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<document ID-DOI="10.11646/zootaxa.5257.1.6" ID-GBIF-Dataset="f96f439c-8ba0-4645-ac22-ce46b40aabae" ID-ISSN="1175-5326" ID-Zenodo-Dep="7765939" ID-ZooBank="E18CE51E-CCD2-4E12-B480-930C79D8B745" checkinTime="1679644040936" checkinUser="plazi" docAuthor="Chen, Hui-Ming, Zheng, Chang-Bin &amp; Jiang, Xuan-Kong" docDate="2023" docId="03B087C80719441A3ACF1BDBD948CC00" docLanguage="en" docName="zootaxa.5257.1.6.pdf" docOrigin="Zootaxa 5257 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5257.1.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epanerchodus wuxi Chen &amp; Zheng &amp; Jiang 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="77" masterDocId="FF89FFB0070544063A581B4CDD5FCA51" masterDocTitle="The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China" masterLastPageNumber="81" masterPageNumber="49" pageNumber="77" updateTime="1679644761978" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Hui-Ming</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-2449-3036</mods:nameIdentifier>
<mods:affiliation>Institute of Biology, Guizhou Academy of Sciences, Guiyang 550009, China &amp; mei 0601 @ 126. com; https: // orcid. org / 0000 - 0002 - 2449 - 3036</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zheng, Chang-Bin</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-2044-5096</mods:nameIdentifier>
<mods:affiliation>Management Center of Yintiaoling National Nature Reserve, Chongqing 405800, China &amp; 552151709 @ qq. com; https: // orcid. org / 0000 - 0002 - 2044 - 5096</mods:affiliation>
<mods:nameIdentifier type="email">552151709@qq.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jiang, Xuan-Kong</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-3506-5894</mods:nameIdentifier>
<mods:affiliation>Institute of Biology, Guizhou Academy of Sciences, Guiyang 550009, China &amp; antoma 93 @ gmail. com; https: // orcid. org / 0000 - 0003 - 3506 - 5894</mods:affiliation>
<mods:nameIdentifier type="email">antoma93@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5257</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>49</mods:start>
<mods:end>81</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5257.1.6</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5257.1.6</mods:identifier>
<mods:identifier type="GBIF-Dataset">f96f439c-8ba0-4645-ac22-ce46b40aabae</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7765939</mods:identifier>
<mods:identifier type="ZooBank">E18CE51E-CCD2-4E12-B480-930C79D8B745</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7765949" ID-Zenodo-Dep="7765949" LSID="urn:lsid:plazi:treatment:03B087C80719441A3ACF1BDBD948CC00" httpUri="http://treatment.plazi.org/id/03B087C80719441A3ACF1BDBD948CC00" lastPageNumber="77" pageId="28" pageNumber="77">
<subSubSection box="[151,494,151,178]" pageId="28" pageNumber="77" type="nomenclature">
<paragraph blockId="28.[151,494,151,213]" box="[151,494,151,178]" pageId="28" pageNumber="77">
<heading bold="true" box="[151,494,151,178]" fontSize="11" level="1" pageId="28" pageNumber="77" reason="1">
<emphasis bold="true" box="[151,494,151,178]" pageId="28" pageNumber="77">
<taxonomicName authority="Chen &amp; Zheng &amp; Jiang, 2023" authorityName="Chen &amp; Zheng &amp; Jiang" authorityYear="2023" box="[151,394,151,178]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="28" pageNumber="77" phylum="Arthropoda" rank="species" species="wuxi" status="sp. nov.">
<emphasis bold="true" box="[151,394,151,178]" italics="true" pageId="28" pageNumber="77">Epanerchodus wuxi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[401,494,151,177]" pageId="28" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,277,187,213]" pageId="28" pageNumber="77" type="description">
<paragraph blockId="28.[151,494,151,213]" box="[151,277,187,213]" pageId="28" pageNumber="77">
