treatments-xml/data/7C/6D/BE/7C6DBE31B35BFFB6DA8FE18AFB1725C7.xml
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<document id="4E93AAE538BD638B40E85EF6C2109E98" ID-DOI="10.5281/zenodo.211147" ID-GBIF-Dataset="c3e315dd-5edd-4c8c-be8e-1d4975fe57ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="211147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460512977999" checkinUser="plazi" docAuthor="Greenfield, David W." docDate="2012" docId="7C6DBE31B35BFFB6DA8FE18AFB1725C7" docLanguage="en" docName="zt03437p031.pdf" docOrigin="Zootaxa 3437" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Colletteichthys flavipinnis Greenfield, 2012, n. sp." docType="treatment" docVersion="8" lastPageNumber="27" masterDocId="8054C649B35AFFB5DA18E11DFF99222B" masterDocTitle="Colletteichthys flavipinnis, a new toadfish species from Sri Lanka and India (Teleostei: Batrachoididae)" masterLastPageNumber="31" masterPageNumber="24" pageNumber="25" updateTime="1698313587033" updateUser="plazi">
<mods:mods id="5FBF988108C65C883414BFDCF5B73D86" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F60252098640D32A17FB1B98E7ACF0B6">
<mods:title id="8D86413E12AA8100A603A6D98F470517">Colletteichthys flavipinnis, a new toadfish species from Sri Lanka and India (Teleostei: Batrachoididae)</mods:title>
</mods:titleInfo>
<mods:name id="13412080B4BF6722A53599DA50013AEF" type="personal">
<mods:role id="B9311EA3F2D94B127925C8350BCAC4DE">
<mods:roleTerm id="072AE75546EBF1192151F3C309E60936">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C953843143A2E35C01B3BDD75A7C4925">Greenfield, David W.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C6AEBAD10DB25F646A7D8E2E5E827F71">text</mods:typeOfResource>
<mods:relatedItem id="5B3A04717F6DD5F5DA40834EC2B27ED1" type="host">
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<mods:title id="213B27255B1B88566F7CD830C87BC6AB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E397D163B0E6D6F565103B38200DE853">
<mods:date id="9BC743B9A95237234008470BB571EACE">2012</mods:date>
<mods:detail id="CFA409D6A158232A5A40266ACDDDC50D" type="volume">
<mods:number id="8A5518F3EBBA3CD8FD5FF9D03ABF8732">3437</mods:number>
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<mods:classification id="D0EAB05F50103A9197B410F13D79135D">journal article</mods:classification>
<mods:identifier id="93532DF6B01B61FE24097CD1F6A70555" type="DOI">10.5281/zenodo.211147</mods:identifier>
<mods:identifier id="ECFC5802230DC97591458F2D1A23617E" type="GBIF-Dataset">c3e315dd-5edd-4c8c-be8e-1d4975fe57ec</mods:identifier>
<mods:identifier id="7BCEA2E60BED31F81EC9687075C21F7F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3DCFEF700089F89C6A27DC88121B6BB0" type="Zenodo-Dep">211147</mods:identifier>
</mods:mods>
<treatment id="7C6DBE31B35BFFB6DA8FE18AFB1725C7" ID-DOI="http://doi.org/10.5281/zenodo.6170713" ID-GBIF-Taxon="119523542" ID-Zenodo-Dep="6170713" LSID="urn:lsid:zoobank.org:act:A1E01D73-2217-47BF-A9C1-60DA3081CA3C" httpUri="http://treatment.plazi.org/id/7C6DBE31B35BFFB6DA8FE18AFB1725C7" lastPageId="3" lastPageNumber="27" pageId="1" pageNumber="25">
<subSubSection id="BCDE5CACB35BFFB4DA8FE18AFED322DD" pageId="1" pageNumber="25" type="nomenclature">
<paragraph id="F47B0F27B35BFFB4DA8FE18AFD82229A" blockId="1.[151,539,151,246]" box="[151,539,151,177]" pageId="1" pageNumber="25">
<heading id="AF33B84BB35BFFB4DA8FE18AFD82229A" bold="true" box="[151,539,151,177]" fontSize="11" level="1" pageId="1" pageNumber="25" reason="1">
<emphasis id="C6B0D335B35BFFB4DA8FE18AFD82229A" bold="true" box="[151,539,151,177]" pageId="1" pageNumber="25">
<taxonomicName id="33C474A4B35BFFB4DA8FE18AFE4B229A" ID-CoL="X48B" box="[151,466,151,177]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="1" pageNumber="25" phylum="Chordata" rank="species" species="flavipinnis" status="sp. nov.">
<emphasis id="C6B0D335B35BFFB4DA8FE18AFE4B229A" bold="true" box="[151,466,151,177]" italics="true" pageId="1" pageNumber="25">Colletteichthys flavipinnis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD836E4EB35BFFB4DBC1E18AFD82229A" box="[473,539,151,177]" pageId="1" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F47B0F27B35BFFB4DA8FE1A7FEF722F9" blockId="1.[151,539,151,246]" box="[151,366,186,210]" pageId="1" pageNumber="25">Yellowfin Toadfish</paragraph>
<paragraph id="F47B0F27B35BFFB4DA8FE1C3FED322DD" blockId="1.[151,539,151,246]" box="[151,330,222,246]" pageId="1" pageNumber="25">
(
<figureCitation id="6CFF13A2B35BFFB4DA87E1C3FE9322DD" box="[159,266,222,246]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1671,1694]" captionStartId-1="2.[151,250,758,781]" captionStartId-2="2.[151,250,1344,1367]" captionTargetBox-0="[151,1436,1022,1647]" captionTargetBox-1="[172,1434,301,735]" captionTargetBox-2="[403,1183,814,1322]" captionTargetId-0="figure@1.[151,1436,1022,1647]" captionTargetId-1="figure@2.[151,1436,301,737]" captionTargetId-2="figure@2.[403,1184,788,1323]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionText-0="FIGURE 1. Freshly collected specimen of Colletteichthys flavipinnis, Tuticorin, India. Photograph by Arumugam. Murugan." captionText-1="FIGURE 2. Freshly collected holotype of Colletteichthys flavipinnis. Photograph by K. K. Bineesh." captionText-2="FIGURE 3. Eye of holotype of Colletteichthys flavipinnis showing single tentacle above it. Photograph by D. W. Greenfield." httpUri-0="https://zenodo.org/record/211148/files/figure.png" httpUri-1="https://zenodo.org/record/211149/files/figure.png" httpUri-2="https://zenodo.org/record/211150/files/figure.png" pageId="1" pageNumber="25">Figs. 13</figureCitation>
,
<figureCitation id="6CFF13A2B35BFFB4DB0EE1C3FEDB22DD" box="[278,322,222,246]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="4.[151,250,1989,2012]" captionStartId-1="5.[151,250,550,573]" captionTargetBox-0="[151,1435,1182,1966]" captionTargetBox-1="[159,1424,203,518]" captionTargetId-0="figure@4.[151,1436,1182,1967]" captionTargetId-1="figure@5.[151,1436,193,529]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 6. Head coloration of freshly collected paratype of Colletteichthys flavipinnis. Photograph by K. K. Bineesh." captionText-1="FIGURE 7. Preserved holotype of Colletteichthys flavipinnis. Photograph by D. W. Greenfield." httpUri-0="https://zenodo.org/record/211153/files/figure.png" httpUri-1="https://zenodo.org/record/211154/files/figure.png" pageId="1" pageNumber="25">67</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BCDE5CACB35BFFB4DA8FE03AFCB5208D" pageId="1" pageNumber="25" type="materials_examined">
<paragraph id="F47B0F27B35BFFB4DA8FE03AFF4F2349" blockId="1.[151,1437,294,967]" pageId="1" pageNumber="25">
<emphasis id="C6B0D335B35BFFB4DA8FE03AFE922314" bold="true" box="[151,267,295,319]" pageId="1" pageNumber="25">
<typeStatus id="2B7FB185B35BFFB4DA8FE03AFE9F2314" box="[151,262,295,319]" pageId="1" pageNumber="25" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="92D597E2B35BFFB4DB0BE03BFEF32315" box="[275,362,294,318]" httpUri="http://grbio.org/cool/mve2-984h" name="See FMRI" pageId="1" pageNumber="25">CMFRI</collectionCode>
GB.9.1.2.1,111.2 mm, off Tuticorin,
<collectingCountry id="8CD34FB7B35BFFB4D90CE03BFCC92315" box="[788,848,294,318]" name="India" pageId="1" pageNumber="25">India</collectingCountry>
, mini trawl, K.K. Bineesh &amp; R. Kumar,
<date id="807A29E7B35BFFB4DF39E03BFF4A2349" pageId="1" pageNumber="25" value="2010-01-22">22 January 2010</date>
.
