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<document id="AE406A246A303DCB64B2B79872F59954" ID-CLB-Dataset="2500" ID-DOI="10.11646/zootaxa.5092.1.2" ID-GBIF-Dataset="5d398a06-dda1-412f-ac7a-855f7ef9a66a" ID-ISSN="1175-5326" ID-Zenodo-Dep="5869488" ID-ZooBank="A546CCCB-6072-434B-B366-1AFB1BE20CD8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1642495732090" checkinUser="plazi" docAuthor="Gill, Anthony C." docDate="2022" docId="1B7A87C4FFBDFFDAB7AB1ADEFE8CFC46" docLanguage="en" docName="zootaxa.5092.1.2.pdf" docOrigin="Zootaxa 5092 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pyronotanthias Gill 2022, new genus" docType="treatment" docVersion="7" lastPageNumber="58" masterDocId="E743FFBCFFB2FFCBB73C186CFFFEFFE3" masterDocTitle="Revised definitions of the anthiadine fish genera Mirolabrichthys Herre and Nemanthias Smith, with description of a new genus (Teleostei: Serranidae)" masterLastPageNumber="66" masterPageNumber="41" pageNumber="56" updateTime="1699302727720" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9BEC3CB81558EAA3AB0E6558702C046D">Revised definitions of the anthiadine fish genera Mirolabrichthys Herre and Nemanthias Smith, with description of a new genus (Teleostei: Serranidae)</mods:title>
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<mods:namePart id="F7183F8779E701F03929CA4CB93BD7DA">Gill, Anthony C.</mods:namePart>
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<mods:affiliation id="AACD5AFCE6153F919C1C7F038D9B4DC6">Macleay Collections, Chau Chak Wing Museum, University Avenue, The University of Sydney, NSW 2006, Australia. Anthony. c. gill @ sydney. edu. au; https: // orcid. org / 0000 - 0002 - 8990 - 3657 &amp; School of Life and Environmental Sciences, The University of Sydney, NSW 2006, Australia. &amp; Ichthyology, Australian Museum Research Institute, 1 William Street, Sydney, New South Wales 2010, Australia.</mods:affiliation>
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<mods:date id="ADF5DE4962B73841B7DFD77741CFC438">2022</mods:date>
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<treatment id="1B7A87C4FFBDFFDAB7AB1ADEFE8CFC46" ID-DOI="http://doi.org/10.5281/zenodo.5883169" ID-GBIF-Taxon="192443538" ID-Zenodo-Dep="5883169" LSID="urn:lsid:plazi:treatment:1B7A87C4FFBDFFDAB7AB1ADEFE8CFC46" httpUri="http://treatment.plazi.org/id/1B7A87C4FFBDFFDAB7AB1ADEFE8CFC46" lastPageId="17" lastPageNumber="58" pageId="15" pageNumber="56">
<subSubSection id="DBC96559FFBDFFC4B7AB1ADEFE24FD2E" box="[151,474,690,717]" pageId="15" pageNumber="56" type="nomenclature">
<paragraph id="936C36D2FFBDFFC4B7AB1ADEFE24FD2E" blockId="15.[151,474,690,717]" box="[151,474,690,717]" pageId="15" pageNumber="56">
<heading id="C82481BEFFBDFFC4B7AB1ADEFE24FD2E" bold="true" box="[151,474,690,717]" fontSize="11" level="1" pageId="15" pageNumber="56" reason="1">
<emphasis id="A1A7EAC0FFBDFFC4B7AB1ADEFE24FD2E" bold="true" box="[151,474,690,717]" pageId="15" pageNumber="56">
<taxonomicName id="54D34D51FFBDFFC4B7AB1ADEFEAFFD2E" authority="Gill, 2022" authorityName="Gill" authorityYear="2022" box="[151,337,690,717]" class="Actinopterygii" family="Serranidae" genus="Pyronotanthias" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="56" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="A1A7EAC0FFBDFFC4B7AB1ADEFEAFFD2E" bold="true" box="[151,337,690,717]" italics="true" pageId="15" pageNumber="56">Pyronotanthias</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BA9457BBFFBDFFC4B6641ADFFE24FD2E" box="[344,474,691,717]" pageId="15" pageNumber="56" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DBC96559FFBDFFC4B7AB1A9BFB29FCEC" box="[151,1239,759,784]" pageId="15" pageNumber="56" type="type_taxon">
<paragraph id="936C36D2FFBDFFC4B7AB1A9BFB29FCEC" blockId="15.[151,1239,759,784]" box="[151,1239,759,784]" pageId="15" pageNumber="56">
<taxonomicName id="54D34D51FFBDFFC4B7AB1A9BFECCFCEC" authority="Gill, 2022" authorityName="Gill" authorityYear="2022" box="[151,306,759,783]" class="Actinopterygii" family="Serranidae" genus="Pyronotanthias" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="56" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="A1A7EAC0FFBDFFC4B7AB1A9BFECCFCEC" box="[151,306,759,783]" italics="true" pageId="15" pageNumber="56">Pyronotanthias</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BA9457BBFFBDFFC4B6051A94FE59FCF3" box="[313,423,760,784]" pageId="15" pageNumber="56" rank="genus">new genus</taxonomicNameLabel>
,
<typeStatus id="4C688870FFBDFFC4B68D1A94FE23FCF3" box="[433,477,760,784]" pageId="15" pageNumber="56">type</typeStatus>
species
<taxonomicName id="54D34D51FFBDFFC4B50F1A9BFC84FCEC" ID-CoL="5VD7Q" authority="Randall &amp; Lubbock" authorityName="Randall &amp; Lubbock" box="[563,890,759,783]" class="Actinopterygii" family="Serranidae" genus="Anthias" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="56" phylum="Chordata" rank="species" species="lori">
<emphasis id="A1A7EAC0FFBDFFC4B50F1A9BFD55FCEC" box="[563,683,759,783]" italics="true" pageId="15" pageNumber="56">Anthias lori</emphasis>
Randall &amp; Lubbock
</taxonomicName>
in
<bibRefCitation id="F7424B23FFBDFFC4B4A71A9BFB2DFCEC" author="Fourmanoir, P. &amp; Laboute, P." box="[923,1235,759,783]" pageId="15" pageNumber="56" refId="ref14020" refString="Fourmanoir, P. &amp; Laboute, P. (1976) Poissons des mers tropicales. Nouvelle Caledonie, Nouvelles Hebrides. Les editions du Pacifique, Papeete, 376 pp." type="book" year="1976">Fourmanoir &amp; Laboute (1976)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DBC96559FFBDFFC4B7AB1B51FBA0FB00" pageId="15" pageNumber="56" type="diagnosis">
