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<document id="CEC28C45B2CB75159EDE8A1F53734044" ID-DOI="10.5281/zenodo.276721" ID-GBIF-Dataset="868b3928-58b9-4b29-b518-d01d7bf2dbea" ID-ISSN="1175-5326" ID-Zenodo-Dep="276721" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460232901816" checkinUser="plazi" docAuthor="Collins, Allen G., Bentlage, Bastian, Lynn, Tara H., Morandini, André C. &amp; Marques, Antonio C." docDate="2011" docId="601F878AFFBAFFBA55F37AC62EC3F65E" docLanguage="en" docName="zt02753p068.pdf" docOrigin="Zootaxa 2753" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tamoya Muller 1859" docType="treatment" docVersion="8" lastPageNumber="58" masterDocId="9C26FFF2FFBEFFBF55647A512A1EF329" masterDocTitle="Naming the Bonaire banded box jelly, Tamoya ohboya, n. sp. (Cnidaria: Cubozoa: Carybdeida: Tamoyidae)" masterLastPageNumber="68" masterPageNumber="53" pageNumber="57" updateTime="1698239106890" updateUser="plazi">
<mods:mods id="B92E203AB89186A0F816278235AC0E98" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="81EBBCB36F6DC2745E5A8F2D7650F3E8">Naming the Bonaire banded box jelly, Tamoya ohboya, n. sp. (Cnidaria: Cubozoa: Carybdeida: Tamoyidae)</mods:title>
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<mods:namePart id="1E87F01C9E8D85173E0A49428B36658E">Collins, Allen G.</mods:namePart>
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<mods:namePart id="706803A8CA3DC64A586C56BFBE0A552C">Bentlage, Bastian</mods:namePart>
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<mods:namePart id="48690A014A6B9001780F8C9E54E25FF8">Lynn, Tara H.</mods:namePart>
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<mods:namePart id="6C9C83CFA6FB7ABB318930BC1278CBD2">Morandini, André C.</mods:namePart>
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<mods:namePart id="F0E47548F2F13A66EBAB968CD9F5C047">Marques, Antonio C.</mods:namePart>
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<mods:title id="E48CACC7049B443E8CBDFCC4FFBF2892">Zootaxa</mods:title>
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<mods:date id="732038C601B07FC1B01D1F801C0D13BA">2011</mods:date>
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<mods:number id="D4E4BA5B0ABD0DB44CBB2C18C73ABE9E">2753</mods:number>
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<mods:identifier id="B8AC6F1322B1C1C76C914C9806726987" type="DOI">10.5281/zenodo.276721</mods:identifier>
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<treatment id="601F878AFFBAFFBA55F37AC62EC3F65E" ID-DOI="http://doi.org/10.5281/zenodo.6194275" ID-GBIF-Taxon="119387555" ID-Zenodo-Dep="6194275" LSID="urn:lsid:plazi:treatment:601F878AFFBAFFBA55F37AC62EC3F65E" httpUri="http://treatment.plazi.org/id/601F878AFFBAFFBA55F37AC62EC3F65E" lastPageId="5" lastPageNumber="58" pageId="4" pageNumber="57">
<subSubSection id="A0AC6517FFBAFFBB55F37AC62B4DF3FB" pageId="4" pageNumber="57" type="nomenclature">
<paragraph id="E809369CFFBAFFBB55F37AC62BEFF398" blockId="4.[151,497,151,210]" box="[151,497,151,177]" pageId="4" pageNumber="57">
<heading id="B34181F0FFBAFFBB55F37AC62BEFF398" bold="true" box="[151,497,151,177]" fontSize="11" level="1" pageId="4" pageNumber="57" reason="1">
<emphasis id="DAC2EA8EFFBAFFBB55F37AC62BEFF398" bold="true" box="[151,497,151,177]" pageId="4" pageNumber="57">
Genus
<taxonomicName id="2FB64D1FFFBAFFBB558A7AC62BEFF398" authority="Muller, 1859" authorityName="Muller" authorityYear="1859" box="[238,497,151,177]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="4" pageNumber="57" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBAFFBB558A7AC62B54F398" bold="true" box="[238,330,151,177]" italics="true" pageId="4" pageNumber="57">Tamoya</emphasis>
<bibRefCitation id="8C274B6DFFBAFFBB54307AC62BEFF398" author="Muller" box="[340,497,151,177]" pageId="4" pageNumber="68" refString="Muller, F. (1859) Zwei neue Quallen von Santa Catharina (Brazilien). Abhandlungen der Naturforschenden Gesellschaft zu Halle - Originalaufsatze aus dem Gebiete der gesamten Naturwissenschaften, 5, 1 - 12 (pls. 1 - 3)." type="journal article" year="1859">Müller, 1859</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="E809369CFFBAFFBB55F37AEB2B4DF3FB" blockId="4.[151,497,151,210]" box="[151,339,186,210]" pageId="4" pageNumber="57">
<figureCitation id="708D2A19FFBAFFBB55F37AEB2B1EF3FB" box="[151,256,186,210]" captionStart="FIGURE 2" captionStartId="4.[151,250,1853,1876]" captionTargetBox="[153,1434,409,1807]" captionTargetId="figure@4.[151,1436,407,1809]" captionTargetPageId="4" captionText="FIGURE 2. Comparison among characters of Tamoya sp. (USNM 1114657) and Carybdea rastonii Haacke, 1887 (USNM 88365). A. Frown-shaped rhopaliar niche ostium of T. sp. B. Heart-shaped rhopaliar niche ostium (only a single upper covering scale present) of C. rastonii. C. Stomach of T. sp. with mesenteries and vertically arranged gastric phacellae. D. Gastric phacellae of T. sp.; note that individual gastric cirri are simple and unbranched. E. Whole specimen of T. sp. cut in half longitudinally. F. Longitudinal cut specimen of C. rastonii with epaulette-like gastric phacellae visible in corners of the shallow stomach. Me = mesentery; Ph = gastric phacellae; Sto = stomach. Scale bars: 10 mm (F &amp; E), 5 mm (C &amp; D), 1 mm (A &amp; B)." httpUri="https://zenodo.org/record/276723/files/figure.png" pageId="4" pageNumber="57">Figures 2</figureCitation>