<figureCitation box="[151,277,187,213]" captionStart-0="FIGURE 32" captionStart-1="FIGURE 33" captionStart-2="FIGURE 34" captionStartId-0="25.[151,250,1727,1751]" captionStartId-1="26.[151,250,1947,1971]" captionStartId-2="27.[151,250,1981,2005]" captionTargetBox-0="[151,1435,181,1704]" captionTargetBox-1="[202,1386,181,1922]" captionTargetBox-2="[198,1396,185,1950]" captionTargetId-0="figure-19@25.[151,1435,181,1704]" captionTargetId-1="figure-20@26.[202,1386,181,1922]" captionTargetId-2="figure-19@27.[190,1397,181,1957]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionText-0="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." captionText-1="FIGURE 33. Epanerchodus wuxi sp. nov., holotype. A. Anterior part of body, ventral view; B. Posterior part of body, ventral view; C. Head, collum and segments 2–4, dorsal view; D. Telson, ventral view. Scale bars: A, B = 1 mm; C, D = 0.5 mm." captionText-2="FIGURE 34. Epanerchodus wuxi sp. nov., holotype. A. Right gonopod, mesal view; B. Right gonopod, lateral view; C. Subapical part of endomere, sub-medial view; D. Apical part of endomere, top view. Scale bars: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7766033" figureDoi-1="http://doi.org/10.5281/zenodo.7766035" figureDoi-2="http://doi.org/10.5281/zenodo.7766037" httpUri-0="https://zenodo.org/record/7766033/files/figure.png" httpUri-1="https://zenodo.org/record/7766035/files/figure.png" httpUri-2="https://zenodo.org/record/7766037/files/figure.png" pageId="28" pageNumber="77">Figs 32–34</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="77" type="materials_examined">
<paragraph blockId="28.[151,1437,259,1618]" pageId="28" pageNumber="77">
<emphasis bold="true" box="[151,337,259,285]" pageId="28" pageNumber="77">Type materials.</emphasis>
<materialsCitation collectingDate="2022-08-14" collectorName="X. K. Jiang &amp; H. M. Chen" country="China" county="Wuxi County" elevation="1677" latitude="31.511314" location="Yintiaoling National Nature Reserve" longLatPrecision="1" longitude="109.82803" municipality="Male" pageId="28" pageNumber="77" specimenCount="1" stateProvince="Chongqing" typeStatus="holotype">
<emphasis bold="true" box="[344,453,259,285]" pageId="28" pageNumber="77">
<typeStatus box="[344,453,259,285]" pageId="28" pageNumber="77">Holotype</typeStatus>
</emphasis>
<collectingMunicipality box="[461,523,259,285]" pageId="28" pageNumber="77">Male</collectingMunicipality>
:
<collectingCountry box="[534,604,259,285]" name="China" pageId="28" pageNumber="77">China</collectingCountry>
,
<collectingRegion box="[616,743,259,285]" country="China" name="Chongqing" pageId="28" pageNumber="77">Chongqing</collectingRegion>
,
<collectingCounty box="[755,908,259,285]" pageId="28" pageNumber="77">Wuxi County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:03B087C80719441A3ACF1BDBD948CC00:8EC660050719441A39CE1A4FD86ACB4F" box="[918,1333,259,286]" country="China" county="Wuxi County" latitude="31.511314" longLatPrecision="1" longitude="109.82803" municipality="Male" name="Yintiaoling National Nature Reserve" pageId="28" pageNumber="77" stateProvince="Chongqing">Yintiaoling National Nature Reserve</location>
,
<location LSID="urn:lsid:plazi:treatment:03B087C80719441A3ACF1BDBD948CC00:8EC660050719441A3F191A4FD8C7CB4C" box="[1345,1432,259,285]" country="China" county="Wuxi County" latitude="31.511314" longLatPrecision="1" longitude="109.82803" municipality="Male" name="Hongqi" pageId="28" pageNumber="77" stateProvince="Chongqing">Hongqi</location>
,
<location LSID="urn:lsid:plazi:treatment:03B087C80719441A3ACF1BDBD948CC00:8EC660050719441A3ACF1A64DC64CB10" box="[151,315,295,322]" country="China" county="Wuxi County" latitude="31.511314" longLatPrecision="1" longitude="109.82803" municipality="Male" name="Longtan Cave" pageId="28" pageNumber="77" stateProvince="Chongqing">Longtan Cave</location>
,
<geoCoordinate box="[329,514,295,322]" degrees="31" direction="north" minutes="30" orientation="latitude" pageId="28" pageNumber="77" precision="1" seconds="40.73" value="31.511314">31°30’40.73” N</geoCoordinate>
,
<geoCoordinate box="[529,724,295,322]" degrees="109" direction="east" minutes="49" orientation="longitude" pageId="28" pageNumber="77" precision="1" seconds="40.93" value="109.82803">109°49’40.93” E</geoCoordinate>
, alt.