<paragraph id="936C36D2FFBDFFC4B7AB1B51FC17FCBB" blockId="15.[151,1001,829,856]" box="[151,1001,829,856]" pageId="15" pageNumber="56">
<emphasis id="A1A7EAC0FFBDFFC4B7AB1B51FEF4FCB4" bold="true" box="[151,266,829,855]" pageId="15" pageNumber="56">Diagnosis</emphasis>
. The following synapomorphy supports monophyly of the genus:
</paragraph>
<paragraph id="936C36D2FFBDFFC4B7AB1BE9FC87FBB0" blockId="15.[151,1436,901,1108]" pageId="15" pageNumber="56">
<emphasis id="A1A7EAC0FFBDFFC4B7AB1BE9FAE5FC7C" box="[151,1307,901,927]" italics="true" pageId="15" pageNumber="56">1. Red stripe or series of blotches below posterior part of dorsal fin and along upper caudal peduncle</emphasis>
. Species of
<taxonomicName id="54D34D51FFBDFFC4B7FA1BC5FE8CFC20" authority="Gill, 2022" authorityName="Gill" authorityYear="2022" box="[198,370,937,963]" class="Actinopterygii" family="Serranidae" genus="Pyronotanthias" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="56" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="A1A7EAC0FFBDFFC4B7FA1BC5FE8CFC20" box="[198,370,937,963]" italics="true" pageId="15" pageNumber="56">Pyronotanthias</emphasis>
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are unusual in having a distinctive bright orange to red stripe or series of blotches beneath the soft dorsal fin and along the upper edge of the caudal peduncle (
<figureCitation id="0BE82A57FFBDFFC4B49B1BA1FC02FC04" box="[935,1020,973,999]" captionStart="FIGURE 10" captionStartId="16.[151,250,1595,1619]" captionTargetBox="[151,1437,417,1571]" captionTargetId="figure-127@16.[151,1437,417,1571]" captionTargetPageId="16" captionText="FIGURE 10. Male Pyronotanthias: A) P. lori, paralectotype, BPBM 13998, 41 mm SL; P. smithvanizi, holotype, BPBM 15606, 58.2 mm SL. Photographs by J.E. Randall." figureDoi="http://doi.org/10.5281/zenodo.5869512" httpUri="https://zenodo.org/record/5869512/files/figure.png" pageId="15" pageNumber="56">Fig. 10</figureCitation>
). Although the stripe or blotches are not always apparent in
<taxonomicName id="54D34D51FFBDFFC4B6F21B9EFD80FBE8" box="[462,638,1009,1035]" class="Actinopterygii" family="Serranidae" genus="Pyronotanthias" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="56" phylum="Chordata" rank="species" species="bimarginatus">
<emphasis id="A1A7EAC0FFBDFFC4B6F21B9EFD80FBE8" box="[462,638,1009,1035]" italics="true" pageId="15" pageNumber="56">P. bimarginatus</emphasis>
</taxonomicName>
,
<taxonomicName id="54D34D51FFBDFFC4B5B21B9EFCD3FBE8" box="[654,813,1010,1035]" class="Actinopterygii" family="Serranidae" genus="Pyronotanthias" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="56" phylum="Chordata" rank="species" species="parvirostris">
<emphasis id="A1A7EAC0FFBDFFC4B5B21B9EFCD3FBE8" box="[654,813,1010,1035]" italics="true" pageId="15" pageNumber="56">P. parvirostris</emphasis>
</taxonomicName>
,
<taxonomicName id="54D34D51FFBDFFC4B4011B9EFC27FBE8" box="[829,985,1009,1035]" class="Actinopterygii" family="Serranidae" genus="Pyronotanthias" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="56" phylum="Chordata" rank="species" species="smithvanizi">
<emphasis id="A1A7EAC0FFBDFFC4B4011B9EFC27FBE8" box="[829,985,1009,1035]" italics="true" pageId="15" pageNumber="56">P. smithvanizi</emphasis>
</taxonomicName>
and
<taxonomicName id="54D34D51FFBDFFC4B32D1B9EFB2EFBE8" box="[1041,1232,1010,1035]" class="Actinopterygii" family="Serranidae" genus="Pyronotanthias" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="56" phylum="Chordata" rank="species" species="unimarginatus">
<emphasis id="A1A7EAC0FFBDFFC4B32D1B9EFB2EFBE8" box="[1041,1232,1010,1035]" italics="true" pageId="15" pageNumber="56">P. unimarginatus</emphasis>
</taxonomicName>
, a very narrow or diffuse orange to red stripe is evident in at least some individuals of the four species, and is particularly obvious in life (
<bibRefCitation id="F7424B23FFBDFFC4B62A1C55FE5DFBB0" author="Kuiter, R. H." box="[278,419,1081,1107]" pageId="15" pageNumber="56" refId="ref16555" refString="Kuiter, R. H. (2004) Basslets Hamlets and Their Relatives. A Comprehensive Guide to Selected Serranidae and Plesiopidae. TMC Publishing, Chorleywood, 216 pp." type="book" year="2004">Kuiter 2004</bibRefCitation>
;
<bibRefCitation id="F7424B23FFBDFFC4B6901C55FD46FBB0" author="Allen, G. R. &amp; Erdmann, M. V." box="[428,696,1081,1107]" pageId="15" pageNumber="56" refId="ref13503" refString="Allen, G. R. &amp; Erdmann, M. V. (2012) Reef Fishes of the East Indies. Volumes I - III. Tropical Reef Research, Perth, 1292 pp." type="book" year="2012">Allen &amp; Erdmann 2012</bibRefCitation>
;
<bibRefCitation id="F7424B23FFBDFFC4B5FE1C56FC93FBB0" author="Tea, Y. K. &amp; Plantard, P. &amp; Greene, B. D." box="[706,877,1081,1108]" pageId="15" pageNumber="56" pagination="1 - 7" refId="ref18013" refString="Tea, Y. K., Plantard, P., &amp; Greene, B. D. (2020) Notes on fishes of the mesophotic reefs of Reunion Island. Journal of the Ocean Science Foundation, 35, 1 - 7." type="journal article" year="2020">
Tea
<emphasis id="A1A7EAC0FFBDFFC4B5CD1C56FCD7FBB0" box="[753,809,1081,1107]" italics="true" pageId="15" pageNumber="56">et al.</emphasis>
2020
</bibRefCitation>
).