</paragraph>
<paragraph id="F47B0F27B35BFFB4DADFE072FA0423D9" blockId="1.[151,1437,294,967]" pageId="1" pageNumber="25">
<emphasis id="C6B0D335B35BFFB4DADFE072FEDF23AC" bold="true" box="[199,326,367,391]" pageId="1" pageNumber="25">
<typeStatus id="2B7FB185B35BFFB4DADFE072FED823AC" box="[199,321,367,391]" pageId="1" pageNumber="25" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="92D597E2B35BFFB4DB56E073FE0623AD" LSID="urn:lsid:biocol.org:col:34871" box="[334,415,366,390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="25">USNM</collectionCode>
221342 (4),
<quantity id="333CA2C2B35BFFB4D82BE073FD7023AD" box="[563,745,366,390]" metricMagnitude="-2" metricUnit="m" metricValue="9.364999999999998" metricValueMax="13.219999999999999" metricValueMin="5.51" pageId="1" pageNumber="25" unit="mm" value="93.64999999999999" valueMax="132.2" valueMin="55.1">55.1132.2 mm</quantity>
,
<collectingCountry id="8CD34FB7B35BFFB4D8EDE073FCFE23AD" box="[757,871,366,390]" name="Sri Lanka" pageId="1" pageNumber="25">Sri Lanka</collectingCountry>
, Kakaithivu, about
<quantity id="333CA2C2B35BFFB4DE5DE073FB0D23AD" box="[1093,1172,366,390]" metricMagnitude="3" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="25" unit="km" value="4.8">4.8 km</quantity>
south of Vaddukkodai, Jaffna area,
<quantity id="333CA2C2B35BFFB4DB39E08FFEF22381" box="[289,363,402,426]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="25" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">23 m</quantity>
, T. Roberts,
<date id="807A29E7B35BFFB4DBE5E08FFD412381" box="[509,728,402,426]" pageId="1" pageNumber="25" value="1970-03-20" valueMax="1970-03-21" valueMin="1970-03-20">2021 March 1970</date>
;
<collectionCode id="92D597E2B35BFFB4D8FDE08FFC822381" LSID="urn:lsid:biocol.org:col:34700" box="[741,795,402,426]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="1" pageNumber="25">CAS</collectionCode>
<quantity id="333CA2C2B35BFFB4D93BE08FFC652381" box="[803,1020,402,426]" metricMagnitude="5" metricUnit="m" metricValue="2.336571047" pageId="1" pageNumber="25" unit="mm" value="2.336571047E8">233657, 104.7 mm</quantity>
, taken with
<collectionCode id="92D597E2B35BFFB4DE90E08FFB402381" LSID="urn:lsid:biocol.org:col:34871" box="[1160,1241,402,426]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="25">USNM</collectionCode>
221342;
<collectionCode id="92D597E2B35BFFB4DF5DE08FFA052381" box="[1349,1436,402,426]" httpUri="http://grbio.org/cool/mve2-984h" name="See FMRI" pageId="1" pageNumber="25">CMFRI</collectionCode>
GB.9.1.2.1.1--2 (2),
<quantity id="333CA2C2B35BFFB4DB64E0ABFDA223E5" box="[380,571,438,462]" metricMagnitude="-1" metricUnit="m" metricValue="1.0819999999999999" metricValueMax="1.148" metricValueMin="1.016" pageId="1" pageNumber="25" unit="mm" value="108.19999999999999" valueMax="114.8" valueMin="101.6">101.6114.8 mm</quantity>
, off Tuticorin,
<collectingCountry id="8CD34FB7B35BFFB4D8F9E0ABFC8423E5" box="[737,797,438,462]" name="India" pageId="1" pageNumber="25">India</collectingCountry>
, mini trawl, K.K. Bineesh &amp; R. Shanis,
<date id="807A29E7B35BFFB4DEC2E0ABFA0123E5" box="[1242,1432,438,462]" pageId="1" pageNumber="25" value="2010-10-17">17 October 2010</date>
;
<collectionCode id="92D597E2B35BFFB4DA8FE0C7FF7723D9" box="[151,238,474,498]" httpUri="http://grbio.org/cool/mve2-984h" name="See FMRI" pageId="1" pageNumber="25">CMFRI</collectionCode>
GB.9.1.2.1.3--13, (11),
<quantity id="333CA2C2B35BFFB4DBE3E0C7FD0623D9" box="[507,671,474,498]" metricMagnitude="-2" metricUnit="m" metricValue="8.8" metricValueMax="9.48" metricValueMin="8.12" pageId="1" pageNumber="25" unit="mm" value="88.0" valueMax="94.8" valueMin="81.2">81.294.8 mm</quantity>
, off Tuticorin,
<collectingCountry id="8CD34FB7B35BFFB4D95DE0C7FC1823D9" box="[837,897,474,498]" name="India" pageId="1" pageNumber="25">India</collectingCountry>
, K.K. Bineesh &amp; R. Shanis,
<date id="807A29E7B35BFFB4DEA6E0C7FA0023D9" box="[1214,1433,474,498]" pageId="1" pageNumber="25" value="2010-11-27">27 November 2010</date>
.
