE675675C4C956302ED1E279C755D" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Mitochondrial haplotype minimum spanning networks of Amphidromus buelowi Fruhstorfer, 1905 A COI and B 16 S rRNA. The size of each circle corresponds to the frequency of that haplotype, also shown as the number in that circle. The bars on the branches indicate the number of mutational steps between haplotypes. Specimen codes correspond to those in Table 1." figureDoi="10.3897/zookeys.1196.112146.figure16" httpUri="https://binary.pensoft.net/fig/1007834" pageId="0" pageNumber="15">16A</figureCitation>
|
||
) and three 16S haplotypes (Fig.
|
||
<figureCitation id="E356C73E7F88BD1D251C2F0EF0E5F2F6" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Mitochondrial haplotype minimum spanning networks of Amphidromus buelowi Fruhstorfer, 1905 A COI and B 16 S rRNA. The size of each circle corresponds to the frequency of that haplotype, also shown as the number in that circle. The bars on the branches indicate the number of mutational steps between haplotypes. Specimen codes correspond to those in Table 1." figureDoi="10.3897/zookeys.1196.112146.figure16" httpUri="https://binary.pensoft.net/fig/1007834" pageId="0" pageNumber="15">16B</figureCitation>
|
||
) of
|
||
<taxonomicName id="B6C2A1BC4D6239A6E64DBF905FDDA78C" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. buelowi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="4A75A745B6F80504FEB3DFAF46CA05BE" italics="true" pageId="0" pageNumber="15">A. buelowi</emphasis>
|
||
</taxonomicName>
|
||
in this study, and the highest numbers of mutational steps in the COI and 16S minimum spanning networks are ten and one, respectively.
|
||
</paragraph>
|
||
<caption id="8875BE686C6971D529F88EC6CCF8F153" doi="10.3897/zookeys.1196.112146.figure16" httpUri="https://binary.pensoft.net/fig/1007834" pageId="0" pageNumber="15" start="Figure 16" startId="F16">
|
||
<paragraph id="C3B735B7521B125B9B2C2C79CB5ADF63" pageId="0" pageNumber="15">
|
||
<emphasis id="E554D155317F7A76BD4432545A1D11A8" bold="true" pageId="0" pageNumber="15">Figure 16.</emphasis>
|
||
Mitochondrial haplotype minimum spanning networks of
|
||
<taxonomicName id="A34C4817637AE90CD13927799A66B593" authorityName="Fruhstorfer" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus buelowi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="EB864ABB1CDB88929EC960BC1726EE24" italics="true" pageId="0" pageNumber="15">Amphidromus buelowi</emphasis>
|
||
</taxonomicName>
|
||
Fruhstorfer, 1905
|
||
<emphasis id="5B3517DA6A6888EA836CC88AD0D21574" bold="true" pageId="0" pageNumber="15">A</emphasis>
|
||
COI and
|
||
<emphasis id="17C1FD7DEBE0E25CD0BFBC4B61FFE540" bold="true" pageId="0" pageNumber="15">B</emphasis>
|
||
16S rRNA. The size of each circle corresponds to the frequency of that haplotype, also shown as the number in that circle. The bars on the branches indicate the number of mutational steps between haplotypes. Specimen codes correspond to those in Table
|
||
<tableCitation id="1B36700F21E1F6A5BC5AEA74DFAF2C82" captionStart="Table 1" captionStartId="T1" captionText="Table 1. List of specimens used in this study with species name, locality details, voucher and GenBank accession numbers. Species numbers correspond to those in Figs 1, 2, and Table 2." httpUri="http://table.plazi.org/id/536020F98423605B1D469F287C9C42E1" pageId="0" pageNumber="15" tableUuid="536020F98423605B1D469F287C9C42E1">1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="76F14307B060B5834783DF7F2FCB98B3" pageId="0" pageNumber="15" type="distribution">
|
||
<paragraph id="64C47698D089A9E605A55D0BBB6FF6A1" pageId="0" pageNumber="15">Distribution.</paragraph>
|
||
<paragraph id="8758651D61E93E4FB83383F0E0DF1CD5" pageId="0" pageNumber="15">The species has a widely disjunct distribution: one in Mount Singgalang, West Sumatra, Indonesia, and some localities in Khanh Hoa and Lam Dong provinces, South Vietnam.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1808F7D5F32F2534E040EF3649BA93E1" pageId="0" pageNumber="15" type="remarks">
|
||
<paragraph id="B5C583E91D7BA39F607D09B90C86248D" pageId="0" pageNumber="15">Remarks.</paragraph>
|
||
<paragraph id="A8CF2C53475449892E9597899EC5D4F2" pageId="0" pageNumber="15">
|
||
This species was originally described by
|
||
<bibRefCitation id="575C082BA6A4D8A53F13C34F432726D2" author="Fruhstorfer, H" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="15" pagination="83 - 84" publicationUrl="https://biostor.org/reference/190426" refId="B15" refString="Fruhstorfer, H, 1905. Ein neuer Amphidromus. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 37: 83 - 84, https://biostor.org/reference/190426" title="Ein neuer Amphidromus." url="https://biostor.org/reference/190426" volume="37" year="1905">Fruhstorfer (1905)</bibRefCitation>
|
||
from four chirally dimorphic specimens from West Sumatra.
