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<document id="F77D369537CDF122F0DC36795B47CDD1" ID-CLB-Dataset="28117" ID-DOI="10.11646/zootaxa.4568.3.8" ID-GBIF-Dataset="6496901c-5528-4fe3-96d6-0f4cf2be9c10" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601770" ID-ZooBank="5882BDAC-02E4-4EEA-9AFF-EE4A509FFDC6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553173246349" checkinUser="plazi" docAuthor="Runyon, Justin B." docDate="2019" docId="BF14B73B0D1BFF98FF30A03A4881FEE8" docLanguage="en" docName="zootaxa.4568.3.8.pdf" docOrigin="Zootaxa 4568 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hurleyella Runyon &amp; Robinson 2010" docType="key" docVersion="9" lastPageNumber="551" masterDocId="432DCF430D19FF9BFFA7A66C4D1DFFCA" masterDocTitle="Two new species of Hurleyella Runyon &amp; Robinson (Diptera: Dolichopodidae), with the first record from the Neotropics" masterLastPageNumber="560" masterPageNumber="548" pageNumber="550" updateTime="1698703610928" updateUser="ExternalLinkService">
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<mods:title id="22CE635164AD96E0CF25EC5AEDF155D5">Two new species of Hurleyella Runyon &amp; Robinson (Diptera: Dolichopodidae), with the first record from the Neotropics</mods:title>
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<treatment id="BF14B73B0D1BFF98FF30A03A4881FEE8" ID-DOI="http://doi.org/10.5281/zenodo.5313971" ID-GBIF-Taxon="154475058" ID-Zenodo-Dep="5313971" LSID="urn:lsid:plazi:treatment:BF14B73B0D1BFF98FF30A03A4881FEE8" httpUri="http://treatment.plazi.org/id/BF14B73B0D1BFF98FF30A03A4881FEE8" lastPageId="3" lastPageNumber="551" pageId="2" pageNumber="550">
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<heading id="6C4AB1410D1BFF99FF30A03A4E44F9BA" bold="true" box="[151,857,1621,1648]" fontSize="11" level="1" pageId="2" pageNumber="550" reason="1">
<emphasis id="05C9DA3F0D1BFF99FF30A03A4E44F9BA" bold="true" box="[151,857,1621,1648]" pageId="2" pageNumber="550">
Key to species of
<taxonomicName id="F0BD7DAE0D1BFF99FECAA0394FEAF9BA" authority="Runyon &amp; Robinson" authorityName="Runyon &amp; Robinson" authorityYear="2010" box="[365,759,1621,1648]" class="Insecta" family="Dolichopodidae" genus="Hurleyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="550" phylum="Arthropoda" rank="genus">
<emphasis id="05C9DA3F0D1BFF99FECAA0394CF0F9A5" bold="true" box="[365,493,1621,1647]" italics="true" pageId="2" pageNumber="550">Hurleyella</emphasis>
Runyon &amp; Robinson
</taxonomicName>
(males)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FA755A60D1BFF98FF30A0F04881FEE8" lastPageId="3" lastPageNumber="551" pageId="2" pageNumber="550" type="key">
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<paragraph id="3702062D0D1BFF99FF30A0F04881F925" blockId="2.[151,1436,1692,2014]" pageId="2" pageNumber="550">
<keyLead id="8C4CA5180D1BFF99FF30A0F04881F925" pageId="2" pageNumber="550">
1 Body length approximately
<quantity id="F045ABC80D1BFF99FE4BA0F04F3BF97B" box="[492,550,1692,1713]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="550" unit="mm" value="1.0">1 mm</quantity>
; wing with vein R
<subScript id="AB3904680D1BFF99FD72A0C64FF7F97C" attach="left" box="[725,746,1706,1718]" fontSize="5" pageId="2" pageNumber="550">2+3</subScript>
unusually short and veins R
<subScript id="AB3904680D1BFF99FC5EA0C64913F97C" attach="left" box="[1017,1038,1706,1718]" fontSize="5" pageId="2" pageNumber="550">4+5</subScript>
and M
<subScript id="AB3904680D1BFF99FBF6A0C64945F97C" attach="left" box="[1105,1112,1706,1718]" fontSize="5" pageId="2" pageNumber="550">1</subScript>
converging to nearly parallel apically (as in
<figureCitation id="AF861AA80D1BFF99FEF2A0D04CB3F91B" box="[341,430,1724,1745]" captionStart="FIGURES 34" captionStartId="2.[151,264,1164,1187]" captionTargetBox="[340,1247,195,1142]" captionTargetId="figure@2.[340,1247,193,1143]" captionTargetPageId="2" captionText="FIGURES 34. Photographs of male wings. 3. Hurleyella belizensis sp. nov.; 4. Hurleyella salina sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/2601774/files/figure.png" pageId="2" pageNumber="550">Figs 3, 4</figureCitation>
); palpus and fore tarsus unmodified; wing without median fuscous band; hypopygium without distinct projections from one side;
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<emphasis id="05C9DA3F0D1BFF99FE45A0B54F54F925" box="[482,585,1753,1775]" italics="true" pageId="2" pageNumber="550">Hurleyella</emphasis>
</taxonomicName>
...................................................................... 2
</keyLead>
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- Without above combination of characters.................................................. other
<taxonomicName id="F0BD7DAE0D1BFF99FAA5A09B4886F8C6" ID-CoL="623XN" authorityName="Latreille" authorityYear="1809" box="[1282,1435,1783,1804]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="550" phylum="Arthropoda" rank="family">Dolichopodidae</taxonomicName>
</keyLead>
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<keyStep id="8C491E880D1BFF99FF30A1784886F80B" pageId="2" pageNumber="550">
<paragraph id="3702062D0D1BFF99FF30A1784881F8AD" blockId="2.[151,1436,1692,2014]" pageId="2" pageNumber="550">
<keyLead id="8C4CA5180D1BFF99FF30A1784881F8AD" pageId="2" pageNumber="550">
2 Coxa I mostly yellow; face and frons covered with dense silver pruinosity (
<figureCitation id="AF861AA80D1BFF99FC1BA1784EEAF8E3" box="[956,1015,1812,1833]" captionStart="FIGURES 56" captionStartId="4.[151,264,1038,1061]" captionTargetBox="[340,1247,555,1017]" captionTargetId="figure@4.[340,1247,555,1017]" captionTargetPageId="4" captionText="FIGURES 56. Photographs of male heads, anterior view. 5. Hurleyella belizensis sp. nov.; 6. Hurleyella salina sp. nov. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2601776/files/figure.png" pageId="2" pageNumber="550">Fig. 6</figureCitation>
); wing with vein M
<subScript id="AB3904680D1BFF99FB1FA14D49A2F8E7" attach="left" box="[1208,1215,1825,1837]" fontSize="5" pageId="2" pageNumber="550">1</subScript>
arching greatly anteriorly on apical half (
<figureCitation id="AF861AA80D1BFF99FE3CA1584CCBF883" box="[411,470,1844,1865]" captionStart="FIGURES 34" captionStartId="2.[151,264,1164,1187]" captionTargetBox="[340,1247,195,1142]" captionTargetId="figure@2.[340,1247,193,1143]" captionTargetPageId="2" captionText="FIGURES 34. Photographs of male wings. 3. Hurleyella belizensis sp. nov.; 4. Hurleyella salina sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/2601774/files/figure.png" pageId="2" pageNumber="550">Fig. 4</figureCitation>
); halter bright yellow (
<figureCitation id="AF861AA80D1BFF99FD17A1584FF6F883" box="[688,747,1844,1865]" captionStart="FIGURES 12" captionStartId="1.[151,264,634,657]" captionTargetBox="[192,1394,198,606]" captionTargetId="figure@1.[192,1394,195,610]" captionTargetPageId="1" captionText="FIGURES 12. Habitus photographs of male holotypes. 1. Hurleyella belizensis sp. nov.; 2. Hurleyella salina sp. nov. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2601772/files/figure.png" pageId="2" pageNumber="550">Fig. 2</figureCitation>
); hypandrium nearly symmetrical, hood-like (
<figureCitation id="AF861AA80D1BFF99FB05A1584818F883" box="[1186,1285,1844,1865]" captionStart="FIGURES 910" captionStartId="6.[151,264,1843,1866]" captionTargetBox="[153,1396,188,1825]" captionTargetId="figure@6.[150,1436,188,1828]" captionTargetPageId="6" captionText="FIGURES 910. Hurleyella salina sp. nov., male terminalia. 9. ventral view; 10. left lateral view. Color codes of structures: light brown—epandrium; red—hypandrium; green—phallus + sperm pump + ejaculatory apodeme; blue—surstylus; orangebacilliform sclerite; purple—subepandrial sclerite; dark gray—abdominal sternite 8 and tergite 7. Abbreviations: epandepandrium; epd lobe—epandrial lobe; fsur—lateroapical flange of surstylus; hf—hypopygial foramen; hypd—hypandrium; hyprct—hypoproct; left v epand lb—left ventral lobe of epandrium; right v epand lb—right ventral lobe of epandrium; sbepand scl—subepandrial sclerite; S—abdominal sternite; T—abdominal tergite. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2601780/files/figure.png" pageId="2" pageNumber="550">Figs 9, 10</figureCitation>
); alkali habitats (
<figureCitation id="AF861AA80D1BFF99FF40A13D4C48F8AC" box="[231,341,1873,1894]" captionStart="FIGURES 1520" captionStartId="8.[151,264,1668,1691]" captionTargetBox="[181,1402,224,1647]" captionTargetId="figure@8.[181,1405,193,1648]" captionTargetPageId="8" captionText="FIGURES 1520. Collection localities and habitats of Hurleyella salina sp. nov. 15, 16. Type locality of H. salina sp. nov. at Deadmans Basin, Montana, USA (46°20.53N 109°22.94W); 16. Specimens of H. salina sp. nov. were found at Deadmans Basin in areas with lots of salt, mud cracks, and a little moisture; 17. Specimens were collected at Piedmont Pond, Montana, USA (45°50.634N 112°06.848W) on mostly dry soil with scattered grasses and a little crusted salt; 18. At Gyp Springs in south central Montana, USA (45°00.36N 108°25.80W), specimens were found on bare soil along lower wall of small canyon where water evaporates and salt forms (left center); 19. At Salt Creek in west central Wyoming, USA (42°28.00N 110°58.07W), H. salina sp. nov. was taken from areas near the creek with lots of salt (lower left); 20. At Fall Creek in eastern Idaho, USA (43°25.48N 111°24.86W), specimens were found on dry part of a travertine mound formed by cold mineral springs with evidence of some salt and other minerals." httpUri="https://zenodo.org/record/2601784/files/figure.png" pageId="2" pageNumber="550">Figs 1520</figureCitation>
) (
<collectingRegion id="F579C8CF0D1BFF99FECEA13D4CBEF8AC" box="[361,419,1873,1894]" country="United States of America" name="Idaho" pageId="2" pageNumber="550">Idaho</collectingRegion>
,
<collectingRegion id="F579C8CF0D1BFF99FE0BA13D4F19F8AC" box="[428,516,1873,1894]" country="United States of America" name="Montana" pageId="2" pageNumber="550">Montana</collectingRegion>
,
<collectingRegion id="F579C8CF0D1BFF99FDA9A13D4F72F8AC" box="[526,623,1873,1894]" country="United States of America" name="Wyoming" pageId="2" pageNumber="550">Wyoming</collectingRegion>
)....................................................
<taxonomicName id="F0BD7DAE0D1BFF99FB56A13E4854F8AD" authority="Runyon, 2019" authorityName="Runyon" authorityYear="2019" box="[1265,1353,1873,1895]" class="Insecta" family="Dolichopodidae" genus="Hurleyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="550" phylum="Arthropoda" rank="species" species="salina" status="sp. nov.">
<emphasis id="05C9DA3F0D1BFF99FB56A13E4854F8AD" box="[1265,1353,1873,1895]" italics="true" pageId="2" pageNumber="550">H. salina</emphasis>
</taxonomicName>
<emphasis id="05C9DA3F0D1BFF99FAE8A13D4881F8AD" bold="true" box="[1359,1436,1873,1895]" pageId="2" pageNumber="550">
<taxonomicNameLabel id="1EFA67440D1BFF99FAE8A13D4881F8AD" box="[1359,1436,1873,1895]" pageId="2" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="3702062D0D1BFF99FF30A1034886F80B" blockId="2.[151,1436,1692,2014]" pageId="2" pageNumber="550">
<keyLead id="8C4CA5180D1BFF99FF30A1034886F80B" pageId="2" pageNumber="550">
- Coxa I brown to black; face and frons covered with brownish pruinosity (as in
<figureCitation id="AF861AA80D1BFF99FC77A1034914F84E" box="[976,1033,1903,1924]" captionStart="FIGURES 56" captionStartId="4.[151,264,1038,1061]" captionTargetBox="[340,1247,555,1017]" captionTargetId="figure@4.[340,1247,555,1017]" captionTargetPageId="4" captionText="FIGURES 56. Photographs of male heads, anterior view. 5. Hurleyella belizensis sp. nov.; 6. Hurleyella salina sp. nov. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/2601776/files/figure.png" pageId="2" pageNumber="550">Fig. 5</figureCitation>
); wing with M
<subScript id="AB3904680D1BFF99FB30A1104983F842" attach="left" box="[1175,1182,1916,1928]" fontSize="5" pageId="2" pageNumber="550">1</subScript>
slightly arching anteriorly only near apex (
<figureCitation id="AF861AA80D1BFF99FEDBA1E24CAAF869" box="[380,439,1934,1955]" captionStart="FIGURES 34" captionStartId="2.[151,264,1164,1187]" captionTargetBox="[340,1247,195,1142]" captionTargetId="figure@2.[340,1247,193,1143]" captionTargetPageId="2" captionText="FIGURES 34. Photographs of male wings. 3. Hurleyella belizensis sp. nov.; 4. Hurleyella salina sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/2601774/files/figure.png" pageId="2" pageNumber="550">Fig. 3</figureCitation>
;
<bibRefCitation id="532C7BDC0D1BFF99FE65A1E24FDCF869" author="Runyon, J. B. &amp; Robinson, H." box="[450,705,1934,1955]" pageId="2" pageNumber="550" pagination="57 - 65" refId="ref7031" refString="Runyon, J. B. &amp; Robinson, H. (2010) Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa, 2400, 57 - 65. https: // doi. org / 10.11646 / zootaxa. 2400.1.6" type="journal article" year="2010">Runyon &amp; Robinson 2010</bibRefCitation>
, fig. 3); halter brown; hypandrium asymmetrical (as in
<figureCitation id="AF861AA80D1BFF99FB70A1E24836F869" box="[1239,1323,1934,1955]" captionStart="FIGURES 78" captionStartId="5.[151,264,1902,1925]" captionTargetBox="[176,1360,242,1881]" captionTargetId="figure@5.[176,1408,193,1891]" captionTargetPageId="5" captionText="FIGURES 78. Hurleyella belizensis sp. nov., male terminalia. 7. ventral view; 8. left lateral view. Abbreviations: epandepandrium; epd lobe—epandrial lobe; fsur—lateroapical flange of surstylus; hyprct—hypoproct; left hypd lobe—left lobe of hypandrium; right hypd lobe—right lobe of hypandrium; sbepand scl—subepandrial sclerite; S—abdominal sternite; Tabdominal tergite. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2601778/files/figure.png" pageId="2" pageNumber="550">Figs 7, 8</figureCitation>
), not hoodlike; habitat not noticeably alkaline....................................................................... 3
</keyLead>
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<paragraph id="3702062D0D1BFF99FF30A1A54881F814" blockId="2.[151,1436,1692,2014]" box="[151,1436,1993,2014]" pageId="2" pageNumber="550">
<keyLead id="8C4CA5180D1BFF99FF30A1A54881F814" box="[151,1436,1993,2014]" pageId="2" pageNumber="550">
3 Surstylus apically with large membranous flange (
<bibRefCitation id="532C7BDC0D1BFF99FD65A1A54EE3F814" author="Runyon, J. B. &amp; Robinson, H." box="[706,1022,1993,2014]" pageId="2" pageNumber="550" pagination="57 - 65" refId="ref7031" refString="Runyon, J. B. &amp; Robinson, H. (2010) Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa, 2400, 57 - 65. https: // doi. org / 10.11646 / zootaxa. 2400.1.6" type="journal article" year="2010">Runyon &amp; Robinson 2010, fig. 5</bibRefCitation>
) (Nearctic)........................ 4
</keyLead>
</paragraph>
<paragraph id="3702062D0D1AFF98FF30A6FB4881FF67" blockId="3.[151,1436,151,290]" box="[151,1436,151,173]" pageId="3" pageNumber="551">
<keyLead id="8C4CA5180D1AFF98FF30A6FB4881FF67" box="[151,1436,151,173]" pageId="3" pageNumber="551">
- Surstylus with very small, inconspicuous membranous flange (
<figureCitation id="AF861AA80D1AFF98FC9DA6FB4E8DFF66" box="[826,912,151,172]" captionStart="FIGURES 78" captionStartId="5.[151,264,1902,1925]" captionTargetBox="[176,1360,242,1881]" captionTargetId="figure@5.[176,1408,193,1891]" captionTargetPageId="5" captionText="FIGURES 78. Hurleyella belizensis sp. nov., male terminalia. 7. ventral view; 8. left lateral view. Abbreviations: epandepandrium; epd lobe—epandrial lobe; fsur—lateroapical flange of surstylus; hyprct—hypoproct; left hypd lobe—left lobe of hypandrium; right hypd lobe—right lobe of hypandrium; sbepand scl—subepandrial sclerite; S—abdominal sternite; Tabdominal tergite. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2601778/files/figure.png" pageId="3" pageNumber="551">Figs 7, 8</figureCitation>
, fsur) (
<collectingCountry id="4FAA46BD0D1AFF98FC72A6FB4908FF66" box="[981,1045,151,172]" name="Belize" pageId="3" pageNumber="551">Belize</collectingCountry>
)..............
<taxonomicName id="F0BD7DAE0D1AFF98FB69A6F44855FF67" authority="Runyon, 2019" authorityName="Runyon" authorityYear="2019" box="[1230,1352,151,173]" class="Insecta" family="Dolichopodidae" genus="Hurleyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="551" phylum="Arthropoda" rank="species" species="belizensis" status="sp. nov.">
<emphasis id="05C9DA3F0D1AFF98FB69A6F44855FF67" box="[1230,1352,151,173]" italics="true" pageId="3" pageNumber="551">H. belizensis</emphasis>
</taxonomicName>
<emphasis id="05C9DA3F0D1AFF98FAE8A6FB4881FF67" bold="true" box="[1359,1436,151,173]" pageId="3" pageNumber="551">
<taxonomicNameLabel id="1EFA67440D1AFF98FAE8A6FB4881FF67" box="[1359,1436,151,173]" pageId="3" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="8C491E880D1AFF98FF30A6D94881FEE8" pageId="3" pageNumber="551">
<paragraph id="3702062D0D1AFF98FF30A6D94881FF2D" blockId="3.[151,1436,151,290]" pageId="3" pageNumber="551">
<keyLead id="8C4CA5180D1AFF98FF30A6D94881FF2D" pageId="3" pageNumber="551">
4 First flagellomere of antenna distinctly wider than narrowest part of face; males with flange of surstylus relatively narrow, tapered (
<bibRefCitation id="532C7BDC0D1AFF98FE93A6BE4FBFFF2D" author="Runyon, J. B. &amp; Robinson, H." box="[308,674,210,231]" pageId="3" pageNumber="551" pagination="57 - 65" refId="ref7031" refString="Runyon, J. B. &amp; Robinson, H. (2010) Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa, 2400, 57 - 65. https: // doi. org / 10.11646 / zootaxa. 2400.1.6" type="journal article" year="2010">Runyon &amp; Robinson 2010, figs 5AB</bibRefCitation>
) (
<collectingRegion id="F579C8CF0D1AFF98FD12A6BE4E1BFF2D" box="[693,774,210,231]" country="United States of America" name="Virginia" pageId="3" pageNumber="551">Virginia</collectingRegion>
).....................
<taxonomicName id="F0BD7DAE0D1AFF98FBA8A6BE4881FF2D" authority="Runyon &amp; Robinson" authorityName="Runyon &amp; Robinson" box="[1039,1436,209,231]" class="Insecta" family="Dolichopodidae" genus="Hurleyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="551" phylum="Arthropoda" rank="species" species="cumberlandensis">
<emphasis id="05C9DA3F0D1AFF98FBA8A6BE49D2FF2D" box="[1039,1231,209,231]" italics="true" pageId="3" pageNumber="551">H. cumberlandensis</emphasis>
Runyon &amp; Robinson
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="3702062D0D1AFF98FF30A6834881FEE8" blockId="3.[151,1436,151,290]" pageId="3" pageNumber="551">
<keyLead id="8C4CA5180D1AFF98FF30A6834881FEE8" pageId="3" pageNumber="551">
- Width of first flagellomere of antenna subequal to narrowest part of face; males with flange of surstylus spatulate (
<bibRefCitation id="532C7BDC0D1AFF98FA92A6834CDFFEE8" author="Runyon, J. B. &amp; Robinson, H." pageId="3" pageNumber="551" pagination="57 - 65" refId="ref7031" refString="Runyon, J. B. &amp; Robinson, H. (2010) Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa, 2400, 57 - 65. https: // doi. org / 10.11646 / zootaxa. 2400.1.6" type="journal article" year="2010">Runyon &amp; Robinson 2010, fig. 5C</bibRefCitation>
) (
<collectingRegion id="F579C8CF0D1AFF98FE71A7614F0CFEE8" box="[470,529,269,290]" country="United States of America" name="Texas" pageId="3" pageNumber="551">Texas</collectingRegion>
).................................................
<taxonomicName id="F0BD7DAE0D1AFF98FBCDA7614881FEE8" authority="Runyon &amp; Robinson" authorityName="Runyon &amp; Robinson" box="[1130,1436,268,290]" class="Insecta" family="Dolichopodidae" genus="Hurleyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="551" phylum="Arthropoda" rank="species" species="brooksi">
<emphasis id="05C9DA3F0D1AFF98FBCDA76149D3FEE8" box="[1130,1230,268,290]" italics="true" pageId="3" pageNumber="551">H. brooksi</emphasis>
Runyon &amp; Robinson
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>