</paragraph>
<paragraph id="F47B0F27B35BFFB4DADFE0E3FCB5208D" blockId="1.[151,1437,294,967]" pageId="1" pageNumber="25">
<emphasis id="C6B0D335B35BFFB4DADFE0E3FE67203C" bold="true" box="[199,510,510,535]" pageId="1" pageNumber="25">Other material examined.</emphasis>
<taxonomicName id="33C474A4B35BFFB4D81DE0E3FCA9203D" box="[517,816,510,534]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="1" pageNumber="25" phylum="Chordata" rank="species" species="dussumieri">
<emphasis id="C6B0D335B35BFFB4D81DE0E3FCA9203D" box="[517,816,510,534]" italics="true" pageId="1" pageNumber="25">Colletteichthys dussumieri</emphasis>
</taxonomicName>
—Syntypes-MNHN A. 4715 and A. 4748;
<collectionCode id="92D597E2B35BFFB4DF00E0E3FAD7203D" LSID="urn:lsid:biocol.org:col:34700" box="[1304,1358,510,534]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="1" pageNumber="25">CAS</collectionCode>
23719 Malabar,
<collectingCountry id="8CD34FB7B35BFFB4DB1BE33FFED92011" box="[259,320,546,570]" name="India" pageId="1" pageNumber="25">India</collectingCountry>
;
<collectionCode id="92D597E2B35BFFB4DB54E33FFE1B2011" LSID="urn:lsid:biocol.org:col:34700" box="[332,386,546,570]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="1" pageNumber="25">CAS</collectionCode>
29743,
<collectingCountry id="8CD34FB7B35BFFB4DBF9E33FFDE12011" box="[481,632,546,570]" name="India" pageId="1" pageNumber="25">Cochin India</collectingCountry>
;
<collectionCode id="92D597E2B35BFFB4D89DE33FFD222011" LSID="urn:lsid:biocol.org:col:34700" box="[645,699,546,570]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="1" pageNumber="25">CAS</collectionCode>
2333658,
<collectingCountry id="8CD34FB7B35BFFB4D92DE33FFC552011" box="[821,972,546,570]" name="India" pageId="1" pageNumber="25">Cochin India</collectingCountry>
; AMS B. 8112, Malabar,
<collectingCountry id="8CD34FB7B35BFFB4DEEBE33FFAA92011" box="[1267,1328,546,570]" name="India" pageId="1" pageNumber="25">India</collectingCountry>
; AMS B. 8113, Malabar
<collectingCountry id="8CD34FB7B35BFFB4DB5DE35BFE1B2075" box="[325,386,582,606]" name="India" pageId="1" pageNumber="25">India</collectingCountry>
.
<taxonomicName id="33C474A4B35BFFB4DB88E35BFD512075" box="[400,712,582,606]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="1" pageNumber="25" phylum="Chordata" rank="species" species="occidentalis">
<emphasis id="C6B0D335B35BFFB4DB88E35BFD512075" box="[400,712,582,606]" italics="true" pageId="1" pageNumber="25">Colletteichthys occidentalis</emphasis>
</taxonomicName>
—Holotype-BPBM 29525,
<collectingCountry id="8CD34FB7B35BFFB4DE1CE35BFBF92075" box="[1028,1120,582,606]" name="Bahrain" pageId="1" pageNumber="25">Bahrain</collectingCountry>
; Paratypes—BPBM 30509,
<collectingCountry id="8CD34FB7B35BFFB4DA8FE377FF6A20A9" box="[151,243,618,642]" name="Bahrain" pageId="1" pageNumber="25">Bahrain</collectingCountry>
;
<collectionCode id="92D597E2B35BFFB4DAE6E376FED520A9" LSID="urn:lsid:biocol.org:col:34697" box="[254,332,619,642]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="1" pageNumber="25">BPBM</collectionCode>
33302,
<collectingCountry id="8CD34FB7B35BFFB4DBBEE377FDA520A9" box="[422,572,618,642]" name="Saudi Arabia" pageId="1" pageNumber="25">Saudi Arabia</collectingCountry>
;
<collectionCode id="92D597E2B35BFFB4D85FE376FD0C20A9" LSID="urn:lsid:biocol.org:col:34697" box="[583,661,619,642]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="1" pageNumber="25">BPBM</collectionCode>
33398,
<collectingCountry id="8CD34FB7B35BFFB4D8F7E377FC1F20A9" box="[751,902,618,642]" name="Saudi Arabia" pageId="1" pageNumber="25">Saudi Arabia</collectingCountry>
;
<collectionCode id="92D597E2B35BFFB4D988E376FC4720A9" LSID="urn:lsid:biocol.org:col:34697" box="[912,990,619,642]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="1" pageNumber="25">BPBM</collectionCode>
41090,
<collectingCountry id="8CD34FB7B35BFFB4DE21E377FB0C20A9" box="[1081,1173,618,642]" name="Bahrain" pageId="1" pageNumber="25">Bahrain</collectingCountry>
;
<collectionCode id="92D597E2B35BFFB4DEB8E376FB7720A9" LSID="urn:lsid:biocol.org:col:34697" box="[1184,1262,619,642]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="1" pageNumber="25">BPBM</collectionCode>
21485, Quatar;
<collectionCode id="92D597E2B35BFFB4DA8FE393FF71208D" LSID="urn:lsid:biocol.org:col:34871" box="[151,232,654,678]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="25">USNM</collectionCode>
147913,
<collectingCountry id="8CD34FB7B35BFFB4DB49E393FE7E208D" box="[337,487,654,678]" name="Saudi Arabia" pageId="1" pageNumber="25">Saudi Arabia</collectingCountry>
;
<collectionCode id="92D597E2B35BFFB4DBEAE393FDB0208D" LSID="urn:lsid:biocol.org:col:34700" box="[498,553,654,678]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="1" pageNumber="25">CAS</collectionCode>
233656,
<collectingCountry id="8CD34FB7B35BFFB4D88AE393FCB1208D" box="[658,808,654,678]" name="Saudi Arabia" pageId="1" pageNumber="25">Saudi Arabia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="BCDE5CACB35BFFB4DADFE3AFFEC921ED" pageId="1" pageNumber="25" type="diagnosis">
<paragraph id="F47B0F27B35BFFB4DADFE3AFFEC921ED" blockId="1.[151,1437,294,967]" pageId="1" pageNumber="25">
<emphasis id="C6B0D335B35BFFB4DADFE3AFFED820E0" bold="true" box="[199,321,690,715]" pageId="1" pageNumber="25">Diagnosis.</emphasis>
A species of
<taxonomicName id="33C474A4B35BFFB4DBF3E3AEFD0A20E0" box="[491,659,691,715]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="1" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="C6B0D335B35BFFB4DBF3E3AEFD0A20E0" box="[491,659,691,715]" italics="true" pageId="1" pageNumber="25">Colletteichthys</emphasis>
</taxonomicName>
due to the following features: three dorsal-fin spines; the body is naked; a funnel-shaped pit at the top of the pectoral-fin axil; lower gill opening well below the lower pectoral-fin base; two subopercular spines; a single tentacle located above the posterior portion of the eye; no maxillary flaps; anterior nasal tentacle not elongate; dorsal-fin rays usually 20, anal-fin rays usually 16.