<quantity box="[784,875,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.677" pageId="28" pageNumber="77" unit="m" value="1677.0">
<elevation box="[784,875,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.677" pageId="28" pageNumber="77" unit="m" value="1677.0">1677 m</elevation>
</quantity>
,
<date box="[890,1076,295,321]" pageId="28" pageNumber="77" value="2022-08-14">
<collectingDate box="[890,1076,295,321]" pageId="28" pageNumber="77" value="2022-08-14">14 August 2022</collectingDate>
</date>
,
<collectorName box="[1090,1214,295,322]" pageId="28" pageNumber="77">X.K. Jiang</collectorName>
&amp;
<collectorName box="[1256,1385,295,322]" pageId="28" pageNumber="77">H.M. Chen</collectorName>
leg.
</materialsCitation>
<materialsCitation box="[151,674,331,358]" collectingDate="2022-08-14" collectorName="X. K. Jiang &amp; H. M. Chen" country="China" county="Wuxi County" elevation="1677" latitude="31.511314" location="Yintiaoling National Nature Reserve" longLatPrecision="1" longitude="109.82803" municipality="Male" pageId="28" pageNumber="77" specimenCount="12" specimenCount-female="12" stateProvince="Chongqing" typeStatus="paratype">
<emphasis bold="true" box="[151,280,331,357]" pageId="28" pageNumber="77">
<typeStatus box="[151,275,331,357]" pageId="28" pageNumber="77">Paratypes</typeStatus>
:
</emphasis>
<specimenCount box="[287,412,331,357]" pageId="28" pageNumber="77" type="female">12 females</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation country="China" elevation="1458" latitude="31.521341" location="Longtandong Cave" longLatPrecision="1" longitude="109.83106" pageId="28" pageNumber="77" specimenCount="1" specimenCount-male="1" stateProvince="Chongqing" typeStatus="paratype">
<specimenCount box="[685,763,331,357]" pageId="28" pageNumber="77" type="male">1 male</specimenCount>
, a manual tunnel near the
<location LSID="urn:lsid:plazi:treatment:03B087C80719441A3ACF1BDBD948CC00:8EC660050719441A3E781A07D9A6CB34" box="[1056,1273,331,357]" country="China" latitude="31.521341" longLatPrecision="1" longitude="109.83106" name="Longtandong Cave" pageId="28" pageNumber="77" stateProvince="Chongqing">Longtandong Cave</location>
,
<geoCoordinate degrees="31" direction="north" minutes="31" orientation="latitude" pageId="28" pageNumber="77" precision="1" seconds="16.83" value="31.521341">31°31’16.83” N</geoCoordinate>
,
<geoCoordinate box="[185,376,367,394]" degrees="109" direction="east" minutes="49" orientation="longitude" pageId="28" pageNumber="77" precision="1" seconds="51.84" value="109.83106">109°49’51.84” E</geoCoordinate>
, alt.
<quantity box="[429,515,367,394]" metricMagnitude="3" metricUnit="m" metricValue="1.458" pageId="28" pageNumber="77" unit="m" value="1458.0">
<elevation box="[429,515,367,394]" metricMagnitude="3" metricUnit="m" metricValue="1.458" pageId="28" pageNumber="77" unit="m" value="1458.0">1458 m</elevation>
</quantity>
, same time and collectors as
<typeStatus box="[839,941,367,393]" pageId="28" pageNumber="77">holotype</typeStatus>
</materialsCitation>
;
<specimenCount box="[951,1050,367,393]" pageId="28" pageNumber="77" type="female">1 female</specimenCount>
, same cave as
<typeStatus box="[1212,1313,367,393]" pageId="28" pageNumber="77">holotype</typeStatus>
,
<date pageId="28" pageNumber="77" value="2022-08-11">11 August 2022</date>
, Q.D. Zhu leg.