</paragraph>
<paragraph id="936C36D2FFBDFFC4B7FB1CEEFBA0FB00" blockId="15.[151,1437,1153,2008]" pageId="15" pageNumber="56">In addition, the following combination of characters distinguishes the genus from other anthiadine genera: dorsal fin X,1417, not incised between spinous and soft portions of fin; pectoral-fin rays 1619, longest few rays bearing serrations; tubed lateral-line scales 4156; posterior rim of orbit with papillae.</paragraph>
</subSubSection>
<subSubSection id="DBC96559FFBDFFDAB7FB1C81FB4DFE32" lastPageId="17" lastPageNumber="58" pageId="15" pageNumber="56" type="description">
<paragraph id="936C36D2FFBDFFC4B7FB1C81FDEEF9E0" blockId="15.[151,1437,1153,2008]" pageId="15" pageNumber="56">
<emphasis id="A1A7EAC0FFBDFFC4B7FB1C81FEA7FAE4" bold="true" box="[199,345,1261,1287]" pageId="15" pageNumber="56">Description.</emphasis>
See
<tableCitation id="DE510369FFBDFFC4B6AE1C81FDF3FAEB" box="[402,525,1261,1288]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStart-4="TABLE 5" captionStartId-0="3.[151,239,151,175]" captionStartId-1="9.[150,238,544,570]" captionStartId-2="12.[151,239,296,320]" captionStartId-3="19.[151,239,1052,1076]" captionStartId-4="20.[151,239,151,175]" captionTargetBox-1="[165,2008,622,1603]" captionTargetBox-2="[166,1381,337,1072]" captionTargetPageId-0="3" captionTargetPageId-2="12" captionTargetPageId-3="19" captionTargetPageId-4="20" captionText-0="TABLE 1. Summary of fin-ray counts of Mirolabrichthys, Nemanthias and Pyronotanthias species. Caudal-fin ray counts are presented as upper procurrent + upper principal + lower principal + lower procurrent (upper branched + lower branched)." captionText-1="TABLE 2. Summary of selected dorsal-, anal- and pectoral-fin characters of Mirolabrichthys, Nemanthias and Pyronotanthias species. Abbreviations: ADPF, anterior dorsal-fin pterygiophore formula; OP, opercle; POP, preopercle." captionText-2="TABLE 3. Summary of selected scalation characters of Mirolabrichthys, Nemanthias and Pyronotanthias species." captionText-3="TABLE 4. Summary of selected post-cranial axial skeleton characters of Mirolabrichthys, Nemanthias and Pyronotanthias species." captionText-4="TABLE 5. Gill-raker counts, vomerine dentition and presence or absence of papillae on the posterior orbital rim in Mirolabrichthys, Nemanthias and Pyronotanthias species." httpUri-0="http://table.plazi.org/id/C7AC665AFFB1FFC8B7AB18FBFEF7FF14" httpUri-1="http://table.plazi.org/id/C7AC665AFFBBFFC2B7AA1A4CFD2CFDBD" httpUri-2="http://table.plazi.org/id/C7AC665AFFBEFFC7B7AB1944FAA7FEA3" httpUri-3="http://table.plazi.org/id/C7AC665AFFA1FFD8B7AB1C70FF12FBBB" pageId="15" pageNumber="56" tableUuid-0="C7AC665AFFB1FFC8B7AB18FBFEF7FF14" tableUuid-1="C7AC665AFFBBFFC2B7AA1A4CFD2CFDBD" tableUuid-2="C7AC665AFFBEFFC7B7AB1944FAA7FEA3" tableUuid-3="C7AC665AFFA1FFD8B7AB1C70FF12FBBB">Tables 15</tableCitation>
for variation among species. Dorsal-fin rays X,1417; dorsal-fin origin varying from above pectoral-fin base to above opercle posterior edge (
<figureCitation id="0BE82A57FFBDFFC4B47C1D7DFC6FFAC8" box="[832,913,1297,1323]" captionStart="FIGURE 10" captionStartId="16.[151,250,1595,1619]" captionTargetBox="[151,1437,417,1571]" captionTargetId="figure-127@16.[151,1437,417,1571]" captionTargetPageId="16" captionText="FIGURE 10. Male Pyronotanthias: A) P. lori, paralectotype, BPBM 13998, 41 mm SL; P. smithvanizi, holotype, BPBM 15606, 58.2 mm SL. Photographs by J.E. Randall." figureDoi="http://doi.org/10.5281/zenodo.5869512" httpUri="https://zenodo.org/record/5869512/files/figure.png" pageId="15" pageNumber="56">Fig. 10</figureCitation>
); primary shaft of first dorsal-fin pterygiophore angled distinctly posterodorsally; ADPF S//3/1+1/1/1/1/1/1, S/S/3/1+1/1/1/1/1/1 or S/S+
<superScript id="64A69B9AFFBDFFC4B3941D59FB41FAA0" attach="left" box="[1192,1215,1333,1359]" fontSize="6" pageId="15" pageNumber="56">Sv</superScript>
/3/1+1/1/1/1/1/1 (= predorsal formula 0//2/1+1, 0/0/2/1+1 or 0/0+0/2/1+1;
<figureCitation id="0BE82A57FFBDFFC4B42F1D35FC93FA90" box="[787,877,1369,1395]" captionStart="FIGURE 3" captionStartId="5.[151,250,823,847]" captionTargetBox="[151,1436,182,799]" captionTargetId="figure-15@5.[151,1436,181,799]" captionTargetPageId="5" captionText="FIGURE 3. Predorsal and anterior portion of dorsal fin of cleared and stained specimens.A) Mirolabrichthys tuka, ACG CS532, 47 mm SL; B) Pyronotanthias lori, ACG CS675, 59 mm SL; C) Nemanthias dispar, ACG CS907, 47 mm SL; D) N. carberryi, AMS I.48957-001, 61 mm SL. Photographs are of the right side (digitally reversed) after removal of the pectoral girdle, scales, and skin. Abbreviations: DP, dorsal-fin pterygiophore; DS, dorsal-fin spines; NS, neural spines; SN, supraneurals. Scale bars = 2 mm. Photographs by A.C. Gill." figureDoi="http://doi.org/10.5281/zenodo.5869498" httpUri="https://zenodo.org/record/5869498/files/figure.png" pageId="15" pageNumber="56">Fig. 3B</figureCitation>
); dorsal-fin pterygiophores in interneural spaces 913 1/1/1+1/1+1/1, 1/1+1/1+1/1/1+1, 1/1+1/1+1/1+1/1 or 1/1+1/1+1/1+1/1+1; terminal dorsal-fin pterygiophore in interneural space 1718; anal-fin rays III,78 (usually III,7); terminal anal-fin pterygiophore in interhaemal space 57; pectoral-fin rays 1619; longest few rays bearing serrations; pelvic-fin rays I,5; caudal-fin rays 711+9+8+7 11; branched caudal-fin rays 7+6.
</paragraph>
<paragraph id="936C36D2FFBDFFC4B7FB1E61FDC8F98C" blockId="15.[151,1437,1153,2008]" pageId="15" pageNumber="56">
Scales moderately small, tubed scales in lateral line 4156; circumpeduncle scales 2130; no auxiliary scales on head or body; lower jaw scaled; basal scaled area on dorsal and anal fins either narrow or broad; scales with peripheral cteni only (
<bibRefCitation id="F7424B23FFBDFFC4B6B11E39FDD7F98C" author="Roberts, C. D." box="[397,553,1621,1647]" pageId="15" pageNumber="56" pagination="60 - 113" refId="ref17501" refString="Roberts, C. D. (1993) Comparative morphology of spined scales and their phylogenetic significance in the Teleostei. Bulletin of Marine Science, 52 (1), 60 - 113." type="journal article" year="1993">Roberts 1993</bibRefCitation>
).
</paragraph>
<paragraph id="936C36D2FFBDFFC4B7FB1E15FB21F954" blockId="15.[151,1437,1153,2008]" pageId="15" pageNumber="56">Greatest body depth 2536% SL; head length 2835% SL; orbit diameter 712% SL; predorsal length 2937% SL; preanal length 5467% SL; prepelvic length 3137% SL; caudal peduncle depth 1316% SL.</paragraph>
<paragraph id="936C36D2FFBDFFDBB7FB1EADFA8CFF1A" blockId="15.[151,1437,1153,2008]" lastBlockId="16.[151,1436,151,393]" lastPageId="16" lastPageNumber="57" pageId="15" pageNumber="56">
Anterior part of upper lip of males weakly to strongly hypertrophied (
<figureCitation id="0BE82A57FFBDFFC4B4E41EADFBD2F938" box="[984,1068,1729,1755]" captionStart="FIGURE 10" captionStartId="16.[151,250,1595,1619]" captionTargetBox="[151,1437,417,1571]" captionTargetId="figure-127@16.[151,1437,417,1571]" captionTargetPageId="16" captionText="FIGURE 10. Male Pyronotanthias: A) P. lori, paralectotype, BPBM 13998, 41 mm SL; P. smithvanizi, holotype, BPBM 15606, 58.2 mm SL. Photographs by J.E. Randall." figureDoi="http://doi.org/10.5281/zenodo.5869512" httpUri="https://zenodo.org/record/5869512/files/figure.png" pageId="15" pageNumber="56">Fig. 10</figureCitation>
); mouth large, oblique, posterior margin of maxilla reaching to point ranging from vertical through posterior edge of pupil to vertical through posterior edge of eye; mouth terminal, becoming inferior in males; supramaxilla absent; premaxilla with 1 or 2 enlarged canines anterolaterally, a band of small conical teeth about 4 or 5 rows wide at symphysis reducing to 1 or 2 rows on sides of jaw, with outer-row teeth much larger, those of posterior half of jaw curved anteriorly; 1 or 2 posterior teeth in band nearest symphysis sometimes enlarged and caniniform, lying almost flat against roof of mouth; dentary with 1 or 2 forward projecting or laterally curved, enlarged canines at front of jaw, followed by band of small conical teeth about 3 or 4 rows wide reducing to 1 or 2 rows posteriorly, outer-row teeth much larger, those of posterior part of jaw curved anteriorly; anterior third of dentary with 13 enlarged, posteriorly curved canines outside band of teeth; vomer with small patch of small teeth; palatine with either a small irregular patch of small teeth or a narrow band of small conical teeth, 25 rows wide at widest point; ectopterygoid, endopterygoid, and tongue edentate.
</paragraph>
<paragraph id="936C36D2FFA2FFDBB7FB196FFC6CFE6A" blockId="16.[151,1436,151,393]" pageId="16" pageNumber="57">Opercle with three flat spines, uppermost indistinct, lowermost below junction with subopercle, middle spine closer to lowermost spine than to uppermost; preopercle finely serrated on posterior, vertical edge; interopercle and subopercle usually smooth or with indistinct crenulae, sometimes with fine serrations; posttemporal smooth or with indistinct serrations or crenulae; posterior rim of orbit with papillae.</paragraph>
<caption id="C7AC665AFFA2FFDBB7AB1E57FDB7F994" ID-DOI="http://doi.org/10.5281/zenodo.5869512" ID-Zenodo-Dep="5869512" httpUri="https://zenodo.org/record/5869512/files/figure.png" pageId="16" pageNumber="57" startId="16.[151,250,1595,1619]" targetBox="[151,1437,417,1571]" targetPageId="16">
<paragraph id="936C36D2FFA2FFDBB7AB1E57FDB7F994" blockId="16.[151,1436,1595,1655]" pageId="16" pageNumber="57">
<emphasis id="A1A7EAC0FFA2FFDBB7AB1E57FEE1F9B0" bold="true" box="[151,287,1595,1619]" pageId="16" pageNumber="57">FIGURE 10.</emphasis>
Male
<taxonomicName id="54D34D51FFA2FFDBB6621E57FE07F9B0" authority="Gill, 2022" authorityName="Gill" authorityYear="2022" box="[350,505,1595,1619]" class="Actinopterygii" family="Serranidae" genus="Pyronotanthias" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="56" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="A1A7EAC0FFA2FFDBB6621E57FE07F9B0" box="[350,505,1595,1619]" italics="true" pageId="16" pageNumber="57">Pyronotanthias</emphasis>
</taxonomicName>
: A)
<taxonomicName id="54D34D51FFA2FFDBB5191E50FD9CF9B0" box="[549,610,1595,1619]" class="Actinopterygii" family="Serranidae" genus="Anthias" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="57" phylum="Chordata" rank="species" species="lori">
<emphasis id="A1A7EAC0FFA2FFDBB5191E50FD9CF9B0" box="[549,610,1595,1619]" italics="true" pageId="16" pageNumber="57">P. lori</emphasis>
</taxonomicName>
, paralectotype, BPBM 13998, 41 mm SL;
<taxonomicName id="54D34D51FFA2FFDBB3331E50FB64F9B0" box="[1039,1178,1595,1619]" class="Actinopterygii" family="Serranidae" genus="Pyronotanthias" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="57" phylum="Chordata" rank="species" species="smithvanizi">
<emphasis id="A1A7EAC0FFA2FFDBB3331E50FB64F9B0" box="[1039,1178,1595,1619]" italics="true" pageId="16" pageNumber="57">P. smithvanizi</emphasis>
</taxonomicName>
, holotype, BPBM 15606, 58.2 mm SL. Photographs by J.E. Randall.
</paragraph>
</caption>
<paragraph id="936C36D2FFA2FFDBB7FB1EC9FAE4F85F" blockId="16.[151,1437,1701,2016]" pageId="16" pageNumber="57">
Paired pharyngobranchials (pb) 1 through 4 present, pb4 cartilaginous; tooth plates present on pb2 through pb4, tooth plate on pb4 small and autogenous; paired epibranchials (eb) 1 through 4 present; uncinate process on eb1 fan-shaped, broadly rimmed with cartilage, and directed posterodorsally; no tooth plate on eb2; small toothplate on eb3 (
<figureCitation id="0BE82A57FFA2FFDBB7F21F7DFECCF8CF" box="[206,306,1809,1836]" captionStart="FIGURE 5" captionStartId="6.[151,250,1812,1836]" captionTargetBox="[151,1436,181,1788]" captionTargetId="figure-18@6.[151,1436,181,1788]" captionTargetPageId="6" captionText="FIGURE 5. Right side dorsal gill arches in dorsal view of: A) Mirolabrichthys tuka, ACG CS532, 47 mm SL; B) Nemanthias bicolor, ACG CS14, 62 mm SL; C) N. carberryi, AMS I.48957-001, 61 mm SL; D) Pyronotanthias lori, ACG CS675, 59 mm SL.Abbreviations: EB1-4, epibranchials 1-4; LP, levator process; PB1-4, pharyngobranchials 1-4; PB4TP, pharyngobranchial 4 tooth plate; UP, uncinate process. Scale bars = 1 mm. Illustrations by A.C. Gill." figureDoi="http://doi.org/10.5281/zenodo.5869502" httpUri="https://zenodo.org/record/5869502/files/figure.png" pageId="16" pageNumber="57">Figs. 5D</figureCitation>
,
<figureCitation id="0BE82A57FFA2FFDBB6011F7DFE9DF8CF" box="[317,355,1809,1836]" captionStart="FIGURE 6" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[179,1409,181,1920]" captionTargetId="figure-15@7.[179,1409,181,1920]" captionTargetPageId="7" captionText="FIGURE 6. Right side dorsal gill arches in ventral view of: A) Mirolabrichthys tuka; B) Nemanthias bicolor; C) N. carberryi; D) Pyronotanthias lori. Specimens, abbreviations, and scale bars as in Fig. 5. Illustrations by A.C. Gill." figureDoi="http://doi.org/10.5281/zenodo.5869504" httpUri="https://zenodo.org/record/5869504/files/figure.png" pageId="16" pageNumber="57">6D</figureCitation>
); paired ceratobranchials (cb) 1 through 5 present; tooth plate on cb5; paired hypobranchials (hb) 1 through 3 present, none with toothplates; median basibranchials (bb) 1 through 4 present, bb4 as cartilage, none with toothplates; gill rakers elongate on anterior/lateral face of first arch, 811 on eb1, uppermost at junction with pb1, and 2027 on cb1 and hb1; shorter club-like rakers or rudiments present on posterior face of eb1, cb1 and hb1, on both anterior and posterior faces of eb23, cb24 and hb2, and on anterior face of hb3, eb4 and cb5.
</paragraph>
<paragraph id="936C36D2FFA2FFDAB7FB1FA9FE64FEFE" blockId="16.[151,1437,1701,2016]" lastBlockId="17.[151,1437,150,933]" lastPageId="17" lastPageNumber="58" pageId="16" pageNumber="57">Vertebrae 10+16; parapophyses present on first caudal vertebra; epineurals present on vertebrae 1 through 1215; ribs present on vertebrae 3 through 1011; preural 2 (pu2) and pu3 haemal spines autogenous; no radial cartilages in caudal skeleton anterior to pu3 haemal and neural spines; parhypural and hypurals autogenous; welldeveloped hypurapophysis on parhypural; single uroneural; 3 epurals; ventral tip of cleithrum with well-developed posteroventral process.</paragraph>
<paragraph id="936C36D2FFA3FFDAB7FB194AFB4DFE32" blockId="17.[151,1437,150,933]" pageId="17" pageNumber="58">
<emphasis id="A1A7EAC0FFA3FFDAB7FB194AFE99FEA3" bold="true" box="[199,359,294,320]" pageId="17" pageNumber="58">Composition.</emphasis>
<taxonomicName id="54D34D51FFA3FFDAB653194BFBA5FEA2" authority="Randall &amp; McCosker, 1982" authorityName="Randall &amp; McCosker" authorityYear="1982" box="[367,1115,295,321]" class="Actinopterygii" family="Serranidae" genus="Anthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="aurulentus" subGenus="Mirolabrichthys">
<emphasis id="A1A7EAC0FFA3FFDAB653194BFE3BFEA2" box="[367,453,295,321]" italics="true" pageId="17" pageNumber="58">Anthias</emphasis>
(
<emphasis id="A1A7EAC0FFA3FFDAB6EB194BFD72FEA2" box="[471,652,295,321]" italics="true" pageId="17" pageNumber="58">Mirolabrichthys</emphasis>
)
<emphasis id="A1A7EAC0FFA3FFDAB5A2194BFCE8FEA2" box="[670,790,295,321]" italics="true" pageId="17" pageNumber="58">aurulentus</emphasis>
<bibRefCitation id="F7424B23FFA3FFDAB423194BFBA5FEA2" author="Randall, J. E. &amp; McCosker, J. E." box="[799,1115,295,321]" pageId="17" pageNumber="58" pagination="59 - 69" refId="ref17267" refString="Randall, J. E. &amp; McCosker, J. E. (1982) Two new serranid fishes of the genus Anthias from the central Pacific. Journal of Aquariculture, 2 (3), 59 - 69." type="journal article" year="1982">Randall &amp; McCosker, 1982</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="54D34D51FFA3FFDAB354194BFECBFE86" authority="Randall, 2011" authorityName="Randall" authorityYear="2011" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="bimarginatus">
<emphasis id="A1A7EAC0FFA3FFDAB354194BFA62FEA2" box="[1128,1436,295,321]" italics="true" pageId="17" pageNumber="58">Pseudanthias bimarginatus</emphasis>
<bibRefCitation id="F7424B23FFA3FFDAB7AB1927FECBFE86" author="Randall, J. E." box="[151,309,331,357]" pageId="17" pageNumber="58" pagination="79 - 87" refId="ref17058" refString="Randall, J. E. (2011) Two new serranid fishes of the genus Pseudanthias from the western Indian Ocean. Smithiana, Publications in Aquatic Biodiversity, Bulletin, 13, 79 - 87." type="journal article" year="2011">Randall, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="54D34D51FFA3FFDAB6021927FBD1FE86" authority="Katayama &amp; Masuda, 1980" authorityName="Katayama &amp; Masuda" authorityYear="1980" box="[318,1071,331,357]" class="Actinopterygii" family="Serranidae" genus="Anthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="flavoguttatus" subGenus="Mirolabrichthys">
<emphasis id="A1A7EAC0FFA3FFDAB6021927FE6AFE86" box="[318,404,331,357]" italics="true" pageId="17" pageNumber="58">Anthias</emphasis>
(
<emphasis id="A1A7EAC0FFA3FFDAB69E1927FDA9FE86" box="[418,599,331,357]" italics="true" pageId="17" pageNumber="58">Mirolabrichthys</emphasis>
)
<emphasis id="A1A7EAC0FFA3FFDAB55A1927FD06FE86" box="[614,760,331,357]" italics="true" pageId="17" pageNumber="58">flavoguttatus</emphasis>
<bibRefCitation id="F7424B23FFA3FFDAB5C11927FBD1FE86" author="Katayama, M. &amp; Masuda, H." box="[765,1071,331,357]" pageId="17" pageNumber="58" pagination="185 - 190" refId="ref16366" refString="Katayama, M. &amp; Masuda, H. (1980) Two new anthiine fishes from Sagami Bay, Japan. Japanese Journal of Ichthyology, 27 (3), 185 - 190." type="journal article" year="1980">Katayama &amp; Masuda, 1980</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="54D34D51FFA3FFDAB3041927FA62FE86" authority="Lubbock &amp; Randall" authorityName="Lubbock &amp; Randall" authorityYear="1976" box="[1080,1436,331,357]" class="Actinopterygii" family="Serranidae" genus="Anthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="lori">
<emphasis id="A1A7EAC0FFA3FFDAB3041927FB45FE86" box="[1080,1211,331,357]" italics="true" pageId="17" pageNumber="58">Anthias lori</emphasis>
Lubbock &amp; Randall
</taxonomicName>
<emphasis id="A1A7EAC0FFA3FFDAB7AB1903FF53FE6B" box="[151,173,367,392]" italics="true" pageId="17" pageNumber="58">in</emphasis>
<bibRefCitation id="F7424B23FFA3FFDAB7891903FDF9FE6A" author="Fourmanoir, P. &amp; Laboute, P." box="[181,519,367,393]" pageId="17" pageNumber="58" refId="ref14020" refString="Fourmanoir, P. &amp; Laboute, P. (1976) Poissons des mers tropicales. Nouvelle Caledonie, Nouvelles Hebrides. Les editions du Pacifique, Papeete, 376 pp." type="book" year="1976">Fourmanoir &amp; Laboute, 1976</bibRefCitation>
,
<taxonomicName id="54D34D51FFA3FFDAB52F1903FB06FE6A" authority="Randall &amp; Lubbock, 1981" authorityName="Randall &amp; Lubbock" authorityYear="1981" box="[531,1272,367,393]" class="Actinopterygii" family="Serranidae" genus="Anthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="parvirostris" subGenus="Mirolabrichthys">
<emphasis id="A1A7EAC0FFA3FFDAB52F1903FD97FE6A" box="[531,617,367,393]" italics="true" pageId="17" pageNumber="58">Anthias</emphasis>
(
<emphasis id="A1A7EAC0FFA3FFDAB5471903FCCEFE6A" box="[635,816,367,393]" italics="true" pageId="17" pageNumber="58">Mirolabrichthys</emphasis>
)
<emphasis id="A1A7EAC0FFA3FFDAB47D1903FC3AFE6B" box="[833,964,367,392]" italics="true" pageId="17" pageNumber="58">parvirostris</emphasis>
<bibRefCitation id="F7424B23FFA3FFDAB4F11903FB06FE6A" author="Randall, J. E. &amp; Lubbock, R." box="[973,1272,367,393]" pageId="17" pageNumber="58" pagination="1 - 27" refId="ref17199" refString="Randall, J. E. &amp; Lubbock, R. (1981) A revision of the serranid fishes of the subgenus Mirolabrichthys (Anthiinae: Anthias), with description of five new species. Contributions in Science (Natural History Museum of Los Angeles County), 333, 1 - 27. https: // doi. org / 10.5962 / p. 241267" type="journal article" year="1981">Randall &amp; Lubbock, 1981</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="54D34D51FFA3FFDAB2391903FDFFFE4E" authority="Randall &amp; Pyle, 2001" authorityName="Randall &amp; Pyle" authorityYear="2001" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="privitera">
<emphasis id="A1A7EAC0FFA3FFDAB2391903FF07FE4F" italics="true" pageId="17" pageNumber="58">Pseudanthias privitera</emphasis>
<bibRefCitation id="F7424B23FFA3FFDAB63F19FFFDFFFE4E" author="Randall, J. E. &amp; Pyle, R. L." box="[259,513,403,429]" pageId="17" pageNumber="58" pagination="19 - 34" refId="ref17407" refString="Randall, J. E. &amp; Pyle, R. L. (2001) Four new serranid fishes of the anthiine genus Pseudanthias from the South Pacific. Raffles Bulletin of Zoology, 49 (1), 19 - 34." type="journal article" year="2001">Randall &amp; Pyle, 2001</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="54D34D51FFA3FFDAB53219FFFB09FE4E" authority="Randall &amp; Lubbock, 1981" authorityName="Randall &amp; Lubbock" authorityYear="1981" box="[526,1271,403,429]" class="Actinopterygii" family="Serranidae" genus="Anthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="smithvanizi" subGenus="Mirolabrichthys">
<emphasis id="A1A7EAC0FFA3FFDAB53219FFFD9AFE4E" box="[526,612,403,429]" italics="true" pageId="17" pageNumber="58">Anthias</emphasis>
(
<emphasis id="A1A7EAC0FFA3FFDAB54B19FFFCD2FE4E" box="[631,812,403,429]" italics="true" pageId="17" pageNumber="58">Mirolabrichthys</emphasis>
)
<emphasis id="A1A7EAC0FFA3FFDAB40319FFFC41FE4E" box="[831,959,403,429]" italics="true" pageId="17" pageNumber="58">smithvanizi</emphasis>
<bibRefCitation id="F7424B23FFA3FFDAB4F419FFFB09FE4E" author="Randall, J. E. &amp; Lubbock, R." box="[968,1271,403,429]" pageId="17" pageNumber="58" pagination="1 - 27" refId="ref17199" refString="Randall, J. E. &amp; Lubbock, R. (1981) A revision of the serranid fishes of the subgenus Mirolabrichthys (Anthiinae: Anthias), with description of five new species. Contributions in Science (Natural History Museum of Los Angeles County), 333, 1 - 27. https: // doi. org / 10.5962 / p. 241267" type="journal article" year="1981">Randall &amp; Lubbock, 1981</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="54D34D51FFA3FFDAB23919FFFD70FE32" authority="Victor, Teitelbaum &amp; Randall, 2020" authorityName="Victor, Teitelbaum &amp; Randall" authorityYear="2020" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="timanoa">
<emphasis id="A1A7EAC0FFA3FFDAB23919FFFF0DFE33" italics="true" pageId="17" pageNumber="58">Pseudanthias timanoa</emphasis>
<bibRefCitation id="F7424B23FFA3FFDAB7C619DBFD70FE32" author="Victor, B. C. &amp; Teitelbaum, A. &amp; Randall, J. E." box="[250,654,439,465]" pageId="17" pageNumber="58" pagination="6 - 15" refId="ref18059" refString="Victor, B. C., Teitelbaum, A. &amp; Randall, J. E. (2020) Pseudanthias timanoa, a new fairy basslet from New Caledonia, South Pacific (Teleostei: Serranidae: Anthiadinae). Journal of the Ocean Science Foundation, 36, 6 - 15." type="journal article" year="2020">Victor, Teitelbaum &amp; Randall, 2020</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="54D34D51FFA3FFDAB5F419DBFB50FE32" authority="Randall, 2011" authorityName="Randall" authorityYear="2011" box="[712,1198,439,465]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="unimarginatus">
<emphasis id="A1A7EAC0FFA3FFDAB5F419DBFBF6FE33" box="[712,1032,439,465]" italics="true" pageId="17" pageNumber="58">Pseudanthias unimarginatus</emphasis>
<bibRefCitation id="F7424B23FFA3FFDAB33319DBFB50FE32" author="Randall, J. E." box="[1039,1198,439,465]" pageId="17" pageNumber="58" pagination="79 - 87" refId="ref17058" refString="Randall, J. E. (2011) Two new serranid fishes of the genus Pseudanthias from the western Indian Ocean. Smithiana, Publications in Aquatic Biodiversity, Bulletin, 13, 79 - 87." type="journal article" year="2011">Randall, 2011</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DBC96559FFA3FFDAB7FB19B7FED4FDDE" pageId="17" pageNumber="58" type="etymology">
<paragraph id="936C36D2FFA3FFDAB7FB19B7FED4FDDE" blockId="17.[151,1437,150,933]" pageId="17" pageNumber="58">
<emphasis id="A1A7EAC0FFA3FFDAB7FB19B7FEB0FE16" bold="true" box="[199,334,475,501]" pageId="17" pageNumber="58">Etymology.</emphasis>
From the Greek πυρ, fire, νοτος, back, and
<taxonomicName id="54D34D51FFA3FFDAB40219B7FC6AFE16" authorityName="Bloch" authorityYear="1792" box="[830,916,475,501]" class="Actinopterygii" family="Serranidae" genus="Anthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="genus">
<emphasis id="A1A7EAC0FFA3FFDAB40219B7FC6AFE16" box="[830,916,475,501]" italics="true" pageId="17" pageNumber="58">Anthias</emphasis>
</taxonomicName>
. It alludes to the characteristic red blotches or stripes on the upper caudal peduncle and beneath the posterior part of the dorsal fin of the included species. Gender is masculine.
