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<document id="609FA90C2085DBC168DF4A7DD760BB83" ID-DOI="10.1649/0010-065X-72.1.166" ID-ISSN="1938-4394" ID-Zenodo-Dep="5380850" ID-ZooBank="urn:lsid:zoobank.org:pub:C0A34919-7D2A-4CBA-ACCB-98F5126C13BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630290938695" checkinUser="diego" docAuthor="Bortoluzzi, Sidnei &amp; Caron, Edilson" docDate="2018" docId="83176A787977B3789FD1CDC8FB608284" docLanguage="en" docName="0010-065X-72.1.166.pdf" docOrigin="The Coleopterists Bulletin 72 (1)" docSource="http://dx.doi.org/10.1649/0010-065x-72.1.166" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Gnathymenus hoppi Bortoluzzi and Caron 2018, new species" docType="treatment" docVersion="3" lastPageNumber="169" masterDocId="7F2E12007976B37C9D14C93CFFC9860E" masterDocTitle="A New Brazilian Species of Gnathymenus Solier (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="170" masterPageNumber="166" pageNumber="166" updateTime="1699216731144" updateUser="plazi">
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<mods:title id="34BD9D53D3ECE63ABECC248EA66FA5B8">A New Brazilian Species of Gnathymenus Solier (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<mods:namePart id="F88B32547E10A54C85A722563B7C405F">Bortoluzzi, Sidnei</mods:namePart>
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<mods:namePart id="18D1665043A32D0CB4F8C525711285FC">Caron, Edilson</mods:namePart>
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<mods:title id="10F8EAADF84D37E593A7A06EDB4020AD">The Coleopterists Bulletin</mods:title>
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<mods:date id="BD5B64EDC50AD219C10A67CAFCDD06B9">2018</mods:date>
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<paragraph id="0B01DB6E7977B37D9FD1CDC8FC12832D" blockId="1.[704,1153,1268,1396]" pageId="1" pageNumber="166">
<heading id="50496C027977B37D9FD1CDC8FC12832D" centered="true" fontSize="9" level="2" pageId="1" pageNumber="166" reason="2">
<emphasis id="39CA077C7977B37D9FD1CDC8FC12832D" bold="true" pageId="1" pageNumber="166">
<taxonomicName id="CCBEA0ED7977B37D9FD1CDC8FBBE8307" ID-CoL="8Y43Z" authority="Bortoluzzi and Caron" authorityName="Bortoluzzi and Caron" authorityYear="2018" box="[709,1143,1268,1289]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="species" species="hoppi" status="sp. nov.">
<emphasis id="39CA077C7977B37D9FD1CDC8FC468307" bold="true" box="[709,911,1268,1289]" italics="true" pageId="1" pageNumber="166">Gnathymenus hoppi</emphasis>
Bortoluzzi and Caron
</taxonomicName>
,
<taxonomicNameLabel id="22F9BA077977B37D9E72CC32FC12832D" box="[870,987,1294,1315]" pageId="1" pageNumber="166" rank="species">new species</taxonomicNameLabel>
</emphasis>
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Zoobank.org/
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urn:lsid:zoobank.org:act:
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(
<figureCitation id="9385C7EB7977B37D9E79CC62FC1A837D" box="[877,979,1374,1396]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="2.[197,236,1266,1285]" captionStartId-1="3.[175,214,1470,1489]" captionTargetBox-0="[173,1187,194,1242]" captionTargetBox-1="[167,1157,210,1452]" captionTargetId-0="figure-213@2.[173,1195,194,1250]" captionTargetId-1="figure-59@3.[164,1159,194,1454]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs. 12. Gnathymenus hoppi. 1) Holotype, male, dorsal view; 2) Habitus, male, ventral view." captionText-1="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi-0="http://doi.org/10.5281/zenodo.5380852" figureDoi-1="http://doi.org/10.5281/zenodo.5380854" httpUri-0="https://zenodo.org/record/5380852/files/figure.png" httpUri-1="https://zenodo.org/record/5380854/files/figure.png" pageId="1" pageNumber="166">Figs. 112</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="43A488E57977B37E9FD1CCB5FE358006" lastPageId="2" lastPageNumber="167" pageId="1" pageNumber="166" type="materials_examined">
<paragraph id="0B01DB6E7977B37E9FD1CCB5FE358006" blockId="1.[685,1173,1417,1730]" lastBlockId="2.[173,659,1337,1730]" lastPageId="2" lastPageNumber="167" pageId="1" pageNumber="166">
<emphasis id="39CA077C7977B37D9FD1CCB5FC928390" bold="true" box="[709,859,1417,1438]" pageId="1" pageNumber="166">Type Material.</emphasis>
<materialsCitation id="BBD6D1337977B37E9E75CCB5FE358006" collectionCode="DZUP, FMNH, MZUSP, AMNH" collectorName="P. Hopp &amp; Tewnty-nine" country="Brazil" county="Res" lastPageId="2" lastPageNumber="167" location="Serra do Itaqui" municipality="Reserva" pageId="1" pageNumber="166" specimenCode="DZUP/268500" specimenCount="5" stateProvince="Para" typeStatus="holotype">
<typeStatus id="D40565CC7977B37D9E75CCB5FC748390" box="[865,957,1417,1438]" pageId="1" pageNumber="166" type="holotype">Holotype</typeStatus>
: pinned, male (
<figureCitation id="9385C7EB7977B37D995ACCB5FB4C8390" box="[1102,1157,1417,1438]" captionStart="Figs" captionStartId="2.[197,236,1266,1285]" captionTargetBox="[173,1187,194,1242]" captionTargetId="figure-213@2.[173,1195,194,1250]" captionTargetPageId="2" captionText="Figs. 12. Gnathymenus hoppi. 1) Holotype, male, dorsal view; 2) Habitus, male, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5380852" httpUri="https://zenodo.org/record/5380852/files/figure.png" pageId="1" pageNumber="166">Fig. 1</figureCitation>
), deposited in
<collectionCode id="6DAF43AB7977B37D9E3FCC9FFCA083B6" box="[811,873,1443,1464]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="1" pageNumber="166" type="University or college">DZUP</collectionCode>
, with labels: 1) “
<collectingCountry id="73A99BFE7977B37D9906CC9FFBA283B6" box="[1042,1131,1443,1464]" name="Brazil" pageId="1" pageNumber="166">BRAZIL</collectingCountry>
:
<collectingRegion id="C97A158C7977B37D9962CC9FFB5A83B6" box="[1142,1171,1443,1464]" country="Brazil" name="Para" pageId="1" pageNumber="166">PR</collectingRegion>
[
<collectingRegion id="C97A158C7977B37D9FA7CC82FD3283DD" box="[691,763,1470,1491]" country="Brazil" name="Parana" pageId="1" pageNumber="166">Paraná</collectingRegion>
]:
<collectingCounty id="E260A3E27977B37D9E1FCC82FCF983DD" box="[779,816,1470,1491]" pageId="1" pageNumber="166">Res</collectingCounty>
[
<collectingMunicipality id="EB6541147977B37D9E21CC82FC4F83DD" box="[821,902,1470,1491]" pageId="1" pageNumber="166">Reserva</collectingMunicipality>
].
