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<document id="A8B768DAD493917740DDF7E3479AB4DC" ID-CLB-Dataset="36828" ID-DOI="10.11646/zootaxa.3824.1.1" ID-GBIF-Dataset="df227efc-13d9-421e-92ed-f9aa417b0cf7" ID-ISSN="1175-5326" ID-Zenodo-Dep="286404" ID-ZooBank="023D78D0-4182-48D2-BAEB-CDA6473CF585" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460443577100" checkinUser="plazi" docAuthor="Webster, Mark &amp; Bohach, Lisa L." docDate="2014" docId="B10C8793FFECFF9D61B5FE0EFCE98041" docLanguage="en" docName="zt03824p066.pdf" docOrigin="Zootaxa 3824 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lochmanolenellus bispinata Harrington 1956" docType="treatment" docVersion="6" lastPageNumber="21" masterDocId="4D35FFEBFFFDFF886122FFE2FFE18379" masterDocTitle="Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia" masterLastPageNumber="66" masterPageNumber="1" pageNumber="18" updateTime="1698333190807" updateUser="ExternalLinkService">
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<mods:title id="DC926BA49F804F1966ACCA1253562EDD">Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia</mods:title>
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<treatment id="B10C8793FFECFF9D61B5FE0EFCE98041" ID-DOI="http://doi.org/10.5281/zenodo.6129718" ID-GBIF-Taxon="127668552" ID-Zenodo-Dep="6129718" LSID="urn:lsid:plazi:treatment:B10C8793FFECFF9D61B5FE0EFCE98041" httpUri="http://treatment.plazi.org/id/B10C8793FFECFF9D61B5FE0EFCE98041" lastPageId="21" lastPageNumber="21" pageId="17" pageNumber="18">
<subSubSection id="71BF650EFFECFF9961B5FE0EFD148728" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="391A3685FFECFF9961B5FE0EFD80817F" blockId="17.[151,609,492,551]" box="[151,609,492,518]" pageId="17" pageNumber="18">
<heading id="625281E9FFECFF9961B5FE0EFD80817F" bold="true" box="[151,609,492,518]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="0BD1EA97FFECFF9961B5FE0EFD80817F" bold="true" box="[151,609,492,518]" pageId="17" pageNumber="18">
<emphasis id="0BD1EA97FFECFF9961B5FE0EFEED817F" bold="true" box="[151,268,492,518]" italics="true" pageId="17" pageNumber="18">Laudonia</emphasis>
<taxonomicName id="FEA54D06FFECFF996031FE0EFD80817F" authority="Harrington, 1956" authorityName="Harrington" authorityYear="1956" box="[275,609,492,518]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFECFF996031FE0EFE60817F" bold="true" box="[275,385,492,518]" italics="true" pageId="17" pageNumber="18">bispinata</emphasis>
<bibRefCitation id="5D344B74FFECFF9960ABFE0EFD80817F" author="Harrington" box="[393,609,492,518]" pageId="17" pageNumber="18" refString="Harrington, H. J. (1956) Olenellidae with advanced cephalic spines. Journal of Paleontology, 30, 56 - 61." type="journal article" year="1956">Harrington, 1956</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="391A3685FFECFF9961B5FDECFE54815E" blockId="17.[151,609,492,551]" box="[151,437,526,551]" pageId="17" pageNumber="18">Figs 3.2, 4.5, 4.6, 5.1–5.5</paragraph>
<paragraph id="391A3685FFECFF9961B5FDB2FC46811F" blockId="17.[151,1436,592,1105]" box="[151,935,592,614]" pageId="17" pageNumber="18">
1956
<emphasis id="0BD1EA97FFECFF9961F2FDB2FE74811F" box="[208,405,592,614]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996015FDB2FE74811F" authority="Harrington" authorityName="Harrington" box="[311,405,592,614]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington, pp. 60, 61, text-fig. 1f, pl. 15, figs. 4, 5.
</paragraph>
<paragraph id="391A3685FFECFF9961B5FD8CFBEE81FD" blockId="17.[151,1436,592,1105]" box="[151,1039,622,645]" pageId="17" pageNumber="18">
1959
<emphasis id="0BD1EA97FFECFF9961F2FD8CFE7481FD" box="[208,405,622,644]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996015FD8CFE7481FD" authority="Harrington" authorityName="Harrington" box="[311,405,622,644]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington; Harrington in Harrington
<emphasis id="0BD1EA97FFECFF99623FFD92FCA881FD" box="[797,841,622,645]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
., p.
<collectingCountry id="41B27615FFECFF996257FD8CFC6981FD" box="[885,904,622,644]" name="Iceland" pageId="17" pageNumber="18">O</collectingCountry>
154, fig. 38c.
</paragraph>
<paragraph id="391A3685FFECFF9961B5FD6FFC1081DA" blockId="17.[151,1436,592,1105]" box="[151,1009,653,675]" pageId="17" pageNumber="18">
1959
<emphasis id="0BD1EA97FFECFF9961F2FD6FFE7481DA" box="[208,405,653,675]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996015FD6FFE7481DA" authority="Harrington" authorityName="Harrington" box="[311,405,653,675]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington; Poulsen in Harrington
<emphasis id="0BD1EA97FFECFF9963DCFD6CFCCA81DA" box="[766,811,653,675]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
., p.
<collectingCountry id="41B27615FFECFF996275FD6FFC8B81DA" box="[855,874,653,675]" name="Iceland" pageId="17" pageNumber="18">O</collectingCountry>
192, fig. 134.
</paragraph>
<paragraph id="391A3685FFECFF9961B5FD4EFD9C81BB" blockId="17.[151,1436,592,1105]" box="[151,637,684,706]" pageId="17" pageNumber="18">
1971
<emphasis id="0BD1EA97FFECFF9961F2FD4EFE7481BB" box="[208,405,684,706]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996015FD4EFE7481BB" authority="Harrington" authorityName="Harrington" box="[311,405,684,706]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington; Hu, p. 79.
</paragraph>
<paragraph id="391A3685FFECFF9961B5FD29FD6F8198" blockId="17.[151,1436,592,1105]" box="[151,654,715,737]" pageId="17" pageNumber="18">
1972
<emphasis id="0BD1EA97FFECFF9961F2FD29FE748198" box="[208,405,715,737]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996015FD29FE748198" authority="Harrington" authorityName="Harrington" box="[311,405,715,737]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington; Fritz, p. 27.
</paragraph>
<paragraph id="391A3685FFECFF9961B5FD0BFD3C8186" blockId="17.[151,1436,592,1105]" box="[151,733,745,767]" pageId="17" pageNumber="18">
1978
<emphasis id="0BD1EA97FFECFF9961F2FD0BFE748186" box="[208,405,745,767]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996015FD0BFE748186" authority="Harrington" authorityName="Harrington" box="[311,405,745,767]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington; McNamara, p. 642.
</paragraph>
<paragraph id="391A3685FFECFF9961B5FCEAFEC78045" blockId="17.[151,1436,592,1105]" pageId="17" pageNumber="18">
1992
<emphasis id="0BD1EA97FFECFF9961F3FCEAFE798067" box="[209,408,776,798]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996018FCEAFE798067" authority="Harrington" authorityName="Harrington" box="[314,408,776,798]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington; Fritz, pp. 5, 7, 13, 16, 17, 26, 27, text-fig. 6c, pl. 12, figs. 4–8, pl. 13, figs. 1–8, pl. 14, figs. 1, 2.
</paragraph>
<paragraph id="391A3685FFECFF9961B5FCA7FCEF8022" blockId="17.[151,1436,592,1105]" box="[151,782,837,859]" pageId="17" pageNumber="18">
1993
<emphasis id="0BD1EA97FFECFF9961F2FCA7FE748022" box="[208,405,837,859]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996015FCA7FE748022" authority="Harrington" authorityName="Harrington" box="[311,405,837,859]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington; Palmer &amp; Repina, p. 24.
</paragraph>
<paragraph id="391A3685FFECFF9961B5FC86FBCF8003" blockId="17.[151,1436,592,1105]" box="[151,1070,868,890]" pageId="17" pageNumber="18">
1997
<emphasis id="0BD1EA97FFECFF9961F2FC86FE748003" box="[208,405,868,890]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996015FC86FE748003" authority="Harrington" authorityName="Harrington" box="[311,405,868,890]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington; Palmer &amp; Repina in
<bibRefCitation id="5D344B74FFECFF9963CAFC86FC3C8003" author="Whittington" box="[744,989,868,890]" pageId="17" pageNumber="18" refString="Whittington, H. B., Chatterton, B. D. E., Speyer, S. E., Fortey, R. A., Owens, R. M., Chang, W. T., Dean, W. T., Jell, P. A., Laurie, J. R., Palmer, A. R., Repina, L. N., Rushton, A. W. A., Shergold, J. H., Clarkson, E. N. K., Wilmot, N. V. &amp; Kelly, S. R. A. (1997) Treatise on Invertebrate Paleontology. Part O. Arthropoda 1. Trilobita, Revised. Volume 1: Introduction, Order Agnostida, Order Redlichiida. Geological Society of America and Univeristy of Kansas, Boulder, Colorado and Lawrence, Kansas, 530 pp." type="book" year="1997">
Whittington
<emphasis id="0BD1EA97FFECFF99624BFC87FC778003" box="[873,918,868,890]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
., 1997
</bibRefCitation>
, p. 412.
</paragraph>
<paragraph id="391A3685FFECFF9961B5FC61FCB580E0" blockId="17.[151,1436,592,1105]" box="[151,852,899,921]" pageId="17" pageNumber="18">
1998
<emphasis id="0BD1EA97FFECFF9961F2FC61FE7480E0" box="[208,405,899,921]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996015FC61FE7480E0" authority="Harrington" authorityName="Harrington" box="[311,405,899,921]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington; Lieberman, pp. 62, 73, fig. 4.2.
</paragraph>
<paragraph id="391A3685FFECFF9961B5FC43FEB980AF" blockId="17.[151,1436,592,1105]" pageId="17" pageNumber="18">
1999
<emphasis id="0BD1EA97FFECFF9961F7FC43FE7E80CE" box="[213,415,929,951]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996063FC43FE7E80CE" authority="Harrington" authorityName="Harrington" box="[321,415,929,951]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington; Lieberman (part), pp. 107, 110, 114, 140 only [not fig. 20.1, =
<taxonomicName id="FEA54D06FFECFF9965C0FC43FEAB80AF" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pentagonalis">
<emphasis id="0BD1EA97FFECFF9965C0FC43FEAB80AF" italics="true" pageId="17" pageNumber="18">Lochmanolenellus pentagonalis</emphasis>
</taxonomicName>
].
</paragraph>
<paragraph id="391A3685FFECFF9961B5FC3CFCD7808D" blockId="17.[151,1436,592,1105]" box="[151,822,990,1012]" pageId="17" pageNumber="18">
1999
<emphasis id="0BD1EA97FFECFF9961F2FC3CFE74808D" box="[208,405,990,1012]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996015FC3CFE74808D" authority="Harrington" authorityName="Harrington" box="[311,405,990,1012]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington; Smith &amp; Lieberman, p. 462.
</paragraph>
<paragraph id="391A3685FFECFF9961B5FC1FFD38876A" blockId="17.[151,1436,592,1105]" box="[151,729,1021,1043]" pageId="17" pageNumber="18">
2002
<emphasis id="0BD1EA97FFECFF9961F2FC1FFE74876A" box="[208,405,1021,1043]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996015FC1FFE74876A" authority="Harrington" authorityName="Harrington" box="[311,405,1021,1043]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington; Lieberman, p. 699.
</paragraph>
<paragraph id="391A3685FFECFF9961B5FBFEFD2D874B" blockId="17.[151,1436,592,1105]" box="[151,716,1052,1074]" pageId="17" pageNumber="18">
2003
<emphasis id="0BD1EA97FFECFF9961F2FBFEFE74874B" box="[208,405,1052,1074]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996015FBFEFE74874B" authority="Harrington" authorityName="Harrington" box="[311,405,1052,1074]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington; Lieberman, p. 63.
</paragraph>
<paragraph id="391A3685FFECFF9961B5FBD9FD148728" blockId="17.[151,1436,592,1105]" box="[151,757,1083,1105]" pageId="17" pageNumber="18">
2003
<emphasis id="0BD1EA97FFECFF9961F2FBD9FE748728" box="[208,405,1083,1105]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF996015FBD9FE748728" authority="Harrington" authorityName="Harrington" box="[311,405,1083,1105]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
Harrington; Jell &amp; Adrain, p. 396.
</paragraph>
</subSubSection>
<subSubSection id="71BF650EFFECFF9961B5FB94FB7686D6" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="391A3685FFECFF9961B5FB94FB7686D6" blockId="17.[151,1437,1142,2031]" pageId="17" pageNumber="18">
<emphasis id="0BD1EA97FFECFF9961B5FB94FEF087F6" bold="true" box="[151,273,1142,1167]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
Genal spine base transversely opposite lateral margin of LA; distal portion of posterior cephalic margin oriented slightly inwards at approximately 6° (range 10° to 3°) relative to an exsagittal line when traced toward base of genal spine. Distance (tr.) between genal spine bases approximately 189% (range 153% to 229%) of glabellar length (sag.), increasing through phase 5 of cephalic development. Angle between inner margin of proximal portion of genal spine and distal portion of posterior cephalic margin approximately 55° (range 50° to 66°). Width of anterior cephalic border (measured anterior to ocular lobes) 118% to 220% of length (exsag.) of LO, increasing through phase 5 of cephalic development. Ocular lobes divergent from exsagittal axis by approximately 52° (range 29° to 76°); posterior tip of ocular lobe transversely opposite lateral margin of L2, S2, or L3. Intergenal ridge with much narrower ridge running along summit, giving ‘ridge-on-ridge’ appearance.
</paragraph>
</subSubSection>
<subSubSection id="71BF650EFFECFF9B61E5FA59FBCE84DF" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="391A3685FFECFF9961E5FA59FEDF8562" blockId="17.[151,1437,1142,2031]" pageId="17" pageNumber="18">
<emphasis id="0BD1EA97FFECFF9961E5FA59FEDA86AD" bold="true" box="[199,315,1467,1492]" pageId="17" pageNumber="18">
<typeStatus id="E61E8827FFECFF9961E5FA59FED786AD" box="[199,310,1467,1492]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="5FB4AE40FFECFF996064FA5EFE4186AA" box="[326,416,1468,1491]" pageId="17" pageNumber="18">KUMIP</collectionCode>
32400. This specimen was not examined first-hand during the present study, although morphometric data were extracted from published photographs (
<bibRefCitation id="5D344B74FFECFF996258FA3CFBA3868E" author="Harrington" box="[890,1090,1502,1527]" pageId="17" pageNumber="18" refString="Harrington, H. J. (1956) Olenellidae with advanced cephalic spines. Journal of Paleontology, 30, 56 - 61." type="journal article" year="1956">Harrington, 1956</bibRefCitation>
, pl. 15, figs. 4, 5;
<bibRefCitation id="5D344B74FFECFF99643EFA3CFF338562" author="Lieberman" pageId="17" pageNumber="18" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
, fig. 4.2).
</paragraph>
<paragraph id="391A3685FFECFF9B61E5F9C4FB18858B" blockId="17.[151,1437,1142,2031]" lastBlockId="19.[151,1437,1645,2030]" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<emphasis id="0BD1EA97FFECFF9961E5F9C4FE638546" bold="true" box="[199,386,1574,1599]" pageId="17" pageNumber="18">Other material.</emphasis>
Detailed qualitative and/or morphometric data were recorded from first-hand examination of the following
<specimenCount id="2FA3FD0CFFECFF996019F9A9FE3B851D" box="[315,474,1611,1636]" pageId="17" pageNumber="18" type="generic">16 specimens</specimenCount>
:
<collectionCode id="5FB4AE40FFECFF9960C5F9A9FDD9851A" LSID="urn:lsid:biocol.org:col:34871" box="[487,568,1611,1635]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
137534,
<collectionCode id="5FB4AE40FFECFF99638AF9A9FD18851A" LSID="urn:lsid:biocol.org:col:34871" box="[680,761,1611,1635]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
137630c (
<specimenCount id="2FA3FD0CFFECFF996254F9A9FBE9851D" box="[886,1032,1611,1636]" pageId="17" pageNumber="18" type="generic">2 specimens</specimenCount>
),
<collectionCode id="5FB4AE40FFECFF99653DF9A9FB91851A" LSID="urn:lsid:biocol.org:col:34871" box="[1055,1136,1611,1635]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
137630d,
<collectionCode id="5FB4AE40FFECFF9965CFF9A9FADF851A" LSID="urn:lsid:biocol.org:col:34871" box="[1261,1342,1611,1635]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
443757 (also figured by
<bibRefCitation id="5D344B74FFECFF99607AF98CFE3885FE" author="Fritz" box="[344,473,1646,1671]" pageId="17" pageNumber="18" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 12, fig. 4),
<collectionCode id="5FB4AE40FFECFF9963B7F98DFD0785FE" LSID="urn:lsid:biocol.org:col:34871" box="[661,742,1647,1671]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
443758 (also figured by
<bibRefCitation id="5D344B74FFECFF99652DF98CFB7185FE" author="Fritz" box="[1039,1168,1646,1671]" pageId="17" pageNumber="18" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 12, fig. 5),
<collectionCode id="5FB4AE40FFECFF996469F98DFA7D85FE" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,1647,1671]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
443759 (also figured by
<bibRefCitation id="5D344B74FFECFF996094F971FDD685D2" author="Fritz" box="[438,567,1683,1708]" pageId="17" pageNumber="18" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 12, fig. 5),
<collectionCode id="5FB4AE40FFECFF9963D3F971FCA385D2" LSID="urn:lsid:biocol.org:col:34871" box="[753,834,1683,1707]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
443760 (also figured by
<bibRefCitation id="5D344B74FFECFF996549F971FB0D85D2" author="Fritz" box="[1131,1260,1683,1708]" pageId="17" pageNumber="18" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 12, fig. 6),
<collectionCode id="5FB4AE40FFECFF9961B5F955FF0985B6" LSID="urn:lsid:biocol.org:col:34871" box="[151,232,1719,1743]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
443761 (also figured by
<bibRefCitation id="5D344B74FFECFF996326F954FD6385B6" author="Fritz" box="[516,642,1718,1743]" pageId="17" pageNumber="18" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 12, fig. 7),
<collectionCode id="5FB4AE40FFECFF996212F955FC6085B6" LSID="urn:lsid:biocol.org:col:34871" box="[816,897,1719,1743]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
443762 (also figured by
<bibRefCitation id="5D344B74FFECFF9965BCF954FAFD85B6" author="Fritz" box="[1182,1308,1718,1743]" pageId="17" pageNumber="18" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 12, fig. 8),
<collectionCode id="5FB4AE40FFECFF99619FF939FEEF858A" LSID="urn:lsid:biocol.org:col:34871" box="[189,270,1755,1779]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
443763 (also figured by
<bibRefCitation id="5D344B74FFECFF99630EF939FD4A858A" author="Fritz" box="[556,683,1755,1780]" pageId="17" pageNumber="18" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 13, fig. 1),
<collectionCode id="5FB4AE40FFECFF996278F939FC4A858A" LSID="urn:lsid:biocol.org:col:34871" box="[858,939,1755,1779]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
443764 (also figured by
<bibRefCitation id="5D344B74FFECFF9965E8F939FAA8858A" author="Fritz" box="[1226,1353,1755,1780]" pageId="17" pageNumber="18" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 13, figs. 2, 3),
<collectionCode id="5FB4AE40FFECFF996037F91DFE86846E" LSID="urn:lsid:biocol.org:col:34871" box="[277,359,1791,1815]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
443765 (also figured by
<bibRefCitation id="5D344B74FFECFF9963A9F91CFCEC846E" author="Fritz" box="[651,781,1790,1815]" pageId="17" pageNumber="18" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 13, figs. 4–7),
<collectionCode id="5FB4AE40FFECFF9962CAF91DFBDB846E" LSID="urn:lsid:biocol.org:col:34871" box="[1000,1082,1791,1815]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
443766 (also figured by
<bibRefCitation id="5D344B74FFECFF99647CF91CFF338442" author="Fritz" pageId="17" pageNumber="18" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 13, fig. 8),
<collectionCode id="5FB4AE40FFECFF9960A1F8C1FE358442" LSID="urn:lsid:biocol.org:col:34871" box="[387,468,1827,1851]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
443768 (also figured by
<bibRefCitation id="5D344B74FFECFF9963D6F8C1FC928442" author="Fritz" box="[756,883,1827,1852]" pageId="17" pageNumber="18" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 14, figs. 1, 2), and
<collectionCode id="5FB4AE40FFECFF99655FF8C1FB588442" LSID="urn:lsid:biocol.org:col:33791" box="[1149,1209,1827,1851]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="17" pageNumber="18">MCZ</collectionCode>
<date id="4D1B1045FFECFF9965E3F8C1FAF58445" box="[1217,1300,1827,1852]" pageId="17" pageNumber="18" value="80-11-06">110680</date>
(tentatively assigned).
<collectionCode id="5FB4AE40FFECFF996033F8A5FEAF8426" LSID="urn:lsid:biocol.org:col:33791" box="[273,334,1863,1887]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="17" pageNumber="18">MCZ</collectionCode>
<date id="4D1B1045FFECFF996075F8A4FE4B8426" box="[343,426,1862,1887]" pageId="17" pageNumber="18" value="80-11-06">110680</date>
was identifed as
<emphasis id="0BD1EA97FFECFF996357F8A5FCB08426" box="[629,849,1863,1887]" italics="true" pageId="17" pageNumber="18">
Laudonia
<taxonomicName id="FEA54D06FFECFF9963CBF8A5FCB08426" box="[745,849,1863,1887]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
by
<bibRefCitation id="5D344B74FFECFF9962A2F8A4FBA78426" author="Lieberman" box="[896,1094,1862,1887]" pageId="17" pageNumber="18" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998</bibRefCitation>
, p. 73; 1999, p. 114), and the anterior advancement of the genal spine bases and general cephalic outline are consistent with this identification. However, the ocular lobes of this specimen are chipped and their length and angle of divergence cannot be determined with precision.
<collectionCode id="5FB4AE40FFECFF9960FFF851FDF884B2" LSID="urn:lsid:biocol.org:col:33791" box="[477,537,1971,1995]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="17" pageNumber="18">MCZ</collectionCode>
<date id="4D1B1045FFECFF99630AF851FD9A84B5" box="[552,635,1971,1996]" pageId="17" pageNumber="18" value="80-11-06">110680</date>
is therefore herein assigned to
<taxonomicName id="FEA54D06FFECFF996523F856FB7A84B2" box="[1025,1179,1971,1995]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFECFF996523F856FB7A84B2" box="[1025,1179,1971,1995]" italics="true" pageId="17" pageNumber="18">La. bispinata</emphasis>
</taxonomicName>
with reservation. An incomplete cephalon
<collectionCode id="5FB4AE40FFECFF9960B6F835FE098496" LSID="urn:lsid:biocol.org:col:34871" box="[404,488,2007,2031]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
443767 (figured by
<bibRefCitation id="5D344B74FFECFF9963F8F834FCBE8496" author="Fritz" box="[730,863,2006,2031]" pageId="17" pageNumber="18" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 14, fig. 1) was also examined first-hand but qualitative and morphometric data were not recorded.
<bibRefCitation id="5D344B74FFEEFF9B63DEF98FFC6285FF" author="Fritz" box="[764,899,1645,1670]" pageId="19" pageNumber="20" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz (1992)</bibRefCitation>
recovered a single cephalon from
<collectionCode id="5FB4AE40FFEEFF9B6428F98CFAA185FF" box="[1290,1344,1646,1670]" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="19" pageNumber="20">GSC</collectionCode>
locality 87891 and ten cephala from
<collectionCode id="5FB4AE40FFEEFF9B60FFF970FDF285D3" box="[477,531,1682,1706]" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="19" pageNumber="20">GSC</collectionCode>
locality 90577, which were not figured and have not been examined during the course of this study.
<bibRefCitation id="5D344B74FFEEFF9B60A6F957FDA885B7" author="Lieberman" box="[388,585,1717,1742]" pageId="19" pageNumber="20" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998</bibRefCitation>
, p. 73; 1999, p. 114) also assigned
<collectionCode id="5FB4AE40FFEEFF9B62FDF954FBFA85B7" LSID="urn:lsid:biocol.org:col:33791" box="[991,1051,1718,1742]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="19" pageNumber="20">MCZ</collectionCode>
<date id="4D1B1045FFEEFF9B6506F957FB9685B7" box="[1060,1143,1717,1742]" pageId="19" pageNumber="20" value="79-11-06">110679</date>
to this species. However,
<collectionCode id="5FB4AE40FFEEFF9B61B5F938FF35858B" LSID="urn:lsid:biocol.org:col:33791" box="[151,212,1754,1778]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="19" pageNumber="20">MCZ</collectionCode>
<date id="4D1B1045FFEEFF9B61F9F93BFECF858B" box="[219,302,1753,1778]" pageId="19" pageNumber="20" value="79-11-06">110679</date>
is an almost complete individual of
<taxonomicName id="FEA54D06FFEEFF9B63E4F938FBCA858B" box="[710,1067,1754,1778]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pentagonalis">
<emphasis id="0BD1EA97FFEEFF9B63E4F938FBCA858B" box="[710,1067,1754,1778]" italics="true" pageId="19" pageNumber="20">Lochmanolenellus pentagonalis</emphasis>
</taxonomicName>
, described herein.
</paragraph>
<caption id="6DDA660DFFEEFF9B61B5FA2DFBFA8538" httpUri="https://zenodo.org/record/286409/files/figure.png" pageId="19" pageNumber="20" targetBox="[161,1426,204,1456]" targetPageId="19">
<paragraph id="391A3685FFEEFF9B61B5FA2DFBFA8538" blockId="19.[151,1436,1487,1601]" pageId="19" pageNumber="20">
<emphasis id="0BD1EA97FFEEFF9B61B5FA2DFEF2869D" bold="true" box="[151,275,1487,1509]" pageId="19" pageNumber="20">FIGURE 5.</emphasis>
Morphologically mature specimens of
<emphasis id="0BD1EA97FFEEFF9B6385FA2DFCE9869C" box="[679,776,1487,1509]" italics="true" pageId="19" pageNumber="20">Laudonia</emphasis>
<taxonomicName id="FEA54D06FFEEFF9B6232FA2DFBC9869C" authority="Harrington, 1956" authorityName="Harrington" authorityYear="1956" box="[784,1064,1487,1509]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFEEFF9B6232FA2DFC8F869C" box="[784,878,1487,1509]" italics="true" pageId="19" pageNumber="20">bispinata</emphasis>
Harrington, 1956
</taxonomicName>
. 1, Cephalon, internal mold; USNM 443761, x3. 2, Cephalon, internal mold; USNM 443765, x3 (see also Fig. 3.2). 3, Cephalon, external mold; USNM 443763, x2. 4, Cephalon, external mold; USNM 443766, x1.5. 5, Thorax and pygidium, internal mold; USNM 443768, x2. All from middle member of Mural Formation at Locality 61k, Mumm Peak, Mount Robson area, Alberta.
</paragraph>
</caption>
<paragraph id="391A3685FFEEFF9B61E5F91FFBCE84DF" blockId="19.[151,1437,1645,2030]" pageId="19" pageNumber="20">
<emphasis id="0BD1EA97FFEEFF9B61E5F91FFEB8846F" bold="true" box="[199,345,1789,1814]" pageId="19" pageNumber="20">Occurrence.</emphasis>
<collectingCountry id="41B27615FFEEFF9B6047F91CFE00846F" box="[357,481,1790,1814]" name="Canada" pageId="19" pageNumber="20">CANADA</collectingCountry>
:
<emphasis id="0BD1EA97FFEEFF9B60D3F91CFBB2846C" box="[497,1107,1789,1813]" italics="true" pageId="19" pageNumber="20">
Mount Robson area, British
<collectingCountry id="41B27615FFEEFF9B6267F91FFC52846C" box="[837,947,1789,1813]" name="United States of America" pageId="19" pageNumber="20">Columbia</collectingCountry>
and Alberta:
</emphasis>
<collectionCode id="5FB4AE40FFEEFF9B6542F91CFB77846F" box="[1120,1174,1790,1814]" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="19" pageNumber="20">GSC</collectionCode>
locality 90577, 138.1 metres above the base of the Mural Formation, Mumm Peak measured section, Alberta (
<bibRefCitation id="5D344B74FFEEFF9B6551F8C3FB158443" author="Fritz" box="[1139,1268,1825,1850]" pageId="19" pageNumber="20" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
). Locality 61k, talus from the middle member of the Mural Formation on the western margin of Mumm Peak, Alberta (see
<bibRefCitation id="5D344B74FFEEFF9B6442F8A7FF3284FB" author="Fritz" pageId="19" pageNumber="20" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
).
<collectionCode id="5FB4AE40FFEEFF9B61C9F888FEC084FB" box="[235,289,1898,1922]" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="19" pageNumber="20">GSC</collectionCode>
locality 87891, 169.2 metres above the base of the Mural Formation, Cinnamon Peak-Whitehorn Mountain section, Mount Robson Provincial Park, British
<collectingCountry id="41B27615FFEEFF9B6207F86FFC7284DF" box="[805,915,1933,1958]" name="United States of America" pageId="19" pageNumber="20">Columbia</collectingCountry>
(
<bibRefCitation id="5D344B74FFEEFF9B6280F86FFBC284DF" author="Fritz" box="[930,1059,1933,1958]" pageId="19" pageNumber="20" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="71BF650EFFEEFF9C61E5F853FF1E80F9" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="discussion">
<paragraph id="391A3685FFEEFF9C61E5F853FD398381" blockId="19.[151,1437,1645,2030]" lastBlockId="20.[151,1437,151,2013]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<emphasis id="0BD1EA97FFEEFF9B61E5F853FEAD84B3" bold="true" box="[199,332,1969,1994]" pageId="19" pageNumber="20">Discussion.</emphasis>
Fritz’s (1992) description of
<emphasis id="0BD1EA97FFEEFF9B63BFF850FC9884B3" box="[669,889,1970,1994]" italics="true" pageId="19" pageNumber="20">
Laudonia
<taxonomicName id="FEA54D06FFEEFF9B6233F850FC9884B3" box="[785,889,1970,1994]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
is generally adequate and, when complemented with the diagnosis provided previously, renders a full redescription of the species unneccesary. Fritz’s (1992) description did not mention the presence of the tropidium, however (Figs 3.2, 5.1, 5.2, 5.3, 5.4)—this omission is corrected herein by including the tropidium within the generic diagnosis. Differences between
<taxonomicName id="FEA54D06FFE9FF9C65F6FF5FFA8B83AD" box="[1236,1386,188,212]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFE9FF9C65F6FF5FFA8B83AD" box="[1236,1386,188,212]" italics="true" pageId="20" pageNumber="21">La. bispinata</emphasis>
</taxonomicName>
and
<taxonomicName id="FEA54D06FFE9FF9C61B5FF03FECD8381" box="[151,300,225,248]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="amputata">
<emphasis id="0BD1EA97FFE9FF9C61B5FF03FECD8381" box="[151,300,225,248]" italics="true" pageId="20" pageNumber="21">La. amputata</emphasis>
</taxonomicName>
are discussed under the latter species.
</paragraph>
<paragraph id="391A3685FFE9FF9C61E5FEE6FF1E80F9" blockId="20.[151,1437,151,2013]" pageId="20" pageNumber="21">
Specimens of
<taxonomicName id="FEA54D06FFE9FF9C6047FEE7FE198265" box="[357,504,260,284]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFE9FF9C6047FEE7FE198265" box="[357,504,260,284]" italics="true" pageId="20" pageNumber="21">La. bispinata</emphasis>
</taxonomicName>
examined herein range from approximately
<quantity id="FE5D9B60FFE9FF9C62C8FEE6FBA18265" box="[1002,1088,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="20" pageNumber="21" unit="mm" value="7.9">7.9 mm</quantity>
to more than
<quantity id="FE5D9B60FFE9FF9C65F8FEE6FAC98265" box="[1242,1320,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="mm" value="25.0">25 mm</quantity>
in sagittal cephalic length, and all represent morphologically mature individuals in phase 5 of cephalic development.
<bibRefCitation id="5D344B74FFE9FF9C6445FEC5FECB821C" author="Fritz" pageId="20" pageNumber="21" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992" yearSuffix="p">Fritz (1992, p. 26)</bibRefCitation>
recovered specimens ranging from
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to
<quantity id="FE5D9B60FFE9FF9C6263FEAEFC78821D" box="[833,921,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="20" pageNumber="21" unit="mm" value="9.5">9.5 mm</quantity>
in cephalic length, but these specimens were not examined herein. The trunk is known from two specimens (USNM 443764 and USNM 443768; Figs 4.5, 4.6, 5.5), both of which preserve the pygidium. (Several thoracic segments are also preserved scattered behind USNM 443766 [Fig. 5.4] and were probably associated with that cephalon in life, but the segments are disarticulated and their specific placement within the thorax cannot be determined.)
<bibRefCitation id="5D344B74FFE9FF9C6259FE3EFBE3828D" author="Fritz" box="[891,1026,476,501]" pageId="20" pageNumber="21" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz (1992)</bibRefCitation>
reported a total count of 22 thoracic segments for the species, although no single known specimen unambiguously preserves this full complement. The posterior-most region of the thorax is not well preserved on USNM 443768 (Fig. 5.5), and the exact number of segments on the specimen cannot be precisely determined. Nineteen thoracic segments are clearly preserved, and an impression of the left pleura and fragmentary axis of at least one more segment is evident behind that. However, the pygidium lies somewhat behind that, and there is sufficient separation between T19 and the pygidium for three segments to have been present. USNM 443764 (Fig. 4.5, 4.6) preserves nineteen segments anterior to the pygidium (with some telescoping) but there is a displacement between the anterior-most preserved segment and the posterior margin of the cephalon, and the anterior few thoracic segments (including T3) are missing. By comparison with USNM 443768, the form of the anterior-most preserved segment on USNM 443764 is consistent with it representing T4, in which case USNM 443764 would have borne 22 thoracic segments (with T1 to T3 now missing).
</paragraph>
</subSubSection>
<subSubSection id="71BF650EFFE9FF9D61E5FC6EFCE98041" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="391A3685FFE9FF9D61E5FC6EFCE98041" blockId="20.[151,1437,151,2013]" lastBlockId="21.[151,1437,151,824]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="0BD1EA97FFE9FF9C61E5FC6EFE4080DD" box="[199,417,908,932]" italics="true" pageId="20" pageNumber="21">
Laudonia
<taxonomicName id="FEA54D06FFE9FF9C601BFC6EFE4080DD" box="[313,417,908,932]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
is the
<typeStatus id="E61E8827FFE9FF9C60CEFC6FFDFD80DC" box="[492,540,909,933]" pageId="20" pageNumber="21">type</typeStatus>
species of the genus, and was included in cladistic analyses of the Olenelloidea by
<bibRefCitation id="5D344B74FFE9FF9C619DFC4DFE6F80B1" author="Lieberman" box="[191,398,943,968]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998)</bibRefCitation>
and of the
<taxonomicName id="FEA54D06FFE9FF9C6302FC4DFD4280B1" box="[544,675,943,968]" class="Trilobita" family="Biceratopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="subFamily" subFamily="Bristoliinae">Bristoliinae</taxonomicName>
by Lieberman (1999). Lieberman’s (1998, 1999) coding for
<taxonomicName id="FEA54D06FFE9FF9C6455FC53FF1E8095" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFE9FF9C6455FC53FF1E8095" italics="true" pageId="20" pageNumber="21">La. bispinata</emphasis>
</taxonomicName>
was apparently based on first-hand examination of three specimens (KUMIP 32400, MCZ
<date id="4D1B1045FFE9FF9C6433FC36FA868094" box="[1297,1383,980,1005]" pageId="20" pageNumber="21" value="79-11-06">110679</date>
, and MCZ
<date id="4D1B1045FFE9FF9C61FCFC15FED48769" box="[222,309,1015,1040]" pageId="20" pageNumber="21" value="80-11-06">110680</date>
), one of which (MCZ
<date id="4D1B1045FFE9FF9C6362FC15FD748769" box="[576,661,1015,1040]" pageId="20" pageNumber="21" value="79-11-06">110679</date>
) is here reassigned to a different genus. The present study reveals several uncertainties, unrecognized polymorphisms, and errors in the character state coding of
<taxonomicName id="FEA54D06FFE9FF9C65C6FBFFFA9D874D" box="[1252,1404,1052,1076]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFE9FF9C65C6FBFFFA9D874D" box="[1252,1404,1052,1076]" italics="true" pageId="20" pageNumber="21">La. bispinata</emphasis>
</taxonomicName>
in Lieberman’s (1998) analysis. LA was coded as expanding dorsally (
<bibRefCitation id="5D344B74FFE9FF9C62ADFBDDFADC8721" author="Lieberman" box="[911,1341,1087,1112]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998, character 10, state 1</bibRefCitation>
), but all available material has experienced compactional deformation and this character should be conservatively coded as state unknown. The preocular furrow (character 13) was coded by
<bibRefCitation id="5D344B74FFE9FF9C62A6FB65FBAE87D9" author="Lieberman" box="[900,1103,1159,1184]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998)</bibRefCitation>
as being oriented inward and backward from the glabellar margin (state 0). However, on many specimens the furrow runs inward and forward from the glabellar margin (state 2; e.g., Fig. 5.2, 5.3), and the character should therefore be coded as polymorphic (states 0, 2). The posterolateral margins of LA (character 17) were coded by
<bibRefCitation id="5D344B74FFE9FF9C62D9FB16FB278675" author="Lieberman" box="[1019,1222,1268,1293]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998)</bibRefCitation>
as being divergent anteriorly (state 1; e.g., KUMIP 32400). However, the posterolateral margins are sub-parallel on many specimens (state 2; e.g., Fig. 5.4), and the character should therefore be coded as polymorphic (states 1, 2). The ocular lobes were incorrectly coded by
<bibRefCitation id="5D344B74FFE9FF9C60E3FABDFCC58601" author="Lieberman" box="[449,804,1375,1400]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998, character 20)</bibRefCitation>
as forming a 10°–20° angle with a sagittal line (state 0), but were correctly coded by Lieberman (1999,
<taxonomicName id="FEA54D06FFE9FF9C6392FA66FCD286E4" box="[688,819,1412,1437]" class="Trilobita" family="Biceratopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="subFamily" subFamily="Bristoliinae">Bristoliinae</taxonomicName>
analysis, character 5) as forming approximately a 40° angle (equivalent to state
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the 1998 analysis): angles of divergence measured during the present study averaged 52° (range 29° to 76°; n = 12). The posterior tips of the ocular lobes were coded by
<bibRefCitation id="5D344B74FFE9FF9C6519FA2EFA7D869D" author="Lieberman" box="[1083,1436,1484,1509]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998, character 23)</bibRefCitation>
as being developed transversely opposite the lateral margins of L2 (state 3; e.g., KUMIP 32400). However, the tips are developed transversely opposite S2 on several specimens (state 4; e.g., Fig. 5.3, 5.4), and the character should therefore be coded as polymorphic (states 3, 4). All available material has experienced compactional deformation, and the original dorsal convexity of the extraocular area (
<bibRefCitation id="5D344B74FFE9FF9C6216F9BEFB76850D" author="Lieberman" box="[820,1175,1628,1653]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998, character 44</bibRefCitation>
) is unknown (this had originally been coded as flattened [state 0] by
<bibRefCitation id="5D344B74FFE9FF9C63BCF99DFC8085E1" author="Lieberman" box="[670,865,1663,1688]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
); similarly, the original ventral flexure of the distal tips of the thoracic pleurae (
<bibRefCitation id="5D344B74FFE9FF9C60F2F946FCC885C5" author="Lieberman" box="[464,809,1700,1725]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998, character 71</bibRefCitation>
) is unknown (this had originally been coded as being in roughly the same dorsoventral plane as the medial portion of the pleurae [state 0] by
<bibRefCitation id="5D344B74FFE9FF9C6565F925FAE88599" author="Lieberman" box="[1095,1289,1735,1760]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
). An anterior ocular line (
<bibRefCitation id="5D344B74FFE9FF9C6002F90EFD61847C" author="Lieberman" box="[288,640,1772,1797]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998, character 46</bibRefCitation>
) was not seen on any specimen examined herein (state 1; this feature had originally been coded as visible [state 0] by
<bibRefCitation id="5D344B74FFE9FF9C639BF8EDFC9C8451" author="Lieberman" box="[697,893,1807,1832]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
). A genal ridge (
<bibRefCitation id="5D344B74FFE9FF9C651BF8EDFA758451" author="Lieberman" box="[1081,1428,1807,1832]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998, character 47</bibRefCitation>
) was not seen on any specimen examined herein (state 1; this feature had originally been coded as prominently developed [state 0] by
<bibRefCitation id="5D344B74FFE9FF9C60BEF8B5FD818409" author="Lieberman" box="[412,608,1879,1904]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
). When traced abaxially, the posterior cephalic margin between the axial furrow and the adgenal angle (
<bibRefCitation id="5D344B74FFE9FF9C6323F89EFC8684EC" author="Lieberman" box="[513,871,1916,1941]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998, character 56</bibRefCitation>
) is oriented at an angle ranging from 3° to 15° (average of approximately 8°; n = 11) posteriorly relative to a transverse line. The posterior cephalic margin is therefore variably more-or-less transverse (state 0, as originally coded by
<bibRefCitation id="5D344B74FFE9FF9C6520F826FB2B84A5" author="Lieberman" box="[1026,1226,1988,2013]" pageId="20" pageNumber="21" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
; e.g., Fig. 5.1) to flexing posteriorly (state 1; e.g., Fig. 5.4). The thorax is not clearly differentiated into a prothorax and opisthothorax (
<bibRefCitation id="5D344B74FFE8FF9D6066FF5EFDED83AD" author="Lieberman" box="[324,524,188,213]" pageId="21" pageNumber="22" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
, character 57, state 1; this had originally been coded as being divided into a prothorax and opisthothorax [state 0] by
<bibRefCitation id="5D344B74FFE8FF9D6342FF3DFCFC8381" author="Lieberman" box="[608,797,223,248]" pageId="21" pageNumber="22" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
—see Terminology section [previous] for a discussion of this character). The posterior margin of every thoracic segment on both USNM 443764 and USNM 443768 is worn and/or not preserved (Figs 4.5, 4.6, 5.5), and the presence or absence of axial nodes on any thoracic segment (
<bibRefCitation id="5D344B74FFE8FF9D61BCFEAEFE1D821C" author="Lieberman" box="[158,508,332,357]" pageId="21" pageNumber="22" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998, character 66</bibRefCitation>
) cannot be determined with confidence (axial nodes had originally been coded as absent [state 1] by
<bibRefCitation id="5D344B74FFE8FF9D605DFE8DFDA682F1" author="Lieberman" box="[383,583,367,392]" pageId="21" pageNumber="22" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
; see also discussion under generic diagnosis). On at least T7 and more posterior segments, the pleural furrows (
<bibRefCitation id="5D344B74FFE8FF9D6344FE76FC2682D4" author="Lieberman" box="[614,967,404,429]" pageId="21" pageNumber="22" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998, character 67</bibRefCitation>
) extend a short distance onto the pleural spines (state 1; they had originally been coded as being confined to the inner pleural region [state 0] by
<bibRefCitation id="5D344B74FFE8FF9D643EFE55FF32828D" author="Lieberman" pageId="21" pageNumber="22" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
). However, state 0 applies to pleural furrows on more anterior segments (excluding T3), and this character as defined is accordingly polymorphic (states 0, 1). The width of the pleural spines on T5 to T8 at their midlength is not more than half of the length (exsag.) of the medial portion of the corresponding inner pleural region (
<bibRefCitation id="5D344B74FFE8FF9D61BCFDA5FE828119" author="Lieberman" box="[158,355,583,608]" pageId="21" pageNumber="22" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
, character 72, state 1; the spine widths had originally been coded as being more than two-thirds of the length of the corresponding inner pleural regions [state 0] by
<bibRefCitation id="5D344B74FFE8FF9D62ABFD8EFBAD81FD" author="Lieberman" box="[905,1100,620,645]" pageId="21" pageNumber="22" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
). Finally, the posterior margin of the pygidium is not well preserved on either USNM 443764 or USNM 443768 (Figs 4.6, 5.5), and whether it bore a median notch (
<bibRefCitation id="5D344B74FFE8FF9D60ABFD56FD0281B5" author="Lieberman" box="[393,739,692,717]" pageId="21" pageNumber="22" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998, character 79</bibRefCitation>
) cannot be determined with confidence. This character should accordingly be coded as uncertain (it had originally been coded as being transverse or weakly convex [state 0] by
<bibRefCitation id="5D344B74FFE8FF9D61B5FD1EFE80806D" author="Lieberman" box="[151,353,764,789]" pageId="21" pageNumber="22" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman, 1998</bibRefCitation>
). Given these coding issues, it remains to be seen whether the phylogenetic placement of
<emphasis id="0BD1EA97FFE8FF9D61B5FCC2FEE28041" box="[151,259,800,824]" italics="true" pageId="21" pageNumber="22">Laudonia</emphasis>
as determined by
<bibRefCitation id="5D344B74FFE8FF9D60EDFCFDFD7B8041" author="Lieberman" box="[463,666,799,824]" pageId="21" pageNumber="22" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998)</bibRefCitation>
is robust.
</paragraph>
</subSubSection>
</treatment>
</document>
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