337 lines
30 KiB
XML
337 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.100.1521" ID-GBIF-Dataset="7744cc07-4452-4e98-9e75-0657c53a4f73" ID-PMC="PMC3131010" ID-Pensoft-Pub="1313-2970-100-37" ID-PubMed="21738406" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-100-37" ModsDocOrigin="ZooKeys 100" ModsDocTitle="Eucamaragnathus desenderi, a new ground beetle species from Africa (Coleoptera, Carabidae)" checkinTime="1451250418697" checkinUser="pensoft" docAuthor="Assmann, Thorsten, Drees, Claudia, Matern, Andrea & Schuldt, Andreas" docDate="2011" docId="44D8FC5B4A415B5582040E0AB8C39222" docLanguage="en" docName="ZooKeys 100: 37-46" docOrigin="ZooKeys 100" docSource="http://dx.doi.org/10.3897/zookeys.100.1521" docTitle="Eucamaragnathus desenderi Assmann, Drees, Matern & Schuldt, sp. n." docType="treatment" docVersion="4" lastPageNumber="42" masterDocId="FFF9C015FFDCFFE15D677E0CFFF0FFAB" masterDocTitle="Eucamaragnathus desenderi, a new ground beetle species from Africa (Coleoptera, Carabidae)" masterLastPageNumber="46" masterPageNumber="37" pageNumber="38" updateTime="1668146586973" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Eucamaragnathus desenderi, a new ground beetle species from Africa (Coleoptera, Carabidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Assmann, Thorsten</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Drees, Claudia</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Matern, Andrea</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schuldt, Andreas</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>100</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>37</mods:start>
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<mods:end>46</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.100.1521</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.100.1521</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-100-37</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152029416" LSID="urn:lsid:zoobank.org:act:7F180D34-C5F8-4D79-AC05-E2F727CAD2E3" httpUri="http://treatment.plazi.org/id/44D8FC5B4A415B5582040E0AB8C39222" lastPageId="5" lastPageNumber="42" pageId="1" pageNumber="38">
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<subSubSection pageId="1" pageNumber="38" type="nomenclature">
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<paragraph pageId="1" pageNumber="38">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:7F180D34-C5F8-4D79-AC05-E2F727CAD2E3" authority="Assmann, Drees, Matern & Schuldt" class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi Assmann, Drees, Matern & Schuldt</taxonomicName>
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<taxonomicNameLabel pageId="1" pageNumber="38">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="38" type="type material">
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<paragraph pageId="1" pageNumber="38">Type material:</paragraph>
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<paragraph pageId="1" pageNumber="38">
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Holotype male: "ZAMBIA NE. 2004 / 240 km SE Mansa / 25 km SE Mukuku / 29.11.
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<normalizedToken originalValue="Snižek">Snizek</normalizedToken>
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,
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<normalizedToken originalValue="Tichý”">Tichy"</normalizedToken>
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(CAS). Paratypes: 13 males and 8 females, same as holotype (CAS, CFA, CST, CSH, CSS, CWR). 2 males and 4 females: "RSA, NW prov. 2001 / Klerksdorp, 20 km W / of Bothaville, Vaal riv. / M.
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<normalizedToken originalValue="Snižek">Snizek</normalizedToken>
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lgt. 12.1." (CAS, CWR).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="38" type="diagnosis">
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<paragraph pageId="1" pageNumber="38">Diagnosis:</paragraph>
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<paragraph pageId="1" pageNumber="38">
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A macropterous species of average size for the
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus castelnaui" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="castelnaui">Eucamaragnathus castelnaui</taxonomicName>
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group, black, pronotum transverse, sides sinuate with posterior angles acute, transverse anterior impression punctulate, transverse posterior impression strongly punctate, elytral striae continued to apex. Habitus see Fig. 1.
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</paragraph>
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<caption pageId="1" pageNumber="38">
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<paragraph pageId="1" pageNumber="38">
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Figure 1.