<location id="0E618DB57977B37D9E82CC82FBE883DD" LSID="urn:lsid:plazi:treatment:83176A787977B3789FD1CDC8FB608284:0E618DB57977B37D9E82CC82FBE883DD" box="[918,1057,1470,1491]" country="Brazil" county="Res" municipality="Reserva" name="Serra do Itaqui" pageId="1" pageNumber="166" stateProvince="Para">Serra do Itaqui</location>
/APA
<location id="0E618DB57977B37D994ACC82FC5183E3" LSID="urn:lsid:plazi:treatment:83176A787977B3789FD1CDC8FB608284:0E618DB57977B37D994ACC82FC5183E3" country="Brazil" county="Res" municipality="Reserva" name="Guaraquecaba Moist Lowland" pageId="1" pageNumber="166" stateProvince="Para">Guaraquecaba Moist Lowland</location>
/
<location id="0E618DB57977B37D9E8BCCE4FC1583E3" LSID="urn:lsid:plazi:treatment:83176A787977B3789FD1CDC8FB608284:0E618DB57977B37D9E8BCCE4FC1583E3" box="[927,988,1496,1517]" country="Brazil" county="Res" municipality="Reserva" name="Forest" pageId="1" pageNumber="166" stateProvince="Para">Forest</location>
, sift.
<location id="0E618DB57977B37D9919CCE4FB8A83E3" LSID="urn:lsid:plazi:treatment:83176A787977B3789FD1CDC8FB608284:0E618DB57977B37D9919CCE4FB8A83E3" box="[1037,1091,1496,1517]" country="Brazil" county="Res" municipality="Reserva" name="Litter" pageId="1" pageNumber="166" stateProvince="Para">Litter</location>
, winkler extr./coll.:
<collectorName id="A64BBEB87977B37D9E02CCCFFCA18006" box="[790,872,1523,1544]" pageId="1" pageNumber="166">P. Hopp</collectorName>
; 24.8-09-9.2007” [white label, printed in black]; 2) “
<specimenCode id="5B1873157977B37D9E6BCF31FBC2802C" box="[895,1035,1549,1570]" collectionCode="DZUP" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="1" pageNumber="166" type="University or college">DZUP/268500</specimenCode>
” [white label, printed in black]; 3) “
<typeStatus id="D40565CC7977B37D9E61CF14FC028033" box="[885,971,1576,1597]" pageId="1" pageNumber="166" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="CCBEA0ED7977B37D9EC6CF14FB448033" authorityName="Bortoluzzi and Caron" authorityYear="2018" box="[978,1165,1576,1597]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="species" species="hoppi">Gnathymenus hoppi</taxonomicName>
/ Bortoluzzi &amp; Caron” [red label, printed in black].
<collectorName id="A64BBEB87977B37D9FB9CF61FCEB807C" box="[685,802,1629,1650]" pageId="1" pageNumber="166">Tewnty-nine</collectorName>
<typeStatus id="D40565CC7977B37D9E3FCF61FC4D807C" box="[811,900,1629,1650]" pageId="1" pageNumber="166" type="paratype">paratypes</typeStatus>
with the same first label as
<typeStatus id="D40565CC7977B37D9FB9CF44FCC88083" box="[685,769,1656,1677]" pageId="1" pageNumber="166" type="holotype">holotype</typeStatus>
, the second label with number sequence from
<collectionCode id="6DAF43AB7977B37D9FC8CFAEFCDE80A9" box="[732,791,1682,1703]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="1" pageNumber="166" type="University or college">DZUP</collectionCode>
, detailed below,and third label as “
<typeStatus id="D40565CC7977B37D9955CFAEFB4480A9" box="[1089,1165,1682,1703]" pageId="1" pageNumber="166" type="paratype">Paratype</typeStatus>
/
<taxonomicName id="CCBEA0ED7977B37D9FB9CF91FCA080CC" authorityName="Bortoluzzi and Caron" authorityYear="2018" box="[685,873,1709,1730]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="species" species="hoppi">Gnathymenus hoppi</taxonomicName>
/Bortoluzzi &amp; Caron” [yellow label, printed in black].
