treatments-xml/data/41/B2/3F/41B23F849F8070D50279E4597A075F83.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.168.51904" ID-GBIF-Dataset="e7345e6b-fa55-474e-9b6e-d6b185121541" ID-PMC="PMC7718216" ID-Pensoft-Pub="1314-2003-168-1" ID-Pensoft-UUID="5F39D34A0DEF5952A2C4E9090C14B498" ID-PubMed="33335445" ModsDocID="1314-2003-168-1" checkinTime="1606572050664" checkinUser="pensoft" docAuthor="Dean, Ellen, Poore, Jennifer, Anguiano-Constante, Marco Antonio, Nee, Michael H., Kang, Hannah, Starbuck, Thomas, Rodrigues, Annamarie &amp; Conner, Matthew" docDate="2020" docId="41B23F849F8070D50279E4597A075F83" docLanguage="en" docName="PhytoKeys 168: 1-333" docOrigin="PhytoKeys 168" docSource="http://dx.doi.org/10.3897/phytokeys.168.51904" docTitle="Lycianthes quichensis (J. M. Coult. &amp; Donn. Sm.) Bitter, Abh. Naturwiss. Verein Bremen 24 [preprint]: 428. 1919" docType="treatment" docVersion="4" id="5F39D34A0DEF5952A2C4E9090C14B498" lastPageNumber="1" masterDocId="5F39D34A0DEF5952A2C4E9090C14B498" masterDocTitle="The genus Lycianthes (Solanaceae, Capsiceae) in Mexico and Guatemala" masterLastPageNumber="333" masterPageNumber="1" pageNumber="1" updateTime="1668140149145" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The genus Lycianthes (Solanaceae, Capsiceae) in Mexico and Guatemala</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dean, Ellen</mods:namePart>
<mods:affiliation>UC Davis Center for Plant Diversity, Plant Sciences M. S. 7, One Shields Ave., Davis, CA 95616, USA</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5986-0027</mods:nameIdentifier>
<mods:nameIdentifier type="email">eadean@ucdavis.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Poore, Jennifer</mods:namePart>
<mods:affiliation>UC Davis Center for Plant Diversity, Plant Sciences M. S. 7, One Shields Ave., Davis, CA 95616, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Anguiano-Constante, Marco Antonio</mods:namePart>
<mods:affiliation>Laboratorio Nacional de Identificacion y Caracterizacion Vegetal (LaniVeg), Consejo Nacional de Ciencia y Tecnologia (CONACyT), Centro Universitario de Ciencias Biologicas y Agropecuarias, Universidad de Guadalajara, Camino Ramon Padilla Sanchez 2100, 45110 Nextipac, Zapopan, Jalisco, Mexico</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-4071-8108</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nee, Michael H.</mods:namePart>
<mods:affiliation>26776 US Hwy 14, Richland Center, WI 53581, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kang, Hannah</mods:namePart>
<mods:affiliation>UC Davis Center for Plant Diversity, Plant Sciences M. S. 7, One Shields Ave., Davis, CA 95616, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Starbuck, Thomas</mods:namePart>
<mods:affiliation>UC Davis Center for Plant Diversity, Plant Sciences M. S. 7, One Shields Ave., Davis, CA 95616, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rodrigues, Annamarie</mods:namePart>
<mods:affiliation>UC Davis Center for Plant Diversity, Plant Sciences M. S. 7, One Shields Ave., Davis, CA 95616, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Conner, Matthew</mods:namePart>
<mods:affiliation>UC Davis Center for Plant Diversity, Plant Sciences M. S. 7, One Shields Ave., Davis, CA 95616, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>168</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>333</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.168.51904</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.168.51904</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-168-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">5F39D34A0DEF5952A2C4E9090C14B498</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="170596548" LSID="urn:lsid:plazi:treatment:41B23F849F8070D50279E4597A075F83" httpUri="http://treatment.plazi.org/id/41B23F849F8070D50279E4597A075F83" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
38 
<taxonomicName authority="(J. M. Coult. &amp; Donn. Sm.) Bitter, Abh. Naturwiss. Verein Bremen 24 [preprint]: 428. 1919" authorityName="(J. M. Coult. &amp; Donn. Sm.) Bitter, Abh. Naturwiss. Verein Bremen 24 [preprint]: 428." authorityYear="1919" class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes quichensis" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="quichensis">Lycianthes quichensis (J.M.Coult. &amp; Donn.Sm.) Bitter, Abh. Naturwiss. Verein Bremen 24 [preprint]: 428. 1919</taxonomicName>
<figureCitation captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Image of herbarium specimen of L. quichensis, Breedlove 40387 (NY). Specimen used with permission from the William and Lynda Steere Herbarium, New York Botanical Garden." figureDoi="10.3897/phytokeys.168.51904.figure86" httpUri="https://binary.pensoft.net/fig/480100" pageId="0" pageNumber="1">Fig. 86</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum quichense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="quichense">Solanum quichense</taxonomicName>
J.M.Coult. &amp; Donn.Sm., Bot. Gaz. 37: 422. 1904. Type: Guatemala. 