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi</taxonomicName>
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sp. n., habitus; holotype.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38" type="description">
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<paragraph pageId="1" pageNumber="38">Description:</paragraph>
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<paragraph pageId="1" pageNumber="38">Body length 8.8 - 10.6 mm; width 3.6 - 4.0 mm (holotype 10 mm and 3.8 mm, respectively).</paragraph>
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<subSection pageId="1" pageNumber="38" type="colour">
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<paragraph pageId="1" pageNumber="38">Colour:</paragraph>
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<paragraph pageId="1" pageNumber="38">Black, without iridescence, not metallic; mandibels, mouth-parts, antennae, and tarsi partly infuscate.</paragraph>
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</subSection>
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<subSection pageId="1" pageNumber="38" type="head">
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<paragraph pageId="1" pageNumber="38">Head</paragraph>
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<paragraph pageId="1" pageNumber="38">(Figs 1 and 2) large, about one fourth less wide than pronotum (HW: 2.0 - 2.4 mm, holotype: 2.3 mm; ratio HW/PW: 0.75 - 0.78). Eyes fairly large, their diameter (seen in dorsal view) about four tenth of head width; protected posteriorly by lateral extension of the cranium. Antennae robust, scape longer than the following 4 antennomeres, antennomeres 5 - 11 with dense and fairly fine setae. Mesal edge of mandibles markedly serrate (mandible teeth triangular shaped). Two pairs of supraorbital furrows. Frons not punctate, except basal close to pronotal anterior margin.</paragraph>
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<caption pageId="1" pageNumber="38">
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<paragraph pageId="1" pageNumber="38">
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Figure 2.
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi</taxonomicName>
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sp. n., basal part of head, pronotum, basal part of elytra; holotype.
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</paragraph>
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</caption>
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</subSection>
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<subSection lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38" type="pronotum">
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<paragraph pageId="1" pageNumber="38">Pronotum</paragraph>
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<paragraph lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38">
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(Fig. 2) transverse (PW: 2.6 - 3.1 mm, holotype: 3.0 mm; PL: 1.9 - 2.2, holotype: 2.0 mm), widest prior to middle (basally of lateral seta). Pronotum at the base broader than at the apex (PAW: 2.2 - 2.6 mm, holotype: 2.5 mm; PBW: 2.3 - 2.8 mm, holotype: 2.7 mm). Anterior margin moderately straight; anterior angles pronounced, but rounded; lateral sides clearly sinuate; posterior angles acute, basal
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<pageBreakToken pageId="2" pageNumber="39" start="start">margin</pageBreakToken>
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curved. Anterior transverse impression sparsely punctulate; lateral beads deep, not punctate; basal transverse impression deep, markedly punctate and connecting basal foveae; basal foveae deep, punctate and delimited externally by a keel-like carina without punctations.
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</paragraph>
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</subSection>
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<subSection pageId="2" pageNumber="39" type="legs">
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<paragraph pageId="2" pageNumber="39">Legs</paragraph>
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<paragraph pageId="2" pageNumber="39">
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(Fig. 1) similar to those found in other
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<taxonomicName lsidName="" pageId="2" pageNumber="39" rank="tribe" tribe="Hiletini">Hiletini</taxonomicName>
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species. Males with small tooth on profemur. Single long guard seta of tarsus 5 much longer than claws. Males with spatulate adhesive setae beneath protarsi 1 - 3 and mesotarsus 1.
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</paragraph>
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</subSection>
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<subSection pageId="2" pageNumber="39" type="elytra">
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<paragraph pageId="2" pageNumber="39">Elytra</paragraph>
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<paragraph pageId="2" pageNumber="39">(Figs 1 and 3) with pronounced humeri, slightly enlarged to the end of the second third (EL: 4.8 - 5.9 mm, holotype: 5.75 mm; EW: 3.3 - 3.9 mm, holotype: 3.7 mm). Basal margin reduced, reaching 6th interval. Scutellar striae short; elytral striae deep and punctate, at the apex less impressed, but well visible; intervals flat, at the apex slightly convex. Discal setae of third stria in punctiform depressions.</paragraph>
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<caption pageId="2" pageNumber="39">
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<paragraph pageId="2" pageNumber="39">
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Figure 3.
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi</taxonomicName>
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sp. n., apex of elytra; paratype.
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</paragraph>
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</caption>
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</subSection>
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<subSection pageId="2" pageNumber="39" type="surface">
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<paragraph pageId="2" pageNumber="39">Surface</paragraph>
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<paragraph pageId="2" pageNumber="39">
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with microsculpture of irregular and weak mesh patterns, meshes mainly transverse; a clear micropunctation on head, pronotum and elytra (20
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification); surface shiny.