<collectionCode id="6DAF43AB7974B37E9CB5CC05FE168340" box="[417,479,1337,1358]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="2" pageNumber="167" type="University or college">DZUP</collectionCode>
numbers 268499, 268502, 268492, 268486, 268491, 268493, 268495, 268496, 268497, 268498, 268501, 268503, 268504, 268512); five
<collectionCode id="6DAF43AB7974B37E9CE5CCB5FDFF8390" box="[497,566,1417,1438]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="167" type="Museum">FMNH</collectionCode>
numbers 268468, 268470, 268471, 268472, 268473; five
<collectionCode id="6DAF43AB7974B37E9DB9CC82FF3583DD" box="[173,252,1470,1491]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="167">MZUSP</collectionCode>
numbers 268474, 268478, 268479, 268481, 268482; and five
<collectionCode id="6DAF43AB7974B37E9CB6CCE4FE2283E3" box="[418,491,1496,1517]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="167">AMNH</collectionCode>
numbers 268484, 268485, 268487, 268589, 268490.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="5FC18BE67974B37E9DD1CDCEFBC1830B" ID-DOI="http://doi.org/10.5281/zenodo.5380852" ID-Zenodo-Dep="5380852" box="[197,1032,1265,1285]" httpUri="https://zenodo.org/record/5380852/files/figure.png" pageId="2" pageNumber="167" startId="2.[197,236,1266,1285]" targetBox="[173,1187,194,1242]" targetPageId="2">
<paragraph id="0B01DB6E7974B37E9DD1CDCEFBC1830B" blockId="2.[197,1032,1265,1285]" box="[197,1032,1265,1285]" pageId="2" pageNumber="167">
<emphasis id="39CA077C7974B37E9DD1CDCEFED2830B" bold="true" box="[197,283,1265,1285]" pageId="2" pageNumber="167">Figs. 12.</emphasis>
<taxonomicName id="CCBEA0ED7974B37E9C24CDCDFE12830A" authorityName="Bortoluzzi and Caron" authorityYear="2018" box="[304,475,1265,1284]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="species" species="hoppi">
<emphasis id="39CA077C7974B37E9C24CDCDFE12830A" box="[304,475,1265,1284]" italics="true" pageId="2" pageNumber="167">Gnathymenus hoppi</emphasis>
</taxonomicName>
.
<emphasis id="39CA077C7974B37E9CF3CDCEFE30830B" bold="true" box="[487,505,1266,1285]" pageId="2" pageNumber="167">1)</emphasis>
Holotype, male, dorsal view;
<emphasis id="39CA077C7974B37E9FEACDCEFCC6830B" bold="true" box="[766,783,1266,1285]" pageId="2" pageNumber="167">2)</emphasis>
Habitus, male, ventral view.
</paragraph>
</caption>
<subSubSection id="43A488E57974B37E9DD1CF31FD4580CC" pageId="2" pageNumber="167" type="diagnosis">
<paragraph id="0B01DB6E7974B37E9DD1CF31FD4580CC" blockId="2.[173,659,1337,1730]" pageId="2" pageNumber="167">
<emphasis id="39CA077C7974B37E9DD1CF31FEE5802C" bold="true" box="[197,300,1549,1570]" pageId="2" pageNumber="167">Diagnosis.</emphasis>
<taxonomicName id="CCBEA0ED7974B37E9C29CF31FDCF802D" authorityName="Bortoluzzi and Caron" authorityYear="2018" box="[317,518,1549,1571]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="species" species="hoppi">
<emphasis id="39CA077C7974B37E9C29CF31FDCF802D" box="[317,518,1549,1571]" italics="true" pageId="2" pageNumber="167">Gnathymenus hoppi</emphasis>
</taxonomicName>
differs from other described species of
<taxonomicName id="CCBEA0ED7974B37E9CF7CF1BFDAF8033" authorityName="Solier" authorityYear="1849" box="[483,614,1575,1597]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="genus">
<emphasis id="39CA077C7974B37E9CF7CF1BFDAF8033" box="[483,614,1575,1597]" italics="true" pageId="2" pageNumber="167">Gnathymenus</emphasis>
</taxonomicName>
by aedeagal shape, which is slender and tapering toward the apex, somewhat S-shaped with apically beaked process, and with a small tooth on the subapical dorsal margin. This species does not fit into any of the groups named by
<bibRefCitation id="6F2FA69F7974B37E9CE6CF91FD4E80CC" author="Herman, L. H." box="[498,647,1709,1730]" pageId="2" pageNumber="167" refId="ref2283" refString="Herman, L. H. 1981. Revision of the subtribe Dolicaonina of the New World, with discussions of phylogeny" type="book" year="1981">Herman (1981)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="43A488E57974B3789FCFCC05FEDF838D" lastPageId="4" lastPageNumber="169" pageId="2" pageNumber="167" type="description">
<paragraph id="0B01DB6E7974B3789FCFCC05FE498755" blockId="2.[707,1194,1336,1730]" lastBlockId="4.[173,660,194,1730]" lastPageId="4" lastPageNumber="169" pageId="2" pageNumber="167">
<emphasis id="39CA077C7974B37E9FCFCC05FC9C8340" bold="true" box="[731,853,1337,1358]" pageId="2" pageNumber="167">Description.</emphasis>
Body length 2.83.0 mm; pronotal width 0.40.5 mm. Body subcylindrical, elongated, and parallel-sided (
<figureCitation id="9385C7EB7974B37E9E6BCC52FC73838D" box="[895,954,1390,1411]" captionStart="Figs" captionStartId="2.[197,236,1266,1285]" captionTargetBox="[173,1187,194,1242]" captionTargetId="figure-213@2.[173,1195,194,1250]" captionTargetPageId="2" captionText="Figs. 12. Gnathymenus hoppi. 1) Holotype, male, dorsal view; 2) Habitus, male, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5380852" httpUri="https://zenodo.org/record/5380852/files/figure.png" pageId="2" pageNumber="167">Fig. 1</figureCitation>
). Dorsal surface glossy. Color brownish. Dorsal integument of head and pronotum with sparse and moderately coarse punctation, dorsal surface of body covered with some long and sparse setae.