<normalizedToken originalValue="Quiché">Quiche</normalizedToken>
: Chiul, Apr 1892, 
<emphasis italics="true" pageId="0" pageNumber="1">E. T. Heyde &amp; E. Lux 3450</emphasis>
(holotype: F [0073142F, acc. # 264727]; isotypes: G [G00379126], GH [GH00077532], M [M-0171931], NY [00139026], US [00624010], US [00027769].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Standl" authorityYear="1940" class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes obliquifolia" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="obliquifolia">Lycianthes obliquifolia</taxonomicName>
Standl., Field Mus. Nat. Hist., Bot. Ser. 22: 101. 1940. Type: 
<normalizedToken originalValue="México">Mexico</normalizedToken>
. Chiapas: 
<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
de 
<normalizedToken originalValue="Tacaná">Tacana</normalizedToken>
, 30 Mar 1939, 
<emphasis italics="true" pageId="0" pageNumber="1">E. Matuda 2938</emphasis>
(holotype: F [0072919F, acc. # 980247]; isotypes: A [00077523], MEXU [MEXU00082111], MICH [1109852], NY [00138706]).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Type.</paragraph>
<paragraph pageId="0" pageNumber="1">
Based on 
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum quichense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="quichense">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum quichense</emphasis>
</taxonomicName>
Coult. &amp; Donn.Sm.
</paragraph>
<caption doi="10.3897/phytokeys.168.51904.figure86" httpUri="https://binary.pensoft.net/fig/480100" pageId="0" pageNumber="1" start="Figure 86" startId="F86">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 86.</emphasis>
Image of herbarium specimen of 
<taxonomicName lsidName="L. quichensis" pageId="0" pageNumber="1" rank="species" species="quichensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. quichensis</emphasis>
</taxonomicName>
, 
<emphasis italics="true" pageId="0" pageNumber="1">Breedlove 40387</emphasis>
(NY). Specimen used with permission from the William and Lynda Steere Herbarium, New York Botanical Garden.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Shrub, 1-5 m tall. Indument of white to pale yellow, uniseriate, multicellular, simple, acute, eglandular, appressed to spreading trichomes 0.1-1 mm long, the larger trichomes becoming flattened upon drying. Stems green when young, sparsely to densely pubescent, not much compressed upon drying in a plant press, brown and woody with age; upper sympodial branching points mostly monochasial, some dichasial. Leaves simple, the leaves of the upper sympodia usually paired and unequal in size, the larger ones with blades 4-15 (25) 
<normalizedToken originalValue="×">x</normalizedToken>
2.5-9 (14) cm, ovate to elliptic (sometimes very widely so), rarely lanceolate, the smaller ones with blades 1.5-9 
<normalizedToken originalValue="×">x</normalizedToken>
1-5 cm, ovate, the blades of both the large and small leaves chartaceous, sparsely to moderately pubescent, denser along the veins and in the abaxial leaf axils, the base cuneate to truncate or rounded, sometimes oblique, the margin entire, usually undulate, the apex acute to acuminate, the petiole to 4 (6) cm long, sometimes absent, the larger leaf blades with 4-7 primary veins on each side of the midvein. Flowers solitary or in groups of 2, axillary, oriented horizontally; peduncles absent; pedicels 20-60 mm long, erect to arching in flower, to 70 mm long (probably longer), deflexed and arching in fruit, sparsely to densely pubescent; calyx 3-4 mm long, 3.5-6 mm in diameter, campanulate, often purplish in color, sparsely to densely pubescent, the margin truncate, with 10 spreading, linear-subulate appendages 3-6 mm long emerging 0.5 mm below the calyx rim; fruiting calyx enlarged, widely bowl-shaped, 2-5 mm long, 7-13 mm in diameter, the appendages not greatly enlarging; corolla 1.2-3 cm long, campanulate to rotate in orientation, entire to shallowly stellate in outline, with abundant interpetalar tissue, adaxially light purple with darker purple ring near the base and green markings at very base, glabrous, abaxially light purple, sometimes paler or white on the lobes, sparsely to densely pubescent with small trichomes near the veins; stamens equal, straight, the filaments 1.5-2 mm long, glabrous, the anthers 5-6 mm long, lanceolate, free of one another, purple, glabrous, poricidal at the tips, the pores ovate, dehiscing toward the style, not opening into longitudinal slits; pistil with glabrous ovary, the style 10-12 mm long, linear, glabrous, the stigma capitate. Fruit a berry, 9-17 mm long, 7-15 mm in diameter, usually ovoid (round), red at maturity (drying dark purple on herbarium sheets), glabrous, lacking sclerotic granules. Seeds 20-40 per fruit, 2.5-3 