<taxonomicName id="33C474A4B35BFFB4DE6EE203FA02211D" box="[1142,1435,798,822]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="1" pageNumber="25" phylum="Chordata" rank="species" species="flavipinnis">
<emphasis id="C6B0D335B35BFFB4DE6EE203FA02211D" box="[1142,1435,798,822]" italics="true" pageId="1" pageNumber="25">Colletteichthys flavipinnis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="33C474A4B35BFFB4DB3DE25EFE582170" box="[293,449,835,859]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="1" pageNumber="25" phylum="Chordata" rank="species" species="dussumieri">
<emphasis id="C6B0D335B35BFFB4DB3DE25EFE582170" box="[293,449,835,859]" italics="true" pageId="1" pageNumber="25">C. dussumieri</emphasis>
</taxonomicName>
in having one (
<figureCitation id="6CFF13A2B35BFFB4D869E25FFD2D2171" box="[625,692,834,858]" captionStart="FIGURE 3" captionStartId="2.[151,250,1344,1367]" captionTargetBox="[403,1183,814,1322]" captionTargetId="figure@2.[403,1184,788,1323]" captionTargetPageId="2" captionText="FIGURE 3. Eye of holotype of Colletteichthys flavipinnis showing single tentacle above it. Photograph by D. W. Greenfield." httpUri="https://zenodo.org/record/211150/files/figure.png" pageId="1" pageNumber="25">Fig. 3</figureCitation>
) instead of two or more (
<figureCitation id="6CFF13A2B35BFFB4D9CCE25FFB812171" box="[980,1048,834,858]" captionStart="FIGURE 4" captionStartId="3.[151,250,1186,1209]" captionTargetBox="[242,1343,392,1179]" captionTargetId="figure@3.[242,1344,383,1179]" captionTargetPageId="3" captionText="FIGURE 4. Eye of Colletteichthys dussumieri (CAS 233658) showing three tentacles above it (Fig. 5 from Greenfield, 2012)." httpUri="https://zenodo.org/record/211151/files/figure.png" pageId="1" pageNumber="25">Fig. 4</figureCitation>
) tentacles above the eye. It differs from both
<taxonomicName id="33C474A4B35BFFB4DB0CE27AFE272155" box="[276,446,870,894]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="1" pageNumber="25" phylum="Chordata" rank="species" species="occidentalis">
<emphasis id="C6B0D335B35BFFB4DB0CE27AFE272155" box="[276,446,870,894]" italics="true" pageId="1" pageNumber="25">C. occidentalis</emphasis>
</taxonomicName>
and
<taxonomicName id="33C474A4B35BFFB4DBE4E27AFD032155" box="[508,666,870,894]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="1" pageNumber="25" phylum="Chordata" rank="species" species="dussumieri">
<emphasis id="C6B0D335B35BFFB4DBE4E27AFD032155" box="[508,666,870,894]" italics="true" pageId="1" pageNumber="25">C. dussumieri</emphasis>
</taxonomicName>
in having yellow-orange fins, and from
<taxonomicName id="33C474A4B35BFFB4DE6DE27AFAB82155" box="[1141,1313,870,894]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="1" pageNumber="25" phylum="Chordata" rank="species" species="occidentalis">
<emphasis id="C6B0D335B35BFFB4DE6DE27AFAB82155" box="[1141,1313,870,894]" italics="true" pageId="1" pageNumber="25">C. occidentalis</emphasis>
</taxonomicName>
in usually having 16 instead of 15 anal-fin rays and 20 instead of 19 dorsal-fin rays, and in having a slightly smaller (12% SL) eye (
<figureCitation id="6CFF13A2B35BFFB4DAE6E2B3FEDB21ED" box="[254,322,942,966]" captionStart="FIGURE 5" captionStartId="4.[151,250,1080,1103]" captionTargetBox="[215,1388,202,1054]" captionTargetId="figure@4.[198,1403,195,1075]" captionTargetPageId="4" captionText="FIGURE 5. Plot of eye size as percent standard length versus standard length of C. occidentalis (blue diamonds) and C. flavipinnis from India (magenta squares) and Sri Lanka (green triangles)." httpUri="https://zenodo.org/record/211152/files/figure.png" pageId="1" pageNumber="25">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="A0BB5FAFB35BFFB4DA8FE79AFA1424B5" box="[151,1421,1671,1694]" httpUri="https://zenodo.org/record/211148/files/figure.png" pageId="1" pageNumber="25" targetBox="[151,1436,1022,1647]" targetPageId="1">
<paragraph id="F47B0F27B35BFFB4DA8FE79AFA1424B5" blockId="1.[151,1421,1671,1694]" box="[151,1421,1671,1694]" pageId="1" pageNumber="25">
<emphasis id="C6B0D335B35BFFB4DA8FE79AFE8A24B6" bold="true" box="[151,275,1671,1694]" pageId="1" pageNumber="25">FIGURE 1.</emphasis>
Freshly collected specimen of
<taxonomicName id="33C474A4B35BFFB4D856E795FC5D24B5" authority="Tuticorin" authorityName="Tuticorin" box="[590,964,1672,1694]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="1" pageNumber="25" phylum="Chordata" rank="species" species="flavipinnis">
<emphasis id="C6B0D335B35BFFB4D856E795FCCE24B5" box="[590,855,1672,1694]" italics="true" pageId="1" pageNumber="25">Colletteichthys flavipinnis</emphasis>
, Tuticorin
</taxonomicName>
, India. Photograph by Arumugam. Murugan.
</paragraph>
</caption>
<subSubSection id="BCDE5CACB35BFFB6DADFE7D8FE2E27A3" lastPageId="3" lastPageNumber="27" pageId="1" pageNumber="25" type="description">
<paragraph id="F47B0F27B35BFFB7DADFE7D8FA042560" blockId="1.[151,1436,1733,2009]" lastBlockId="2.[151,1437,1411,2012]" lastPageId="2" lastPageNumber="26" pageId="1" pageNumber="25">
<emphasis id="C6B0D335B35BFFB4DADFE7D8FEC024F5" bold="true" box="[199,345,1733,1758]" pageId="1" pageNumber="25">Description.</emphasis>
Dorsal-fin elements
<date id="807A29E7B35BFFB4D85DE7D8FD1F24F6" box="[581,646,1733,1757]" pageId="1" pageNumber="25" value="1920-03">III-20</date>
(1920, usually 20); anal-fin rays 16 (1416, usually 16); pectoral-fin rays 22 (2022, usually 21); vertebrae 28; head length 35.6 (33.439.3, 36.8%); head width 31.0 (25.232.4, 28.4%); head depth 22.5 (19.523.6, 21.7%); bony interorbital width 3.7 (3.55.0, 4.3%); fleshy interorbital width 15.7 (12.015.7, 13.7%); orbit diameter 7.0 (6.89.3, 7.9%); snout length 5.8 (5.26.8, 5.8%); upper jaw length 18.3 (17.219.7, 18.5%); mouth width at rictus 22.2 (18.224.1, 21.0%); first predorsal-fin distance 36.7 (32.438.7, 35.6%); second predorsal-fin distance 48.9 (44.449.9, 47.3%); preanal-fin distance 62.3 (60.564.7, 62.3%); greatest body depth 22.2 (22.226.7, 25.2%); caudal-peduncle depth 9.6 (8.910.3, 9.7%); caudalpeduncle length 5.9 (5.98.9, 7.3%); first dorsal-fin base length 9.6 (7.212.4, 10.3%); second dorsal-fin base Head moderately depressed and of moderate width, eyes medium in size and not raised above head profile, interorbital area almost flat. A funnel-shaped pit at top of pectoral-fin axil, with glandular tissue inside and extending from ventral pit margin onto axil and onto side under pectoral fin. No scales. Three lateral lines, each pore with two short skin flaps: the upper one with 38 pores originating above opercle, running posteriorly in a straight line to the caudal peduncle; middle lateral line with three scattered pores along midline of side; lower line with 21 pores, running from under pectoral-fin base along anal-fin base to end of anal fin. Three rows of sensory pores with flaps across head: the posteriormost with eight pores in front of the first dorsal fin running from opercular spine to opercular spine; the second with nine pores at middle of head from preopercle to preopercle; the third, four pores with larger flaps behind the eyes. This third row could be connected to a line of pores that run behind and then under the eye. A single tentacle above eye posterior to pupil. Posterior nostrils relatively large oval openings, anterior nostrils tubular with opening on the end and lacking tentacles on the tube. Two small pores, each with a set of flaps centered on snout between posterior nostrils. A row of pores with flaps on end of snout below anterior nostrils above upper lip.
<collectingCountry id="8CD34FB7B358FFB7D811E62EFDAF2560" box="[521,566,1843,1867]" name="India" pageId="2" pageNumber="26">End</collectingCountry>
of maxilla with a small cirrus. Ventral margin of dentary with numerous cirri.
</paragraph>
<paragraph id="F47B0F27B358FFB7DA8FE18AFDFC22DC" pageId="2" pageNumber="26">
<table id="86C4FD87B358004ADA8FE18AFA0522DC" box="[151,1436,151,247]" gridcols="3" gridrows="3" pageId="2" pageNumber="26">
<tr id="4AF40D65B358004ADA8FE18AFA052284" box="[151,1436,151,175]" gridrow="0" pageId="2" pageNumber="26">
<th id="09256419B358004ADA8FE18AFD752284" box="[151,748,151,175]" gridcol="0" gridrow="0" pageId="2" pageNumber="26">length 48.7 (44.049.8, 47.5%); anal-fin base length</th>
<th id="09256419B358004AD8ECE18AFC362284" box="[756,943,151,175]" gridcol="1" gridrow="0" pageId="2" pageNumber="26">34.5 (27.034.5,</th>
<th id="09256419B358004AD9AFE18AFA052284" box="[951,1436,151,175]" gridcol="2" gridrow="0" pageId="2" pageNumber="26">31.3%); caudal-fin length 20.9 (19.923.8,</th>
</tr>
<tr id="4AF40D65B358004ADA8FE1A1FA0522FF" box="[151,1436,188,212]" gridrow="1" pageId="2" pageNumber="26">
<th id="09256419B358004ADA8FE1A1FD7522FF" box="[151,748,188,212]" gridcol="0" gridrow="1" pageId="2" pageNumber="26">21.6%); pectoral-fin length 23.9 (18.024.5, 21.6%);</th>
<td id="09256419B358004AD8ECE1A1FC3622FF" box="[756,943,188,212]" gridcol="1" gridrow="1" pageId="2" pageNumber="26">pelvic-fin length</td>
<td id="09256419B358004AD9AFE1A1FA0522FF" box="[951,1436,188,212]" gridcol="2" gridrow="1" pageId="2" pageNumber="26">24.5 (20.927.1, 24.7%); distance between</td>
</tr>
<tr id="4AF40D65B358004ADA8FE1C2FA0522DC" box="[151,1436,223,247]" gridrow="2" pageId="2" pageNumber="26" rowspan-1="1" rowspan-2="1">
<th id="09256419B358004ADA8FE1C2FD7522DC" box="[151,748,223,247]" gridcol="0" gridrow="2" pageId="2" pageNumber="26">pelvic-fin bases 11.9 (10.214.6, 12.1%).</th>
</tr>
</table>
</paragraph>
<caption id="A0BB5FAFB358FFB7DA8FE3EBFB1E2127" box="[151,1159,758,781]" httpUri="https://zenodo.org/record/211149/files/figure.png" pageId="2" pageNumber="26" targetBox="[172,1434,301,735]" targetPageId="2">
<paragraph id="F47B0F27B358FFB7DA8FE3EBFB1E2127" blockId="2.[151,1159,758,781]" box="[151,1159,758,781]" pageId="2" pageNumber="26">
<emphasis id="C6B0D335B358FFB7DA8FE3EBFE8A2120" bold="true" box="[151,275,758,781]" pageId="2" pageNumber="26">FIGURE 2.</emphasis>
Freshly collected holotype of
<taxonomicName id="33C474A4B358FFB7D85FE3EBFCC92127" box="[583,848,758,780]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="2" pageNumber="26" phylum="Chordata" rank="species" species="flavipinnis">
<emphasis id="C6B0D335B358FFB7D85FE3EBFCC92127" box="[583,848,758,780]" italics="true" pageId="2" pageNumber="26">Colletteichthys flavipinnis</emphasis>
</taxonomicName>
. Photograph by K.K. Bineesh.
</paragraph>
</caption>
<caption id="A0BB5FAFB358FFB7DA8FE45DFA1F277C" box="[151,1414,1344,1367]" httpUri="https://zenodo.org/record/211150/files/figure.png" pageId="2" pageNumber="26" targetBox="[403,1183,814,1322]" targetPageId="2">
<paragraph id="F47B0F27B358FFB7DA8FE45DFA1F277C" blockId="2.[151,1414,1344,1367]" box="[151,1414,1344,1367]" pageId="2" pageNumber="26">
<emphasis id="C6B0D335B358FFB7DA8FE45DFE8B277D" bold="true" box="[151,274,1344,1367]" pageId="2" pageNumber="26">FIGURE 3.</emphasis>
Eye of holotype of
<taxonomicName id="33C474A4B358FFB7DBC4E45CFD7F277C" box="[476,742,1345,1367]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="2" pageNumber="26" phylum="Chordata" rank="species" species="flavipinnis">
<emphasis id="C6B0D335B358FFB7DBC4E45CFD7F277C" box="[476,742,1345,1367]" italics="true" pageId="2" pageNumber="26">Colletteichthys flavipinnis</emphasis>
</taxonomicName>
showing single tentacle above it. Photograph by D.W. Greenfield.
</paragraph>
</caption>
<paragraph id="F47B0F27B358FFB6DADFE64AFB1E234F" blockId="2.[151,1437,1411,2012]" lastBlockId="3.[151,1437,151,356]" lastPageId="3" lastPageNumber="27" pageId="2" pageNumber="26">
<emphasis id="C6B0D335B358FFB7DADFE64AFDFC255B" bold="true" box="[199,613,1879,1904]" pageId="2" pageNumber="26">
Color of freshly collected
<typeStatus id="2B7FB185B358FFB7DBE6E645FDFC255B" box="[510,613,1880,1904]" pageId="2" pageNumber="26" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="6CFF13A2B358FFB7D86DE64AFD252544" box="[629,700,1879,1903]" captionStart="FIGURE 2" captionStartId="2.[151,250,758,781]" captionTargetBox="[172,1434,301,735]" captionTargetId="figure@2.[151,1436,301,737]" captionTargetPageId="2" captionText="FIGURE 2. Freshly collected holotype of Colletteichthys flavipinnis. Photograph by K. K. Bineesh." httpUri="https://zenodo.org/record/211149/files/figure.png" pageId="2" pageNumber="26">Fig. 2</figureCitation>
). Background color of head and body light brown, overlaid with dark brown to black markings. A dark band crossing the caudal peduncle, the posterior edge curved anteriorly in a semicircle. Dorsal half of sides crossed by three dark bands that angle forward: the posteriormost one under soft dorsal-fin rays 1115; middle band under rays 59; anterior band under rays 12, angling forward under pectoral fin. Ventral half of sides with five narrow dark bars angling forward: two under posteriormost dorsal bar; third in space between posterior and middle dorsal bars; and fourth and fifth under anterior dorsal bar. A dark bar across back through first dorsal fin. Top and sides of head with dark mottling, a narrow light brown band across interorbital space. Upper and lower jaws dark brown, crossed by five light brown bars, end of maxilla light brown. Pupil of eye black, iris light brown with dark spoke-like markings. All fins yellow-orange, second dorsal and anal fins with dark bars extending out from body. Faint dark bands on pectoral and caudal fins.
</paragraph>
<caption id="A0BB5FAFB359FFB6DA8FE5BFFA012692" box="[151,1432,1186,1209]" httpUri="https://zenodo.org/record/211151/files/figure.png" pageId="3" pageNumber="27" targetBox="[242,1343,392,1179]" targetPageId="3">
<paragraph id="F47B0F27B359FFB6DA8FE5BFFA012692" blockId="3.[151,1432,1186,1209]" box="[151,1432,1186,1209]" pageId="3" pageNumber="27">
<emphasis id="C6B0D335B359FFB6DA8FE5BFFE8A2693" bold="true" box="[151,275,1186,1209]" pageId="3" pageNumber="27">FIGURE 4.</emphasis>
Eye of
<taxonomicName id="33C474A4B359FFB6DB7AE5BEFDF62692" box="[354,623,1187,1209]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="dussumieri">
<emphasis id="C6B0D335B359FFB6DB7AE5BEFDF62692" box="[354,623,1187,1209]" italics="true" pageId="3" pageNumber="27">Colletteichthys dussumieri</emphasis>
</taxonomicName>
(CAS 233658) showing three tentacles above it (Fig. 5 from Greenfield, 2012).
</paragraph>
</caption>
<paragraph id="F47B0F27B359FFB6DADFE5C2FBA3276B" blockId="3.[151,1436,1247,2029]" pageId="3" pageNumber="27">
<emphasis id="C6B0D335B359FFB6DADFE5C2FDF726D3" bold="true" box="[199,622,1247,1272]" pageId="3" pageNumber="27">
Color of freshly collected
<typeStatus id="2B7FB185B359FFB6DBE0E5FDFDF726D3" box="[504,622,1248,1272]" pageId="3" pageNumber="27" type="paratype">paratypes</typeStatus>
</emphasis>
(
<figureCitation id="6CFF13A2B359FFB6D865E5FDFD5826D3" box="[637,705,1248,1272]" captionStart="FIGURE 6" captionStartId="4.[151,250,1989,2012]" captionTargetBox="[151,1435,1182,1966]" captionTargetId="figure@4.[151,1436,1182,1967]" captionTargetPageId="4" captionText="FIGURE 6. Head coloration of freshly collected paratype of Colletteichthys flavipinnis. Photograph by K. K. Bineesh." httpUri="https://zenodo.org/record/211153/files/figure.png" pageId="3" pageNumber="27">Fig. 6</figureCitation>
). Coloration as in
<typeStatus id="2B7FB185B359FFB6D996E5FDFC6926D3" box="[910,1008,1248,1272]" pageId="3" pageNumber="27" type="holotype">holotype</typeStatus>
except that some individuals have red on the snout, cheek, top of head and nape as follows: The area from the posterior nostrils forward to end of snout; a relatively large patch behind each eye; various smaller spots on nape and on cheek.
</paragraph>
<paragraph id="F47B0F27B359FFB6DADFE456FE2E27A3" blockId="3.[151,1436,1247,2029]" pageId="3" pageNumber="27">
<emphasis id="C6B0D335B359FFB6DADFE456FD80274F" bold="true" box="[199,537,1355,1380]" pageId="3" pageNumber="27">
Color of preserved
<typeStatus id="2B7FB185B359FFB6DBAAE451FD80274F" box="[434,537,1356,1380]" pageId="3" pageNumber="27" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="6CFF13A2B359FFB6D831E451FDF6274F" box="[553,623,1356,1380]" captionStart="FIGURE 7" captionStartId="5.[151,250,550,573]" captionTargetBox="[159,1424,203,518]" captionTargetId="figure@5.[151,1436,193,529]" captionTargetPageId="5" captionText="FIGURE 7. Preserved holotype of Colletteichthys flavipinnis. Photograph by D. W. Greenfield." httpUri="https://zenodo.org/record/211154/files/figure.png" pageId="3" pageNumber="27">Fig. 7</figureCitation>
). Color as in fresh specimen except that background is light cream and dark markings are brown.
</paragraph>
</subSubSection>
<subSubSection id="BCDE5CACB359FFB6DADFE48EFA142787" box="[199,1421,1427,1452]" pageId="3" pageNumber="27" type="distribution">
<paragraph id="F47B0F27B359FFB6DADFE48EFA142787" blockId="3.[151,1436,1247,2029]" box="[199,1421,1427,1452]" pageId="3" pageNumber="27">
<emphasis id="C6B0D335B359FFB6DADFE48EFEC02787" bold="true" box="[199,345,1427,1452]" pageId="3" pageNumber="27">Distribution</emphasis>
. Known only from the Tuticorin area of
<collectingCountry id="8CD34FB7B359FFB6D93AE489FCC52787" box="[802,860,1428,1452]" name="India" pageId="3" pageNumber="27">India</collectingCountry>
and the adjacent Jaffna area of
<collectingCountry id="8CD34FB7B359FFB6DEA6E489FAB52787" box="[1214,1324,1428,1452]" name="Sri Lanka" pageId="3" pageNumber="27">Sri Lanka</collectingCountry>
(
<figureCitation id="6CFF13A2B359FFB6DF23E489FA192787" box="[1339,1408,1428,1452]" captionStart="FIGURE 8" captionStartId="5.[151,250,1990,2013]" captionTargetBox="[267,1321,627,1940]" captionTargetId="figure@5.[254,1341,594,1966]" captionTargetPageId="5" captionText="FIGURE 8. Distribution of species of Colletteichthys. Circles = C. occidentalis, stars = C. dussumieri, squares = C. flavipinnis." httpUri="https://zenodo.org/record/211155/files/figure.png" pageId="3" pageNumber="27">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BCDE5CACB359FFB6DADFE4A5FD722513" pageId="3" pageNumber="27" type="etymology">
<paragraph id="F47B0F27B359FFB6DADFE4A5FD8827DF" blockId="3.[151,1436,1247,2029]" pageId="3" pageNumber="27">
<emphasis id="C6B0D335B359FFB6DADFE4A5FED727FB" bold="true" box="[199,334,1464,1488]" pageId="3" pageNumber="27">Etymology.</emphasis>
The specific epithet is an adjective from the Latin
<emphasis id="C6B0D335B359FFB6D98CE4A5FC4927FB" box="[916,976,1464,1488]" italics="true" pageId="3" pageNumber="27">flavis</emphasis>
(yellow) and
<emphasis id="C6B0D335B359FFB6DE68E4A4FB3627FB" box="[1136,1199,1465,1488]" italics="true" pageId="3" pageNumber="27">pinna</emphasis>
(fin) referring to the yellow-orange fins of the species.
</paragraph>
<paragraph id="F47B0F27B359FFB6DADFE4E2FB3B24E7" blockId="3.[151,1436,1247,2029]" pageId="3" pageNumber="27">
<emphasis id="C6B0D335B359FFB6DADFE4E2FEFF2433" bold="true" box="[199,358,1535,1560]" pageId="3" pageNumber="27">Comparisons</emphasis>
.
<taxonomicName id="33C474A4B359FFB6DB6BE71DFD052433" box="[371,668,1536,1560]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="dussumieri">
<emphasis id="C6B0D335B359FFB6DB6BE71DFD052433" box="[371,668,1536,1560]" italics="true" pageId="3" pageNumber="27">Colletteichthys dussumieri</emphasis>
</taxonomicName>
has two or three tentacles above the eye (
<bibRefCitation id="905572D6B359FFB6DE72E71DFAB42433" author="Greenfield" box="[1130,1325,1536,1560]" pageId="3" pageNumber="27" refString="Greenfield, D. W. (2012) Colletteichthys occidentalis, a new toadfish species from the Arabian Peninsula and northern Arabian Sea (Teleostei: Batrachoididae). Zootaxa, 3165, 64 - 68." type="journal article" year="2012">Greenfield, 2012</bibRefCitation>
), whereas
<taxonomicName id="33C474A4B359FFB6DA8FE738FEB72417" box="[151,302,1572,1596]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="flavipinnis">
<emphasis id="C6B0D335B359FFB6DA8FE738FEB72417" box="[151,302,1572,1596]" italics="true" pageId="3" pageNumber="27">C. flavipinnis</emphasis>
</taxonomicName>
has only one, and
<taxonomicName id="33C474A4B359FFB6DBE6E738FD0D2417" box="[510,660,1572,1596]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="flavipinnis">
<emphasis id="C6B0D335B359FFB6DBE6E738FD0D2417" box="[510,660,1572,1596]" italics="true" pageId="3" pageNumber="27">C. flavipinnis</emphasis>
</taxonomicName>
has yellow fins whereas
<taxonomicName id="33C474A4B359FFB6D9B5E738FBD02417" box="[941,1097,1572,1596]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="dussumieri">
<emphasis id="C6B0D335B359FFB6D9B5E738FBD02417" box="[941,1097,1572,1596]" italics="true" pageId="3" pageNumber="27">C. dussumieri</emphasis>
</taxonomicName>
has dark fins, sometimes with yellow tinges (
<figureCitation id="6CFF13A2B359FFB6DB23E755FEE6244B" box="[315,383,1608,1632]" captionStart="FIGURE 9" captionStartId="6.[151,250,682,705]" captionTargetBox="[158,1436,222,650]" captionTargetId="figure@6.[151,1436,193,661]" captionTargetPageId="6" captionText="FIGURE 9. Freshly collected specimen of Colletteichthys dussumieri, Cochin estuary, India. Photograph by K. K. Bineesh." httpUri="https://zenodo.org/record/211156/files/figure.png" pageId="3" pageNumber="27">Fig. 9</figureCitation>
). Whereas some specimens of
<taxonomicName id="33C474A4B359FFB6D8CFE754FCF7244B" box="[727,878,1608,1632]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="flavipinnis">
<emphasis id="C6B0D335B359FFB6D8CFE754FCF7244B" box="[727,878,1608,1632]" italics="true" pageId="3" pageNumber="27">C. flavipinnis</emphasis>
</taxonomicName>
have red on the top of the head, the second author has never seen this coloration in
<taxonomicName id="33C474A4B359FFB6D808E770FD3724AF" box="[528,686,1644,1668]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="dussumieri">
<emphasis id="C6B0D335B359FFB6D808E770FD3724AF" box="[528,686,1644,1668]" italics="true" pageId="3" pageNumber="27">C. dussumieri</emphasis>
</taxonomicName>
.
<taxonomicName id="33C474A4B359FFB6D8A5E771FC6C24AF" box="[701,1013,1644,1668]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="occidentalis">
<emphasis id="C6B0D335B359FFB6D8A5E771FC6C24AF" box="[701,1013,1644,1668]" italics="true" pageId="3" pageNumber="27">Colletteichthys occidentalis</emphasis>
</taxonomicName>
also has only one tentacle above the eye, but differs in having gray fins with white lines instead of yellow, usually 15 instead of 16 anal-fin rays and 19 instead of 20 dorsal-fin rays, and in having a larger eye for similar sized individuals (
<figureCitation id="6CFF13A2B359FFB6DE48E7A9FB0D24E7" box="[1104,1172,1716,1740]" captionStart="FIGURE 5" captionStartId="4.[151,250,1080,1103]" captionTargetBox="[215,1388,202,1054]" captionTargetId="figure@4.[198,1403,195,1075]" captionTargetPageId="4" captionText="FIGURE 5. Plot of eye size as percent standard length versus standard length of C. occidentalis (blue diamonds) and C. flavipinnis from India (magenta squares) and Sri Lanka (green triangles)." httpUri="https://zenodo.org/record/211152/files/figure.png" pageId="3" pageNumber="27">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="F47B0F27B359FFB6DADFE7C5FD722513" blockId="3.[151,1436,1247,2029]" pageId="3" pageNumber="27">
The COI sequences of
<taxonomicName id="33C474A4B359FFB6DBC5E7C5FC9124DB" box="[477,776,1752,1776]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="flavipinnis">
<emphasis id="C6B0D335B359FFB6DBC5E7C5FC9124DB" box="[477,776,1752,1776]" italics="true" pageId="3" pageNumber="27">Colletteichthys flavipinnis</emphasis>
</taxonomicName>
and
<taxonomicName id="33C474A4B359FFB6D952E7C4FC7524DB" box="[842,1004,1752,1776]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="dussumieri">
<emphasis id="C6B0D335B359FFB6D952E7C4FC7524DB" box="[842,1004,1752,1776]" italics="true" pageId="3" pageNumber="27">C. dussumieri</emphasis>
</taxonomicName>
showed a clear barcode split (4.4% divergence) congruent with morphological differences (
<figureCitation id="6CFF13A2B359FFB6D909E7E1FCFF253F" box="[785,870,1788,1812]" captionStart="FIGURE 10" captionStartId="6.[151,250,1517,1540]" captionTargetBox="[195,1383,769,1448]" captionTargetId="figure@6.[178,1397,755,1460]" captionTargetPageId="6" captionText="FIGURE 10. Neighbor Joining tree of Cytochrome oxidase I (COI) gene sequences derived from Colletteichthys species using K 2 P distances." httpUri="https://zenodo.org/record/211157/files/figure.png" pageId="3" pageNumber="27">Fig. 10</figureCitation>
). Very low (0.5%) intraspecific divergences were found in six specimens of
<emphasis id="C6B0D335B359FFB6DBA5E63DFD722513" box="[445,747,1824,1848]" italics="true" pageId="3" pageNumber="27">
<taxonomicName id="33C474A4B359FFB6DBA5E63DFD7F2513" box="[445,742,1824,1848]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="flavipinnis">Colletteichthys flavipinnis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BCDE5CACB359FFB6DADFE659FB1725C7" pageId="3" pageNumber="27" type="discussion">
<paragraph id="F47B0F27B359FFB6DADFE659FB1725C7" blockId="3.[151,1436,1247,2029]" pageId="3" pageNumber="27">
<emphasis id="C6B0D335B359FFB6DADFE659FEA22577" bold="true" box="[199,315,1860,1884]" pageId="3" pageNumber="27">Remarks.</emphasis>
<taxonomicName id="33C474A4B359FFB6DB52E659FDE12577" box="[330,632,1860,1884]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="flavipinnis">
<emphasis id="C6B0D335B359FFB6DB52E659FDE12577" box="[330,632,1860,1884]" italics="true" pageId="3" pageNumber="27">Colletteichthys flavipinnis</emphasis>
</taxonomicName>
appears to be a smaller species than
<taxonomicName id="33C474A4B359FFB6DE44E658FA992577" box="[1116,1280,1860,1884]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="dussumieri">
<emphasis id="C6B0D335B359FFB6DE44E658FA992577" box="[1116,1280,1860,1884]" italics="true" pageId="3" pageNumber="27">C. dussumieri</emphasis>
</taxonomicName>
. The largest specimens of
<taxonomicName id="33C474A4B359FFB6DB2BE674FE5225AB" box="[307,459,1896,1920]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="flavipinnis">
<emphasis id="C6B0D335B359FFB6DB2BE674FE5225AB" box="[307,459,1896,1920]" italics="true" pageId="3" pageNumber="27">C. flavipinnis</emphasis>
</taxonomicName>
collected to date are
<quantity id="333CA2C2B359FFB6D8A7E675FCAD25AB" box="[703,820,1896,1920]" metricMagnitude="-1" metricUnit="m" metricValue="1.148" pageId="3" pageNumber="27" unit="mm" value="114.8">114.8 mm</quantity>
. One of our specimens (
<quantity id="333CA2C2B359FFB6DE50E675FB2825AB" box="[1096,1201,1896,1920]" metricMagnitude="-2" metricUnit="m" metricValue="8.77" pageId="3" pageNumber="27" unit="mm" value="87.7">87.7 mm</quantity>
) is gravid with large eggs, whereas S. Roja (pers. comm.) has found that the minimum size at maturity for
<taxonomicName id="33C474A4B359FFB6DE46E690FB63258F" box="[1118,1274,1932,1956]" class="Actinopterygii" family="Batrachoididae" genus="Colletteichthys" kingdom="Animalia" order="Batrachoidiformes" pageId="3" pageNumber="27" phylum="Chordata" rank="species" species="dussumieri">
<emphasis id="C6B0D335B359FFB6DE46E690FB63258F" box="[1118,1274,1932,1956]" italics="true" pageId="3" pageNumber="27">C. dussumieri</emphasis>
</taxonomicName>
is
<quantity id="333CA2C2B359FFB6DF04E691FAE5258F" box="[1308,1404,1932,1956]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="27" unit="mm" value="130.0">130 mm</quantity>
. A specimen
<quantity id="333CA2C2B359FFB6DB10E6ADFEC125E3" box="[264,344,1968,1992]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="27" unit="mm" value="80.0">80 mm</quantity>
contained a single crab in its stomach with a carapace width of
<quantity id="333CA2C2B359FFB6DE34E6ADFB0A25E3" box="[1068,1171,1968,1992]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="3" pageNumber="27" unit="mm" value="17.7">17.7 mm</quantity>
. We do not know if red coloration on top of the head in some specimens may be related to their reproductive state.
</paragraph>
</subSubSection>
</treatment>
</document>