<emphasis id="39CA077C7974B37E9EE5CCE4FBE683E3" bold="true" box="[1009,1071,1496,1517]" pageId="2" pageNumber="167">Head:</emphasis>
Transverse, approximately 1.1 times as wide as long, slightly widened behind eyes and with parallel sides; eyes small, not prominent, postocular area slightly longer than length of eyes. Labrum with U-shaped median emargination (
<figureCitation id="9385C7EB7974B37E9E45CF61FC59807C" box="[849,912,1629,1650]" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="2" pageNumber="167">Fig. 3</figureCitation>
). Mandibles symmetrical, 2 teeth on inner margin of each (
<figureCitation id="9385C7EB7974B37E9EF0CF44FBD78083" box="[996,1054,1656,1677]" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="2" pageNumber="167">Fig. 4</figureCitation>
). Labium with palpomere II longer than 3, long setae at apical margin (
<figureCitation id="9385C7EB7974B37E9E0CCF91FC9F80CC" box="[792,854,1709,1730]" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="2" pageNumber="167">Fig. 5</figureCitation>
); glossa split into 2 lobes. Gular 168 sutures separated, gular plate elongated (
<figureCitation id="9385C7EB7975B37F9F27CF61FDBB807C" box="[563,626,1629,1650]" captionStart="Figs" captionStartId="2.[197,236,1266,1285]" captionTargetBox="[173,1187,194,1242]" captionTargetId="figure-213@2.[173,1195,194,1250]" captionTargetPageId="2" captionText="Figs. 12. Gnathymenus hoppi. 1) Holotype, male, dorsal view; 2) Habitus, male, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5380852" httpUri="https://zenodo.org/record/5380852/files/figure.png" pageId="3" pageNumber="168">Fig. 2</figureCitation>
). distinctly tapering posteriad. Prosternum transverse, Neck width 0.6 width of head.
<emphasis id="39CA077C7975B37F9CD2CF44FDD18083" bold="true" box="[454,536,1656,1677]" pageId="3" pageNumber="168">Thorax:</emphasis>
Pronotum with short, triangular prosternal process. Scutellum strongly convex, 1.08 times as long as wide, slightly triangular and long. Elytra together shorter and more wider than head (
<figureCitation id="9385C7EB7975B37F9C5CCF91FE4A80CC" box="[328,387,1709,1730]" captionStart="Figs" captionStartId="2.[197,236,1266,1285]" captionTargetBox="[173,1187,194,1242]" captionTargetId="figure-213@2.[173,1195,194,1250]" captionTargetPageId="2" captionText="Figs. 12. Gnathymenus hoppi. 1) Holotype, male, dorsal view; 2) Habitus, male, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5380852" httpUri="https://zenodo.org/record/5380852/files/figure.png" pageId="3" pageNumber="168">Fig. 1</figureCitation>
); widest in anterior half, slender than pronotum, slightly wider than long. Hind wings not developed (apterous).
<emphasis id="39CA077C7972B3789F3CC9FEFD5A86D9" bold="true" box="[552,659,194,215]" pageId="4" pageNumber="169">Abdomen:</emphasis>
Slightly wider at segments VIIVIII, apical segment tapering. Segments IIIVII with 2 pairs of paratergites, segments IIIVI with paratergites parallel to each other, segment VII with paratergites separated in cross-section.
</paragraph>
<caption id="5FC18BE67975B37F9DBBCC82FC0D8016" ID-DOI="http://doi.org/10.5281/zenodo.5380854" ID-Zenodo-Dep="5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="3" pageNumber="168" startId="3.[175,214,1470,1489]" targetBox="[167,1157,210,1452]" targetPageId="3">
<paragraph id="0B01DB6E7975B37F9DBBCC82FC0D8016" blockId="3.[151,1173,1469,1560]" pageId="3" pageNumber="168">
<emphasis id="39CA077C7975B37F9DBBCC82FEDA83DF" bold="true" box="[175,275,1469,1489]" pageId="3" pageNumber="168">Figs. 312.</emphasis>
<taxonomicName id="CCBEA0ED7975B37F9C3CCC81FE1C83DE" authorityName="Bortoluzzi and Caron" authorityYear="2018" box="[296,469,1469,1488]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="hoppi">
<emphasis id="39CA077C7975B37F9C3CCC81FE1C83DE" box="[296,469,1469,1488]" italics="true" pageId="3" pageNumber="168">Gnathymenus hoppi</emphasis>
</taxonomicName>
.
<emphasis id="39CA077C7975B37F9CF0CC82FE3C83DF" bold="true" box="[484,501,1470,1489]" pageId="3" pageNumber="168">3)</emphasis>
Labrum, dorsal view;
<emphasis id="39CA077C7975B37F9FD1CC82FD1F83DF" bold="true" box="[709,726,1470,1489]" pageId="3" pageNumber="168">4)</emphasis>
Mandibles, dorsal view;
<emphasis id="39CA077C7975B37F9EA8CC82FC0483DF" bold="true" box="[956,973,1470,1489]" pageId="3" pageNumber="168">5)</emphasis>
Labium and mentum, ventral view;
<emphasis id="39CA077C7975B37F9C05CCEAFEEB83E7" bold="true" box="[273,290,1494,1513]" pageId="3" pageNumber="168">6)</emphasis>
Sternum VII, male, ventral view;
<emphasis id="39CA077C7975B37F9F47CCEAFDAC83E7" bold="true" box="[595,613,1494,1513]" pageId="3" pageNumber="168">7)</emphasis>
Tergum VIII, male, dorsal view;
<emphasis id="39CA077C7975B37F9E84CCEAFC6883E7" bold="true" box="[912,929,1494,1513]" pageId="3" pageNumber="168">8)</emphasis>
Sternum VII, male, ventral view;
<emphasis id="39CA077C7975B37F9DDACCD1FF16800E" bold="true" box="[206,223,1517,1536]" pageId="3" pageNumber="168">9)</emphasis>
Terga IX+X, male, dorsal view;
<emphasis id="39CA077C7975B37F9F1CCCD1FDED800E" bold="true" box="[520,548,1517,1536]" pageId="3" pageNumber="168">10)</emphasis>
Aedeagus, lateral view, paramere outlined by dashed line;
<emphasis id="39CA077C7975B37F993ECCD1FB8C800E" bold="true" box="[1066,1093,1517,1536]" pageId="3" pageNumber="168">11)</emphasis>
Sternum VII, female, ventral view;
<emphasis id="39CA077C7975B37F9C6FCF39FE598016" bold="true" box="[379,400,1541,1560]" pageId="3" pageNumber="168">12</emphasis>
) Abdominal segments IX (right arrow) + X (left arrow), female.