<normalizedToken originalValue="×">x</normalizedToken>
2-2.5 mm, compressed but not flat, ridged on one side or near the center, irregular in outline (shallowly crescent-shaped, triangular, or semi-circular), orange-brown, the surface reticulum with a serpentine to honeycomb pattern with deep luminae, appearing pitted, with fibrils protruding from the cell walls.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="chromosome number">
<paragraph pageId="0" pageNumber="1">Chromosome number.</paragraph>
<paragraph pageId="0" pageNumber="1">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">
Mexico (Chiapas) and Guatemala (Chimaltenango, Huehuetenango, Quetzaltenango, 
<normalizedToken originalValue="Quiché">Quiche</normalizedToken>
, 
<normalizedToken originalValue="Sacatepéquez">Sacatepequez</normalizedToken>
, San Marcos, 
<normalizedToken originalValue="Sololá">Solola</normalizedToken>
, 
<normalizedToken originalValue="Totonicapán">Totonicapan</normalizedToken>
), tropical moist forests or thickets in cloud forest and oak-pine forest with 
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Chiranthodendron" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chiranthodendron" order="Malvales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Chiranthodendron</emphasis>
</taxonomicName>
, 
<taxonomicName class="Magnoliopsida" family="Symplocaceae" genus="Symplocos" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Symplocos" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Symplocos</emphasis>
</taxonomicName>
, 
<taxonomicName class="Magnoliopsida" family="Winteraceae" genus="Drimys" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drimys" order="Canellales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Drimys</emphasis>
</taxonomicName>
, and 
<taxonomicName class="Magnoliopsida" family="Clethraceae" genus="Clethra" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Clethra" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Clethra</emphasis>
</taxonomicName>
(in Guatemala often in 
<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Cupressus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cupressus" order="Pinales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Cupressus</emphasis>
</taxonomicName>
or 
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Abies" order="Pinales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Abies</emphasis>
</taxonomicName>
forests), 2200-3900 m in elevation (Fig. 
<figureCitation captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Map of geographic distribution of L. quichensis based on herbarium specimen data." figureDoi="10.3897/phytokeys.168.51904.figure87" httpUri="https://binary.pensoft.net/fig/480101" pageId="0" pageNumber="1">87</figureCitation>
).
</paragraph>
<caption doi="10.3897/phytokeys.168.51904.figure87" httpUri="https://binary.pensoft.net/fig/480101" pageId="0" pageNumber="1" start="Figure 87" startId="F87">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 87.</emphasis>
Map of geographic distribution of 
<taxonomicName lsidName="L. quichensis" pageId="0" pageNumber="1" rank="species" species="quichensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. quichensis</emphasis>
</taxonomicName>
based on herbarium specimen data.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="common names and uses">
<paragraph pageId="0" pageNumber="1">Common names and uses.</paragraph>
<paragraph pageId="0" pageNumber="1">
Guatemala. Chilete, choshel, coxel, flor de rosa, quilete, tomatillo blanco (
<bibRefCitation author="Gentry, Jr JL" journalOrPublisher="Taxon" pageId="0" pageNumber="1" refId="B56" refString="Gentry, Jr JL, Standley, PC, 1974. Flora of Guatemala. Fieldiana: Botany 24 (part 10) no. 1-2. Field Museum of Natural History, Chicago." title="Flora of Guatemala. Fieldiana: Botany 24 (part 10) no. 1 - 2. Field Museum of Natural History, Chicago." year="1974">Gentry and Standley 1974</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="phenology">
<paragraph pageId="0" pageNumber="1">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="1">Flowering specimens and specimens with mature fruits have been collected during most months of the year. Many herbarium specimens of this species have flowers with open corollas, indicating that the flowers must stay open until at least noon, if not after.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="preliminary conservation status">
<paragraph pageId="0" pageNumber="1">Preliminary conservation status.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes quichensis" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="quichensis">