</paragraph>
</subSubSection>
<subSubSection box="[199,1321,439,466]" pageId="28" pageNumber="77" type="etymology">
<paragraph blockId="28.[151,1437,259,1618]" box="[199,1321,439,466]" pageId="28" pageNumber="77">
<emphasis bold="true" box="[199,334,439,465]" pageId="28" pageNumber="77">Etymology.</emphasis>
The specific name is a noun in apposition, derived from the
<typeStatus box="[1014,1062,440,466]" pageId="28" pageNumber="77">type</typeStatus>
locality, Wuxi County.
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="77" type="diagnosis">
<paragraph blockId="28.[151,1437,259,1618]" pageId="28" pageNumber="77">
<emphasis bold="true" box="[199,322,475,501]" pageId="28" pageNumber="77">Diagnosis.</emphasis>
The new species is similar to
<taxonomicName authority="Liu &amp; Golovatch, 2018" authorityName="Liu &amp; Golovatch" authorityYear="2018" box="[663,1156,475,501]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" kingdom="Animalia" order="Polydesmida" pageId="28" pageNumber="77" phylum="Arthropoda" rank="species" species="latus">
<emphasis box="[663,885,475,501]" italics="true" pageId="28" pageNumber="77">Epanerchodus latus</emphasis>
<bibRefCitation author="Liu, W. X. &amp; Golovatch, S. I." box="[893,1156,475,501]" pageId="28" pageNumber="77" pagination="1" refId="ref10616" refString="Liu, W. X. &amp; Golovatch, S. I. (2018) The millipede genus Epanerchodus Attems, 1901 in continental China, with descriptions of seven new cavernicolous species (Diplopoda, Polydesmida, Polydesmidae). Zootaxa, 4459 (1), 053 - 084. https: // doi. org / 10.11646 / zootaxa. 4459.1.2" type="journal article" year="2018">
Liu &amp; Golovatch,
<quantity box="[1100,1156,475,501]" metricMagnitude="1" metricUnit="m" metricValue="5.125719999999999" pageId="28" pageNumber="77" unit="in" value="2018.0">2018</quantity>
</bibRefCitation>
</taxonomicName>
in having the gonopodal femorite with a finger-shaped process distally and a process medially, the endomere carrying a subapical lobe, but distinguished by the broad subapical endomere with a spiny process and a row of apical denticles (
<figureCitation box="[1256,1340,547,573]" captionStart="FIGURE 34" captionStartId="27.[151,250,1981,2005]" captionTargetBox="[198,1396,185,1950]" captionTargetId="figure-19@27.[190,1397,181,1957]" captionTargetPageId="27" captionText="FIGURE 34. Epanerchodus wuxi sp. nov., holotype. A. Right gonopod, mesal view; B. Right gonopod, lateral view; C. Subapical part of endomere, sub-medial view; D. Apical part of endomere, top view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766037" httpUri="https://zenodo.org/record/7766037/files/figure.png" pageId="28" pageNumber="77">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="77" type="description">
<paragraph blockId="28.[151,1437,259,1618]" pageId="28" pageNumber="77">
<emphasis bold="true" box="[199,347,583,609]" pageId="28" pageNumber="77">Description:</emphasis>
Length of
<typeStatus box="[473,571,583,609]" pageId="28" pageNumber="77">holotype</typeStatus>
<quantity box="[579,661,583,610]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="28" pageNumber="77" unit="mm" value="22.0">22 mm</quantity>
, male
<typeStatus box="[735,842,584,610]" pageId="28" pageNumber="77">paratypes</typeStatus>
<quantity box="[850,932,583,610]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="28" pageNumber="77" unit="mm" value="22.0">22 mm</quantity>
, female
<typeStatus box="[1029,1136,584,610]" pageId="28" pageNumber="77">paratypes</typeStatus>
<quantity box="[1144,1268,583,610]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.7" metricValueMin="2.0" pageId="28" pageNumber="77" unit="mm" value="28.5" valueMax="37.0" valueMin="20.0">20–37 mm</quantity>
; width of midbody pro- and metazona in
<typeStatus box="[458,556,619,645]" pageId="28" pageNumber="77">holotype</typeStatus>
<quantity box="[563,648,619,646]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="28" pageNumber="77" unit="mm" value="1.6">1.6 mm</quantity>
and