</paragraph>
</caption>
<paragraph id="0B01DB6E7972B3789DD1C85DFEFC8201" blockId="4.[173,660,194,1730]" pageId="4" pageNumber="169">
<emphasis id="39CA077C7972B3789DD1C85DFF378778" bold="true" box="[197,254,353,374]" pageId="4" pageNumber="169">Male.</emphasis>
Sterna IIIVII with set of coarse setae near center (
<figureCitation id="9385C7EB7972B3789DEDC840FEF0879F" box="[249,313,380,401]" captionStart="Figs" captionStartId="2.[197,236,1266,1285]" captionTargetBox="[173,1187,194,1242]" captionTargetId="figure-213@2.[173,1195,194,1250]" captionTargetPageId="2" captionText="Figs. 12. Gnathymenus hoppi. 1) Holotype, male, dorsal view; 2) Habitus, male, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5380852" httpUri="https://zenodo.org/record/5380852/files/figure.png" pageId="4" pageNumber="169">Fig. 2</figureCitation>
); setae increasing in quantity and changing position from sternum III to VI, sternum III with fewer setae on posterior margin, sternum VI with many setae medially; coarse setae sternum VII sparser than on the preceding segment, but located more toward anterior margin (
<figureCitation id="9385C7EB7972B3789CC1CB3DFDDB8418" box="[469,530,513,534]" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="4" pageNumber="169">Fig. 6</figureCitation>
); tergum VII with posterior margin truncate; sternum VII with posterior margin emarginated (
<figureCitation id="9385C7EB7972B3789CC7CB0AFDC78445" box="[467,526,566,587]" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="4" pageNumber="169">Fig. 6</figureCitation>
); tergum VIII with posterior margin projected, rounded at middle (
<figureCitation id="9385C7EB7972B3789DA0CB57FF38848E" box="[180,241,619,640]" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="4" pageNumber="169">Fig. 7</figureCitation>
); sternum VIII with conspicuously narrow and deep median incision (
<figureCitation id="9385C7EB7972B3789CA1CBBAFE268495" box="[437,495,646,667]" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="4" pageNumber="169">Fig. 8</figureCitation>
) that nearly reaches anterior margin of sternum; tergum IX with short ventral struts contiguous on anterior half (
<figureCitation id="9385C7EB7972B3789F78CB87FF7284E4" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="4" pageNumber="169">Fig. 9</figureCitation>
); tergum X elongated and trapezoidal, barely extending beyond apex of tergum IX (
<figureCitation id="9385C7EB7972B3789F31CBCCFDA9850B" box="[549,608,752,773]" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="4" pageNumber="169">Fig. 9</figureCitation>
), anteriorly fused with tergum IX, but demarcated by an internal ridge; sternum IX elongated, anterior half submembranous, posterior half more sclerotized and pigmented; posterior margin truncate. Aedeagus very slender, tapering toward apex in ventral view; apical half somewhat S-shaped with apically beaked process and with a small tooth on subapical dorsal margin in lateral view (
<figureCitation id="9385C7EB7972B3789CC1CAF9FDD785D4" box="[469,542,965,986]" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="4" pageNumber="169">Fig. 10</figureCitation>
); parameres small and short (
<figureCitation id="9385C7EB7972B3789C5ECAE3FE5985FA" box="[330,400,991,1012]" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="4" pageNumber="169">Fig. 10</figureCitation>
, dashed line), with 2 apical setae on each.
</paragraph>
<paragraph id="0B01DB6E7972B3789DD1CD28FEDF838D" blockId="4.[173,660,194,1730]" pageId="4" pageNumber="169">
<emphasis id="39CA077C7972B3789DD1CD28FEDD8227" bold="true" box="[197,276,1044,1065]" pageId="4" pageNumber="169">Female.</emphasis>
Sterna IIIVII without set of coarse setae; tergum and sternum VII with posterior margin truncate; tergum VIII with posterior margin slightly projected; sternum VIII triangular, posterior margin rounded (
<figureCitation id="9385C7EB7972B3789C19CD43FE9F829A" box="[269,342,1151,1172]" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="4" pageNumber="169">Fig. 11</figureCitation>
), with minute fringes on apex; tergum IX without ventral struts and contiguous on anterior third; tergum X oblong (
<figureCitation id="9385C7EB7972B3789CC9CD88FDEB82C7" box="[477,546,1204,1225]" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="4" pageNumber="169">Fig. 12</figureCitation>
, left arrow), extended beyond apex of tergum IX, anteriorly fused with tergum IX; extra ventral plate between sternum VIII and tergum IX (
<figureCitation id="9385C7EB7972B3789CD3CC38FDC68317" box="[455,527,1284,1305]" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="4" pageNumber="169">Fig. 12</figureCitation>
, right arrow), but the extra plate does not correspond to the same genital appendage mentioned for
<taxonomicName id="CCBEA0ED7972B3789F56CC04FEF58366" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="39CA077C7972B3789F56CC04FF258367" italics="true" pageId="4" pageNumber="169">Gnathymenus</emphasis>
species
</taxonomicName>
by
<bibRefCitation id="6F2FA69F7972B3789C7DCC6FFDCE8366" author="Herman, L. H." box="[361,519,1363,1384]" pageId="4" pageNumber="169" refId="ref2283" refString="Herman, L. H. 1981. Revision of the subtribe Dolicaonina of the New World, with discussions of phylogeny" type="book" year="1981">Herman (1981)</bibRefCitation>
. Spermatheca not found.
</paragraph>
</subSubSection>
<subSubSection id="43A488E57972B3789DD1CCB5FEFB83DD" pageId="4" pageNumber="169" type="distribution">
<paragraph id="0B01DB6E7972B3789DD1CCB5FEFB83DD" blockId="4.[173,660,194,1730]" pageId="4" pageNumber="169">
<emphasis id="39CA077C7972B3789DD1CCB5FE8F8390" bold="true" box="[197,326,1417,1438]" pageId="4" pageNumber="169">Distribution.</emphasis>
The species is known only from the
<typeStatus id="D40565CC7972B3789DB9CC9FFF1F83B6" box="[173,214,1443,1464]" pageId="4" pageNumber="169">type</typeStatus>
locality in east
<collectingRegion id="C97A158C7972B3789C73CC9FFE6383B6" box="[359,426,1443,1464]" country="Brazil" name="Parana" pageId="4" pageNumber="169">Paraná</collectingRegion>
, Guaraqueçaba, Serra do Itaqui,
<collectingCountry id="73A99BFE7972B3789DE5CC82FEE783DD" box="[241,302,1470,1491]" name="Brazil" pageId="4" pageNumber="169">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="43A488E57972B3789DD1CCE4FB608284" pageId="4" pageNumber="169" type="etymology">
<paragraph id="0B01DB6E7972B3789DD1CCE4FDEA8006" blockId="4.[173,660,194,1730]" pageId="4" pageNumber="169">
<emphasis id="39CA077C7972B3789DD1CCE4FEFE83E3" bold="true" box="[197,311,1496,1517]" pageId="4" pageNumber="169">Etymology.</emphasis>
This species name honors Dr. Philipp W. Hopp, the collector of the
<typeStatus id="D40565CC7972B3789CDECCCFFDD68006" box="[458,543,1523,1544]" pageId="4" pageNumber="169" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="0B01DB6E7972B3789DD1CF31FD918056" blockId="4.[173,660,194,1730]" pageId="4" pageNumber="169">
<emphasis id="39CA077C7972B3789DD1CF31FEBB802C" bold="true" box="[197,370,1549,1570]" pageId="4" pageNumber="169">Biological Notes.</emphasis>
The specimens of
<taxonomicName id="CCBEA0ED7972B3789F2ECF31FD5A802D" authorityName="Bortoluzzi and Caron" authorityYear="2018" box="[570,659,1549,1571]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="hoppi">
<emphasis id="39CA077C7972B3789F2ECF31FD5A802D" box="[570,659,1549,1571]" italics="true" pageId="4" pageNumber="169">G. hoppi</emphasis>
</taxonomicName>
were collected from sifted leaf litter in moist lowland forest by Winkler extracting technique.
</paragraph>
<paragraph id="0B01DB6E7972B3789DD1CF61FC4787A5" blockId="4.[173,660,194,1730]" lastBlockId="4.[707,1194,193,586]" pageId="4" pageNumber="169">
<emphasis id="39CA077C7972B3789DD1CF61FEBA807C" bold="true" box="[197,371,1629,1650]" pageId="4" pageNumber="169">Additional Notes.</emphasis>
<taxonomicName id="CCBEA0ED7972B3789C63CF60FDFD807C" authorityName="Bortoluzzi and Caron" authorityYear="2018" box="[375,564,1628,1650]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="hoppi">
<emphasis id="39CA077C7972B3789C63CF60FDFD807C" box="[375,564,1628,1650]" italics="true" pageId="4" pageNumber="169">Gnathymenus hoppi</emphasis>
</taxonomicName>
is the 10
<superScript id="FCCB76267972B3789F91CF64FD5A8068" attach="none" box="[645,659,1624,1638]" fontSize="6" pageId="4" pageNumber="169">th</superScript>
species recorded in
<collectingCountry id="73A99BFE7972B3789C91CF44FE0B8083" box="[389,450,1656,1677]" name="Brazil" pageId="4" pageNumber="169">Brazil</collectingCountry>
. According Newton and Caron (2017), the following species are recorded from
<collectingCountry id="73A99BFE7972B3789C4ECF91FE5E80CC" box="[346,407,1709,1730]" name="Brazil" pageId="4" pageNumber="169">Brazil</collectingCountry>
:
<taxonomicName id="CCBEA0ED7972B3789CA4CF90FCA086D9" authority="Herman" authorityName="Herman" authorityYear="1981" box="[432,873,194,1730]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="bobelus">
<emphasis id="39CA077C7972B3789CA4CF90FD5A80CC" box="[432,659,1708,1730]" italics="true" pageId="4" pageNumber="169">Gnathymenus bobelus</emphasis>
Herman (
<collectingRegion id="C97A158C7972B3789E09C9FEFCAB86D9" box="[797,866,194,215]" country="Brazil" name="Parana" pageId="4" pageNumber="169">Paraná</collectingRegion>
)
</taxonomicName>
,
<taxonomicName id="CCBEA0ED7972B3789E60C9FDFB6086D9" authority="Herman" authorityName="Herman" authorityYear="1981" box="[884,1193,193,215]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="gomphus">
<emphasis id="39CA077C7972B3789E60C9FDFB9C86D9" box="[884,1109,193,215]" italics="true" pageId="4" pageNumber="169">Gnathymenus gomphus</emphasis>
Herman
</taxonomicName>
(S~ ao Paulo),
<taxonomicName id="CCBEA0ED7972B3789E29C9E7FBB086FF" authority="(Bernhauer)" baseAuthorityName="Bernhauer" baseAuthorityYear="1927" box="[829,1145,219,241]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="klimai">
<emphasis id="39CA077C7972B3789E29C9E7FBC886FF" box="[829,1025,219,241]" italics="true" pageId="4" pageNumber="169">Gnathymenus klimai</emphasis>
(Bernhauer)
</taxonomicName>
(
<collectingRegion id="C97A158C7972B3789991C9E0FCEF8702" country="Brazil" name="Rio de Janeiro" pageId="4" pageNumber="169">Rio de Janeiro</collectingRegion>
),
<taxonomicName id="CCBEA0ED7972B3789E20C9CAFB9F8702" authority="Herman" authorityName="Herman" authorityYear="1981" box="[820,1110,246,268]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="plancus">
<emphasis id="39CA077C7972B3789E20C9CAFBCC8702" box="[820,1029,246,268]" italics="true" pageId="4" pageNumber="169">Gnathymenus plancus</emphasis>
Herman
</taxonomicName>
(
<collectingRegion id="C97A158C7972B3789975C9CBFCC08728" country="Brazil" name="Distrito Federal" pageId="4" pageNumber="169">Federal District</collectingRegion>
and
<collectingRegion id="C97A158C7972B3789E21C82DFCB08728" box="[821,889,273,294]" country="Brazil" name="Parana" pageId="4" pageNumber="169">Paraná</collectingRegion>
),
<taxonomicName id="CCBEA0ED7972B3789E9CC82CFB608728" authority="Herman" authorityName="Herman" authorityYear="1981" box="[904,1193,272,294]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="radulus">
<emphasis id="39CA077C7972B3789E9CC82CFB918728" box="[904,1112,272,294]" italics="true" pageId="4" pageNumber="169">Gnathymenus radulus</emphasis>
Herman
</taxonomicName>
(S~ ao Paulo),
<taxonomicName id="CCBEA0ED7972B3789E5EC817FB60874F" authority="Herman" authorityName="Herman" authorityYear="1981" box="[842,1193,299,321]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="scoliodontus">
<emphasis id="39CA077C7972B3789E5EC817FB98874F" box="[842,1105,299,321]" italics="true" pageId="4" pageNumber="169">Gnathymenus scoliodontus</emphasis>
Herman
</taxonomicName>
(S~ ao Paulo and
<collectingRegion id="C97A158C7972B3789E61C87AFBC48755" box="[885,1037,326,347]" country="Brazil" name="Santa Catarina" pageId="4" pageNumber="169">Santa Catarina</collectingRegion>
),
<taxonomicName id="CCBEA0ED7972B3789932C87AFC928778" authority="Herman" authorityName="Herman" authorityYear="1981" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="39CA077C7972B3789932C87AFCCE8778" italics="true" pageId="4" pageNumber="169">Gnathymenus setosus</emphasis>
Herman
</taxonomicName>
(S~ ao Paulo),
<taxonomicName id="CCBEA0ED7972B3789EF4C85CFCC6879F" authority="Herman" authorityName="Herman" authorityYear="1981" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="umbus">
<emphasis id="39CA077C7972B3789EF4C85CFB608778" box="[992,1193,352,374]" italics="true" pageId="4" pageNumber="169">Gnathymenus umbus</emphasis>
Herman
</taxonomicName>
(S~ ao Paulo), and
<taxonomicName id="CCBEA0ED7972B3789ECFC847FCC687A5" authority="Herman" authorityName="Herman" authorityYear="1981" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="zarzus">
<emphasis id="39CA077C7972B3789ECFC847FB60879F" box="[987,1193,379,401]" italics="true" pageId="4" pageNumber="169">Gnathymenus zarzus</emphasis>
Herman
</taxonomicName>
(S~ ao Paulo).
</paragraph>
<paragraph id="0B01DB6E7972B3789FCFC88DFC828444" blockId="4.[707,1194,193,586]" pageId="4" pageNumber="169">
Using Herman s (1981) key to
<emphasis id="39CA077C7972B3789917C88CFB4F87C8" box="[1027,1158,432,454]" italics="true" pageId="4" pageNumber="169">Ganthymenus</emphasis>
,
<taxonomicName id="CCBEA0ED7972B3789986C88CFD3087EE" authorityName="Bortoluzzi and Caron" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="hoppi">
<emphasis id="39CA077C7972B3789986C88CFD3087EE" italics="true" pageId="4" pageNumber="169">G. hoppi</emphasis>
</taxonomicName>
keys to couplet 73. Below is a proposed modification of the key. The updated key does not include species described after 1981 from
<collectingCountry id="73A99BFE7972B378994ECB3CFB60841B" box="[1114,1193,512,533]" name="Ecuador" pageId="4" pageNumber="169">Ecuador</collectingCountry>
by
<bibRefCitation id="6F2FA69F7972B3789FF0CB27FCA3843E" author="Assing, V." box="[740,874,539,560]" pageId="4" pageNumber="169" pagination="1541 - 1547" refId="ref2217" refString="Assing, V. 2013. Two new species and a new record of Dolicaonina from Ecuador (Coleoptera: Staphylinidae: Paederinae). Linzer Biologische Beitrage 45: 1541 - 1547." type="journal article" year="2013">Assing (2013</bibRefCitation>
,
<bibRefCitation id="6F2FA69F7972B3789E63CB27FC63843E" author="Assing, V." box="[887,938,539,560]" pageId="4" pageNumber="169" pagination="1119 - 1132" refId="ref2251" refString="Assing, V. 2015. New species and additional records of Dolicaonina from Ecuador (Coleoptera: Staphylinidae: Paederinae). Linzer Biologische Beitrage 47: 1119 - 1132." type="journal article" year="2015">2015</bibRefCitation>
),
<bibRefCitation id="6F2FA69F7972B3789EABCB27FBB0843E" author="Rougemont, G. De." box="[959,1145,539,560]" pageId="4" pageNumber="169" pagination="251 - 253" refId="ref2397" refString="M. Ross-Nickoll. 2010. Recovery of litter inhab- Rougemont, G. De. 2014. A new species of Gnathymenus iting beetle assemblages during forest regeneration Solier (Staphylinidae: Paederinae, Dolicaonina) in the Atlantic forest of Southern Brazil. Insect from Ecuador. Entomologist's Monthly Magazine Conservation and Diversity 3: 103 - 113. 150: 251 - 253." type="journal article" year="2014">Rougemont (2014)</bibRefCitation>
, and
<bibRefCitation id="6F2FA69F7972B3789FD7CB09FC8E8444" author="Irmler, U." box="[707,839,565,586]" pageId="4" pageNumber="169" refId="ref2464" refString="Irmler, U. 2015. New species and new records of the Neotropical" type="book" year="2015">Irmler (2015)</bibRefCitation>
.