<emphasis italics="true" pageId="0" pageNumber="1">Lycianthes quichensis</emphasis>
</taxonomicName>
is a rare species of Mexico and Guatemala, represented by 38 collections and occurring in seven protected areas. The EOO is 22,627.703 km2, and the AOO is 148 km2. Based on the 
<bibRefCitation author="IUCN" journalOrPublisher="The Linnean Society of London, Academic Press, London" pageId="0" pageNumber="1" publicationUrl="http://jr.iucnredlist.org/documents/redlistGuidelines" refId="B67" refString="IUCN, 2019. Guidelines for using the IUCN red list categories and criteria. version 12. Prepared by the Standards and Petitions Subcommittee in February 2019. http://jr.iucnredlist.org/documents/redlistGuidelines" title="Guidelines for using the IUCN red list categories and criteria. version 12. Prepared by the Standards and Petitions Subcommittee in February 2019." url="http://jr.iucnredlist.org/documents/redlistGuidelines" year="2019">IUCN (2019)</bibRefCitation>
criteria, the preliminary assessment category is Endangered (EN).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="discussion">
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycianthes quichensis" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="quichensis">
<emphasis italics="true" pageId="0" pageNumber="1">Lycianthes quichensis</emphasis>
</taxonomicName>
is an attractive shrub that horticulturalists are trying to bring into cultivation and is sometimes available in seed catalogues. It has relatively large purple flowers with distinctive dark purple and green markings and widely ovate leaves. It has sometimes been confused with 
<taxonomicName lsidName="L. pilifera" pageId="0" pageNumber="1" rank="species" species="pilifera">
<emphasis italics="true" pageId="0" pageNumber="1">L. pilifera</emphasis>
</taxonomicName>
, a species of Oaxaca, Mexico. The two species differ in fruit color (
<taxonomicName lsidName="L. pilifera" pageId="0" pageNumber="1" rank="species" species="pilifera">
<emphasis italics="true" pageId="0" pageNumber="1">L. pilifera</emphasis>
</taxonomicName>
has purple-black fruits) and pubescence (the trichomes of 
<taxonomicName lsidName="L. pilifera" pageId="0" pageNumber="1" rank="species" species="pilifera">
<emphasis italics="true" pageId="0" pageNumber="1">L. pilifera</emphasis>
</taxonomicName>
are darker and more tubular-conical upon drying than those of 
<taxonomicName lsidName="L. quichensis" pageId="0" pageNumber="1" rank="species" species="quichensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. quichensis</emphasis>
</taxonomicName>
which are pale yellow to white and flattened upon drying) (
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.425.3.6" author="Dean, E" journalOrPublisher="Phytologia" pageId="0" pageNumber="1" refId="B44" refString="Dean, E, McNair, DM, Castillo-Campos, G, Starbuck, T, Aguiano-Constante, MA, Mawdsley, K, Veliz-Perez, M, Archila, F, 2019b. Identification of the Mexican species of Lycianthes series Piliferae (Capsiceae, Solanaceae) and the rediscovery of Lycianthescaeciliae. Phytotaxa 425(3): 163-189. https://doi.org/10.11646/phytotaxa.425.3.6" title="Identification of the Mexican species of Lycianthes series Piliferae (Capsiceae, Solanaceae) and the rediscovery of Lycianthescaeciliae. Phytotaxa 425 (3): 163 - 189." url="https://doi.org/10.11646/phytotaxa.425.3.6" year="2019 b">Dean et al. 2019b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Representative specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Guatemala. Chimaltenango</emphasis>
: 
<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
de Acatenango, [
<geoCoordinate degrees="14.5255" direction="north" orientation="latitude" precision="5" value="14.5255">14.5255</geoCoordinate>
, 
<geoCoordinate degrees="90.8753" direction="west" orientation="longitude" precision="5" value="-90.8753">-90.8753</geoCoordinate>
], 2800 m, 22 Apr 1999, 
<emphasis italics="true" pageId="0" pageNumber="1">
M. 
<normalizedToken originalValue="Véliz">Veliz</normalizedToken>
7020
</emphasis>
(BIGU, MEXU). 
<emphasis bold="true" pageId="0" pageNumber="1">Huehuetenango</emphasis>
: Mpio. La Libertad, 
<normalizedToken originalValue="Peña">Pena</normalizedToken>
Blanca, 
<geoCoordinate degrees="15.5075" direction="north" orientation="latitude" precision="5" value="15.5075">15.5075</geoCoordinate>
, 
<geoCoordinate degrees="91.9158" direction="west" orientation="longitude" precision="5" value="-91.9158">-91.9158</geoCoordinate>
, 3193 m, 14 Dec 2000, 
<emphasis italics="true" pageId="0" pageNumber="1">
M. 
<normalizedToken originalValue="Véliz">Veliz</normalizedToken>
10846
</emphasis>
(BIGU, CAS). 