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</paragraph>
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</subSection>
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<subSection pageId="2" pageNumber="39" type="male genitalia">
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<paragraph pageId="2" pageNumber="39">Male genitalia</paragraph>
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<paragraph pageId="2" pageNumber="39">(Figs 4 and 5). Median lobe with ostium dextral. Both parameres multisetiferous, the setae of the narrow right paramere are longer than those of the broad left one.</paragraph>
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<caption pageId="2" pageNumber="39">
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<paragraph pageId="2" pageNumber="39">
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Figure 4.
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi</taxonomicName>
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sp. n., male genitalia, left lateral aspect of median lobe (aedeagus); paratype.
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</paragraph>
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</caption>
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<caption pageId="2" pageNumber="39">
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<paragraph pageId="2" pageNumber="39">
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Figure 5.
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi</taxonomicName>
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sp. n., male genitalia, ventral aspect of parameres; paratype.
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</paragraph>
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</caption>
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</subSection>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="40" pageId="2" pageNumber="39" type="comparisons">
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<paragraph pageId="2" pageNumber="39">Comparisons:</paragraph>
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<paragraph lastPageId="3" lastPageNumber="40" pageId="2" pageNumber="39">
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Due to form of mandible teeth and long single guard seta of last tarsomere the new species belongs to the genus
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">Eucamaragnathus</taxonomicName>
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Jeannel, 1937. The small tooth of profemora in males, the dextral position of the ostium of the aed
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<pageBreakToken pageId="3" pageNumber="40" start="start">eagus</pageBreakToken>
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and elytral striae continued to the apex place the new species in the
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus castelnaui" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="castelnaui">Eucamaragnathus castelnaui</taxonomicName>
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(
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<normalizedToken originalValue="Bocandé">Bocande</normalizedToken>
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, 1849) group (cf.
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<bibRefCitation author="Erwin, TL" journalOrPublisher="Systematic Entomology" pageId="6" pageNumber="43" pagination="405 - 451" title="The Hiletini, an ancient and enigmatic tribe of Carabidae with a pantropical distribution (Coleoptera)." url="10.111/j.1365–3113.1985.tb00149.x" volume="10" year="1985">Erwin and Stork 1985</bibRefCitation>
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) which is exclusively distributed in Africa.
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</paragraph>
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<paragraph pageId="3" pageNumber="40">
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The new species is similar to
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus castelnaui" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="castelnaui">Eucamaragnathus castelnaui</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus fissipennis" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="fissipennis">Eucamaragnathus fissipennis</taxonomicName>
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(Ancey, 1882). The best character to separate
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi</taxonomicName>
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sp. n. from the nominate species of the group is the shape of the pronotum and especially the weak punctation of the pronotal anterior impression which is markedly punctate in
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus castelnaui" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="castelnaui">Eucamaragnathus castelnaui</taxonomicName>
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. In comparison to the other species of the group,
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi</taxonomicName>
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sp. n. has acute pronotal anterior angles, but they are less produced than in
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus oxygonus" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="oxygonus">Eucamaragnathus oxygonus</taxonomicName>
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Chaudoir, 1861. Moreover the median lobe, especially its internal sac structures, of
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi</taxonomicName>
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sp. n. differs from all other species of the given group. From
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus fissipennis" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="fissipennis">Eucamaragnathus fissipennis</taxonomicName>
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the new species can be easily distinguished by stronger punctation of posterior transverse impressions of pronotum (Figs 2 and 6), stronger punctation of elytral striae, which are weaker at the apex, but still well visible (Figs 3 and 7) and a microsculpture with stronger punctation.
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</paragraph>
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<caption pageId="3" pageNumber="40">
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<paragraph pageId="3" pageNumber="40">
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Figure 6.
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus fissipennis" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="fissipennis">Eucamaragnathus fissipennis</taxonomicName>
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, basal part of elytra.