</paragraph>
<paragraph id="0B01DB6E7972B3789FD7CB5DFB6084F5" blockId="4.[707,1194,609,1162]" pageId="4" pageNumber="169">
72(69). Right side of aedeagus with cylindrical, dorsolaterally directed process just distad of middle and flattened, laterally directed process at about apical quarter [see
<bibRefCitation id="6F2FA69F7972B378994BCB8DFC8D84EF" author="Herman, L. H." pageId="4" pageNumber="169" refId="ref2283" refString="Herman, L. H. 1981. Revision of the subtribe Dolicaonina of the New World, with discussions of phylogeny" type="book" year="1981">Herman 1981</bibRefCitation>
, figs. 457, 458];
<collectingCountry id="73A99BFE7972B3789EF1CBF0FB8384EF" box="[997,1098,716,737]" name="Costa Rica" pageId="4" pageNumber="169">Costa Rica</collectingCountry>
............ ..........................
<taxonomicName id="CCBEA0ED7972B3789ECFCBDAFB6084F5" authorityName="Herman" authorityYear="1981" box="[987,1193,742,763]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="lirellus">
<emphasis id="39CA077C7972B3789ECFCBDAFB6084F5" bold="true" box="[987,1193,742,763]" italics="true" pageId="4" pageNumber="169">Gnathymenus lirellus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="0B01DB6E7972B3789FD7CA3DFB60853E" blockId="4.[707,1194,609,1162]" pageId="4" pageNumber="169">- Aedeagus not as described and illustrated above ...................................... 73A</paragraph>
<paragraph id="0B01DB6E7972B3789FD7CA0AFB6085BB" blockId="4.[707,1194,609,1162]" pageId="4" pageNumber="169">
73A. Aedeagus long and slender, with short, hooked subapical process extending dorsally, strongly uncinated apically (
<figureCitation id="9385C7EB7972B3789943CA57FB54858E" box="[1111,1181,875,896]" captionStart="Figs" captionStartId="3.[175,214,1470,1489]" captionTargetBox="[167,1157,210,1452]" captionTargetId="figure-59@3.[164,1159,194,1454]" captionTargetPageId="3" captionText="Figs. 312. Gnathymenus hoppi. 3) Labrum, dorsal view; 4) Mandibles, dorsal view; 5) Labium and mentum, ventral view; 6) Sternum VII, male, ventral view; 7) Tergum VIII, male, dorsal view; 8) Sternum VII, male, ventral view; 9) Terga IX+X, male, dorsal view; 10) Aedeagus, lateral view, paramere outlined by dashed line; 11) Sternum VII, female, ventral view; 12) Abdominal segments IX (right arrow) + X (left arrow), female." figureDoi="http://doi.org/10.5281/zenodo.5380854" httpUri="https://zenodo.org/record/5380854/files/figure.png" pageId="4" pageNumber="169">Fig. 10</figureCitation>
);
<collectingCountry id="73A99BFE7972B3789E01CABAFC878595" box="[789,846,902,923]" name="Brazil" pageId="4" pageNumber="169">Brazil</collectingCountry>
............................................. .........
<emphasis id="39CA077C7972B3789E71CA9CFB6085BB" bold="true" box="[869,1193,928,949]" pageId="4" pageNumber="169">
<taxonomicName id="CCBEA0ED7972B3789E71CA9CFBE085BB" authority="Bortoluzzi &amp; Caron, 2018" authorityName="Bortoluzzi &amp; Caron" authorityYear="2018" box="[869,1065,928,949]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="hoppi" status="sp. nov.">
<emphasis id="39CA077C7972B3789E71CA9CFBE085BB" bold="true" box="[869,1065,928,949]" italics="true" pageId="4" pageNumber="169">Gnathymenus hoppi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="22F9BA077972B3789922CA9CFB6085BB" box="[1078,1193,928,949]" pageId="4" pageNumber="169" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="0B01DB6E7972B3789FD7CA87FB6085DE" blockId="4.[707,1194,609,1162]" box="[707,1193,955,976]" pageId="4" pageNumber="169">- Aedeagus not as described above .... 73B</paragraph>
<paragraph id="0B01DB6E7972B3789FD0CAE9FB608234" blockId="4.[707,1194,609,1162]" pageId="4" pageNumber="169">
73B. Aedeagus, in lateral view, with short apical and longer subapical hooked processes [see Herman, fig. 189];
<collectingCountry id="73A99BFE7972B378995CCD36FB608211" box="[1096,1193,1034,1055]" name="Colombia" pageId="4" pageNumber="169">Colombia</collectingCountry>
. .. .. .. .. ... .. .. .. ..
<taxonomicName id="CCBEA0ED7972B3789EDCCD19FB608234" authorityName="Herman" authorityYear="1981" box="[968,1193,1061,1082]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="speccus">
<emphasis id="39CA077C7972B3789EDCCD19FB608234" bold="true" box="[968,1193,1061,1082]" italics="true" pageId="4" pageNumber="169">Gnathymenus speccus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="0B01DB6E7972B3789FD7CD03FB608284" blockId="4.[707,1194,609,1162]" pageId="4" pageNumber="169">
- Aedeagus gradually tapered to acute apex [see
<bibRefCitation id="6F2FA69F7972B3789E44CD66FC2B8261" author="Herman, L. H." box="[848,994,1114,1135]" pageId="4" pageNumber="169" refId="ref2283" refString="Herman, L. H. 1981. Revision of the subtribe Dolicaonina of the New World, with discussions of phylogeny" type="book" year="1981">Herman 1981</bibRefCitation>
, figs. 141, 142];
<collectingCountry id="73A99BFE7972B3789E01CD49FCBA8284" box="[789,883,1141,1162]" name="Colombia" pageId="4" pageNumber="169">Colombia</collectingCountry>
...........
<taxonomicName id="CCBEA0ED7972B3789EF4CD49FB608284" authorityName="Herman" authorityYear="1981" box="[992,1193,1141,1162]" class="Insecta" family="Staphylinidae" genus="Gnathymenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="garus">
<emphasis id="39CA077C7972B3789EF4CD49FB608284" bold="true" box="[992,1193,1141,1162]" italics="true" pageId="4" pageNumber="169">Gnathymenus garus</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>