<quantity box="[702,790,619,646]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="28" pageNumber="77" unit="mm" value="3.1">3.1 mm</quantity>
, respectively, another male
<typeStatus box="[1101,1197,620,646]" pageId="28" pageNumber="77">paratype</typeStatus>
<quantity box="[1204,1290,619,646]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="77" unit="mm" value="1.5">1.5 mm</quantity>
and 3.0 mm, in female
<typeStatus box="[262,369,656,682]" pageId="28" pageNumber="77">paratypes</typeStatus>
up to
<quantity box="[439,573,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.9" metricValueMin="1.2" pageId="28" pageNumber="77" unit="mm" value="1.5499999999999998" valueMax="1.9" valueMin="1.2">1.2–1.9 mm</quantity>
and
<quantity box="[626,763,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.5" metricValueMin="2.4" pageId="28" pageNumber="77" unit="mm" value="2.95" valueMax="3.5" valueMin="2.4">2.4–3.5 mm</quantity>
. Live specimens whitish to light yellow and semitransparent (
<figureCitation box="[159,264,691,717]" captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Fig. 32A</figureCitation>
), in alcohol light yellow to light brown (
<figureCitation box="[717,855,691,717]" captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Figs 32B, C</figureCitation>
,
<figureCitation box="[867,898,691,717]" captionStart="FIGURE 33" captionStartId="26.[151,250,1947,1971]" captionTargetBox="[202,1386,181,1922]" captionTargetId="figure-20@26.[202,1386,181,1922]" captionTargetPageId="26" captionText="FIGURE 33. Epanerchodus wuxi sp. nov., holotype. A. Anterior part of body, ventral view; B. Posterior part of body, ventral view; C. Head, collum and segments 2–4, dorsal view; D. Telson, ventral view. Scale bars: A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7766035" httpUri="https://zenodo.org/record/7766035/files/figure.png" pageId="28" pageNumber="77">33</figureCitation>
).
</paragraph>
<paragraph blockId="28.[151,1437,259,1618]" pageId="28" pageNumber="77">
Adults with 20 segments. Width of
<typeStatus box="[590,692,727,753]" pageId="28" pageNumber="77">holotype</typeStatus>
: head (1.8) &lt;&lt; collum (2.7) &lt;segment 2 (2.8) = 3 = 4 = 5 &lt;6 (2.9)– 8 &lt;9 (3.1) –16, thereafter segments 17 (2.9)–19 (1.7). Body gradually tapering posteriorly towards telson (
<figureCitation box="[1345,1431,763,789]" captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Figs 32</figureCitation>
,
<figureCitation box="[151,182,799,825]" captionStart="FIGURE 33" captionStartId="26.[151,250,1947,1971]" captionTargetBox="[202,1386,181,1922]" captionTargetId="figure-20@26.[202,1386,181,1922]" captionTargetPageId="26" captionText="FIGURE 33. Epanerchodus wuxi sp. nov., holotype. A. Anterior part of body, ventral view; B. Posterior part of body, ventral view; C. Head, collum and segments 2–4, dorsal view; D. Telson, ventral view. Scale bars: A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7766035" httpUri="https://zenodo.org/record/7766035/files/figure.png" pageId="28" pageNumber="77">33</figureCitation>
). Head covered with sparse pubescence throughout, vertigial sulcus apparent (
<figureCitation box="[1069,1211,799,826]" captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Figs 32A, B</figureCitation>
;
<figureCitation box="[1224,1309,799,825]" captionStart="FIGURE 33" captionStartId="26.[151,250,1947,1971]" captionTargetBox="[202,1386,181,1922]" captionTargetId="figure-20@26.[202,1386,181,1922]" captionTargetPageId="26" captionText="FIGURE 33. Epanerchodus wuxi sp. nov., holotype. A. Anterior part of body, ventral view; B. Posterior part of body, ventral view; C. Head, collum and segments 2–4, dorsal view; D. Telson, ventral view. Scale bars: A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7766035" httpUri="https://zenodo.org/record/7766035/files/figure.png" pageId="28" pageNumber="77">33A, C</figureCitation>
). Antennae long and slender, clavate,
<emphasis box="[444,513,836,861]" italics="true" pageId="28" pageNumber="77">in situ</emphasis>