A, C–E
</paragraph>
</subSubSection>
<subSubSection id="A0AC6517FFBAFFBA55F37B532BF2F235" lastPageId="5" lastPageNumber="58" pageId="4" pageNumber="57" type="description">
<paragraph id="E809369CFFBAFFBB55F37B532E95F233" blockId="4.[151,1436,258,355]" box="[151,1163,258,283]" pageId="4" pageNumber="57">
<emphasis id="DAC2EA8EFFBAFFBB55F37B532B31F232" bold="true" box="[151,303,258,283]" pageId="4" pageNumber="57">Valid species</emphasis>
:
<taxonomicName id="2FB64D1FFFBAFFBB545A7B5528AAF233" authority="Muller, 1859" authorityName="Muller" authorityYear="1859" box="[318,692,258,283]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="4" pageNumber="57" phylum="Cnidaria" rank="species" species="haplonema">
<emphasis id="DAC2EA8EFFBAFFBB545A7B552806F232" box="[318,536,259,283]" italics="true" pageId="4" pageNumber="57">Tamoya haplonema</emphasis>
<bibRefCitation id="8C274B6DFFBAFFBB57447B5328AAF233" author="Muller" box="[544,692,258,282]" pageId="4" pageNumber="68" refString="Muller, F. (1859) Zwei neue Quallen von Santa Catharina (Brazilien). Abhandlungen der Naturforschenden Gesellschaft zu Halle - Originalaufsatze aus dem Gebiete der gesamten Naturwissenschaften, 5, 1 - 12 (pls. 1 - 3)." type="journal article" year="1859">Müller, 1859</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="2FB64D1FFFBAFFBB57A47B55296EF232" box="[704,880,259,283]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="4" pageNumber="57" phylum="Cnidaria" rank="species" species="ohboya" status="sp. nov.">
<emphasis id="DAC2EA8EFFBAFFBB57A47B55296EF232" box="[704,880,259,283]" italics="true" pageId="4" pageNumber="57">Tamoya ohboya</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C1F157F5FFBAFFBB561B7B5229A3F232" box="[895,957,259,283]" pageId="4" pageNumber="57" rank="species">
<emphasis id="DAC2EA8EFFBAFFBB561B7B5229A3F232" bold="true" box="[895,957,259,283]" pageId="4" pageNumber="57">n. sp.</emphasis>
</taxonomicNameLabel>
, described herein.
</paragraph>
<paragraph id="E809369CFFBAFFBB55A17B772F08F24B" blockId="4.[151,1436,258,355]" pageId="4" pageNumber="57">
<emphasis id="DAC2EA8EFFBAFFBB55A17B772945F216" bold="true" box="[197,859,294,319]" pageId="4" pageNumber="57">Insufficiently known to be considered valid species</emphasis>
:
<taxonomicName id="2FB64D1FFFBAFFBB56177B792E25F217" box="[883,1083,294,319]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="4" pageNumber="57" phylum="Cnidaria" rank="species" species="bursaria">
<emphasis id="DAC2EA8EFFBAFFBB56177B792E25F217" box="[883,1083,294,319]" italics="true" pageId="4" pageNumber="57">Tamoya bursaria</emphasis>
</taxonomicName>
(Lesson, 1829 as
<emphasis id="DAC2EA8EFFBAFFBB50497B762B1DF24A" italics="true" pageId="4" pageNumber="57">Bursarius cythereae</emphasis>
);
<taxonomicName id="2FB64D1FFFBAFFBB547F7B1D2881F24B" authority="Lesson, 1829" authorityName="Lesson" authorityYear="1829" box="[283,671,330,355]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="4" pageNumber="57" phylum="Cnidaria" rank="species" species="gargantua">
<emphasis id="DAC2EA8EFFBAFFBB547F7B1D2BF3F24A" box="[283,493,332,355]" italics="true" pageId="4" pageNumber="57">Tamoya gargantua</emphasis>
(Lesson, 1829)
</taxonomicName>
;
<taxonomicName id="2FB64D1FFFBAFFBB57CF7B1D2E01F24B" authority="Southcott, 1967" authorityName="Southcott" authorityYear="1967" box="[683,1055,330,355]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="4" pageNumber="57" phylum="Cnidaria" rank="species" species="haeckeli">
<emphasis id="DAC2EA8EFFBAFFBB57CF7B1D2978F24A" box="[683,870,331,355]" italics="true" pageId="4" pageNumber="57">Tamoya haeckeli</emphasis>
Southcott, 1967
</taxonomicName>
=
<taxonomicName id="2FB64D1FFFBAFFBB51587B1D2F11F24A" box="[1084,1295,332,355]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="4" pageNumber="57" phylum="Cnidaria" rank="species" species="gargantua">
<emphasis id="DAC2EA8EFFBAFFBB51587B1D2F11F24A" box="[1084,1295,332,355]" italics="true" pageId="4" pageNumber="57">Tamoya gargantua</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="BCC96614FFBAFFBB55F37D6C2F18F4C4" httpUri="https://zenodo.org/record/276723/files/figure.png" pageId="4" pageNumber="57" targetBox="[153,1434,409,1807]" targetPageId="4">
<paragraph id="E809369CFFBAFFBB55F37D6C2F18F4C4" blockId="4.[151,1437,1853,2029]" pageId="4" pageNumber="57">
<emphasis id="DAC2EA8EFFBAFFBB55F37D6C2B0BF47B" bold="true" box="[151,277,1853,1876]" pageId="4" pageNumber="57">FIGURE 2.</emphasis>
Comparison among characters of
<taxonomicName id="2FB64D1FFFBAFFBB571B7D6F28D2F47A" box="[639,716,1854,1875]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="4" pageNumber="57" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBAFFBB571B7D6F28D2F47A" box="[639,716,1854,1875]" italics="true" pageId="4" pageNumber="57">Tamoya</emphasis>
</taxonomicName>
sp. (USNM 1114657) and
<taxonomicName id="2FB64D1FFFBAFFBB568A7D6C2F5CF47A" authority="Haacke, 1887" authorityName="Haacke" authorityYear="1887" box="[1006,1346,1853,1875]" class="Cubozoa" family="Carybdeidae" genus="Carybdea" kingdom="Animalia" order="Cubomedusae" pageId="4" pageNumber="57" phylum="Cnidaria" rank="species" species="rastonii">