|
||
<bibRefCitation id="5779D85A235FB7954B43FF34CD839ADC" author="Fruhstorfer, H" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="15" pagination="83 - 84" publicationUrl="https://biostor.org/reference/190426" refId="B15" refString="Fruhstorfer, H, 1905. Ein neuer Amphidromus. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 37: 83 - 84, https://biostor.org/reference/190426" title="Ein neuer Amphidromus." url="https://biostor.org/reference/190426" volume="37" year="1905">Fruhstorfer (1905)</bibRefCitation>
|
||
also indicated that there was a similar species collected on the way to the Lang-Bian plateau, ~ 120 km inland from the coast in southern Vietnam. He sent one specimen from this locality to O.F. von
|
||
<normalizedToken id="17D0FD08B6B847B79009C534BC2C6DCB" originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
, who did not describe or taxonomically treat this specimen any further. Later,
|
||
<bibRefCitation id="02BDCADB8EC254D0CCCCB0304CB295A5" author="Haas, F" journalOrPublisher="Senckenbergiana" pageId="0" pageNumber="15" pagination="94 - 98" refId="B19" refString="Haas, F, 1934. Neue Landschnecken des Senckenberg-Museums. Senckenbergiana 16: 94 - 98" title="Neue Landschnecken des Senckenberg-Museums." volume="16" year="1934">Haas (1934)</bibRefCitation>
|
||
, recognising that there were some differences in shell characters to
|
||
<taxonomicName id="C472B0C5F330E2D1814E76C108210964" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. buelowi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="B09EA4E67D3049968EE1E9A7687CC47C" italics="true" pageId="0" pageNumber="15">A. buelowi</emphasis>
|
||
</taxonomicName>
|
||
, described this particular specimen (now deposited in SMF) as a new species,
|
||
<taxonomicName id="4BEAEDE23F6422F96BBEEA9DF104FB00" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. asper" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asper">
|
||
<emphasis id="0738CB4CD337914696DD36AA4C9E4D7D" italics="true" pageId="0" pageNumber="15">A. asper</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="43797F4E329EF406EFC8B9C24F1DC4B8" author="Thach, NN" journalOrPublisher="48 HrBooks Company, Ohio, USA" pageId="0" pageNumber="15" refId="B72" refString="Thach, NN, 2016. Vietnamese new mollusks with 59 new species. 48HrBooks Company, Ohio, USA" title="Vietnamese new mollusks with 59 new species." year="2016">Thach (2016)</bibRefCitation>
|
||
also described a similar species,
|
||
<taxonomicName id="18FE51786AF37594F9866DF3EBAA2F37" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. franzhuberi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="franzhuberi">
|
||
<emphasis id="2613D435A16F37E7499860C378B4ABFD" italics="true" pageId="0" pageNumber="15">A. franzhuberi</emphasis>
|
||
</taxonomicName>
|
||
from the border of Nha Trang, Vietnam, which is described to differ from
|
||
<taxonomicName id="14F61D6F7848861D4034722A7DDFDF5D" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. buelowi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="518C7C1F2679620AF2DC432054866531" italics="true" pageId="0" pageNumber="15">A. buelowi</emphasis>
|
||
</taxonomicName>
|
||
in having a broader shell shape, more swollen body whorls, a less excavated base, a more inflated spire, a rounded anterior end of the outer lip, and monomorphic dextrality (just from four type series). However,
|
||
<bibRefCitation id="AF95BC90A446516F1BC8633F7E007B83" author="Thach, NN" journalOrPublisher="48 HrBooks Company, Ohio, USA" pageId="0" pageNumber="15" refId="B72" refString="Thach, NN, 2016. Vietnamese new mollusks with 59 new species. 48HrBooks Company, Ohio, USA" title="Vietnamese new mollusks with 59 new species." year="2016">Thach (2016)</bibRefCitation>
|
||
did not compare with
|
||
<taxonomicName id="210BCAD744AF330442F93CE9C6D9BADF" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. asper" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asper">
|
||
<emphasis id="47A347D45F4BD2A3EFB4FB70FD32C0F0" italics="true" pageId="0" pageNumber="15">A. asper</emphasis>
|
||
</taxonomicName>
|
||
from the nearby area. In this study, the samples from Nha Trang exhibit dimorphic chirality (the specimen lot containing both sinistral and dextral shell coiling; Fig.