reaching behind segment 4. Collum fan-shaped, with irregular sparse setae, and a faint lateral incision/denticle on each side (
<figureCitation box="[633,774,871,898]" captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Figs 32A, B</figureCitation>
;
<figureCitation box="[787,837,871,897]" captionStart="FIGURE 33" captionStartId="26.[151,250,1947,1971]" captionTargetBox="[202,1386,181,1922]" captionTargetId="figure-20@26.[202,1386,181,1922]" captionTargetPageId="26" captionText="FIGURE 33. Epanerchodus wuxi sp. nov., holotype. A. Anterior part of body, ventral view; B. Posterior part of body, ventral view; C. Head, collum and segments 2–4, dorsal view; D. Telson, ventral view. Scale bars: A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7766035" httpUri="https://zenodo.org/record/7766035/files/figure.png" pageId="28" pageNumber="77">33C</figureCitation>
). Paraterga distinct and broad (
<figureCitation box="[1186,1273,871,897]" captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Figs 32</figureCitation>
,
<figureCitation box="[1285,1316,871,897]" captionStart="FIGURE 33" captionStartId="26.[151,250,1947,1971]" captionTargetBox="[202,1386,181,1922]" captionTargetId="figure-20@26.[202,1386,181,1922]" captionTargetPageId="26" captionText="FIGURE 33. Epanerchodus wuxi sp. nov., holotype. A. Anterior part of body, ventral view; B. Posterior part of body, ventral view; C. Head, collum and segments 2–4, dorsal view; D. Telson, ventral view. Scale bars: A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7766035" httpUri="https://zenodo.org/record/7766035/files/figure.png" pageId="28" pageNumber="77">33</figureCitation>
), midbody metaterga about 1.9 times as wide as prozonae. All paraterga slightly upturned dorsally above a faintly convex dorsum (
<figureCitation box="[213,297,943,969]" captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Fig. 32</figureCitation>
). Anterior edge of metaterga forming a shoulder (
<figureCitation box="[852,938,943,969]" captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Figs 32</figureCitation>
,
<figureCitation box="[950,981,943,969]" captionStart="FIGURE 33" captionStartId="26.[151,250,1947,1971]" captionTargetBox="[202,1386,181,1922]" captionTargetId="figure-20@26.[202,1386,181,1922]" captionTargetPageId="26" captionText="FIGURE 33. Epanerchodus wuxi sp. nov., holotype. A. Anterior part of body, ventral view; B. Posterior part of body, ventral view; C. Head, collum and segments 2–4, dorsal view; D. Telson, ventral view. Scale bars: A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7766035" httpUri="https://zenodo.org/record/7766035/files/figure.png" pageId="28" pageNumber="77">33</figureCitation>
). Caudolateral corners of paraterga from rings 2 protruded toward back-inner past tergal margin, subsequent ones acuminate, especially on rings 18–19 (
<figureCitation captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Figs 32</figureCitation>
,
<figureCitation box="[194,225,1015,1041]" captionStart="FIGURE 33" captionStartId="26.[151,250,1947,1971]" captionTargetBox="[202,1386,181,1922]" captionTargetId="figure-20@26.[202,1386,181,1922]" captionTargetPageId="26" captionText="FIGURE 33. Epanerchodus wuxi sp. nov., holotype. A. Anterior part of body, ventral view; B. Posterior part of body, ventral view; C. Head, collum and segments 2–4, dorsal view; D. Telson, ventral view. Scale bars: A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7766035" httpUri="https://zenodo.org/record/7766035/files/figure.png" pageId="28" pageNumber="77">33</figureCitation>
). Integument shining, translucent, prozonae very delicately alveolate (
<figureCitation box="[1020,1108,1015,1041]" captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Figs 32</figureCitation>
,
<figureCitation box="[1121,1171,1015,1041]" captionStart="FIGURE 33" captionStartId="26.[151,250,1947,1971]" captionTargetBox="[202,1386,181,1922]" captionTargetId="figure-20@26.[202,1386,181,1922]" captionTargetPageId="26" captionText="FIGURE 33. Epanerchodus wuxi sp. nov., holotype. A. Anterior part of body, ventral view; B. Posterior part of body, ventral view; C. Head, collum and segments 2–4, dorsal view; D. Telson, ventral view. Scale bars: A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7766035" httpUri="https://zenodo.org/record/7766035/files/figure.png" pageId="28" pageNumber="77">33C</figureCitation>
). Three or four notches at lateral margins of paraterga (
<figureCitation box="[500,586,1051,1077]" captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Figs 32</figureCitation>
,
<figureCitation box="[598,629,1051,1077]" captionStart="FIGURE 33" captionStartId="26.[151,250,1947,1971]" captionTargetBox="[202,1386,181,1922]" captionTargetId="figure-20@26.[202,1386,181,1922]" captionTargetPageId="26" captionText="FIGURE 33. Epanerchodus wuxi sp. nov., holotype. A. Anterior part of body, ventral view; B. Posterior part of body, ventral view; C. Head, collum and segments 2–4, dorsal view; D. Telson, ventral view. Scale bars: A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7766035" httpUri="https://zenodo.org/record/7766035/files/figure.png" pageId="28" pageNumber="77">33</figureCitation>
). Constriction between pro- and metazonae narrow, shallow and smooth (
<figureCitation box="[159,245,1087,1113]" captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Fig. 32</figureCitation>
). Metatergal sculpture faint, with three irregular transverse rows of setigerous polygonal bosses (
<figureCitation box="[1339,1424,1087,1113]" captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Fig. 32</figureCitation>
). Sulcus between front and middle rows of setae a little deeper than that between middle and caudal rows (
<figureCitation box="[1339,1424,1123,1149]" captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Fig. 32</figureCitation>
). Tergal setae very short, sparse.
</paragraph>
<paragraph blockId="28.[151,1437,259,1618]" pageId="28" pageNumber="77">
Pore formula normal: 5, 7, 9, 10, 12, 13, 15–19. Ozopores large and evident, located between last and penultimate marginal incisions. Epiproct short, conical, slightly surpassing paraprocts caudally, pre-apical lateral papillae small, with two long setae (
<figureCitation box="[456,559,1267,1293]" captionStart="FIGURE 32" captionStartId="25.[151,250,1727,1751]" captionTargetBox="[151,1435,181,1704]" captionTargetId="figure-19@25.[151,1435,181,1704]" captionTargetPageId="25" captionText="FIGURE 32. Epanerchodus wuxi sp. nov., holotype. A. Living specimen; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766033" httpUri="https://zenodo.org/record/7766033/files/figure.png" pageId="28" pageNumber="77">Fig. 32C</figureCitation>
). Hypoproct lunate, with two setigerous conical papillae at caudal margin (
<figureCitation captionStart="FIGURE 33" captionStartId="26.[151,250,1947,1971]" captionTargetBox="[202,1386,181,1922]" captionTargetId="figure-20@26.[202,1386,181,1922]" captionTargetPageId="26" captionText="FIGURE 33. Epanerchodus wuxi sp. nov., holotype. A. Anterior part of body, ventral view; B. Posterior part of body, ventral view; C. Head, collum and segments 2–4, dorsal view; D. Telson, ventral view. Scale bars: A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7766035" httpUri="https://zenodo.org/record/7766035/files/figure.png" pageId="28" pageNumber="77">Fig. 33B, C</figureCitation>
). Sterna sparsely setose, cross-shaped impressions shallow (
<figureCitation box="[912,1049,1303,1329]" captionStart="FIGURE 33" captionStartId="26.[151,250,1947,1971]" captionTargetBox="[202,1386,181,1922]" captionTargetId="figure-20@26.[202,1386,181,1922]" captionTargetPageId="26" captionText="FIGURE 33. Epanerchodus wuxi sp. nov., holotype. A. Anterior part of body, ventral view; B. Posterior part of body, ventral view; C. Head, collum and segments 2–4, dorsal view; D. Telson, ventral view. Scale bars: A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7766035" httpUri="https://zenodo.org/record/7766035/files/figure.png" pageId="28" pageNumber="77">Fig. 33A, B</figureCitation>
). Legs long and slender, about 2.0 times as long as body ring height.
</paragraph>
<paragraph blockId="28.[151,1437,259,1618]" pageId="28" pageNumber="77">
Coxa of gonopod (
<figureCitation box="[411,494,1375,1401]" captionStart="FIGURE 34" captionStartId="27.[151,250,1981,2005]" captionTargetBox="[198,1396,185,1950]" captionTargetId="figure-19@27.[190,1397,181,1957]" captionTargetPageId="27" captionText="FIGURE 34. Epanerchodus wuxi sp. nov., holotype. A. Right gonopod, mesal view; B. Right gonopod, lateral view; C. Subapical part of endomere, sub-medial view; D. Apical part of endomere, top view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7766037" httpUri="https://zenodo.org/record/7766037/files/figure.png" pageId="28" pageNumber="77">Fig. 34</figureCitation>
) large and squarish, as usual. Prefemur densely setose and nearly half the length of telopodite. Femorite with a clivus (
<emphasis bold="true" box="[543,563,1411,1437]" pageId="28" pageNumber="77">cl</emphasis>
), a process (
<emphasis bold="true" box="[705,735,1411,1437]" pageId="28" pageNumber="77">p2</emphasis>
) medially and a finger-shaped process (
<emphasis bold="true" box="[1181,1211,1411,1437]" pageId="28" pageNumber="77">p1</emphasis>
) distally. Endomere (
<emphasis bold="true" box="[161,189,1447,1473]" pageId="28" pageNumber="77">en</emphasis>
) strongly curved, broad subapical, with a small subapical curved lobe (
<emphasis bold="true" box="[1009,1017,1447,1473]" pageId="28" pageNumber="77">l</emphasis>
), a row of apical denticles (
<emphasis bold="true" box="[1341,1357,1447,1473]" pageId="28" pageNumber="77">d</emphasis>
) and a spiny protuberance medially (
<emphasis bold="true" box="[489,515,1483,1509]" pageId="28" pageNumber="77">sp</emphasis>
). Seminal groove starting mesally, distally recurved laterad near base of
<emphasis bold="true" box="[1343,1373,1483,1509]" pageId="28" pageNumber="77">p1</emphasis>
, then run into an accessory seminal chamber, the latter opening on a hairy pulvillus, which makes a clear-cut distolateral loop. Exomere absent.
</paragraph>
</subSubSection>
<subSubSection box="[199,1047,1591,1618]" pageId="28" pageNumber="77" type="distribution">
<paragraph blockId="28.[151,1437,259,1618]" box="[199,1047,1591,1618]" pageId="28" pageNumber="77">
<emphasis bold="true" box="[199,353,1591,1617]" pageId="28" pageNumber="77">Distribution.</emphasis>
Known only from the
<typeStatus box="[608,656,1592,1618]" pageId="28" pageNumber="77">type</typeStatus>
locality, Wuxi,
<collectingRegion box="[834,961,1591,1617]" country="China" name="Chongqing" pageId="28" pageNumber="77">Chongqing</collectingRegion>
,
<collectingCountry box="[973,1044,1591,1617]" name="China" pageId="28" pageNumber="77">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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