<emphasis bold="true" pageId="0" pageNumber="1">Quetzaltenango</emphasis>
: Cumbre de Alaska, [
<geoCoordinate degrees="14.7560" direction="north" orientation="latitude" precision="5" value="14.756">14.7560</geoCoordinate>
, 
<geoCoordinate degrees="91.4705" direction="west" orientation="longitude" precision="5" value="-91.4705">-91.4705</geoCoordinate>
], 3100 m, 10 Sep 1999, 
<emphasis italics="true" pageId="0" pageNumber="1">
M. 
<normalizedToken originalValue="Véliz">Veliz</normalizedToken>
7285
</emphasis>
(BIGU, MEXU). 
<emphasis bold="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="Quiché">Quiche</normalizedToken>
</emphasis>
: Chiul, Apr 1892, 
<emphasis italics="true" pageId="0" pageNumber="1">E.T. Heyde &amp; E. Lux 3450</emphasis>
(F, GH, M, NY, US). 
<emphasis bold="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="Sacatepéquez">Sacatepequez</normalizedToken>
</emphasis>
: 
<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
de Agua, 
<geoCoordinate degrees="14.4769" direction="north" orientation="latitude" precision="5" value="14.4769">14.4769</geoCoordinate>
, 
<geoCoordinate degrees="90.7322" direction="west" orientation="longitude" precision="5" value="-90.7322">-90.7322</geoCoordinate>
, 2852 m, 6 Feb 2006, 
<emphasis italics="true" pageId="0" pageNumber="1">
M. 
<normalizedToken originalValue="Véliz">Veliz</normalizedToken>
16671
</emphasis>
(BIGU, TEX). 
<emphasis bold="true" pageId="0" pageNumber="1">San Marcos</emphasis>
: Mpio. San 
<normalizedToken originalValue="José">Jose</normalizedToken>
Ojetenam, [
<geoCoordinate degrees="15.2342" direction="north" orientation="latitude" precision="5" value="15.2342">15.2342</geoCoordinate>
, 
<geoCoordinate degrees="91.9736" direction="west" orientation="longitude" precision="5" value="-91.9736">-91.9736</geoCoordinate>
], 3100 m, 26 Nov 2009, 
<emphasis italics="true" pageId="0" pageNumber="1">
F. 
<normalizedToken originalValue="Pérez">Perez</normalizedToken>
18
</emphasis>
(BIGU). 
<emphasis bold="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="Sololá">Solola</normalizedToken>
</emphasis>
: San Pedro La Laguna, 
<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
San Pedro, ladera noreste del 
<normalizedToken originalValue="volcán">volcan</normalizedToken>
, 
<geoCoordinate degrees="14.6572" direction="north" orientation="latitude" precision="5" value="14.6572">14.6572</geoCoordinate>
, 
<geoCoordinate degrees="91.2664" direction="west" orientation="longitude" precision="5" value="-91.2664">-91.2664</geoCoordinate>
, 3006 m, 28 Jan 2005, 
<emphasis italics="true" pageId="0" pageNumber="1">P. Pardo 27</emphasis>
(CAS). 
<emphasis bold="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="Totonicapán">Totonicapan</normalizedToken>
</emphasis>
: 
<normalizedToken originalValue="María">Maria</normalizedToken>
<normalizedToken originalValue="Tecún">Tecun</normalizedToken>
, [
<geoCoordinate degrees="14.8341" direction="north" orientation="latitude" precision="5" value="14.8341">14.8341</geoCoordinate>
, 
<geoCoordinate degrees="91.2174" direction="west" orientation="longitude" precision="5" value="-91.2174">-91.2174</geoCoordinate>
], 3000-3600 m, 12--23 Jan 1966, 
<emphasis italics="true" pageId="0" pageNumber="1">A. Molina R. 16399</emphasis>
(CAS-DS, NY). 
<emphasis bold="true" pageId="0" pageNumber="1">Mexico. Chiapas</emphasis>
: Mpio. Chamula, Tzontehuitz, 
<geoCoordinate degrees="16.8108" direction="north" orientation="latitude" precision="5" value="16.8108">16.8108</geoCoordinate>
, 
<geoCoordinate degrees="92.5775" direction="west" orientation="longitude" precision="5" value="-92.5775">-92.5775</geoCoordinate>
, 2740 m, 20 Apr 1999, 
<emphasis italics="true" pageId="0" pageNumber="1">
L.Y. 
<normalizedToken originalValue="Domínguez-Torres">Dominguez-Torres</normalizedToken>
70
</emphasis>
(DAV, MEXU).
</paragraph>
</subSubSection>
</treatment>
</document>