|
||
<figureCitation id="8AFB798ADDD4E36C98402AEA57063074" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Shells of Amphidromus buelowi Fruhstorfer, 1905 A lectotype of " Amphidromus buelowi " (NHMUK 1910.12.30.98) B holotype of " Amphidromus asper " (SMF 7762) C holotype of " Amphidromus franzhuberi " (MNHN-IM- 2000 - 31892) D specimen from Padang Soekeli, Singalang, Indonesia (RBINS I. G. 10591) E specimen from Mount Singgalang, Sepuluh Koto, Tanah Datar Regency, West Sumatra, Indonesia (NMNS- 8764 - 025) F specimen from Lang-Biang, Annam, Vietnam (RBINS I. G. 10591) G specimen from Lang-Biang plateau, Lac Duong, Lam Dong, Vietnam (NMNS- 8764 - 027) H, I specimen from Nha Trang, Khanh Hoa, Vietnam (NMNS- 8764 - 030, NMNS- 8764 - 031). Credit: H. Taylor, NHM (A), M. Caballer, MNHN (C), RBINS (D, F)." figureDoi="10.3897/zookeys.1196.112146.figure15" httpUri="https://binary.pensoft.net/fig/1007833" pageId="0" pageNumber="15">15H, I</figureCitation>
|
||
), and upon examining the type specimens of
|
||
<taxonomicName id="DB820C61C290F9B153D9AC61CC513CCD" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. asper" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asper">
|
||
<emphasis id="791F40FB8494DCD6AE5697A0F717F7FD" italics="true" pageId="0" pageNumber="15">A. asper</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="674D8BC1BCCB5EDAD50F595D55A3872B" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. franzhuberi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="franzhuberi">
|
||
<emphasis id="A72B4959215A9CAC682B96FFEEE69684" italics="true" pageId="0" pageNumber="15">A. franzhuberi</emphasis>
|
||
</taxonomicName>
|
||
, they agree well with the type specimen of
|
||
<taxonomicName id="2BF6EF7ADCEE626E4A69DC2DE537ECCF" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. buelowi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="9D1CFD8BC7B24677C8A0FE3E58420E36" italics="true" pageId="0" pageNumber="15">A. buelowi</emphasis>
|
||
</taxonomicName>
|
||
in having the common diagnostic traits of a distinct twisted columella plait, a prominent umbilical hump, and a distinct apertural notch. The molecular phylogeny also revealed that all specimens from Mount Singgalang, West Sumatra, Indonesia, and Lang-Biang plateau and Nha Trang, Vietnam belong to the same clade. The mutational steps between Indonesian and Vietnamese specimens are only ten and one in the COI and 16S haplotype networks, respectively (Fig.
|
||
<figureCitation id="695199BCCDA0028A6A007EFCB4A32F05" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Mitochondrial haplotype minimum spanning networks of Amphidromus buelowi Fruhstorfer, 1905 A COI and B 16 S rRNA. The size of each circle corresponds to the frequency of that haplotype, also shown as the number in that circle. The bars on the branches indicate the number of mutational steps between haplotypes. Specimen codes correspond to those in Table 1." figureDoi="10.3897/zookeys.1196.112146.figure16" httpUri="https://binary.pensoft.net/fig/1007834" pageId="0" pageNumber="15">16</figureCitation>
|
||
). Based on the phylogenetic analyses and the common morphological diagnostics, we therefore treat
|
||
<taxonomicName id="22BD777530B09EC66CEE8C64CCACE9BE" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. asper" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asper">
|
||
<emphasis id="41092B874F88AD9200E4CB92AC7C67EB" italics="true" pageId="0" pageNumber="15">A. asper</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6A45A87437DE560420A70B33D47C5C98" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. franzhuberi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="franzhuberi">
|
||
<emphasis id="FE75EA04F84C3F6E6BF19C89BCD6F607" italics="true" pageId="0" pageNumber="15">A. franzhuberi</emphasis>
|
||
</taxonomicName>
|
||
as junior subjective synonyms of
|
||
<taxonomicName id="13C4898C0C5688067BF55AC507BB79E7" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. buelowi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="FF9905D4687860E607ADDD95C21FCD58" italics="true" pageId="0" pageNumber="15">A. buelowi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="175EA249AE34FD513C0BDF0B262D4719" pageId="0" pageNumber="15">
|
||
<bibRefCitation id="0910C734FB80B0E4B91090E418C810EE" author="Buelow, C" journalOrPublisher="Nachrichtsblatt der Deutschen Malakologischen Gesellschaft" pageId="0" pageNumber="15" pagination="78 - 83" publicationUrl="https://biostor.org/reference/190425" refId="B7" refString="Buelow, C, 1905. Einige Seltenheiten aus meiner Sammlung. Nachrichtsblatt der Deutschen Malakologischen Gesellschaft 37: 78 - 83, https://biostor.org/reference/190425" title="Einige Seltenheiten aus meiner Sammlung." url="https://biostor.org/reference/190425" volume="37" year="1905">
|
||
<normalizedToken id="A786B0667DC5650A0238EC0184B82658" originalValue="Bülow">Buelow</normalizedToken>
|
||
(1905)
|
||
</bibRefCitation>
|
||
introduced the monotypic subgenus
|
||
<taxonomicName id="48112867940AA1CAF2A4C858A63F212C" authorityName="Bülow" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus (Goniodromus)" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="subGenus" subGenus="Goniodromus">Amphidromus Goniodromus</taxonomicName>
|
||
to include
|
||
<taxonomicName id="EE1288A47D16C6F393FE32195074941B" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. buelowi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="928997B3BF125B2206722FCEEB047E6F" italics="true" pageId="0" pageNumber="15">A. buelowi</emphasis>
|
||
</taxonomicName>
|
||
, based on a less ovate aperture with an apertural notch projecting anteriorly. Later,
|
||
<bibRefCitation id="9D60580588B39F9FA04667C07696F246" DOI="https://doi.org/10.5962/bhl.title.2832" author="Laidlaw, FF" journalOrPublisher="Zoology" pageId="0" pageNumber="15" pagination="507 - 677" refId="B34" refString="Laidlaw, FF, Solem, A, 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana. Zoology 41: 507 - 677, DOI: https://doi.org/10.5962/bhl.title.2832" title="The land snail genus Amphidromus: A synoptic catalogue. Fieldiana." url="https://doi.org/10.5962/bhl.title.2832" volume="41" year="1961">Laidlaw and Solem (1961)</bibRefCitation>
|
||
, although with doubt, listed
|
||
<taxonomicName id="88B581B268B1F35B812C24A3A5DD2C1E" class="Gastropoda" family="Camaenidae" genus="Goniodromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Goniodromus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="genus">
|
||
<emphasis id="FEB93E3F4E26A359F935EE78F91E5680" italics="true" pageId="0" pageNumber="15">Goniodromus</emphasis>
|
||
</taxonomicName>
|
||
as one of the three subgenera of
|
||
<taxonomicName id="CBBC262283AD66DDF24FEE1BC18DE5E6" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="genus">
|
||
<emphasis id="92F1E5C9869A2A9B8ADBE582B52C8A05" italics="true" pageId="0" pageNumber="15">Amphidromus</emphasis>
|
||
</taxonomicName>
|
||
, and included two more species,
|
||
<taxonomicName id="4110362B613345A56E17ADEC17BFC089" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. asper" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asper">
|
||
<emphasis id="E1E375B7DD8D627068A8C340E03AF0F9" italics="true" pageId="0" pageNumber="15">A. asper</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="535EA9CA31BF669F11E886760555B29A" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. mirandus" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="mirandus">
|
||
<emphasis id="B8774D3ABA6F5EF08BF5656DAE39C61E" italics="true" pageId="0" pageNumber="15">A. mirandus</emphasis>
|
||
</taxonomicName>
|
||
Bavay & Dautzenberg, 1912. Another species,
|
||
<taxonomicName id="DD9A9DA871F5C3B3FBB56ABCE2C0B880" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. thachi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="thachi">
|
||
<emphasis id="26F7DDCE2C90CCE3416C06445DDF8E6E" italics="true" pageId="0" pageNumber="15">A. thachi</emphasis>
|
||
</taxonomicName>
|
||
, also possesses an aperture with prominent anterior notch (Fig.
|
||
<figureCitation id="4AE760778F3DE138916D6EE5E1099E31" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Shells of Amphidromus thachi Huber, 2015 A holotype of " Amphidromus thachi " (RBINS MT. 3381) B, C specimens from fin de la route de Hon Ba (chalets de Yersin), Commune de Suoi Cat, Province de Khanh Hoa, Vietnam (MNHN- IM- 214 - 6873) D specimen from Vinh Thanh, Binh Dinh, Vietnam (NMNS- 8764 - 267) E Specimen from Buon Don, Dak Lak, Vietnam (NMNS- 8764 - 271) F specimen from Da Lat, Lam Dong, Vietnam (NMNS- 8764 - 272) G, H specimens from Lac Duong, Lam Dong, Vietnam (NMNS- 8764 - 265, NMNS- 8764 - 264). Credit: T. Backeljau and S. Yves, RBINS (A), B. Pall-Gergely (B, C)." figureDoi="10.3897/zookeys.1196.112146.figure17" httpUri="https://binary.pensoft.net/fig/1007835" pageId="0" pageNumber="15">17</figureCitation>
|
||
). However, these three species,
|
||
<taxonomicName id="7D2C6F6950104BA67B0F2C5622D4E65F" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. buelowi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="buelowi">
|
||
<emphasis id="76F3B90D64A96C4BACD48F0D2363D4A1" italics="true" pageId="0" pageNumber="15">A. buelowi</emphasis>
|
||
</taxonomicName>
|
||
(and its synonyms
|
||
<taxonomicName id="252AEF5A31A554158C9B7B2FC9960BD0" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. asper" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="asper">
|
||
<emphasis id="665B78797767671201F76FF1E2F27716" italics="true" pageId="0" pageNumber="15">A. asper</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="224216B7C223D9923845948EC203CA8F" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. franzhuberi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="franzhuberi">
|
||
<emphasis id="5FD675851D1D54E733A26EB98834164B" italics="true" pageId="0" pageNumber="15">A. franzhuberi</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName id="1BA36C81B82324FE2DEB3796D185D471" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. thachi" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="thachi">
|
||
<emphasis id="52CFEBD5A1B418DBF7997AFFF0CA1C96" italics="true" pageId="0" pageNumber="15">A. thachi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="88B1FCAABCF3116D1DE486F126B24EBE" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. mirandus" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="mirandus">
|
||
<emphasis id="C969FAFE76B905294356493CA5E8AACF" italics="true" pageId="0" pageNumber="15">A. mirandus</emphasis>
|
||
</taxonomicName>
|
||
did not together form a clade (Fig.