<emphasis id="DAC2EA8EFFBAFFBB568A7D6C2EB4F47A" box="[1006,1194,1853,1875]" italics="true" pageId="4" pageNumber="57">Carybdea rastonii</emphasis>
Haacke, 1887
</taxonomicName>
(USNM 88365). A. Frown-shaped rhopaliar niche ostium of
<emphasis id="DAC2EA8EFFBAFFBB57C17D0C28A9F45B" box="[677,695,1885,1906]" italics="true" pageId="4" pageNumber="57">T.</emphasis>
sp. B. Heart-shaped rhopaliar niche ostium (only a single upper covering scale present) of
<taxonomicName id="2FB64D1FFFBAFFBB54277D2A2BB0F4B8" box="[323,430,1915,1937]" class="Cubozoa" family="Carybdeidae" genus="Carybdea" kingdom="Animalia" order="Cubomedusae" pageId="4" pageNumber="57" phylum="Cnidaria" rank="species" species="rastonii">
<emphasis id="DAC2EA8EFFBAFFBB54277D2A2BB0F4B8" box="[323,430,1915,1937]" italics="true" pageId="4" pageNumber="57">C. rastonii</emphasis>
</taxonomicName>
. C. Stomach of
<emphasis id="DAC2EA8EFFBAFFBB57367D2D287AF4B8" box="[594,612,1916,1937]" italics="true" pageId="4" pageNumber="57">T.</emphasis>
sp. with mesenteries and vertically arranged gastric phacellae. D. Gastric phacellae of
<emphasis id="DAC2EA8EFFBAFFBB55B27DCA2AF6F499" box="[214,232,1947,1968]" italics="true" pageId="4" pageNumber="57">T.</emphasis>
sp.; note that individual gastric cirri are simple and unbranched. E. Whole specimen of
<emphasis id="DAC2EA8EFFBAFFBB51057DCA2E6DF499" box="[1121,1139,1947,1968]" italics="true" pageId="4" pageNumber="57">T.</emphasis>
sp. cut in half longitudinally. F. Longitudinal cut specimen of
<taxonomicName id="2FB64D1FFFBAFFBB54867DE82851F4E7" box="[482,591,1977,1998]" class="Cubozoa" family="Carybdeidae" genus="Carybdea" kingdom="Animalia" order="Cubomedusae" pageId="4" pageNumber="57" phylum="Cnidaria" rank="species" species="rastonii">
<emphasis id="DAC2EA8EFFBAFFBB54867DE82851F4E7" box="[482,591,1977,1998]" italics="true" pageId="4" pageNumber="57">C. rastonii</emphasis>
</taxonomicName>
with epaulette-like gastric phacellae visible in corners of the shallow stomach. Me = mesentery; Ph = gastric phacellae; Sto = stomach. Scale bars: 10 mm (F &amp; E), 5 mm (C &amp; D), 1 mm (A &amp; B).
</paragraph>
</caption>
<paragraph id="E809369CFFBBFFBA55A17AC62BF2F235" blockId="5.[151,1437,151,1400]" pageId="5" pageNumber="58">
<emphasis id="DAC2EA8EFFBBFFBA55A17AC62BDDF399" bold="true" box="[197,451,151,176]" pageId="5" pageNumber="58">Other named species</emphasis>
:
<taxonomicName id="2FB64D1FFFBBFFBA54B17AC82943F386" authority="Haeckel, 1880" authorityName="Haeckel" authorityYear="1880" box="[469,861,151,176]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="species" species="prismatica">
<emphasis id="DAC2EA8EFFBBFFBA54B17AC828B3F399" box="[469,685,153,176]" italics="true" pageId="5" pageNumber="58">Tamoya prismatica</emphasis>
<bibRefCitation id="8C274B6DFFBBFFBA57DC7AC62943F386" author="Haeckel" box="[696,861,151,175]" pageId="5" pageNumber="68" refString="Haeckel E. (1880) System der Acraspeden - Zweite Halfte des Systems der Medusen. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena, Jena, 672 pp." type="book" year="1880">Haeckel, 1880</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="2FB64D1FFFBBFFBA56E47AC82E43F399" box="[896,1117,152,176]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="species" species="haplonema">
<emphasis id="DAC2EA8EFFBBFFBA56E47AC82E43F399" box="[896,1117,152,176]" italics="true" pageId="5" pageNumber="58">Tamoya haplonema</emphasis>
</taxonomicName>
(see
<bibRefCitation id="8C274B6DFFBBFFBA51FA7AC92F33F386" author="Mayer" box="[1182,1325,151,176]" pageId="5" pageNumber="68" refString="Mayer, A. G. (1910) Medusae of the World, Volume III. The Scyphomedusae. Carnegie Institution of Washington Publication 109 (3), 499 - 735." type="journal article" year="1910">Mayer 1910</bibRefCitation>
);
<taxonomicName id="2FB64D1FFFBBFFBA50207AC82BA0F3FD" authority="Muller, 1859" authorityName="Muller" authorityYear="1859" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="species" species="quadrumana">
<emphasis id="DAC2EA8EFFBBFFBA50207AC82B38F3FD" italics="true" pageId="5" pageNumber="58">Tamoya quadrumana</emphasis>
<bibRefCitation id="8C274B6DFFBBFFBA54497AED2BA0F3FD" author="Muller" box="[301,446,188,212]" pageId="5" pageNumber="68" refString="Muller, F. (1859) Zwei neue Quallen von Santa Catharina (Brazilien). Abhandlungen der Naturforschenden Gesellschaft zu Halle - Originalaufsatze aus dem Gebiete der gesamten Naturwissenschaften, 5, 1 - 12 (pls. 1 - 3)." type="journal article" year="1859">Müller, 1859</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="2FB64D1FFFBBFFBA54B87AED2907F3FD" box="[476,793,188,212]" class="Cubozoa" family="Chiropsalmidae" genus="Chiropsalmus" kingdom="Animalia" order="Chirodropida" pageId="5" pageNumber="58" phylum="Cnidaria" rank="species" species="quadrumanus">
<emphasis id="DAC2EA8EFFBBFFBA54B87AED2907F3FD" box="[476,793,188,212]" italics="true" pageId="5" pageNumber="58">Chiropsalmus quadrumanus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="8C274B6DFFBBFFBA56307AED29F1F3FD" author="Agassiz" box="[852,1007,188,212]" pageId="5" pageNumber="67" refString="Agassiz, L. (1862) C ontributions to the natural history of the United States of America - Second monograph 4. Little, Brown &amp; Co., Boston, 380 pp." type="book" year="1862">Agassiz 1862</bibRefCitation>
);
<taxonomicName id="2FB64D1FFFBBFFBA51677AEC2F89F3FD" authority="Kishinouye, 1910" authorityName="Kishinouye" authorityYear="1910" box="[1027,1431,188,212]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="species" species="virulenta">
<emphasis id="DAC2EA8EFFBBFFBA51677AEC2EDBF3FD" box="[1027,1221,188,212]" italics="true" pageId="5" pageNumber="58">Tamoya virulenta</emphasis>
Kishinouye, 1910
</taxonomicName>
, known in
<collectingCountry id="90A1760CFFBBFFBA546C7AB12B56F3DE" box="[264,328,224,247]" name="Japan" pageId="5" pageNumber="58">Japan</collectingCountry>
as “Hikurage”, is considered valid, but is a species of
<taxonomicName id="2FB64D1FFFBBFFBA56D77AB12E38F3D1" box="[947,1062,224,248]" class="Cubozoa" family="Tamoyidae" genus="Morbakka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA56D77AB12E38F3D1" box="[947,1062,224,248]" italics="true" pageId="5" pageNumber="58">Morbakka</emphasis>
</taxonomicName>
in the newly erected family
<taxonomicName id="2FB64D1FFFBBFFBA500F7A8E2BF6F235" authority="Bentlage et al. 2010" authorityName="Bentlage et al." authorityYear="2010" class="Cubozoa" family="Carukiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carybdeida" pageId="5" pageNumber="58" phylum="Cnidaria" rank="family">
Carukiidae (
<bibRefCitation id="8C274B6DFFBBFFBA559C7B552BFEF235" author="Bentlage" box="[248,480,260,284]" pageId="5" pageNumber="67" refString="Bentlage, B., Cartwright, P., Yanagihara, A. A., Lewis, C., Richards, G. S. &amp; Collins A. G. (2010) Evolution of box jellyfish (Cnidaria: Cubozoa), a group of highly toxic invertebrates. Proceedings of the Royal Society B, 277, 493 - 501." type="journal article" year="2010">
Bentlage
<emphasis id="DAC2EA8EFFBBFFBA54007B542B8BF235" box="[356,405,260,284]" italics="true" pageId="5" pageNumber="58">et al</emphasis>
. 2010
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A0AC6517FFBBFFBA55A17B76298FF24D" pageId="5" pageNumber="58" type="diagnosis">
<paragraph id="E809369CFFBBFFBA55A17B76298FF24D" blockId="5.[151,1437,151,1400]" pageId="5" pageNumber="58">
<emphasis id="DAC2EA8EFFBBFFBA55A17B762B26F269" bold="true" box="[197,312,295,320]" pageId="5" pageNumber="58">Diagnosis</emphasis>
(following
<bibRefCitation id="8C274B6DFFBBFFBA54A17B762877F216" author="Haeckel" box="[453,617,295,319]" pageId="5" pageNumber="68" refString="Haeckel E. (1880) System der Acraspeden - Zweite Halfte des Systems der Medusen. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena, Jena, 672 pp." type="book" year="1880">Haeckel 1880</bibRefCitation>
):
<taxonomicName id="2FB64D1FFFBBFFBA57E57B7628E3F216" box="[641,765,295,319]" class="Cubozoa" family="Tamoyidae" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="family">Tamoyidae</taxonomicName>
with deep stomach connected to the subumbrella with 4 prominent perradial mesenteries. Vertical phacellae in the interradii.
</paragraph>
</subSubSection>
<subSubSection id="A0AC6517FFBBFFBA55A37B212EECF2AE" box="[199,1266,367,392]" pageId="5" pageNumber="58" type="materials_examined">
<paragraph id="E809369CFFBBFFBA55A37B212EECF2AE" blockId="5.[151,1437,151,1400]" box="[199,1266,367,392]" pageId="5" pageNumber="58">
<emphasis id="DAC2EA8EFFBBFFBA55A37B212B7AF2A1" bold="true" box="[199,356,367,392]" pageId="5" pageNumber="58">
<typeStatus id="370D883EFFBBFFBA55A37B212B1CF2A1" box="[199,258,368,392]" pageId="5" pageNumber="58">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="2FB64D1FFFBBFFBA540F7B2028FEF2AE" authority="Muller, 1859" authorityName="Muller" authorityYear="1859" box="[363,736,367,392]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="species" species="haplonema">
<emphasis id="DAC2EA8EFFBBFFBA540F7B202859F2A1" box="[363,583,368,392]" italics="true" pageId="5" pageNumber="58">Tamoya haplonema</emphasis>
<bibRefCitation id="8C274B6DFFBBFFBA57297B3E28FEF2AE" author="Muller" box="[589,736,367,391]" pageId="5" pageNumber="68" refString="Muller, F. (1859) Zwei neue Quallen von Santa Catharina (Brazilien). Abhandlungen der Naturforschenden Gesellschaft zu Halle - Originalaufsatze aus dem Gebiete der gesamten Naturwissenschaften, 5, 1 - 12 (pls. 1 - 3)." type="journal article" year="1859">Müller, 1859</bibRefCitation>
</taxonomicName>
, by subsequent designation by
<bibRefCitation id="8C274B6DFFBBFFBA51597B3E2EF0F2AE" author="Haeckel" box="[1085,1262,367,391]" pageId="5" pageNumber="68" refString="Haeckel E. (1880) System der Acraspeden - Zweite Halfte des Systems der Medusen. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena, Jena, 672 pp." type="book" year="1880">Haeckel (1880)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A0AC6517FFBBFFBA55A17BC52EC3F65E" pageId="5" pageNumber="58" type="discussion">
<paragraph id="E809369CFFBBFFBA55A17BC529AFF0A9" blockId="5.[151,1437,151,1400]" pageId="5" pageNumber="58">
<emphasis id="DAC2EA8EFFBBFFBA55A17BC52B25F285" bold="true" box="[197,315,404,428]" pageId="5" pageNumber="58">Remarks:</emphasis>