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</paragraph>
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</caption>
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<paragraph pageId="3" pageNumber="40">
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From
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus bocandei" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="bocandei">Eucamaragnathus bocandei</taxonomicName>
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(Alluaud, 1914), which forms an own species group, the new species differs by its strong punctation of pronotal posterior impression and from
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus suberbiei" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="suberbiei">Eucamaragnathus suberbiei</taxonomicName>
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(Alluaud, 1914) it can be separated by the size of tooth on ventral surface of profemur in males.
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</paragraph>
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<paragraph pageId="3" pageNumber="40">
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For better distinction we present an identification key for the known members of the African
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="genus">Eucamaragnathus</taxonomicName>
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species (see below).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="40" type="etymology">
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<paragraph pageId="3" pageNumber="40">Etymology:</paragraph>
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<paragraph pageId="3" pageNumber="40">
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It gives us great pleasure to dedicate this species to the memory of Konjev Desender, the well known Belgian carabidologist who recently deceased. We had many scientific meetings, excursions and productive collaborations with him, and we will honor his memory. An obituary is given by
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<bibRefCitation author="Loevei, GL" journalOrPublisher="ZooKeys" pageId="6" pageNumber="43" pagination="1 - 36" title="In memoriam: Konjev Desender (1956 - 2008) and Jean-Pierre Maelfait (1950 - 2009). In: Kotze DJ, Assmann T, Noordijk J, Turin H, Vermeulen R (Eds) Carabid Beetles as Bioindicators: Biogeographical, Ecological and Environmental Studies." volume="100" year="2011">
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<normalizedToken originalValue="Lövei">Loevei</normalizedToken>
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(2011)
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</bibRefCitation>
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including a list of his publications.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40" type="distribution">
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<paragraph pageId="3" pageNumber="40">Distribution:</paragraph>
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<paragraph lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40">
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Up to now
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<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi</taxonomicName>
|
||
sp. n. is only known from the two sites in Zambia and South Africa. The population from Zambia (close to the border to Congo) lies in the tropical part of Africa fitting well to the main distribution area of the tribe in tropical Africa. In contrast, Bothaville in South Africa, the other site from where
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi</taxonomicName>
|
||
sp. n. is known, is located between the 27th and 28th degrees of southern latitude, doubtless in the subtropical realm, and seems to be the most southern known
|
||
<pageBreakToken pageId="4" pageNumber="41" start="start">record</pageBreakToken>
|
||
of a
|
||
<taxonomicName lsidName="" pageId="4" pageNumber="41" rank="tribe" tribe="Hiletini">Hiletini</taxonomicName>
|
||
species in Africa (and worldwide). The wide distribution of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi</taxonomicName>
|
||
sp. n. in Africa is not unusual for a
|
||
<taxonomicName lsidName="" pageId="4" pageNumber="41" rank="tribe" tribe="Hiletini">Hiletini</taxonomicName>
|
||
species (cf. the large distribution areas of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Hiletus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hiletus alluaudi" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="alluaudi">Hiletus alluaudi</taxonomicName>
|
||
(Jeannel, 1937) and
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus fissipennis" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="fissipennis">Eucamaragnathus fissipennis</taxonomicName>
|
||
,
|
||
<bibRefCitation author="Erwin, TL" journalOrPublisher="Systematic Entomology" pageId="6" pageNumber="43" pagination="405 - 451" title="The Hiletini, an ancient and enigmatic tribe of Carabidae with a pantropical distribution (Coleoptera)." url="10.111/j.1365–3113.1985.tb00149.x" volume="10" year="1985">Erwin and Stork 1985</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="4" pageNumber="41">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi</taxonomicName>
|
||
sp. n. seems to co-occur with
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus fissipennis" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="fissipennis">Eucamaragnathus fissipennis</taxonomicName>
|
||
which is distributed in tropical East Africa and south-eastern Africa.
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus oxygonus" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="oxygonus">Eucamaragnathus oxygonus</taxonomicName>
|
||
is known only from one locality in South Africa. All other African species of the genus
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus">Eucamaragnathus</taxonomicName>
|
||
show - so far known - an allopatric distribution (
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus suberbiei" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="suberbiei">Eucamaragnathus suberbiei</taxonomicName>
|
||
is an endemic of Madagascar,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus castelnaui" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="castelnaui">Eucamaragnathus castelnaui</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus bocandei" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="bocandei">Eucamaragnathus bocandei</taxonomicName>
|
||
occur exclusively in tropical western Africa,
|
||
<bibRefCitation author="Erwin, TL" journalOrPublisher="Systematic Entomology" pageId="6" pageNumber="43" pagination="405 - 451" title="The Hiletini, an ancient and enigmatic tribe of Carabidae with a pantropical distribution (Coleoptera)." url="10.111/j.1365–3113.1985.tb00149.x" volume="10" year="1985">Erwin and Stork 1985</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="41" type="habitat">
|
||
<paragraph pageId="4" pageNumber="41">Habitat:</paragraph>
|
||
<paragraph pageId="4" pageNumber="41">
|
||
The specimens were caught at light and habitat preferences are therefore unknown. Together with the holotype of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi</taxonomicName>
|
||
sp. n., a single
|
||
<taxonomicName lsidName="" pageId="4" pageNumber="41" rank="tribe" tribe="Hiletini">Hiletini</taxonomicName>
|
||
specimen of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Hiletus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hiletus katanganus" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="katanganus">Hiletus katanganus</taxonomicName>
|
||
Basilewsky, 1948 has been found. We compared this specimen of the rarely recorded species with the type material preserved in the Africa Museum (collection of Basilewsky) and detected morphological differences. Without more material (especially males) it seems to be impossible to assign specimens conclusively to this species (see also the note in
|
||
<bibRefCitation author="Erwin, TL" journalOrPublisher="Systematic Entomology" pageId="6" pageNumber="43" pagination="405 - 451" title="The Hiletini, an ancient and enigmatic tribe of Carabidae with a pantropical distribution (Coleoptera)." url="10.111/j.1365–3113.1985.tb00149.x" volume="10" year="1985">Erwin and Stork 1985</bibRefCitation>
|
||
: 431).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="5" lastPageNumber="42" pageId="4" pageNumber="41" type="key to the african species of eucamaragnathus jeannel">
|
||
<paragraph pageId="4" pageNumber="41">
|
||
Key to the African species of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus">Eucamaragnathus</taxonomicName>
|
||
Jeannel
|
||
</paragraph>
|
||
<paragraph pageId="4" pageNumber="41">This new identification key is based on the one presented by Erwin & Stork (1985), but it is modified and illustrated additionally.</paragraph>
|
||
<paragraph lastPageId="5" lastPageNumber="42" pageId="4" pageNumber="41">
|
||
<table lastPageId="5" lastPageNumber="42" pageId="4" pageNumber="41">
|
||
<tr pageId="4" pageNumber="41">
|
||
<td colspan="1" pageId="4" pageNumber="41" rowspan="1">Fig. 7</td>
|
||
<td colspan="1" pageId="4" pageNumber="41" rowspan="1">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus fissipennis" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="fissipennis">Eucamaragnathus fissipennis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="42">
|
||
<td colspan="1" pageId="5" pageNumber="42" rowspan="1">
|
||
<pageBreakToken pageId="5" pageNumber="42" start="start">Fig</pageBreakToken>
|
||
. 3
|
||
</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="42">
|
||
<td colspan="1" pageId="5" pageNumber="42" rowspan="1">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus suberbiei" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="suberbiei">Eucamaragnathus suberbiei</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="42">
|
||
<td colspan="1" pageId="5" pageNumber="42" rowspan="1">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus oxygonus" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="oxygonus">Eucamaragnathus oxygonus</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="42">
|
||
<td colspan="1" pageId="5" pageNumber="42" rowspan="1">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus bocandei" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="bocandei">Eucamaragnathus bocandei</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="42">
|
||
<td colspan="1" pageId="5" pageNumber="42" rowspan="1">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus castelnaui" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="castelnaui">Eucamaragnathus castelnaui</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="42">
|
||
<td colspan="1" pageId="5" pageNumber="42" rowspan="1">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus desenderi" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="desenderi">Eucamaragnathus desenderi</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption pageId="5" pageNumber="42">
|
||
<paragraph pageId="5" pageNumber="42">
|
||
Figure 7.
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Eucamaragnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucamaragnathus fissipennis" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="fissipennis">Eucamaragnathus fissipennis</taxonomicName>
|
||
, apex of elytra.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |