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111 lines
25 KiB
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<document id="5DC16653146038310BB53000E9542FF8" ID-DOI="10.1080/14772019.2021.1978005" ID-ISSN="1478-0941" ID-ZooBank="31F0D48F-C1DA-406E-A811-1F5937ED19F4" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1711485377302" checkinUser="felipe" docAuthor="Lockwood, Jeremy A. F., Martill, David M. & Maidment, Susannah C. R." docDate="2021" docId="03FC4C3C6D63FFF486A8FE391FBE2CD7" docLanguage="en" docName="JSystPalaeontol.19.12.847-888.pdf" docOrigin="Journal of Systematic Palaeontology 19 (12)" docSource="http://dx.doi.org/10.1080/14772019.2021.1978005" docStyle="DocumentStyle:6400638016C1F4AE0D0B6502E032D571.1:JSystPalaeontol.2016-.journal_article.closed" docStyleId="6400638016C1F4AE0D0B6502E032D571" docStyleName="JSystPalaeontol.2016-.journal_article.closed" docStyleVersion="1" docTitle="Brighstoneus simmondsi Lockwood & Martill & Maidment 2021, gen. et sp. nov." docType="treatment" docVersion="2" lastPageNumber="851" masterDocId="FFC534446D66FFF1864DFF851C2C2D28" masterDocTitle="A new hadrosauriform dinosaur from the Wessex Formation, Wealden Group (Early Cretaceous), of the Isle of Wight, southern England" masterLastPageNumber="888" masterPageNumber="847" pageNumber="851" updateTime="1712838945022" updateUser="diego">
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<mods:title id="8EF10920A03191260E2E3CBE148593BF">A new hadrosauriform dinosaur from the Wessex Formation, Wealden Group (Early Cretaceous), of the Isle of Wight, southern England</mods:title>
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<mods:namePart id="2D61C296F7A219DD481D6909A30101D6">Lockwood, Jeremy A. F.</mods:namePart>
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<mods:affiliation id="68EFA047A14F174EE806074BF2A691BA">School of Environment, Geography and Geosciences, University of Portsmouth PO 1 2 DT, UK; & Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="86ACCA68E4F2A0E8B058BFDD06AF0652">Martill, David M.</mods:namePart>
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<mods:affiliation id="FE030A2EAAEAD3E9506EA3286E00EC52">School of Environment, Geography and Geosciences, University of Portsmouth PO 1 2 DT, UK;</mods:affiliation>
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<mods:namePart id="DC1438A9DE8851E626386411FCCD4337">Maidment, Susannah C. R.</mods:namePart>
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<mods:affiliation id="631A048263DB78C504A4C8B7B8DA010D">Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:title id="94F83CF959D03626EC3CB764C6CC1714">Journal of Systematic Palaeontology</mods:title>
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<mods:date id="418E1868EE6DAF7300A2B9FE6160B566">2021</mods:date>
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<taxonomicName id="4C5586A96D63FFF486A8FE391DD42CFC" authority="Lockwood & Martill & Maidment, 2021" authorityName="Lockwood & Martill & Maidment" authorityYear="2021" box="[229,504,444,468]" genus="Brighstoneus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="851" phylum="Chordata" rank="species" species="simmondsi" status="gen. et sp. nov.">
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<emphasis id="B92121386D63FFF486A8FE391DD42CFC" bold="true" box="[229,504,444,468]" italics="true" pageId="5" pageNumber="851">Brighstoneus simmondsi</emphasis>
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(
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<figureCitation id="136EE1AF6D63FFF48731FE671DC42CD1" box="[380,488,481,505]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-10="Figure 13" captionStart-11="Figure 14" captionStart-12="Figure 15" captionStart-13="Figure 16" captionStart-14="Figure 17" captionStart-15="Figure 18" captionStart-16="Figure 19" captionStart-17="Figure 20" captionStart-18="Figure 21" captionStart-19="Figure 22" captionStart-2="Figure 5" captionStart-3="Figure 6" captionStart-4="Figure 7" captionStart-5="Figure 8" captionStart-6="Figure 9" captionStart-7="Figure 10" captionStart-8="Figure 11" captionStart-9="Figure 12" captionStartId-0="6.[159,226,509,530]" captionStartId-1="7.[159,226,1618,1639]" captionStartId-10="19.[159,226,967,988]" captionStartId-11="20.[159,226,1399,1420]" captionStartId-12="21.[159,226,1609,1630]" captionStartId-13="23.[159,226,1256,1277]" captionStartId-14="24.[159,226,1261,1282]" captionStartId-15="26.[159,226,1648,1669]" captionStartId-16="28.[159,226,931,952]" captionStartId-17="30.[844,911,820,841]" captionStartId-18="31.[159,226,815,836]" captionStartId-19="32.[159,226,1046,1067]" captionStartId-2="9.[159,226,1213,1234]" captionStartId-3="10.[159,226,1206,1227]" captionStartId-4="11.[844,911,915,936]" captionStartId-5="12.[159,226,1337,1358]" captionStartId-6="14.[159,226,1467,1488]" captionStartId-7="15.[844,911,736,757]" captionStartId-8="16.[159,226,1576,1597]" captionStartId-9="18.[159,226,1039,1060]" captionTargetBox-0="[179,776,198,487]" captionTargetBox-1="[160,1465,188,1601]" captionTargetBox-10="[160,1465,189,947]" captionTargetBox-11="[159,1465,190,1379]" captionTargetBox-12="[160,1465,189,1589]" captionTargetBox-13="[159,1466,192,1238]" captionTargetBox-14="[159,1466,190,1243]" captionTargetBox-15="[167,1463,201,1631]" captionTargetBox-16="[159,1466,188,914]" captionTargetBox-17="[846,1457,188,803]" captionTargetBox-18="[162,1465,191,793]" captionTargetBox-19="[159,1465,200,1029]" captionTargetBox-2="[159,1466,189,1196]" captionTargetBox-3="[159,1466,188,1188]" captionTargetBox-4="[845,1464,188,898]" captionTargetBox-5="[159,1466,188,1320]" captionTargetBox-6="[159,1466,188,1448]" captionTargetBox-7="[846,1463,189,719]" captionTargetBox-8="[161,1465,188,1558]" captionTargetBox-9="[159,1466,188,1022]" captionTargetId-0="figure-779@6.[160,780,188,491]" captionTargetId-1="figure-78@7.[159,1466,188,1601]" captionTargetId-10="figure-396@19.[159,1466,188,949]" captionTargetId-11="figure-189@20.[159,1466,188,1382]" captionTargetId-12="figure-58@21.[159,1466,188,1592]" captionTargetId-13="figure-261@23.[159,1466,188,1239]" captionTargetId-14="figure-269@24.[159,1466,188,1243]" captionTargetId-15="figure-0@26.[159,1466,188,1631]" captionTargetId-16="figure-411@28.[159,1466,188,914]" captionTargetId-17="figure-651@30.[846,1465,188,803]" captionTargetId-18="figure-479@31.[159,1466,188,798]" captionTargetId-19="figure-308@32.[159,1466,188,1029]" captionTargetId-2="figure-285@9.[159,1466,188,1196]" captionTargetId-3="figure-306@10.[159,1466,188,1188]" captionTargetId-4="figure-646@11.[845,1465,188,898]" captionTargetId-5="figure-273@12.[159,1466,188,1320]" captionTargetId-6="figure-143@14.[159,1466,188,1449]" captionTargetId-7="figure-696@15.[846,1466,188,719]" captionTargetId-8="figure-68@16.[159,1466,188,1559]" captionTargetId-9="figure-369@18.[159,1466,188,1022]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-10="19" captionTargetPageId-11="20" captionTargetPageId-12="21" captionTargetPageId-13="23" captionTargetPageId-14="24" captionTargetPageId-15="26" captionTargetPageId-16="28" captionTargetPageId-17="30" captionTargetPageId-18="31" captionTargetPageId-19="32" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="12" captionTargetPageId-6="14" captionTargetPageId-7="15" captionTargetPageId-8="16" captionTargetPageId-9="18" captionText-0="Figure 3. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), right dorsal process of premaxilla in A, lateral and B, medial views. C, cross-section at anterior end. Abbreviations: gr, groove for anterodorsal process of nasal; ins, internarial septum; lat, lateral; med, medial; na, external naris. Scale bar= 50 mm." captionText-1="Figure 4. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), left maxilla in A, lateral, B, dorsal, C, medial and D, ventral views. Abbreviations: adp, anterodorsal process; aod, antorbital depression; ap, ascending process; avp, anteroventral process; eps, ectopterygoid shelf; frag, fragment of bone, probably erroneously placed; jp, jugal process; ms, medial shelf; pmg, premaxillary groove; sf, special foramina. Scale bar= 100 mm." captionText-10="Figure 13. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), dorsal vertebrae series arranged in probable anterior to posterior order. A–H, left lateral views of dorsals A–H; AƖ, dorsal A anterior view; BƖ, dorsal B anterior view; CƖ dorsal C anterior view; DƖ, dorsal D posterior view; EƖ, dorsal E anterior, view; FƖ, dorsal F anterior view; GƖ, dorsal G anterior view; HƖ, dorsal H posterior view. Scale bar= 100mm." captionText-11="Figure 14. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), anterior dorsal vertebrae. A–C, ‘dorsal A’, probably D1 in A, right lateral, B, anterior and C, posterior views. D–F, ‘dorsal B’, probably D2 in D, right lateral, E, posterior and F, anterior views. Abbreviations: bf, extraneous bone fragment; nc, neural canal; ns, neural spine; pa, parapophysis; poz, postzygapophysis; prz, prezygapophysis; tp, transverse process. Scale bar= 50 mm." captionText-12="Figure 15. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), dorsal vertebrae. A–E, 'dorsal C’ in A, right lateral, B, dorsal (of neural spine), C, anterior, D, ventral and E, posterior views. F, close-up of fracture site on neural spine of ‘dorsal E’. G, fragment of pathological dorsal neural spine. H, section of neural spine with bite marks. Abbreviations: afc, anterior surface of centrum; ca, callus; ch, channel; gr, groove; nc, neural canal; ns, neural spine; os, ostosis; ot, ossified tendons; pa, parapophysis; pfc, posterior surface of centrum; poz, postzygapophysis; prz, prezygapophysis; tm, tooth marks; tp, transverse process; vk, ventral keel. Scale bars= 100mm (A–E) and 50 mm (F–H)." captionText-13="Figure 16. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), dorsal ribs. A, proximal section of right rib in posterior view; B, specimen ‘A’ in anterior view (image reversed); C, close-up of F showing possible bioerosion; D, posterior series rib head with distal transverse process; E, anterior series, left rib, anterior view and EƖ, dorsal view; F, middle series (image reversed), left rib, posterior view and FƖ, dorsal view; G, middle series (image reversed), left rib, posterior view and GƖ, dorsal view; H, posterior series, right rib, posterior view and HƖ dorsal view. Abbreviations: be, bioerosion; cap, capitulum; ef, elliptical foramen; fr, fracture; rr, rounded ridge; rug, rugosity; se, spatulate end; sg, shallow groove; sr, sharp ridge; tp, transverse process; tub, tuberculum. Scale bars= 50mm (A, B, D, E’, F’, G’, H’), 100 mm (E–H) and 5 mm (C)." captionText-14="Figure 17. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), sacrum in A, right lateral, B, posterior and C, ventral views. Abbreviations: da, depressed area; er, eroded area; ns, neural spine; nsf, neural spine fragments in presumed position; pfc, posterior surface of centrum; poz, postzygapophysis; s, sacral vertebra; sr, sacral rib; sy, sacral yoke; vk, ventral keel. Scale bars = 100 mm." captionText-15="Figure 18. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), caudal vertebrae. A, ‘caudal A’ in left lateral view; B, ‘caudal A’ in posterior view; C, ‘caudal B’ in right lateral view; D, ‘caudal B’ in posterior view; E, ‘caudal C’ in left lateral view; F, ‘caudal C’ in anterior view; G, ‘caudal D’ in anterior view; H, ‘caudal D’ in left lateral view; I, ‘caudal D’ in posterior view; J, ‘caudal D’ in dorsal view; K, ‘caudal D’ in ventral view; L, ‘caudal E’ in left lateral view; M, ‘caudal E’ in anterior view; N, ‘caudal E’ in ventral view; O, ‘caudal F’ in right lateral view; P, ‘caudal F’ in posterior view. Abbreviations: ahf, anterior haemapophyseal facet; lr, lateral ridge; ns, neural spine; phf, posterior haemapophyseal facet; plp, pleurapophysis (caudal rib); poz, postzygapophysis; prz, prezygapophysis; sb, small boss. Scale bars= 100mm." captionText-16="Figure 19. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), ilia. A, preacetabular process of right ilium in medial view; B, cross-section of base of A. C–F, left ilium in C, dorsal, D, medial, E, lateral and F, ventral views. G, H, right ilium in G, lateral and H, medial views. Abbreviations: ac, acetabulum; bs, brevis shelf; dms, dorsomedial surface; er, eroded area; ftp, facet for transverse process; gr, groove; ip, ischiadic peduncle; lp, lateral process; poap, postacetabular process; pp, pubic peduncle; pr, primary ridge; prap, preacetabular process; sr, secondary ridge. Scale bars = 100 mm." captionText-17="Figure 20. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), ischium in A, lateral, B, dorsal and C, medial views. Abbreviations: ac, acetabular component; er, eroded areas; gr, lateral groove; ip, iliac peduncle; op, obturator process; pp, pubic peduncle. Scale bar= 100mm." captionText-18="Figure 21. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), possible fragment of anterior pubic process in A, lateral, B, medial, C, anterior and D, posterior views. Views assume that the distal end is to the left. Abbreviation: py, pyrite. Scale bar= 50 mm." captionText-19="Figure 22. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), right femur in A, anterior, B, medial, C, posterior, D, lateral, E, dorsal and F, ventral views. Reconstruction of distal femur shown in A is based on Mantellisaurus atherfieldensis (Norman 1986). Abbreviations: 4t, fourth trochanter; at, anterior trochanter; dr, diagonal ridge; eig, extensor intercondylar groove; es, eroded surface; fh, femoral head; fig, flexor intercondylar groove; gt, greater trochanter; mc, medial condyle. Scale bar= 100 mm." captionText-2="Figure 5. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), left nasal in A, ventromedial (internal surface), B, dorsolateral (external surface), C, ventral, D, dorsal, E, lateral (in life position) and F, anterior views. Abbreviations: adp, anterodorsal process; avp, anteroventral process; bev, bevelled surface; bull, nasal bulla; grdp, groove for dorsal process of the premaxilla; grmp, groove for maxillary process of the premaxilla; na, external naris. Scale bar= 100mm." captionText-3="Figure 6. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344). Left (A, B) and right (C–F) jugals in A, lateral, B, medial, C, medial, D, lateral, E, ventral and F, dorsal views. Abbreviations: bu, buttress; epf, ectopterygoid facet; ls, lacrimal suture; ltf, lateral temporal fenestra; mxs, maxillary suture; orb, orbit; os, orbital shelf; pop, postorbital process; qjs, area of suture with the quadratojugal; re, rolled edge. Scale bar= 50 mm." captionText-4="Figure 7. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), left palpebral in A, ventral, B, medial, C, dorsal and D, lateral views. Abbreviations: apf, articular surface with prefrontal; mp, medial process; rug, rugosity. Scale bar = 50 mm." captionText-5="Figure 8. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), predentary in A, dorsal, B, anterior, C, left lateral, D, ventral and E, right lateral views. Abbreviations: d, denticle; fo, foramina; vg, vascular groove; vp, ventral process. Scale bar= 50mm." captionText-6="Figure 9. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), right (A–E) and left (F, G) dentaries in A, ventral, B, medial, C, lateral, D, posterior, E, dorsal, F, dorsal and G, medial views. Abbreviations: af, adductor fossa; bs, buccal shelf; cp, coronoid process; dc, dentary crown; fo, foramen; gpd, groove for predentary; mc, Meckelian canal; sym, symphysis. Scale bar = 100 mm." captionText-7="Figure 10. Preliminary reconstructions of the skulls of A, Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344) and B, Mantellisaurus atherfieldensis (NHMUK PV R5764) in lateral view. Shaded areas represent material present in the holotypes. Scale bar= 100 mm." captionText-8="Figure 11. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), maxillary crowns. A, right crown in buccal view; B, left crown in buccal view; C, left broken crown dorsal view; D, close-up of A showing mammillae on denticles; E, left broken crown in ventral view; F, ventral view of A; G, left crown in ventral view; H, lingual view of B showing primary and accessory ridges. Arrow labelled b = buccal primary ridge, unlabelled = lingual ridge. Abbreviations: ar, accessory ridge; d, denticle; da, damaged area; m, mammilla; pr, primary ridge. Scale bars= 5 mm (A, B, C, E, F, H) and 2 mm (D, G)." captionText-9="Figure 12. Brighstoneus simmondsi gen. et sp. nov. (MIWG 6344), crowns from right dentary (posterior is to the right). A, dentary crown in lingual view; B, close-up of A showing denticles; C, dentary crown in lingual view; D, close-up of denticle from A showing mammillae. Abbreviations: ar, accessory ridge; d, marginal denticle; ma, mammilla; pr, primary ridge; sr, secondary ridge. Scale bars= 10 mm (A), 5 mm (B, C) and 1mm (D)." pageId="5" pageNumber="851">Figs 3–22</figureCitation>
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,
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)
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||
<emphasis id="B92121386D63FFF486D2FDB71D0D2F61" bold="true" box="[159,289,562,585]" pageId="5" pageNumber="851">Etymology.</emphasis>
|
||
The specific name honours Mr Keith Simmonds who made the discovery of the specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34FAEA16D63FFF486D2FD071D742EF2" pageId="5" pageNumber="851" type="materials_examined">
|
||
<paragraph id="8BEAFD2A6D63FFF486D2FD071D742EF2" blockId="5.[159,781,642,986]" pageId="5" pageNumber="851">
|
||
<materialsCitation id="3B3DF7776D63FFF486D2FD071D742EF2" collectionCode="MIWG" pageId="5" pageNumber="851" specimenCode="MIWG 6344" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B92121386D63FFF486D2FD071D212FB1" bold="true" box="[159,269,642,665]" pageId="5" pageNumber="851">
|
||
<typeStatus id="54EE43886D63FFF486D2FD071D252FB1" box="[159,265,642,665]" pageId="5" pageNumber="851" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="DBF355516D63FFF48755FD071D842FB1" box="[280,424,642,665]" collectionCode="MIWG" country="United Kingdom" httpUri="http://grbio.org/cool/9n8r-c13h" name="Museum of he Isle of Wight Geology" pageId="5" pageNumber="851">MIWG 6344</specimenCode>
|
||
, a partial skeleton composed of the following elements: dorsal process of right premaxilla; both maxillae; both jugals; left palpebral; left nasal; both dentaries; predentary; one transitional dorsal and seven dorsal vertebrae; sacrum; six caudal vertebrae; dorsal ribs, nine from the left side and five from the right side; both ilia; right ischium; possible prepubic process; and the right femur. Some parts of the same individual (including two dorsal vertebrae and other fragments) remain in private ownership and are not described herein.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34FAEA16D63FFF486D2FC761FBE2CD7" pageId="5" pageNumber="851" type="diagnosis">
|
||
<paragraph id="8BEAFD2A6D63FFF486D2FC761ED92803" blockId="5.[159,782,1011,1901]" pageId="5" pageNumber="851">
|
||
<emphasis id="B92121386D63FFF486D2FC761D382922" bold="true" box="[159,276,1011,1034]" pageId="5" pageNumber="851">Diagnosis.</emphasis>
|
||
Differs from all other iguanodontians by possessing the following autapomorphies and unique combination of characters (autapomorphies indicated with an asterisk): maxillary crowns possessing both a primary ridge and mesially placed accessory ridges on the lingual surface‡; nasal expanded postnarially to produce a modest nasal bulla with convex lateral walls‡. A character combination of at least 28 dentary tooth positions in a dentary with one active crown and one replacement tooth for each position and non-parallel alveolar septa.
|
||
</paragraph>
|
||
<paragraph id="8BEAFD2A6D63FFF486F4FAB11FBE2CD7" blockId="5.[159,782,1011,1901]" lastBlockId="5.[844,1467,189,511]" pageId="5" pageNumber="851">
|
||
In addition,
|
||
<taxonomicName id="4C5586A96D63FFF48704FAB11DFA2863" authorityName="Lockwood & Martill & Maidment" authorityYear="2021" box="[329,470,1332,1355]" genus="Brighstoneus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="851" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92121386D63FFF48704FAB11DFA2863" box="[329,470,1332,1355]" italics="true" pageId="5" pageNumber="851">Brighstoneus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from other Barremian–Aptian Wealden Group iguanodontians by possession of the following combination of features: ratio of precoronoid length of the dentary to minimum depth> 6.0; coronoid process projects at approximately 90
|
||
<superScript id="7C2050626D63FFF486F4FA551CEE28C9" attach="left" box="[185,194,1488,1505]" fontSize="7" pageId="5" pageNumber="851">ǫ</superScript>
|
||
with respect to the dorsal margin of dentary; bilobed ‘heart-shaped’ ventral predentary process with prominent anterior denticles with concave mesial and distal edges; posteriorly positioned maxillary ascending process in lateral view with length of the anterior section approximately twice the length of the posterior section; prominent anterodorsal process present on maxilla; anterior (maxillary) process of jugal relatively long (60% of overall length) and tapers distally to form a triangular ending; ventral section of the posterior margin of the jugal (heel) projects posteriorly to form a spur-like feature; in dorsal view the jugal is straight; ventral border overlapped by maxillary process of premaxilla; anteroventral nasal process forms posteroventral margin of external narial opening; ventral surface of ischiadic peduncle of ilium parallel with the anteroventral margin of the postacetabular process; ischiadic peduncle of ilium has flat lateral wall with no pronounced posterolateral boss; ventral shelf at base of iliac preacetabular process weakly developed; dorsoventrally deep preacetabular process with little or no axial twist; deep and short iliac central plate with ratio of depth to length> 1.2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |