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<mods:title id="9560A39FE8BA00210775F4C322EC1BE5">Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia</mods:title>
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<mods:namePart id="D374BC0D0265AA1932A0CFE085D42955">Bohach, Lisa L.</mods:namePart>
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<treatment id="B10C8793FFE8FF9061B5FC6EFD2D81B5" ID-DOI="http://doi.org/10.5281/zenodo.6129720" ID-GBIF-Taxon="127668553" ID-Zenodo-Dep="6129720" LSID="urn:lsid:plazi:treatment:B10C8793FFE8FF9061B5FC6EFD2D81B5" httpUri="http://treatment.plazi.org/id/B10C8793FFE8FF9061B5FC6EFD2D81B5" lastPageId="24" lastPageNumber="25" pageId="21" pageNumber="22">
<subSubSection id="71BF650EFFE8FF9D61B5FC6EFD6F87BD" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="391A3685FFE8FF9D61B5FC6EFDF280DF" blockId="21.[151,531,908,967]" box="[151,531,908,934]" pageId="21" pageNumber="22">
<heading id="625281E9FFE8FF9D61B5FC6EFDF280DF" bold="true" box="[151,531,908,934]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="0BD1EA97FFE8FF9D61B5FC6EFDF280DF" bold="true" box="[151,531,908,934]" pageId="21" pageNumber="22">
<emphasis id="0BD1EA97FFE8FF9D61B5FC6EFEED80DF" bold="true" box="[151,268,908,934]" italics="true" pageId="21" pageNumber="22">Laudonia</emphasis>
<taxonomicName id="FEA54D06FFE8FF9D6031FC6EFDF280DF" authority="Fritz, 1992" authorityName="Fritz" authorityYear="1992" box="[275,531,908,934]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="amputata">
<emphasis id="0BD1EA97FFE8FF9D6031FC6EFE6580DF" bold="true" box="[275,388,908,934]" italics="true" pageId="21" pageNumber="22">amputata</emphasis>
<bibRefCitation id="5D344B74FFE8FF9D60AEFC6EFDF280DF" author="Fritz" box="[396,531,908,934]" pageId="21" pageNumber="22" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="391A3685FFE8FF9D61B5FC4CFF3880BE" blockId="21.[151,531,908,967]" box="[151,217,942,967]" pageId="21" pageNumber="22">
<figureCitation id="A19E2A00FFE8FF9D61B5FC4CFF3880BE" box="[151,217,942,967]" captionStart="FIGURE 6" captionStartId="22.[151,250,1066,1088]" captionTargetBox="[482,1111,203,1031]" captionTargetId="figure@22.[462,1125,193,1045]" captionTargetPageId="22" captionText="FIGURE 6. Morphologically mature specimen of Laudonia amputata Fritz, 1992. Cephalon and incomplete thorax, latex peel of external mold; USNM 443752, x 4. From middle member of Mural Formation at Locality 61 k, Mumm Peak, Mount Robson area, Alberta." httpUri="https://zenodo.org/record/286410/files/figure.png" pageId="21" pageNumber="22">Fig. 6</figureCitation>
</paragraph>
<paragraph id="391A3685FFE8FF9D61B5FC14FD078752" blockId="21.[151,1055,1014,1221]" pageId="21" pageNumber="22">
1992
<emphasis id="0BD1EA97FFE8FF9D61F2FC14FE768774" box="[208,407,1014,1037]" italics="true" pageId="21" pageNumber="22">
Laudonia
<taxonomicName id="FEA54D06FFE8FF9D6015FC1AFE768774" authority="Fritz" authorityName="Fritz" box="[311,407,1016,1037]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="amputata">amputata</taxonomicName>
</emphasis>
Fritz, pp. 2627, text-fig. 6d, pl. 11, figs. 15, pl. 12, figs.
<date id="4D1B1045FFE8FF9D62CEFC14FF2B8752" pageId="21" pageNumber="22" value="1993-03-01">13. 1993</date>
<emphasis id="0BD1EA97FFE8FF9D61F2FBF7FE768752" box="[208,407,1045,1067]" italics="true" pageId="21" pageNumber="22">
Laudonia
<taxonomicName id="FEA54D06FFE8FF9D6015FBF4FE768752" authority="Fritz" authorityName="Fritz" box="[311,407,1046,1067]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="amputata">amputata</taxonomicName>
</emphasis>
Fritz; Palmer &amp; Repina, fig. 4.4.
</paragraph>
<paragraph id="391A3685FFE8FF9D61B5FBD6FCC58711" blockId="21.[151,1055,1014,1221]" pageId="21" pageNumber="22">
1997
<emphasis id="0BD1EA97FFE8FF9D61F2FBD6FE768733" box="[208,407,1076,1098]" italics="true" pageId="21" pageNumber="22">
Laudonia
<taxonomicName id="FEA54D06FFE8FF9D6015FBD7FE768733" authority="Fritz" authorityName="Fritz" box="[311,407,1077,1098]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="amputata">amputata</taxonomicName>
</emphasis>
Fritz; Palmer &amp; Repina in Whittington
<emphasis id="0BD1EA97FFE8FF9D6209FBD7FCB98733" box="[811,856,1076,1098]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
., p. 412, fig. 259.2. 1999
<emphasis id="0BD1EA97FFE8FF9D61F2FBB0FE768711" box="[208,407,1106,1128]" italics="true" pageId="21" pageNumber="22">
Laudonia
<taxonomicName id="FEA54D06FFE8FF9D6015FBB1FE768711" authority="Fritz" authorityName="Fritz" box="[311,407,1107,1128]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="amputata">amputata</taxonomicName>
</emphasis>
Fritz; Lieberman, pp. 3, 107, 114, 115.
</paragraph>
<paragraph id="391A3685FFE8FF9D61B5FB93FD1987FE" blockId="21.[151,1055,1014,1221]" box="[151,760,1137,1159]" pageId="21" pageNumber="22">
1999
<emphasis id="0BD1EA97FFE8FF9D61F2FB93FE7687FE" box="[208,407,1137,1159]" italics="true" pageId="21" pageNumber="22">
Laudonia
<taxonomicName id="FEA54D06FFE8FF9D6015FB90FE7687FE" authority="Fritz" authorityName="Fritz" box="[311,407,1138,1159]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="amputata">amputata</taxonomicName>
</emphasis>
Fritz; Smith &amp; Lieberman, p. 462.
</paragraph>
<paragraph id="391A3685FFE8FF9D61B5FB72FD7A87DF" blockId="21.[151,1055,1014,1221]" box="[151,667,1168,1190]" pageId="21" pageNumber="22">
2002
<emphasis id="0BD1EA97FFE8FF9D61F2FB72FE7687DF" box="[208,407,1168,1190]" italics="true" pageId="21" pageNumber="22">
Laudonia
<taxonomicName id="FEA54D06FFE8FF9D6015FB73FE7687DF" authority="Fritz" authorityName="Fritz" box="[311,407,1169,1190]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="amputata">amputata</taxonomicName>
</emphasis>
Fritz; Lieberman, p. 699.
</paragraph>
<paragraph id="391A3685FFE8FF9D61B5FB4CFD6F87BD" blockId="21.[151,1055,1014,1221]" box="[151,654,1198,1221]" pageId="21" pageNumber="22">
2003
<emphasis id="0BD1EA97FFE8FF9D61F2FB4CFE7687BC" box="[208,407,1198,1221]" italics="true" pageId="21" pageNumber="22">
Laudonia
<taxonomicName id="FEA54D06FFE8FF9D6015FB52FE7687BC" authority="Fritz" authorityName="Fritz" box="[311,407,1200,1221]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="amputata">amputata</taxonomicName>
</emphasis>
Fritz; Lieberman, p. 63.
</paragraph>
</subSubSection>
<subSubSection id="71BF650EFFE8FF9D61B5FB13FCCE850B" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="391A3685FFE8FF9D61B5FB13FCDD857F" blockId="21.[151,1436,1265,1938]" pageId="21" pageNumber="22">
<emphasis id="0BD1EA97FFE8FF9D61B5FB13FEF08673" bold="true" box="[151,273,1265,1290]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
Genal spine base transversely opposite lateral margin of L3 or S3; orientation of distal portion of posterior cephalic margin ranging from 4° inwards to 7° outwards relative to an exsagittal line when traced toward base of genal spine. Distance (tr.) between genal spine bases approximately 162% (range 140% to 174%) of glabellar length (sag.), increasing slightly through phase 5 of cephalic development. Angle between inner margin of proximal portion of genal spine and distal portion of posterior cephalic margin approximately 45° (range 42° to 49°). Width of anterior cephalic border (measured anterior to ocular lobes) 104% to 156% of length (exsag.) of LO. Ocular lobes divergent from exsagittal axis by approximately 15° (range 11° to 22°); posterior tip of ocular lobe transversely opposite lateral margin of L1 or distal tip of S1.
</paragraph>
<paragraph id="391A3685FFE8FF9D61E5F9F3FCCE850B" blockId="21.[151,1436,1265,1938]" pageId="21" pageNumber="22">
<emphasis id="0BD1EA97FFE8FF9D61E5F9F3FEA58553" bold="true" box="[199,324,1553,1578]" pageId="21" pageNumber="22">
<typeStatus id="E61E8827FFE8FF9D61E5F9F3FEDE8553" box="[199,319,1553,1578]" pageId="21" pageNumber="22" type="lectotype">Lectotype</typeStatus>
.
</emphasis>
USNM 443754 (figured by
<bibRefCitation id="5D344B74FFE8FF9D63ABF9F3FCE98553" author="Fritz" box="[649,776,1553,1578]" pageId="21" pageNumber="22" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 11, fig. 4;
<bibRefCitation id="5D344B74FFE8FF9D6292F9F3FB218553" author="Palmer" box="[944,1216,1553,1578]" pageId="21" pageNumber="22" refString="Palmer, A. R. &amp; Repina, L. N. (1993) Through a glass darkly: Taxonomy, phylogeny, and biostratigraphy of the Olenellina. University of Kansas Paleontological Contributions, New Series, 3, 1 - 35." type="journal article" year="1993">Palmer &amp; Repina, 1993</bibRefCitation>
, fig. 4.4; Palmer &amp; Repina in
<bibRefCitation id="5D344B74FFE8FF9D602FF9D7FDC18537" author="Whittington" box="[269,544,1589,1614]" pageId="21" pageNumber="22" refString="Whittington, H. B., Chatterton, B. D. E., Speyer, S. E., Fortey, R. A., Owens, R. M., Chang, W. T., Dean, W. T., Jell, P. A., Laurie, J. R., Palmer, A. R., Repina, L. N., Rushton, A. W. A., Shergold, J. H., Clarkson, E. N. K., Wilmot, N. V. &amp; Kelly, S. R. A. (1997) Treatise on Invertebrate Paleontology. Part O. Arthropoda 1. Trilobita, Revised. Volume 1: Introduction, Order Agnostida, Order Redlichiida. Geological Society of America and Univeristy of Kansas, Boulder, Colorado and Lawrence, Kansas, 530 pp." type="book" year="1997">
Whittington
<emphasis id="0BD1EA97FFE8FF9D60BEF9D4FE2E8534" box="[412,463,1589,1613]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
., 1997
</bibRefCitation>
, fig. 259.2), designated by Lieberman (1999, p. 115). This specimen had been labelled USNM 137533d prior to the study by
<bibRefCitation id="5D344B74FFE8FF9D6383F9BBFCCA850B" author="Fritz" box="[673,811,1625,1650]" pageId="21" pageNumber="22" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz (1992)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="71BF650EFFE8FF9E61E5F99FFBF9862A" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="391A3685FFE8FF9D61E5F99FFDC184EB" blockId="21.[151,1436,1265,1938]" pageId="21" pageNumber="22">
<emphasis id="0BD1EA97FFE8FF9D61E5F99FFE6385EF" bold="true" box="[199,386,1661,1686]" pageId="21" pageNumber="22">Other material.</emphasis>
Detailed qualitative and/or morphometric data were recorded from first-hand examination of the
<typeStatus id="E61E8827FFE8FF9D61E6F943FECA85C3" box="[196,299,1697,1722]" pageId="21" pageNumber="22" type="lectotype">lectotype</typeStatus>
(see previous) plus the following nine specimens:
<collectionCode id="5FB4AE40FFE8FF9D62A4F940FC3685C3" LSID="urn:lsid:biocol.org:col:34871" box="[902,983,1698,1722]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22">USNM</collectionCode>
137533,
<collectionCode id="5FB4AE40FFE8FF9D656BF940FB7B85C3" LSID="urn:lsid:biocol.org:col:34871" box="[1097,1178,1698,1722]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22">USNM</collectionCode>
137533c, and
<collectionCode id="5FB4AE40FFE8FF9D6469F940FA7D85C3" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,1698,1722]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22">USNM</collectionCode>
137533f (all stored in the biological collections);
<collectionCode id="5FB4AE40FFE8FF9D639FF924FCEF85A7" LSID="urn:lsid:biocol.org:col:34871" box="[701,782,1734,1758]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22">USNM</collectionCode>
443752 (also figured by
<bibRefCitation id="5D344B74FFE8FF9D6504F927FB4285A7" author="Fritz" box="[1062,1187,1733,1758]" pageId="21" pageNumber="22" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 11, fig. 1);
<collectionCode id="5FB4AE40FFE8FF9D6469F924FA7D85A7" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,1734,1758]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22">USNM</collectionCode>
443753 (also figured by
<bibRefCitation id="5D344B74FFE8FF9D608CF90BFDCC847B" author="Fritz" box="[430,557,1769,1794]" pageId="21" pageNumber="22" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 11, figs. 2, 3);
<collectionCode id="5FB4AE40FFE8FF9D6227F908FCB7847B" LSID="urn:lsid:biocol.org:col:34871" box="[773,854,1770,1794]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22">USNM</collectionCode>
443755 (also figured by
<bibRefCitation id="5D344B74FFE8FF9D6557F90BFB15847B" author="Fritz" box="[1141,1268,1769,1794]" pageId="21" pageNumber="22" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 11, fig. 5);
<collectionCode id="5FB4AE40FFE8FF9D61B5F8ECFF09845F" LSID="urn:lsid:biocol.org:col:34871" box="[151,232,1806,1830]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22">USNM</collectionCode>
443756 (also figured by
<bibRefCitation id="5D344B74FFE8FF9D6307F8EFFD48845F" author="Fritz" box="[549,681,1805,1830]" pageId="21" pageNumber="22" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 12, figs. 13); an unnumbered specimen in the biological collections of the
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; and
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113773.
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113773 is an incomplete cephalon (part and counterpart) partially obscured by matrix, but shows a morphology consistent with
<emphasis id="0BD1EA97FFE8FF9D62C5F8B7FB288414" box="[999,1225,1877,1901]" italics="true" pageId="21" pageNumber="22">
Laudonia
<taxonomicName id="FEA54D06FFE8FF9D657DF8B4FB288414" box="[1119,1225,1878,1901]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="amputata">amputata</taxonomicName>
</emphasis>
; this specimen is tentatively assigned to this species.
</paragraph>
<caption id="6DDA660DFFEBFF9E61B5FBC8FEC08704" httpUri="https://zenodo.org/record/286410/files/figure.png" pageId="22" pageNumber="23" targetBox="[482,1111,203,1031]" targetPageId="22">
<paragraph id="391A3685FFEBFF9E61B5FBC8FEC08704" blockId="22.[151,1436,1066,1149]" pageId="22" pageNumber="23">
<emphasis id="0BD1EA97FFEBFF9E61B5FBC8FEF28746" bold="true" box="[151,275,1066,1088]" pageId="22" pageNumber="23">FIGURE 6.</emphasis>
Morphologically mature specimen of
<emphasis id="0BD1EA97FFEBFF9E63BBFBC8FD1B8739" box="[665,762,1066,1088]" italics="true" pageId="22" pageNumber="23">Laudonia</emphasis>
<taxonomicName id="FEA54D06FFEBFF9E6222FBC9FC388739" authority="Fritz, 1992" authorityName="Fritz" authorityYear="1992" box="[768,985,1066,1088]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="amputata">
<emphasis id="0BD1EA97FFEBFF9E6222FBC9FC818739" box="[768,864,1067,1088]" italics="true" pageId="22" pageNumber="23">amputata</emphasis>
Fritz, 1992
</taxonomicName>
. Cephalon and incomplete thorax, latex peel of external mold; USNM 443752, x4. From middle member of Mural Formation at Locality 61k, Mumm Peak, Mount Robson area, Alberta.
</paragraph>
</caption>
<paragraph id="391A3685FFEBFF9E61E5FB48FBF9862A" blockId="22.[151,1436,1194,2011]" pageId="22" pageNumber="23">
<emphasis id="0BD1EA97FFEBFF9E61E5FB48FEB887BA" bold="true" box="[199,345,1194,1219]" pageId="22" pageNumber="23">Occurrence.</emphasis>
<collectingCountry id="41B27615FFEBFF9E6046FB49FE0187BA" box="[356,480,1195,1219]" name="Canada" pageId="22" pageNumber="23">CANADA</collectingCountry>
:
<emphasis id="0BD1EA97FFEBFF9E60CDFB49FBA887BB" box="[495,1097,1194,1218]" italics="true" pageId="22" pageNumber="23">
Mount Robson area, British
<collectingCountry id="41B27615FFEBFF9E621FFB48FC4A87BB" box="[829,939,1194,1218]" name="United States of America" pageId="22" pageNumber="23">Columbia</collectingCountry>
and Alberta:
</emphasis>
Locality 61k, talus from the middle member of the Mural Formation on the western margin of Mumm Peak, Alberta (see
<bibRefCitation id="5D344B74FFEBFF9E6582FB2CFAFF879E" author="Fritz" box="[1184,1310,1230,1255]" pageId="22" pageNumber="23" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
).
<emphasis id="0BD1EA97FFEBFF9E6413FB2CFEE08673" italics="true" pageId="22" pageNumber="23">
Laudonia
<taxonomicName id="FEA54D06FFEBFF9E61B5FB11FEE08673" box="[151,257,1267,1290]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="amputata">amputata</taxonomicName>
</emphasis>
was not found in situ during Fritzs (1992) restudy of Walcotts (1913) “61k fauna”, and the precise stratigraphic provenance of the species remains uncertain. Nevertheless, it is assumed to occur within the lower portion of the Dyeran Stage, Waucoban Series (
<bibRefCitation id="5D344B74FFEBFF9E6389FAD8FBED862A" author="Fritz" box="[683,1036,1338,1363]" pageId="22" pageNumber="23" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992" yearSuffix="p">Fritz, 1992, p. 27; see previous</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="71BF650EFFEBFF9E61E5FABCFD498536" pageId="22" pageNumber="23" type="discussion">
<paragraph id="391A3685FFEBFF9E61E5FABCFD498536" blockId="22.[151,1436,1194,2011]" pageId="22" pageNumber="23">
<emphasis id="0BD1EA97FFEBFF9E61E5FABCFEAD860E" bold="true" box="[199,332,1374,1399]" pageId="22" pageNumber="23">Discussion.</emphasis>
The original description of
<emphasis id="0BD1EA97FFEBFF9E63A4FABCFC82860F" box="[646,867,1374,1398]" italics="true" pageId="22" pageNumber="23">
Laudonia
<taxonomicName id="FEA54D06FFEBFF9E63DBFABDFC82860F" box="[761,867,1375,1398]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="amputata">amputata</taxonomicName>
</emphasis>
was based on examination of five specimens from Walcotts locality 61k (
<bibRefCitation id="5D344B74FFEBFF9E6087FA60FDC886E2" author="Fritz" box="[421,553,1410,1435]" pageId="22" pageNumber="23" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
). All of those specimens, plus five additional specimens (listed previously), were studied first-hand during the present work. Cephalic lengths of the suite of specimens examined herein range from
<quantity id="FE5D9B60FFEBFF9E61FFFA28FEDD869B" box="[221,316,1482,1507]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="22" pageNumber="23" unit="mm" value="6.4">6.4 mm</quantity>
to
<quantity id="FE5D9B60FFEBFF9E6050FA29FE00869B" box="[370,481,1483,1507]" metricMagnitude="-2" metricUnit="m" metricValue="3.82" pageId="22" pageNumber="23" unit="mm" value="38.2">38.2 mm</quantity>
, and all represent morphologically mature individuals in phase 5 of cephalic development. Fritzs (1992) description of
<taxonomicName id="FEA54D06FFEBFF9E63AEFA0DFCC7857F" box="[652,806,1519,1542]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="amputata">
<emphasis id="0BD1EA97FFEBFF9E63AEFA0DFCC7857F" box="[652,806,1519,1542]" italics="true" pageId="22" pageNumber="23">La. amputata</emphasis>
</taxonomicName>
is generally adequate and only a species diagnosis is provided herein. However, the presence of the tropidium (
<figureCitation id="A19E2A00FFEBFF9E62A8F9F0FC398552" box="[906,984,1554,1579]" captionStart="FIGURE 6" captionStartId="22.[151,250,1066,1088]" captionTargetBox="[482,1111,203,1031]" captionTargetId="figure@22.[462,1125,193,1045]" captionTargetPageId="22" captionText="FIGURE 6. Morphologically mature specimen of Laudonia amputata Fritz, 1992. Cephalon and incomplete thorax, latex peel of external mold; USNM 443752, x 4. From middle member of Mural Formation at Locality 61 k, Mumm Peak, Mount Robson area, Alberta." httpUri="https://zenodo.org/record/286410/files/figure.png" pageId="22" pageNumber="23">Fig. 6</figureCitation>
)—here included within the generic diagnosis—was not mentioned by
<bibRefCitation id="5D344B74FFEBFF9E6338F9D4FD458536" author="Fritz" box="[538,676,1590,1615]" pageId="22" pageNumber="23" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz (1992)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="71BF650EFFEBFF9061E5F9B8FD2D81B5" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="reference_group">
<paragraph id="391A3685FFEBFF9061E5F9B8FC0E82A9" blockId="22.[151,1436,1194,2011]" lastBlockId="24.[151,1437,151,717]" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
<emphasis id="0BD1EA97FFEBFF9E61E5F9B8FE44850B" box="[199,421,1626,1650]" italics="true" pageId="22" pageNumber="23">
Laudonia
<taxonomicName id="FEA54D06FFEBFF9E6019F9B9FE44850B" box="[315,421,1627,1650]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="amputata">amputata</taxonomicName>
</emphasis>
differs most obviously from
<taxonomicName id="FEA54D06FFEBFF9E63D6F9B9FC68850B" box="[756,905,1626,1650]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFEBFF9E63D6F9B9FC68850B" box="[756,905,1626,1650]" italics="true" pageId="22" pageNumber="23">La. bispinata</emphasis>
</taxonomicName>
by having ocular lobes that are proportionally longer (posterior tips developed transversely opposite the lateral margin of L1 or rarely S
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<taxonomicName id="FEA54D06FFEBFF9E65D9F99DFA7485EF" box="[1275,1429,1663,1686]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="amputata">
<emphasis id="0BD1EA97FFEBFF9E65D9F99DFA7485EF" box="[1275,1429,1663,1686]" italics="true" pageId="22" pageNumber="23">La. amputata</emphasis>
</taxonomicName>
, opposite the lateral margins of L2, S2, or L
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<taxonomicName id="FEA54D06FFEBFF9E638AF941FCDA85C3" box="[680,827,1698,1722]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFEBFF9E638AF941FCDA85C3" box="[680,827,1698,1722]" italics="true" pageId="22" pageNumber="23">La. bispinata</emphasis>
</taxonomicName>
; Fig. 7.1) and less strongly divergent (Fig. 7.2), and in having genal spines that are less anteriorly advanced (bases developed transversely opposite the lateral margin of L3 or S
<quantity id="FE5D9B60FFEBFF9E61CFF909FEF8847A" box="[237,281,1770,1795]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="22" pageNumber="23" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="FEA54D06FFEBFF9E6003F909FE59847B" box="[289,440,1771,1794]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="amputata">
<emphasis id="0BD1EA97FFEBFF9E6003F909FE59847B" box="[289,440,1771,1794]" italics="true" pageId="22" pageNumber="23">La. amputata</emphasis>
</taxonomicName>
, transversely opposite the lateral margin of LA in
<taxonomicName id="FEA54D06FFEBFF9E62DAF909FB6D847B" box="[1016,1164,1770,1794]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFEBFF9E62DAF909FB6D847B" box="[1016,1164,1770,1794]" italics="true" pageId="22" pageNumber="23">La. bispinata</emphasis>
</taxonomicName>
; Fig. 7.3) and that flare less strongly laterally from the distal portion of the posterior cephalic margin (Fig. 7.4). The species also differ in orientation of the distal portion of the posterior cephalic margin (Fig. 7.5): on all specimens of
<taxonomicName id="FEA54D06FFEBFF9E6426F8D1FA7A8433" box="[1284,1435,1842,1866]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFEBFF9E6426F8D1FA7A8433" box="[1284,1435,1842,1866]" italics="true" pageId="22" pageNumber="23">La. bispinata</emphasis>
</taxonomicName>
examined herein, the margin is consistently oriented slightly inward when traced towards the base of the genal spine; while the margin of
<taxonomicName id="FEA54D06FFEBFF9E60E0F899FDB684EB" box="[450,599,1915,1938]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="amputata">
<emphasis id="0BD1EA97FFEBFF9E60E0F899FDB684EB" box="[450,599,1915,1938]" italics="true" pageId="22" pageNumber="23">La. amputata</emphasis>
</taxonomicName>
is typically directed very slightly outward when traced towards the base of the genal spine—on only the largest known specimen of
<taxonomicName id="FEA54D06FFEBFF9E623DF87DFC5784CF" box="[799,950,1951,1974]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="amputata">
<emphasis id="0BD1EA97FFEBFF9E623DF87DFC5784CF" box="[799,950,1951,1974]" italics="true" pageId="22" pageNumber="23">La. amputata</emphasis>
</taxonomicName>
is the margin directed inward and forward (USNM 443753; see
<bibRefCitation id="5D344B74FFEBFF9E60AAF820FDE884A2" author="Fritz" box="[392,521,1986,2011]" pageId="22" pageNumber="23" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz, 1992</bibRefCitation>
, pl. 11, fig. 2), and on that specimen the inward orientation is not as strong as in large specimens of
<taxonomicName id="FEA54D06FFE5FF90605DFF7BFDF683C9" box="[383,535,152,176]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFE5FF90605DFF7BFDF683C9" box="[383,535,152,176]" italics="true" pageId="24" pageNumber="25">La. bispinata</emphasis>
</taxonomicName>
. Relative to
<taxonomicName id="FEA54D06FFE5FF906395FF7BFCB183C9" box="[695,848,152,176]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFE5FF906395FF7BFCB183C9" box="[695,848,152,176]" italics="true" pageId="24" pageNumber="25">La. bispinata</emphasis>
</taxonomicName>
,
<taxonomicName id="FEA54D06FFE5FF906241FF7BFC1F83C9" box="[867,1022,153,176]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="amputata">
<emphasis id="0BD1EA97FFE5FF906241FF7BFC1F83C9" box="[867,1022,153,176]" italics="true" pageId="24" pageNumber="25">La. amputata</emphasis>
</taxonomicName>
also typically has a proportionally slightly narrower anterior cephalic border (Fig. 7.6) and a proportionally narrower (tr.) extraocular area (Fig. 7.7, 7.8), although the two species show some overlap in values for those traits. The intergenal ridge of
<taxonomicName id="FEA54D06FFE5FF906427FF03FA7D8381" box="[1285,1436,225,248]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="amputata">
<emphasis id="0BD1EA97FFE5FF906427FF03FA7D8381" box="[1285,1436,225,248]" italics="true" pageId="24" pageNumber="25">La. amputata</emphasis>
</taxonomicName>
lacks the ridge-on-ridge structure that is developed in
<taxonomicName id="FEA54D06FFE5FF906226FEE7FC798265" box="[772,920,260,284]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFE5FF906226FEE7FC798265" box="[772,920,260,284]" italics="true" pageId="24" pageNumber="25">La. bispinata</emphasis>
</taxonomicName>
. In the thorax, the inner pleural regions of
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<emphasis id="0BD1EA97FFE5FF906455FEE7FEE08239" italics="true" pageId="24" pageNumber="25">La. amputata</emphasis>
</taxonomicName>
are slightly proportionally narrower (tr.) relative to width (tr.) of the axis than are those of
<taxonomicName id="FEA54D06FFE5FF906425FECBFA7D8239" box="[1287,1436,296,320]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFE5FF906425FECBFA7D8239" box="[1287,1436,296,320]" italics="true" pageId="24" pageNumber="25">La. bispinata</emphasis>
</taxonomicName>
(presumably related to the interspecific difference in width of the extraocular area), and the pleural spines of
<taxonomicName id="FEA54D06FFE5FF906455FEAFFEE082F1" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="amputata">
<emphasis id="0BD1EA97FFE5FF906455FEAFFEE082F1" italics="true" pageId="24" pageNumber="25">La. amputata</emphasis>
</taxonomicName>
are slightly shorter than the spines on equivalent segments of
<taxonomicName id="FEA54D06FFE5FF906298FE93FBAC82F1" box="[954,1101,368,392]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFE5FF906298FE93FBAC82F1" box="[954,1101,368,392]" italics="true" pageId="24" pageNumber="25">La. bispinata</emphasis>
</taxonomicName>
.
<bibRefCitation id="5D344B74FFE5FF906579FE8DFB0382F1" author="Fritz" box="[1115,1250,367,392]" pageId="24" pageNumber="25" refString="Fritz, W. H. (1992) Walcott's Lower Cambrian olenellid trilobites collection 61 k, Mount Robson area, Canadian Rocky Mountains. Geological Survey of Canada Bulletin, 432, 1 - 65." type="journal article" year="1992">Fritz (1992)</bibRefCitation>
claimed that the species also differed in the degree of constriction of the glabella at S1, although such a supposed difference was not supported by the morphometric analyses conducted herein (data not shown).
</paragraph>
<caption id="6DDA660DFFEAFF9F61B5F92BFB74848B" httpUri="https://zenodo.org/record/286411/files/figure.png" pageId="23" pageNumber="24" targetBox="[368,1217,215,1699]" targetPageId="23">
<paragraph id="391A3685FFEAFF9F61B5F92BFB74848B" blockId="23.[151,1437,1737,2034]" pageId="23" pageNumber="24">
<emphasis id="0BD1EA97FFEAFF9F61B5F92BFEF485A7" bold="true" box="[151,277,1737,1759]" pageId="23" pageNumber="24">FIGURE 7.</emphasis>
Bivariate scatterplots depicting intraspecific variation and interspecific disparity in various aspects of cephalic morphology among morphologically mature
<emphasis id="0BD1EA97FFEAFF9F6340F905FCC98584" box="[610,808,1767,1789]" italics="true" pageId="23" pageNumber="24">
Laudonia
<taxonomicName id="FEA54D06FFEAFF9F63E8F905FCC98584" box="[714,808,1767,1789]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="bispinata">bispinata</taxonomicName>
</emphasis>
and
<taxonomicName id="FEA54D06FFEAFF9F6279F90BFC028587" box="[859,995,1769,1790]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="amputata">
<emphasis id="0BD1EA97FFEAFF9F6279F90BFC028587" box="[859,995,1769,1790]" italics="true" pageId="23" pageNumber="24">La. amputata</emphasis>
</taxonomicName>
. 1, Proportional length of the ocular lobes. 2, Divergence of the ocular lobe from the exsagittal axis. 3, Proportional anterior advancement of the genal spine bases, measured as the ratio of the exsagittal distance between the posterior limit of the sagittal axis and the base of the genal spine (genal spine advancement) to sagittal glabella length. 4, Degree of lateral flaring of the genal spine, quantified as the angle between the inner margin of the proximal portion of the genal spine and the distal portion of the posterior cephalic margin (distal PCM). 5, Orientation of the distal portion of the posterior cephalic margin (distal PCM). 6, Proportional width of the anterior cephalic border (measured anterior to the contact of the ocular lobes with LA). 7, Proportional cephalic width, measured as the ratio of the transverse distance between the genal spine bases to glabella length. 8, Proportional cephalic width, measured as the ratio of the transverse distance between the intergenal spine bases to glabella length.
</paragraph>
</caption>
<paragraph id="391A3685FFE5FF9061E5FE3EFD2D81B5" blockId="24.[151,1437,151,717]" pageId="24" pageNumber="25">
<emphasis id="0BD1EA97FFE5FF9061E5FE3EFE46828D" box="[199,423,476,500]" italics="true" pageId="24" pageNumber="25">
Laudonia
<taxonomicName id="FEA54D06FFE5FF90601FFE3FFE46828D" box="[317,423,477,500]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="amputata">amputata</taxonomicName>
</emphasis>
was not included in the cladistic analysis of the Olenelloidea presented by
<bibRefCitation id="5D344B74FFE5FF906401FE3EFF048161" author="Lieberman" pageId="24" pageNumber="25" refString="Lieberman, B. S. (1998) Cladistic analysis of the Early Cambrian olenelloid trilobites. Journal of Paleontology, 72, 59 - 78." type="journal article" year="1998">Lieberman (1998)</bibRefCitation>
. However, the species was subsequently used as an outgroup taxon in the cladistic analysis of
<taxonomicName id="FEA54D06FFE5FF906438FE1DFA7C8161" box="[1306,1437,511,536]" class="Trilobita" family="Biceratopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subFamily" subFamily="Bristoliinae">Bristoliinae</taxonomicName>
by Lieberman (1999), where it was assumed to form a clade with
<taxonomicName id="FEA54D06FFE5FF9062A2FDC7FBF58145" box="[896,1044,548,572]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFE5FF9062A2FDC7FBF58145" box="[896,1044,548,572]" italics="true" pageId="24" pageNumber="25">La. bispinata</emphasis>
</taxonomicName>
. Character states for
<taxonomicName id="FEA54D06FFE5FF906424FDC7FA7D8145" box="[1286,1436,549,572]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="amputata">
<emphasis id="0BD1EA97FFE5FF906424FDC7FA7D8145" box="[1286,1436,549,572]" italics="true" pageId="24" pageNumber="25">La. amputata</emphasis>
</taxonomicName>
were coded correctly, with the possible exception of character 7. Lieberman (1999,
<taxonomicName id="FEA54D06FFE5FF906561FDA5FB278119" box="[1091,1222,583,608]" class="Trilobita" family="Biceratopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subFamily" subFamily="Bristoliinae">Bristoliinae</taxonomicName>
analysis, character 7) coded
<taxonomicName id="FEA54D06FFE5FF906026FD8FFE7C81FD" box="[260,413,621,644]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="amputata">
<emphasis id="0BD1EA97FFE5FF906026FD8FFE7C81FD" box="[260,413,621,644]" italics="true" pageId="24" pageNumber="25">La. amputata</emphasis>
</taxonomicName>
as lacking a “faint ventral depression across entire region where ocular lobe hits frontal lobe”. However, if this faint ventral depression refers to a transocular furrow, then it is mis-coded. (The same holds true for coding of that character for
<taxonomicName id="FEA54D06FFE5FF90630BFD57FD5D81B5" box="[553,700,692,716]" class="Trilobita" family="Biceratopsidae" genus="Lochmanolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bispinata">
<emphasis id="0BD1EA97FFE5FF90630BFD57FD5D81B5" box="[553,700,692,716]" italics="true" pageId="24" pageNumber="25">La. bispinata</emphasis>
</taxonomicName>
.)
</paragraph>
</subSubSection>
</treatment>
</document>