<bibRefCitation id="8C274B6DFFBBFFBA54257BC52BC0F285" author="Muller" box="[321,478,404,428]" pageId="5" pageNumber="68" refString="Muller, F. (1859) Zwei neue Quallen von Santa Catharina (Brazilien). Abhandlungen der Naturforschenden Gesellschaft zu Halle - Originalaufsatze aus dem Gebiete der gesamten Naturwissenschaften, 5, 1 - 12 (pls. 1 - 3)." type="journal article" year="1859">Müller (1859)</bibRefCitation>
erected the genus
<taxonomicName id="2FB64D1FFFBBFFBA57C97BC4291CF285" box="[685,770,405,428]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA57C97BC4291CF285" box="[685,770,405,428]" italics="true" pageId="5" pageNumber="58">Tamoya</emphasis>
</taxonomicName>
for two cubozoan species from Santa Catarina state (southern
<collectingCountry id="90A1760CFFBBFFBA55A67BE62B15F2E6" box="[194,267,439,463]" name="Brazil" pageId="5" pageNumber="58">Brazil</collectingCountry>
),
<taxonomicName id="2FB64D1FFFBBFFBA547A7BE82BE6F2F9" box="[286,504,440,464]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="species" species="haplonema">
<emphasis id="DAC2EA8EFFBBFFBA547A7BE82BE6F2F9" box="[286,504,440,464]" italics="true" pageId="5" pageNumber="58">Tamoya haplonema</emphasis>
</taxonomicName>
and
<taxonomicName id="2FB64D1FFFBBFFBA57547BE82900F2F9" box="[560,798,440,464]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="species" species="quadrumana">
<emphasis id="DAC2EA8EFFBBFFBA57547BE82900F2F9" box="[560,798,440,464]" italics="true" pageId="5" pageNumber="58">Tamoya quadrumana</emphasis>
</taxonomicName>
(=
<taxonomicName id="2FB64D1FFFBBFFBA56227BE92E9AF2F9" box="[838,1156,440,464]" class="Cubozoa" family="Chiropsalmidae" genus="Chiropsalmus" kingdom="Animalia" order="Chirodropida" pageId="5" pageNumber="58" phylum="Cnidaria" rank="species" species="quadrumanus">
<emphasis id="DAC2EA8EFFBBFFBA56227BE92E9AF2F9" box="[838,1156,440,464]" italics="true" pageId="5" pageNumber="58">Chiropsalmus quadrumanus</emphasis>
</taxonomicName>
), which today we recognize as representatives of separate orders. At the time, however, Müller believed that the two were so similar that he could provide just a single description, pointing out the specific differences throughout his manuscript. He distinguished
<taxonomicName id="2FB64D1FFFBBFFBA547078742B77F115" box="[276,361,549,572]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA547078742B77F115" box="[276,361,549,572]" italics="true" pageId="5" pageNumber="58">Tamoya</emphasis>
</taxonomicName>
from
<taxonomicName id="2FB64D1FFFBBFFBA54D778752928F115" authority="Peron &amp; Lesueur, 1810" authorityName="Peron &amp; Lesueur" authorityYear="1810" box="[435,822,548,572]" class="Cubozoa" family="Carybdeidae" genus="Carybdea" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA54D77875283FF115" box="[435,545,548,572]" italics="true" pageId="5" pageNumber="58">Carybdea</emphasis>
Péron &amp; Lesueur, 1810
</taxonomicName>
, the only other cubozoan genus described at the time, based on the morphology of the bell margin, stomach extensions, origination of the tentacle bases, presence or absence of a sphincter, whether the gastric cirri pointed towards the stomach or the bell, and whether or not the cirri were hollow. These characters are somewhat nebulous and have been widely disregarded by subsequent workers (e.g.,
<bibRefCitation id="8C274B6DFFBBFFBA55BE78E52B7EF1E5" author="Claus" box="[218,352,692,716]" pageId="5" pageNumber="67" refString="Claus, C. (1878) Uber Charybdea marsupialis. Arbeiten aus dem Zoologischen Institute der Universitat Wien und der Zoologischen Station in Triest, 2, 1 - 56 (pls. 1 - 5)." type="journal article" year="1878">Claus 1878</bibRefCitation>
,
<bibRefCitation id="8C274B6DFFBBFFBA540A78E52811F1E5" author="Haeckel" box="[366,527,692,716]" pageId="5" pageNumber="68" refString="Haeckel E. (1880) System der Acraspeden - Zweite Halfte des Systems der Medusen. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena, Jena, 672 pp." type="book" year="1880">Haeckel 1880</bibRefCitation>
,
<bibRefCitation id="8C274B6DFFBBFFBA577A78E428BEF1E5" author="Maas" box="[542,672,692,716]" pageId="5" pageNumber="68" refString="Maas, O. (1903) Die Scyphomedusen der Siboga-Expedition. Siboga-Expeditie XI, Boekhandel en Drukkerij voorheen E. J. Brill, Leiden, 91 pp." type="book" year="1903">Maas 1903</bibRefCitation>
,
<bibRefCitation id="8C274B6DFFBBFFBA57CB78E42923F1E5" author="Mayer" box="[687,829,692,717]" pageId="5" pageNumber="68" refString="Mayer, A. G. (1910) Medusae of the World, Volume III. The Scyphomedusae. Carnegie Institution of Washington Publication 109 (3), 499 - 735." type="journal article" year="1910">Mayer 1910</bibRefCitation>
,
<bibRefCitation id="8C274B6DFFBBFFBA562878E429C0F1E5" author="Kramp" box="[844,990,692,716]" pageId="5" pageNumber="68" refString="Kramp, P. L. (1961) Synopsis of the medusae of the world. Journal of the Marine Biological Association of the United Kingdom, 40, 1 - 469." type="journal article" year="1961">Kramp 1961</bibRefCitation>
,
<bibRefCitation id="8C274B6DFFBBFFBA568A78E52F58F1E5" author="Gershwin" box="[1006,1350,692,716]" pageId="5" pageNumber="68" refString="Gershwin, L. &amp; Alderslade, P. (2005) A new genus and species of box jellyfish (Cubozoa: Carybdeidae) from tropical Australian waters. The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 21, 27 - 36." type="journal article" year="2005">Gershwin &amp; Alderslade 2005</bibRefCitation>
).
<bibRefCitation id="8C274B6DFFBBFFBA503878E52AFFF1C6" author="Claus" pageId="5" pageNumber="67" refString="Claus, C. (1878) Uber Charybdea marsupialis. Arbeiten aus dem Zoologischen Institute der Universitat Wien und der Zoologischen Station in Triest, 2, 1 - 56 (pls. 1 - 5)." type="journal article" year="1878">Claus (1878)</bibRefCitation>
discussed the generic diagnosis presented by
<bibRefCitation id="8C274B6DFFBBFFBA579978862980F1C6" author="Muller" box="[765,926,727,751]" pageId="5" pageNumber="68" refString="Muller, F. (1859) Zwei neue Quallen von Santa Catharina (Brazilien). Abhandlungen der Naturforschenden Gesellschaft zu Halle - Originalaufsatze aus dem Gebiete der gesamten Naturwissenschaften, 5, 1 - 12 (pls. 1 - 3)." type="journal article" year="1859">Müller (1859)</bibRefCitation>
in detail and considered that the characters invoked either do not differ substantially from those of
<taxonomicName id="2FB64D1FFFBBFFBA567778AD299FF03D" box="[787,897,764,788]" class="Cubozoa" family="Carybdeidae" genus="Carybdea" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA567778AD299FF03D" box="[787,897,764,788]" italics="true" pageId="5" pageNumber="58">Carybdea</emphasis>
</taxonomicName>
or are uninterpretable, and thus concluded that the designation of a new genus for the specimens from
<collectingCountry id="90A1760CFFBBFFBA5674794E294AF01E" box="[784,852,799,823]" name="Brazil" pageId="5" pageNumber="58">Brazil</collectingCountry>
was not warranted. However,
<bibRefCitation id="8C274B6DFFBBFFBA51D5794E2F5DF01E" author="Claus" box="[1201,1347,799,823]" pageId="5" pageNumber="67" refString="Claus, C. (1878) Uber Charybdea marsupialis. Arbeiten aus dem Zoologischen Institute der Universitat Wien und der Zoologischen Station in Triest, 2, 1 - 56 (pls. 1 - 5)." type="journal article" year="1878">Claus (1878)</bibRefCitation>
did not actually inspect the specimens studied by
<bibRefCitation id="8C274B6DFFBBFFBA570879152912F075" author="Muller" box="[620,780,836,860]" pageId="5" pageNumber="68" refString="Muller, F. (1859) Zwei neue Quallen von Santa Catharina (Brazilien). Abhandlungen der Naturforschenden Gesellschaft zu Halle - Originalaufsatze aus dem Gebiete der gesamten Naturwissenschaften, 5, 1 - 12 (pls. 1 - 3)." type="journal article" year="1859">Müller (1859)</bibRefCitation>
, but simply relied on Müller’s description, which arguably contains several shortcomings regarding erection of the genus
<taxonomicName id="2FB64D1FFFBBFFBA5637793829B6F0A9" box="[851,936,873,896]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA5637793829B6F0A9" box="[851,936,873,896]" italics="true" pageId="5" pageNumber="58">Tamoya</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="E809369CFFBBFFBA55A179DD2EC3F65E" blockId="5.[151,1437,151,1400]" pageId="5" pageNumber="58">
Clearly,
<taxonomicName id="2FB64D1FFFBBFFBA544279DC2808F08D" box="[294,534,908,932]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="species" species="quadrumana">
<emphasis id="DAC2EA8EFFBBFFBA544279DC2808F08D" box="[294,534,908,932]" italics="true" pageId="5" pageNumber="58">Tamoya quadrumana</emphasis>
</taxonomicName>
is a chirodropid (a clade of cubozoans not recognized at the time of Müller’s description) and so distinct from any carybdeid that it was soon moved into its own genus,
<taxonomicName id="2FB64D1FFFBBFFBA51E879E12F36F0E1" box="[1164,1320,944,968]" class="Cubozoa" family="Chiropsalmidae" genus="Chiropsalmus" kingdom="Animalia" order="Chirodropida" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA51E879E12F36F0E1" box="[1164,1320,944,968]" italics="true" pageId="5" pageNumber="58">Chiropsalmus</emphasis>
</taxonomicName>
, by
<bibRefCitation id="8C274B6DFFBBFFBA503D79E128B5F0C5" author="Agassiz" pageId="5" pageNumber="67" refString="Agassiz, L. (1862) C ontributions to the natural history of the United States of America - Second monograph 4. Little, Brown &amp; Co., Boston, 380 pp." type="book" year="1862">
Agassiz (1862: 174; as
<taxonomicName id="2FB64D1FFFBBFFBA540779852881F0C5" box="[355,671,980,1004]" class="Cubozoa" family="Chiropsalmidae" genus="Chiropsalmus" kingdom="Animalia" order="Chirodropida" pageId="5" pageNumber="58" phylum="Cnidaria" rank="species" species="quadrumanus">
<emphasis id="DAC2EA8EFFBBFFBA540779852881F0C5" box="[355,671,980,1004]" italics="true" pageId="5" pageNumber="58">Chiropsalmus quadrumanus</emphasis>
</taxonomicName>
)
</bibRefCitation>
. Subsequently,
<bibRefCitation id="8C274B6DFFBBFFBA563D79852E69F0C5" author="Haeckel" box="[857,1143,980,1004]" pageId="5" pageNumber="68" refString="Haeckel E. (1880) System der Acraspeden - Zweite Halfte des Systems der Medusen. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena, Jena, 672 pp." type="book" year="1880">Haeckel (1880: 445–446)</bibRefCitation>
established the order
<taxonomicName id="2FB64D1FFFBBFFBA500E79852B1AF726" class="Cubozoa" kingdom="Animalia" order="Chirodropida" pageId="5" pageNumber="58" phylum="Cnidaria" rank="order">Chirodropida</taxonomicName>
(family
<taxonomicName id="2FB64D1FFFBBFFBA540A79A6280EF726" box="[366,528,1015,1039]" class="Cubozoa" family="Chirodropidae" kingdom="Animalia" order="Chirodropida" pageId="5" pageNumber="58" phylum="Cnidaria" rank="family">Chirodropidae</taxonomicName>
at the time) to accommodate
<taxonomicName id="2FB64D1FFFBBFFBA561C79A92E0AF739" box="[888,1044,1016,1040]" class="Cubozoa" family="Chiropsalmidae" genus="Chiropsalmus" kingdom="Animalia" order="Chirodropida" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA561C79A92E0AF739" box="[888,1044,1016,1040]" italics="true" pageId="5" pageNumber="58">Chiropsalmus</emphasis>
</taxonomicName>
and his new genus
<taxonomicName id="2FB64D1FFFBBFFBA506C79A92F8BF739" box="[1288,1429,1016,1040]" class="Cubozoa" family="Chirodropidae" genus="Chirodropus" kingdom="Animalia" order="Chirodropida" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA506C79A92F8BF739" box="[1288,1429,1016,1040]" italics="true" pageId="5" pageNumber="58">Chirodropus</emphasis>
</taxonomicName>
.
<bibRefCitation id="8C274B6DFFBBFFBA55F37E4D2BA6F71D" author="Haeckel" box="[151,440,1052,1076]" pageId="5" pageNumber="68" refString="Haeckel E. (1880) System der Acraspeden - Zweite Halfte des Systems der Medusen. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena, Jena, 672 pp." type="book" year="1880">Haeckel (1880: 442–443)</bibRefCitation>
, in disagreement with
<bibRefCitation id="8C274B6DFFBBFFBA57D07E4D2959F71D" author="Claus" box="[692,839,1052,1076]" pageId="5" pageNumber="67" refString="Claus, C. (1878) Uber Charybdea marsupialis. Arbeiten aus dem Zoologischen Institute der Universitat Wien und der Zoologischen Station in Triest, 2, 1 - 56 (pls. 1 - 5)." type="journal article" year="1878">Claus (1878)</bibRefCitation>
, also modified the diagnosis of
<taxonomicName id="2FB64D1FFFBBFFBA51CF7E4C2F1EF71D" box="[1195,1280,1053,1076]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA51CF7E4C2F1EF71D" box="[1195,1280,1053,1076]" italics="true" pageId="5" pageNumber="58">Tamoya</emphasis>
</taxonomicName>
to contain all those species that possess a stomach that protrudes deep into the subumbrellar cavity and is connected to the subumbrella by mesenterial bands (
<figureCitation id="708D2A19FFBBFFBA57677E352859F755" box="[515,583,1124,1148]" captionStart="FIGURE 2" captionStartId="4.[151,250,1853,1876]" captionTargetBox="[153,1434,409,1807]" captionTargetId="figure@4.[151,1436,407,1809]" captionTargetPageId="4" captionText="FIGURE 2. Comparison among characters of Tamoya sp. (USNM 1114657) and Carybdea rastonii Haacke, 1887 (USNM 88365). A. Frown-shaped rhopaliar niche ostium of T. sp. B. Heart-shaped rhopaliar niche ostium (only a single upper covering scale present) of C. rastonii. C. Stomach of T. sp. with mesenteries and vertically arranged gastric phacellae. D. Gastric phacellae of T. sp.; note that individual gastric cirri are simple and unbranched. E. Whole specimen of T. sp. cut in half longitudinally. F. Longitudinal cut specimen of C. rastonii with epaulette-like gastric phacellae visible in corners of the shallow stomach. Me = mesentery; Ph = gastric phacellae; Sto = stomach. Scale bars: 10 mm (F &amp; E), 5 mm (C &amp; D), 1 mm (A &amp; B)." httpUri="https://zenodo.org/record/276723/files/figure.png" pageId="5" pageNumber="58">Fig. 2</figureCitation>
C &amp; E). Furthermore, the gastric cirri of
<taxonomicName id="2FB64D1FFFBBFFBA517F7E342F6FF755" authority="Haeckel (1880)" authorityName="Haeckel" authorityYear="1880" box="[1051,1393,1124,1148]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA517F7E342E6EF755" box="[1051,1136,1125,1148]" italics="true" pageId="5" pageNumber="58">Tamoya</emphasis>
sensu
<bibRefCitation id="8C274B6DFFBBFFBA51A67E352F6FF755" author="Haeckel" box="[1218,1393,1124,1148]" pageId="5" pageNumber="68" refString="Haeckel E. (1880) System der Acraspeden - Zweite Halfte des Systems der Medusen. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena, Jena, 672 pp." type="book" year="1880">Haeckel (1880)</bibRefCitation>
</taxonomicName>
are arranged in four interradial bands running along the stomach walls from the upper portion of the stomach down towards the manubrium (
<figureCitation id="708D2A19FFBBFFBA54D97EFD281BF7ED" box="[445,517,1196,1220]" captionStart="FIGURE 2" captionStartId="4.[151,250,1853,1876]" captionTargetBox="[153,1434,409,1807]" captionTargetId="figure@4.[151,1436,407,1809]" captionTargetPageId="4" captionText="FIGURE 2. Comparison among characters of Tamoya sp. (USNM 1114657) and Carybdea rastonii Haacke, 1887 (USNM 88365). A. Frown-shaped rhopaliar niche ostium of T. sp. B. Heart-shaped rhopaliar niche ostium (only a single upper covering scale present) of C. rastonii. C. Stomach of T. sp. with mesenteries and vertically arranged gastric phacellae. D. Gastric phacellae of T. sp.; note that individual gastric cirri are simple and unbranched. E. Whole specimen of T. sp. cut in half longitudinally. F. Longitudinal cut specimen of C. rastonii with epaulette-like gastric phacellae visible in corners of the shallow stomach. Me = mesentery; Ph = gastric phacellae; Sto = stomach. Scale bars: 10 mm (F &amp; E), 5 mm (C &amp; D), 1 mm (A &amp; B)." httpUri="https://zenodo.org/record/276723/files/figure.png" pageId="5" pageNumber="58">Fig. 2</figureCitation>
C, D &amp; E). As
<bibRefCitation id="8C274B6DFFBBFFBA57A57EFD296DF7ED" author="Haeckel" box="[705,883,1196,1220]" pageId="5" pageNumber="68" refString="Haeckel E. (1880) System der Acraspeden - Zweite Halfte des Systems der Medusen. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena, Jena, 672 pp." type="book" year="1880">Haeckel (1880)</bibRefCitation>
pointed out, the stomach of
<taxonomicName id="2FB64D1FFFBBFFBA51A87EFC2F3FF7ED" box="[1228,1313,1197,1220]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA51A87EFC2F3FF7ED" box="[1228,1313,1197,1220]" italics="true" pageId="5" pageNumber="58">Tamoya</emphasis>
</taxonomicName>
differs so strongly from the stomach of
<taxonomicName id="2FB64D1FFFBBFFBA54887E812845F7C1" box="[492,603,1232,1256]" class="Cubozoa" family="Carybdeidae" genus="Carybdea" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA54887E812845F7C1" box="[492,603,1232,1256]" italics="true" pageId="5" pageNumber="58">Carybdea</emphasis>
</taxonomicName>
that taxonomic distinction is warranted. For comparison, the stomach of
<taxonomicName id="2FB64D1FFFBBFFBA55F37EA52B18F625" box="[151,262,1268,1292]" class="Cubozoa" family="Carybdeidae" genus="Carybdea" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA55F37EA52B18F625" box="[151,262,1268,1292]" italics="true" pageId="5" pageNumber="58">Carybdea</emphasis>
</taxonomicName>
is shallow, the gastric phacellae cluster in the corners of the stomach, and mesenteries are absent (
<figureCitation id="708D2A19FFBBFFBA500A7EA52AB8F606" captionStart="FIGURE 2" captionStartId="4.[151,250,1853,1876]" captionTargetBox="[153,1434,409,1807]" captionTargetId="figure@4.[151,1436,407,1809]" captionTargetPageId="4" captionText="FIGURE 2. Comparison among characters of Tamoya sp. (USNM 1114657) and Carybdea rastonii Haacke, 1887 (USNM 88365). A. Frown-shaped rhopaliar niche ostium of T. sp. B. Heart-shaped rhopaliar niche ostium (only a single upper covering scale present) of C. rastonii. C. Stomach of T. sp. with mesenteries and vertically arranged gastric phacellae. D. Gastric phacellae of T. sp.; note that individual gastric cirri are simple and unbranched. E. Whole specimen of T. sp. cut in half longitudinally. F. Longitudinal cut specimen of C. rastonii with epaulette-like gastric phacellae visible in corners of the shallow stomach. Me = mesentery; Ph = gastric phacellae; Sto = stomach. Scale bars: 10 mm (F &amp; E), 5 mm (C &amp; D), 1 mm (A &amp; B)." httpUri="https://zenodo.org/record/276723/files/figure.png" pageId="5" pageNumber="58">Fig. 2</figureCitation>
F).
<bibRefCitation id="8C274B6DFFBBFFBA55A97F482BBDF606" author="Mayer" box="[205,419,1303,1329]" pageId="5" pageNumber="68" refString="Mayer, A. G. (1910) Medusae of the World, Volume III. The Scyphomedusae. Carnegie Institution of Washington Publication 109 (3), 499 - 735." type="journal article" year="1910">Mayer (1910: 512)</bibRefCitation>
considered both
<taxonomicName id="2FB64D1FFFBBFFBA57037F4828A2F619" box="[615,700,1305,1328]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA57037F4828A2F619" box="[615,700,1305,1328]" italics="true" pageId="5" pageNumber="58">Tamoya</emphasis>
</taxonomicName>
and
<taxonomicName id="2FB64D1FFFBBFFBA579C7F492978F619" box="[760,870,1304,1328]" class="Cubozoa" family="Carybdeidae" genus="Carybdea" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA579C7F492978F619" box="[760,870,1304,1328]" italics="true" pageId="5" pageNumber="58">Carybdea</emphasis>
</taxonomicName>
to be valid, though he posited that the distinction between the two may be based on intergrading characters. Following
<bibRefCitation id="8C274B6DFFBBFFBA56F97F6D2E53F67D" author="Haeckel" box="[925,1101,1340,1364]" pageId="5" pageNumber="68" refString="Haeckel E. (1880) System der Acraspeden - Zweite Halfte des Systems der Medusen. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena, Jena, 672 pp." type="book" year="1880">Haeckel (1880)</bibRefCitation>
, the genus
<taxonomicName id="2FB64D1FFFBBFFBA51AD7F6C2F00F67D" box="[1225,1310,1341,1364]" class="Cubozoa" family="Tamoyidae" genus="Tamoya" kingdom="Animalia" order="Cubomedusae" pageId="5" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="DAC2EA8EFFBBFFBA51AD7F6C2F00F67D" box="[1225,1310,1341,1364]" italics="true" pageId="5" pageNumber="58">Tamoya</emphasis>
</taxonomicName>
persists, is in widespread use, and is consistent with molecular phylogenetic analyses (
<bibRefCitation id="8C274B6DFFBBFFBA56867F0E2ECCF65E" author="Bentlage" box="[994,1234,1375,1400]" pageId="5" pageNumber="67" refString="Bentlage, B., Cartwright, P., Yanagihara, A. A., Lewis, C., Richards, G. S. &amp; Collins A. G. (2010) Evolution of box jellyfish (Cnidaria: Cubozoa), a group of highly toxic invertebrates. Proceedings of the Royal Society B, 277, 493 - 501." type="journal article" year="2010">
Bentlage
<emphasis id="DAC2EA8EFFBBFFBA512A7F302E9EF651" box="[1102,1152,1376,1400]" italics="true" pageId="5" pageNumber="58">et al</emphasis>
., 2010
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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