</paragraph>
</subSubSection>
<subSubSection id="DBC96559FFA3FFDAB7FB1A2BFCEFFD66" pageId="17" pageNumber="58" type="discussion">
<paragraph id="936C36D2FFA3FFDAB7FB1A2BFCEFFD66" blockId="17.[151,1437,150,933]" pageId="17" pageNumber="58">
<emphasis id="A1A7EAC0FFA3FFDAB7FB1A2BFEC5FD82" bold="true" box="[199,315,583,609]" pageId="17" pageNumber="58">Remarks.</emphasis>
<bibRefCitation id="F7424B23FFA3FFDAB67D1A2BFDF8FD82" author="Gill, A. C. &amp; Pogonoski, J. J. &amp; Johnson, J. W. &amp; Tea, Y. K." box="[321,518,583,609]" pageId="17" pageNumber="58" pagination="49 - 82" refId="ref14518" refString="Gill, A. C., Pogonoski, J. J., Johnson, J. W. &amp; Tea, Y. K. (2021 a) Three new species of Australian anthiadine fishes, with comments on the monophyly of Pseudanthias (Teleostei: Serranidae). Zootaxa, 4996 (1), 49 - 82. https: // doi. org / 10.11646 / zootaxa. 4996.1.2" type="journal article" year="2021">
Gill
<emphasis id="A1A7EAC0FFA3FFDAB64E1A2BFE57FD82" box="[370,425,583,609]" italics="true" pageId="17" pageNumber="58">et al.</emphasis>
(2021a)
</bibRefCitation>
included three species in their analysis of
<emphasis id="A1A7EAC0FFA3FFDAB4E41A2BFBF6FD82" box="[984,1032,583,609]" italics="true" pageId="17" pageNumber="58">COI</emphasis>
sequences,
<taxonomicName id="54D34D51FFA3FFDAB3B01A2BFAE1FD82" authorityName="Randall &amp; McCosker" authorityYear="1982" box="[1164,1311,583,609]" class="Actinopterygii" family="Serranidae" genus="Anthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="aurulentus">
<emphasis id="A1A7EAC0FFA3FFDAB3B01A2BFAE1FD82" box="[1164,1311,583,609]" italics="true" pageId="17" pageNumber="58">P. aurulentus</emphasis>
</taxonomicName>
,
<taxonomicName id="54D34D51FFA3FFDAB2171A2BFA90FD82" authorityName="Lubbock &amp; Randall" authorityYear="1976" box="[1323,1390,583,609]" class="Actinopterygii" family="Serranidae" genus="Anthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="lori">
<emphasis id="A1A7EAC0FFA3FFDAB2171A2BFA90FD82" box="[1323,1390,583,609]" italics="true" pageId="17" pageNumber="58">P. lori</emphasis>
</taxonomicName>
and
<taxonomicName id="54D34D51FFA3FFDAB7AB1A07FEF1FD67" authorityName="Victor, Teitelbaum &amp; Randall" authorityYear="2020" box="[151,271,619,644]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="timanoa">
<emphasis id="A1A7EAC0FFA3FFDAB7AB1A07FEF1FD67" box="[151,271,619,644]" italics="true" pageId="17" pageNumber="58">P. timanoa</emphasis>
</taxonomicName>
, and retrieved them as a monophyletic group.
</paragraph>
</subSubSection>
<subSubSection id="DBC96559FFA3FFDAB7FB1AE2FE8CFC46" pageId="17" pageNumber="58" type="materials_examined">
<paragraph id="936C36D2FFA3FFDAB7FB1AE2FE8CFC46" blockId="17.[151,1437,150,933]" pageId="17" pageNumber="58">
<emphasis id="A1A7EAC0FFA3FFDAB7FB1AE2FE4FFD4B" bold="true" box="[199,433,654,680]" pageId="17" pageNumber="58">Material examined.</emphasis>
<taxonomicName id="54D34D51FFA3FFDAB6851AE3FCC3FD4A" authority="AMS I." authorityName="AMS I." box="[441,829,655,681]" class="Actinopterygii" family="Serranidae" genus="Pyronotanthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="aurulentus">
<emphasis id="A1A7EAC0FFA3FFDAB6851AE3FD1AFD4A" box="[441,740,655,681]" italics="true" pageId="17" pageNumber="58">Pyronotanthias aurulentus</emphasis>
AMS I.
</taxonomicName>
22467-001 (1W,X,
<typeStatus id="4C688870FFA3FFDAB3261AE3FB83FD4A" box="[1050,1149,655,681]" pageId="17" pageNumber="58" type="paratype">paratype</typeStatus>
,
<quantity id="542B9B37FFA3FFDAB3B61AE3FB27FD4A" box="[1162,1241,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="17" pageNumber="58" unit="mm" value="29.0">29 mm</quantity>
SL),
<collectionCode id="F5C2AE17FFA3FFDAB2251AE3FAB1FD4A" box="[1305,1359,655,681]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="17" pageNumber="58">ZRC</collectionCode>
61164 (1W,X, 70.0 mm SL);
<taxonomicName id="54D34D51FFA3FFDAB6AF1ADFFD62FD2E" authority="USNM" authorityName="USNM" box="[403,668,691,717]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="bimarginatus">
<emphasis id="A1A7EAC0FFA3FFDAB6AF1ADFFDBDFD2E" box="[403,579,691,717]" italics="true" pageId="17" pageNumber="58">P. bimarginatus</emphasis>
USNM
</taxonomicName>
398058 (1X,
<typeStatus id="4C688870FFA3FFDAB40B1ADFFC64FD2E" box="[823,922,691,717]" pageId="17" pageNumber="58" type="paratype">paratype</typeStatus>
,
<quantity id="542B9B37FFA3FFDAB49A1ADFFBF4FD2E" box="[934,1034,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="3.6799999999999997" pageId="17" pageNumber="58" unit="mm" value="36.8">36.8 mm</quantity>
SL),
<collectionCode id="F5C2AE17FFA3FFDAB3751ADFFB81FD2E" box="[1097,1151,691,717]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="17" pageNumber="58">ZRC</collectionCode>
62122 (6W,X,
<quantity id="542B9B37FFA3FFDAB2101ADEFF3DFD12" metricMagnitude="-2" metricUnit="m" metricValue="4.375" metricValueMax="4.529999999999999" metricValueMin="4.22" pageId="17" pageNumber="58" unit="mm" value="43.75" valueMax="45.3" valueMin="42.2">42.245.3 mm</quantity>
SL);
<taxonomicName id="54D34D51FFA3FFDAB6391ABBFE0DFD12" authority="ZRC" authorityName="ZRC" box="[261,499,727,753]" class="Actinopterygii" family="Serranidae" genus="Anthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="flavoguttatus">
<emphasis id="A1A7EAC0FFA3FFDAB6391ABBFE4AFD12" box="[261,436,727,753]" italics="true" pageId="17" pageNumber="58">P. flavoguttatus</emphasis>
ZRC
</taxonomicName>
62054 (2W,X,
<quantity id="542B9B37FFA3FFDAB5981ABAFCB7FD12" box="[676,841,726,753]" metricMagnitude="-2" metricUnit="m" metricValue="3.495" metricValueMax="3.65" metricValueMin="3.34" pageId="17" pageNumber="58" unit="mm" value="34.95" valueMax="36.5" valueMin="33.4">33.436.5 mm</quantity>
SL),
<collectionCode id="F5C2AE17FFA3FFDAB4B61ABBFC3EFD12" box="[906,960,727,753]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="17" pageNumber="58">ZRC</collectionCode>
62119 (2W,X, 27.0
<quantity id="542B9B37FFA3FFDAB38C1ABBFAEBFD12" box="[1200,1301,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="3.17" pageId="17" pageNumber="58" unit="mm" value="31.7">31.7 mm</quantity>
SL);
<taxonomicName id="54D34D51FFA3FFDAB26B1ABBFF02FCF6" authority="ACG CS" authorityName="ACG CS" class="Actinopterygii" family="Serranidae" genus="Anthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="lori">
<emphasis id="A1A7EAC0FFA3FFDAB26B1ABBFA62FD12" box="[1367,1436,727,753]" italics="true" pageId="17" pageNumber="58">P. lori</emphasis>
ACG CS
</taxonomicName>
675 (3
<collectionCode id="F5C2AE17FFA3FFDAB67F1A97FE97FCF6" box="[323,361,763,789]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="17" pageNumber="58">CS</collectionCode>
,
<quantity id="542B9B37FFA3FFDAB6491A97FE11FCF6" box="[373,495,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" metricValueMax="6.2" metricValueMin="4.0" pageId="17" pageNumber="58" unit="mm" value="51.0" valueMax="62.0" valueMin="40.0">4062 mm</quantity>
SL), ACG
<collectionCode id="F5C2AE17FFA3FFDAB5531A97FD6EFCF6" box="[623,656,763,789]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="17" pageNumber="58">CS</collectionCode>
934 (1
<collectionCode id="F5C2AE17FFA3FFDAB5EA1A97FD00FCF6" box="[726,766,763,789]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="17" pageNumber="58">CS</collectionCode>
,
<quantity id="542B9B37FFA3FFDAB4371A97FCA4FCF6" box="[779,858,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="17" pageNumber="58" unit="mm" value="39.0">39 mm</quantity>
SL), AMS I.18563-001 (1W,X,
<typeStatus id="4C688870FFA3FFDAB3FA1A97FA9EFCF6" box="[1222,1376,763,789]" pageId="17" pageNumber="58" type="paralectotype">paralectotype</typeStatus>
,
<quantity id="542B9B37FFA3FFDAB2571A96FF3DFCDA" metricMagnitude="-2" metricUnit="m" metricValue="3.35" pageId="17" pageNumber="58" unit="mm" value="33.5">33.5 mm</quantity>
SL),
<collectionCode id="F5C2AE17FFA3FFDAB63D1B73FEACFCDA" box="[257,338,799,825]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="58" type="Museum">USNM</collectionCode>
216922 (2X,
<typeStatus id="4C688870FFA3FFDAB6D11B73FDA6FCDA" box="[493,600,799,825]" pageId="17" pageNumber="58" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="54D34D51FFA3FFDAB5411B73FBC2FCDA" authority="Burgess, 1977" authorityName="Burgess" authorityYear="1977" box="[637,1084,799,825]" class="Actinopterygii" family="Serranidae" genus="Mirolabrichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="imeldae">
<emphasis id="A1A7EAC0FFA3FFDAB5411B73FC6CFCDA" box="[637,914,799,825]" italics="true" pageId="17" pageNumber="58">Mirolabrichthys imeldae</emphasis>
<bibRefCitation id="F7424B23FFA3FFDAB4A41B73FBC2FCDA" author="Burgess, W. E." box="[920,1084,799,825]" pageId="17" pageNumber="58" pagination="39 - 42" refId="ref13978" refString="Burgess, W. E. (1977) Mirolabrichthys immeldae, a new species of anthiid fish from the Philippines (Perciformes: Serranidae). Tropical Fish Hobbyist, 25 (10), 39 - 42, 44." type="journal article" year="1977">Burgess, 1977</bibRefCitation>
</taxonomicName>
),
<collectionCode id="F5C2AE17FFA3FFDAB3731B73FB5EFCDA" box="[1103,1184,799,825]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="58" type="Museum">USNM</collectionCode>
216923 (1X,
<typeStatus id="4C688870FFA3FFDAB2061B73FA62FCDA" box="[1338,1436,799,825]" pageId="17" pageNumber="58" type="holotype">holotype</typeStatus>
of
<taxonomicName id="54D34D51FFA3FFDAB7841B2FFEC6FCBE" authorityName="Burgess" authorityYear="1977" box="[184,312,835,861]" class="Actinopterygii" family="Serranidae" genus="Mirolabrichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="imeldae">
<emphasis id="A1A7EAC0FFA3FFDAB7841B2FFEC6FCBE" box="[184,312,835,861]" italics="true" pageId="17" pageNumber="58">M. imeldae</emphasis>
</taxonomicName>
);
<taxonomicName id="54D34D51FFA3FFDAB66E1B2FFDD0FCBE" authority="AMS I." authorityName="AMS I." box="[338,558,835,861]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="privitera">
<emphasis id="A1A7EAC0FFA3FFDAB66E1B2FFE2CFCBF" box="[338,466,835,860]" italics="true" pageId="17" pageNumber="58">P. privitera</emphasis>
AMS I.
</taxonomicName>
39928-001 (1W,X,
<typeStatus id="4C688870FFA3FFDAB4321B2FFC8FFCBE" box="[782,881,835,861]" pageId="17" pageNumber="58" type="paratype">paratype</typeStatus>
,
<quantity id="542B9B37FFA3FFDAB4421B2EFC1AFCBE" box="[894,996,834,861]" metricMagnitude="-2" metricUnit="m" metricValue="4.45" pageId="17" pageNumber="58" unit="mm" value="44.5">44.5 mm</quantity>
SL);
<taxonomicName id="54D34D51FFA3FFDAB3141B2FFADEFCBE" authority="AMS I." authorityName="AMS I." box="[1064,1312,835,861]" class="Actinopterygii" family="Serranidae" genus="Anthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="smithvanizi">
<emphasis id="A1A7EAC0FFA3FFDAB3141B2FFB38FCBE" box="[1064,1222,835,861]" italics="true" pageId="17" pageNumber="58">P. smithvanizi</emphasis>
AMS I.
</taxonomicName>
18603-001 (2W,X,
<typeStatus id="4C688870FFA3FFDAB7CF1B0BFE9FFC62" box="[243,353,871,897]" pageId="17" pageNumber="58" type="paratype">paratypes</typeStatus>
,
<quantity id="542B9B37FFA3FFDAB6531B0BFE15FC62" box="[367,491,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="17" pageNumber="58" unit="mm" value="40.5" valueMax="42.0" valueMin="39.0">3942 mm</quantity>
SL), AMS I.20436-009 (1
<collectionCode id="F5C2AE17FFA3FFDAB4201B0BFCBAFC62" box="[796,836,871,897]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="17" pageNumber="58">CS</collectionCode>
,
<quantity id="542B9B37FFA3FFDAB46F1B0BFC58FC62" box="[851,934,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="17" pageNumber="58" unit="mm" value="37.0">37 mm</quantity>
SL);
<taxonomicName id="54D34D51FFA3FFDAB4D71B0BFB53FC62" authority="AMS" authorityName="AMS" box="[1003,1197,871,897]" class="Actinopterygii" family="Serranidae" genus="Pseudanthias" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="58" phylum="Chordata" rank="species" species="timanoa">
<emphasis id="A1A7EAC0FFA3FFDAB4D71B0BFB98FC63" box="[1003,1126,871,896]" italics="true" pageId="17" pageNumber="58">P. timanoa</emphasis>
AMS
</taxonomicName>
unregistered (3W,X, 63.475.0 mm SL).
</paragraph>
</subSubSection>
</treatment>
</document>