|
||
<figureCitation id="58D1256F78FE0D322F7896479A1FAFEE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bayesian phylogeny of Amphidromus spp. based on mitochondrial COI and 16 S genes. Nodal support values are given as SH-aLRT / aBayes / ultra-fast bootstrap (IQ-TREE, ML) / posterior probability (MrBayes, BI). An asterisk on each branch indicates a clade with all well-supported values (SH-aLRT ≥ 80 %, aBayes ≥ 0.95, BS ≥ 95 %, PP ≥ 0.95). Species numbers correspond to those in Fig. 1 and Tables 1, 2." figureDoi="10.3897/zookeys.1196.112146.figure2" httpUri="https://binary.pensoft.net/fig/1007820" pageId="0" pageNumber="15">2</figureCitation>
|
||
; C-TL, unpublished data), revealing that an apertural anterior notch is not a shared derived character. Thus, the subgenus
|
||
<taxonomicName id="19950A450472C35FD8AF73934D57830B" authorityName="Bülow" authorityYear="1905" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus (Goniodromus)" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="subGenus" subGenus="Goniodromus">Amphidromus Goniodromus</taxonomicName>
|
||
is regarded herein as a junior subjective synonym of the subgenus
|
||
<taxonomicName id="A2908C60AF377C757C39F7E6F01AD5A2" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus (Amphidromus)" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="subGenus" subGenus="Amphidromus">Amphidromus Amphidromus</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="66DE68B701A4B28936DABA584A64C1EC" doi="10.3897/zookeys.1196.112146.figure17" httpUri="https://binary.pensoft.net/fig/1007835" pageId="0" pageNumber="15" start="Figure 17" startId="F17">
|
||
<paragraph id="27EA4457DDA72F30B44D2D15EC480EB5" pageId="0" pageNumber="15">
|
||
<emphasis id="4DD9F5AC1E34473516C82BC36737CD94" bold="true" pageId="0" pageNumber="15">Figure 17.</emphasis>
|
||
Shells of
|
||
<taxonomicName id="FCF97E2985E5079C2CD37E52D1E5854B" authorityName="Huber" authorityYear="2015" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus thachi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="thachi">
|
||
<emphasis id="7D5798BE031804B4364D205BEBC16ACA" italics="true" pageId="0" pageNumber="15">Amphidromus thachi</emphasis>
|
||
</taxonomicName>
|
||
Huber, 2015
|
||
<emphasis id="A379B13BECF2BD38EEFCEA60817CED96" bold="true" pageId="0" pageNumber="15">A</emphasis>
|
||
holotype of "
|
||
<taxonomicName id="DE535C14207DE0E073A5AF33B2EC5080" authorityName="Huber" authorityYear="2015" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" lsidName="Amphidromus thachi" order="Stylommatophora" pageId="0" pageNumber="15" phylum="Mollusca" rank="species" species="thachi">
|
||
<emphasis id="73606BD6CF8A64555C504BD5926D40FE" italics="true" pageId="0" pageNumber="15">Amphidromus thachi</emphasis>
|
||
</taxonomicName>
|
||
" (RBINS MT.3381)
|
||
<emphasis id="A5A5E9D0E6ECCEBB66136E055EF99B55" bold="true" pageId="0" pageNumber="15">B, C</emphasis>
|
||
specimens from fin de la route de Hon Ba (chalets de Yersin), Commune de Suoi Cat, Province de Khanh Hoa, Vietnam (MNHN- IM-214-6873)
|
||
<emphasis id="3E30F18F37BEDE9830898A44D16420D0" bold="true" pageId="0" pageNumber="15">D</emphasis>
|
||
specimen from Vinh Thanh, Binh Dinh, Vietnam (NMNS-8764-267)
|
||
<emphasis id="4D93E05A91B368E19A051E16A8017025" bold="true" pageId="0" pageNumber="15">E</emphasis>
|
||
Specimen from Buon Don, Dak Lak, Vietnam (NMNS-8764-271)
|
||
<emphasis id="3B5F85FCB30DC9852ED2ABA6ADA092A1" bold="true" pageId="0" pageNumber="15">F</emphasis>
|
||
specimen from Da Lat, Lam Dong, Vietnam (NMNS-8764-272)
|
||
<emphasis id="0A1AA5ECFD3326B8FDC4234BC8BB7234" bold="true" pageId="0" pageNumber="15">G, H</emphasis>
|
||
specimens from Lac Duong, Lam Dong, Vietnam (NMNS-8764-265, NMNS-8764-264). Credit: T. Backeljau and S. Yves, RBINS (
|
||
<emphasis id="0DAC962C3F1C490202A52D877325E9FA" bold="true" pageId="0" pageNumber="15">A</emphasis>
|
||
), B.
|
||
<normalizedToken id="9B0B8079C6FD818A3AF5887FE638DA0D" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
||
(
|
||
<emphasis id="4844AA6058646446CF567AE4071331B5" bold="true" pageId="0" pageNumber="15">B, C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |