220 lines
36 KiB
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220 lines
36 KiB
XML
<document ID-DOI="10.11646/zootaxa.5239.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7634797" ID-ZooBank="BF1FF837-56D5-4829-8D46-E821D9D31AB3" checkinTime="1676281666712" checkinUser="plazi" docAuthor="Framenau, Volker W., Castanheira, Pedro De S. & Yoo, Jung-Sun" docDate="2023" docId="193AC81C0004FFFCFF3CFF1F1DAD6695" docLanguage="en" docName="zootaxa.5239.3.1.pdf" docOrigin="Zootaxa 5239 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5239.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Artoriinae Framenau 2007" docType="key" docVersion="3" lastPageNumber="307" masterDocId="E503B0640002FFFAFFABFF8818306231" masterDocTitle="The artoriine wolf spiders of Australia: the new genus Kochosa and a key to genera (Araneae: Lycosidae)" masterLastPageNumber="357" masterPageNumber="301" pageNumber="307" updateTime="1676313737328" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:titleInfo>
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<mods:title>The artoriine wolf spiders of Australia: the new genus Kochosa and a key to genera (Araneae: Lycosidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Framenau, Volker W.</mods:namePart>
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<mods:affiliation>Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150. & Department of Terrestrial Zoology, Western Australian Museum, Welshpool DC, Western Australia 6986, Australia & Zoological Museum Hamburg, Leibnitz Institute for the Analysis of Biodiversity Change (LIB), Centre for Taxonomy & Morphology,</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Castanheira, Pedro De S.</mods:namePart>
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<mods:affiliation>Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150. & https // orcid. org / 0000 - 0002 - 0623 - 1622</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Yoo, Jung-Sun</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-3243-2006</mods:nameIdentifier>
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<mods:affiliation>Animal Resources Division, National Institute of Biological Resources, Environmental Research Complex, Incheon, 22689, Korea https // orcid. org / 0000 - 0002 - 3243 - 2006</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-10</mods:number>
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<mods:detail type="volume">
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<mods:number>5239</mods:number>
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<mods:number>3</mods:number>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5239.3.1</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5239.3.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:193AC81C0004FFFCFF3CFF1F1DAD6695" httpUri="http://treatment.plazi.org/id/193AC81C0004FFFCFF3CFF1F1DAD6695" lastPageNumber="307" pageId="6" pageNumber="307">
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Key to the genera of
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<taxonomicName authority="Framenau, 2007" authorityName="Framenau" authorityYear="2007" box="[409,541,151,177]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="307" phylum="Arthropoda" rank="subFamily" subFamily="Artoriinae">Artoriinae</taxonomicName>
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in
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<collectingCountry box="[579,696,151,177]" name="Australia" pageId="6" pageNumber="307">Australia</collectingCountry>
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</emphasis>
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<subSubSection pageId="6" pageNumber="307" type="discussion">
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<paragraph blockId="6.[151,1437,223,538]" pageId="6" pageNumber="307">
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Male
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<taxonomicName authorityName="Framenau" authorityYear="2007" box="[215,330,223,249]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="307" phylum="Arthropoda" rank="subFamily" subFamily="Artoriinae">Artoriinae</taxonomicName>
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differ from those of the other three Australian subfamilies of wolf spiders, Lycosinae, Venoniinae and Zoicinae by the presence of a basoembolic apophysis in the male pedipalp (Framenau 2007). There is no character or character combination currently known that unequivocally identifies females to be part of the subfamily. However,
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<taxonomicName authorityName="Framenau" authorityYear="2007" box="[267,382,331,357]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="307" phylum="Arthropoda" rank="subFamily" subFamily="Artoriinae">Artoriinae</taxonomicName>
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lack the elongated basal segment of the posteriors spinnerets of the Venoniinae (i.e.,
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<bibRefCitation author="Framenau, V. W. & Yoo, J. - S." pageId="6" pageNumber="307" pagination="185 - 202" refId="ref35396" refString="Framenau, V. W. & Yoo, J. - S. (2006) Systematics of the new Australian wolf spider genus Tuberculosa (Araneae, Lycosidae). Invertebrate Systematics, 20, 185 - 202. https: // doi. org / 10.1071 / IS 05036" type="journal article" year="2006">Yoo & Framenau 2006</bibRefCitation>
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,
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<bibRefCitation author="Framenau, V. W." box="[341,530,367,394]" pageId="6" pageNumber="307" pagination="1 - 20" refId="ref34878" refString="Framenau, V. W. (2006 a) Revision of the Australian wolf spider genus Anomalosa Roewer, 1960 (Araneae: Lycosidae). Zootaxa, 1304 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 1304.1.1" type="journal article" year="2006">Framenau 2006a</bibRefCitation>
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) and are larger than the members of the Zoicinae which are less than
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<quantity box="[1314,1379,367,394]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="307" unit="mm" value="2.0">2 mm</quantity>
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long (i.e.,
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<bibRefCitation author="McKay, R. J." box="[213,368,403,430]" pageId="6" pageNumber="307" pagination="231 - 235" refId="ref36499" refString="McKay, R. J. (1979) The wolf spiders of Australia (Araneae: Lycosidae): 10. A new species of the genus Flanona Simon. Memoirs of the Queensland Museum, 19, 231 - 235." type="journal article" year="1979">McKay 1979</bibRefCitation>
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).
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<taxonomicName authorityName="Framenau" authorityYear="2007" box="[389,504,403,429]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="307" phylum="Arthropoda" rank="subFamily" subFamily="Artoriinae">Artoriinae</taxonomicName>
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generally differ from the Lycosinae by smaller size (but sizes overlap) and the presence of a light, lanceolate median cardiac mark on the abdomen, whereas Lycosinae are much more varied in the abdominal colouration. In
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<collectingCountry box="[485,591,475,501]" name="Australia" pageId="6" pageNumber="307">Australia</collectingCountry>
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, most
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<taxonomicName authorityName="Framenau" authorityYear="2007" box="[660,775,475,501]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="307" phylum="Arthropoda" rank="subFamily" subFamily="Artoriinae">Artoriinae</taxonomicName>
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are vagrant, often diurnal hunters, whereas most Lycosinae live in burrows and are nocturnal.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="307" type="key">
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<keyStep pageId="6" pageNumber="307">
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<paragraph blockId="6.[151,1437,580,1189]" pageId="6" pageNumber="307">
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<keyLead pageId="6" pageNumber="307">
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1. Tegular apophysis of the male pedipalp a more or less basally bent, short hook (
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<figureCitation box="[988,1065,580,602]" captionStart="FIGURE 2" captionStartId="7.[151,250,1888,1912]" captionTargetBox="[163,1418,190,1849]" captionTargetId="figure-17@7.[150,1436,181,1864]" captionTargetPageId="7" captionText="FIGURE 2A–I. Key to the genera of Australian Artoriinae Framenau, 2007. A, Tetralycosa oraria (L. Koch, 1876), male pedipalp ventral; B, T. alteripa (McKay, 1976), epigyne ventral; C, T. eyrei (Hickman, 1944), epigyne ventral; D, T. arabanae Framenau, Gotch & Austin, 2006, epigyne ventral; E, T. oraria, epigyne ventral; F, G, generic frontal views of Lycosidae; H, Kochosa australia sp. nov., male dorsal view; I, K. timwintoni sp. nov., male dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7634803" httpUri="https://zenodo.org/record/7634803/files/figure.png" pageId="6" pageNumber="307">Fig. 2A</figureCitation>
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), epigyne with two anterior lobes (
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<figureCitation captionStart="FIGURE 2" captionStartId="7.[151,250,1888,1912]" captionTargetBox="[163,1418,190,1849]" captionTargetId="figure-17@7.[150,1436,181,1864]" captionTargetPageId="7" captionText="FIGURE 2A–I. Key to the genera of Australian Artoriinae Framenau, 2007. A, Tetralycosa oraria (L. Koch, 1876), male pedipalp ventral; B, T. alteripa (McKay, 1976), epigyne ventral; C, T. eyrei (Hickman, 1944), epigyne ventral; D, T. arabanae Framenau, Gotch & Austin, 2006, epigyne ventral; E, T. oraria, epigyne ventral; F, G, generic frontal views of Lycosidae; H, Kochosa australia sp. nov., male dorsal view; I, K. timwintoni sp. nov., male dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7634803" httpUri="https://zenodo.org/record/7634803/files/figure.png" pageId="6" pageNumber="307">Fig. 2B</figureCitation>
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), an open cavity with anterior notch (
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<figureCitation box="[619,688,609,631]" captionStart="FIGURE 2" captionStartId="7.[151,250,1888,1912]" captionTargetBox="[163,1418,190,1849]" captionTargetId="figure-17@7.[150,1436,181,1864]" captionTargetPageId="7" captionText="FIGURE 2A–I. Key to the genera of Australian Artoriinae Framenau, 2007. A, Tetralycosa oraria (L. Koch, 1876), male pedipalp ventral; B, T. alteripa (McKay, 1976), epigyne ventral; C, T. eyrei (Hickman, 1944), epigyne ventral; D, T. arabanae Framenau, Gotch & Austin, 2006, epigyne ventral; E, T. oraria, epigyne ventral; F, G, generic frontal views of Lycosidae; H, Kochosa australia sp. nov., male dorsal view; I, K. timwintoni sp. nov., male dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7634803" httpUri="https://zenodo.org/record/7634803/files/figure.png" pageId="6" pageNumber="307">Fig 2C</figureCitation>
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) or as in
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<figureCitation box="[779,886,609,631]" captionStart="FIGURE 2" captionStartId="7.[151,250,1888,1912]" captionTargetBox="[163,1418,190,1849]" captionTargetId="figure-17@7.[150,1436,181,1864]" captionTargetPageId="7" captionText="FIGURE 2A–I. Key to the genera of Australian Artoriinae Framenau, 2007. A, Tetralycosa oraria (L. Koch, 1876), male pedipalp ventral; B, T. alteripa (McKay, 1976), epigyne ventral; C, T. eyrei (Hickman, 1944), epigyne ventral; D, T. arabanae Framenau, Gotch & Austin, 2006, epigyne ventral; E, T. oraria, epigyne ventral; F, G, generic frontal views of Lycosidae; H, Kochosa australia sp. nov., male dorsal view; I, K. timwintoni sp. nov., male dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7634803" httpUri="https://zenodo.org/record/7634803/files/figure.png" pageId="6" pageNumber="307">Figs 2D–E</figureCitation>
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; all species with a beige to light brown basic colouration although some body parts can be darker (
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<figureCitation box="[632,743,639,661]" captionStart="FIGURE 1" captionStartId="4.[151,250,1820,1844]" captionTargetBox="[183,1405,179,1794]" captionTargetId="figure-24@4.[183,1405,179,1794]" captionTargetPageId="4" captionText="FIGURE 1A–H. Life images of members of the subfamily Artoriinae Framenau, 2007. A, Artoria cingulipes Simon, 1909, male (Western Australia); B, Artoriopsis expolita (L. Koch, 1877), female (Western Australia); C, Diahogna martensii (Karsch, 1878), female (Tasmania); D, Kangarosa tristicula (L. Koch, 1877), female (no live image for genus available); E, Tetralycosa oraria (L. Koch, 1876), male (Western Australia); F, Tetralycosa baudinettei Framenau & Hudson, 2017, male (Western Australia); G, Kochosa australia sp. nov., female (Victoria); H, same species, male (Victoria) (both MV K-14336). All images © V.W. Framenau, except C, John Douglas; G, H, Joseph Schubert." figureDoi="http://doi.org/10.5281/zenodo.7634799" httpUri="https://zenodo.org/record/7634799/files/figure.png" pageId="6" pageNumber="307">Figs 1C–D</figureCitation>
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); all species in genus halophile (salt lakes and pans, samphire, coastal, Artesian springs).............................................................................
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<taxonomicName authorityName="Roewer" authorityYear="1960" box="[1324,1436,668,690]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[1324,1436,668,690]" italics="true" pageId="6" pageNumber="307">Tetralycosa</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph blockId="6.[151,1437,580,1189]" pageId="6" pageNumber="307">
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<keyLead pageId="6" pageNumber="307">
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- Tegular apophysis of variable shape, if with terminal hook (e.g.,
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<figureCitation box="[846,921,697,719]" captionStart="FIGURE 3" captionStartId="8.[152,255,1806,1830]" captionTargetBox="[292,1310,189,1765]" captionTargetId="figure-24@8.[278,1310,181,1782]" captionTargetPageId="8" captionText="FIGURE 3A–L. Key to the genera of Australian Artoriinae Framenau, 2007. A, Artoriopsis expolita, male, dorsal; B, A. bogabilla Framenau & Douglas, 2021, male, dorsal; Artoria cingulipes Simon, 1909, male, dorsal; D, Kangarosa tristicula (L. Koch, 1877), male pedipalp, ventral; E, Diahogna exculta (L. Koch, 1876), male pedipalp, ventral; F, D. martensii (Karsch, 1878), male pedipalp, ventral;G, K. tristicula, epigyne ventral;H, K. ludwigi Framenau, 2010; epigyne ventral; I, K. alboguttulata (L. Koch, 1878), epigyne ventral; J, D. exculta, epigyne ventral; D. hildegardae Framenau, 2006, epigyne ventral; D. martensii, epigyne ventral." figureDoi="http://doi.org/10.5281/zenodo.7634805" httpUri="https://zenodo.org/record/7634805/files/figure.png" pageId="6" pageNumber="307">Fig. 3D</figureCitation>
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), base longer; epigyne without two anterior lobes and not as in
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<figureCitation box="[313,418,727,749]" captionStart="FIGURE 2" captionStartId="7.[151,250,1888,1912]" captionTargetBox="[163,1418,190,1849]" captionTargetId="figure-17@7.[150,1436,181,1864]" captionTargetPageId="7" captionText="FIGURE 2A–I. Key to the genera of Australian Artoriinae Framenau, 2007. A, Tetralycosa oraria (L. Koch, 1876), male pedipalp ventral; B, T. alteripa (McKay, 1976), epigyne ventral; C, T. eyrei (Hickman, 1944), epigyne ventral; D, T. arabanae Framenau, Gotch & Austin, 2006, epigyne ventral; E, T. oraria, epigyne ventral; F, G, generic frontal views of Lycosidae; H, Kochosa australia sp. nov., male dorsal view; I, K. timwintoni sp. nov., male dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7634803" httpUri="https://zenodo.org/record/7634803/files/figure.png" pageId="6" pageNumber="307">Figs 2B–E</figureCitation>
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; not halophile........................................................................ 2
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="6" pageNumber="307">
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<paragraph blockId="6.[151,1437,580,1189]" box="[151,1436,756,778]" pageId="6" pageNumber="307">
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<keyLead box="[151,1436,756,778]" pageId="6" pageNumber="307">
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2. Row of anterior eyes narrower than row of posterior median eyes (
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<figureCitation box="[865,936,756,778]" captionStart="FIGURE 2" captionStartId="7.[151,250,1888,1912]" captionTargetBox="[163,1418,190,1849]" captionTargetId="figure-17@7.[150,1436,181,1864]" captionTargetPageId="7" captionText="FIGURE 2A–I. Key to the genera of Australian Artoriinae Framenau, 2007. A, Tetralycosa oraria (L. Koch, 1876), male pedipalp ventral; B, T. alteripa (McKay, 1976), epigyne ventral; C, T. eyrei (Hickman, 1944), epigyne ventral; D, T. arabanae Framenau, Gotch & Austin, 2006, epigyne ventral; E, T. oraria, epigyne ventral; F, G, generic frontal views of Lycosidae; H, Kochosa australia sp. nov., male dorsal view; I, K. timwintoni sp. nov., male dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7634803" httpUri="https://zenodo.org/record/7634803/files/figure.png" pageId="6" pageNumber="307">Fig. 2F</figureCitation>
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)....................................... 3
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</keyLead>
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</paragraph>
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<paragraph blockId="6.[151,1437,580,1189]" box="[151,1436,785,807]" pageId="6" pageNumber="307">
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<keyLead box="[151,1436,785,807]" pageId="6" pageNumber="307">
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- Row of anterior eyes as wide or wider than row of posterior median eyes (
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<figureCitation box="[938,1013,785,807]" captionStart="FIGURE 2" captionStartId="7.[151,250,1888,1912]" captionTargetBox="[163,1418,190,1849]" captionTargetId="figure-17@7.[150,1436,181,1864]" captionTargetPageId="7" captionText="FIGURE 2A–I. Key to the genera of Australian Artoriinae Framenau, 2007. A, Tetralycosa oraria (L. Koch, 1876), male pedipalp ventral; B, T. alteripa (McKay, 1976), epigyne ventral; C, T. eyrei (Hickman, 1944), epigyne ventral; D, T. arabanae Framenau, Gotch & Austin, 2006, epigyne ventral; E, T. oraria, epigyne ventral; F, G, generic frontal views of Lycosidae; H, Kochosa australia sp. nov., male dorsal view; I, K. timwintoni sp. nov., male dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7634803" httpUri="https://zenodo.org/record/7634803/files/figure.png" pageId="6" pageNumber="307">Fig. 2G</figureCitation>
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)................................. 5
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="6" pageNumber="307">
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<paragraph blockId="6.[151,1437,580,1189]" pageId="6" pageNumber="307">
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<keyLead pageId="6" pageNumber="307">
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3. Abdomen generally uniformly dark with a very distinct light cardiac mark (sometimes reduced to two distinct spots) that is surrounded posteriorly by dark margin increasing the contrast of the cardiac mark (
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<figureCitation box="[1023,1132,844,866]" captionStart="FIGURE 1" captionStartId="4.[151,250,1820,1844]" captionTargetBox="[183,1405,179,1794]" captionTargetId="figure-24@4.[183,1405,179,1794]" captionTargetPageId="4" captionText="FIGURE 1A–H. Life images of members of the subfamily Artoriinae Framenau, 2007. A, Artoria cingulipes Simon, 1909, male (Western Australia); B, Artoriopsis expolita (L. Koch, 1877), female (Western Australia); C, Diahogna martensii (Karsch, 1878), female (Tasmania); D, Kangarosa tristicula (L. Koch, 1877), female (no live image for genus available); E, Tetralycosa oraria (L. Koch, 1876), male (Western Australia); F, Tetralycosa baudinettei Framenau & Hudson, 2017, male (Western Australia); G, Kochosa australia sp. nov., female (Victoria); H, same species, male (Victoria) (both MV K-14336). All images © V.W. Framenau, except C, John Douglas; G, H, Joseph Schubert." figureDoi="http://doi.org/10.5281/zenodo.7634799" httpUri="https://zenodo.org/record/7634799/files/figure.png" pageId="6" pageNumber="307">Figs 1G, H</figureCitation>
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,
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<figureCitation box="[1142,1192,844,865]" captionStart="FIGURE 2" captionStartId="7.[151,250,1888,1912]" captionTargetBox="[163,1418,190,1849]" captionTargetId="figure-17@7.[150,1436,181,1864]" captionTargetPageId="7" captionText="FIGURE 2A–I. Key to the genera of Australian Artoriinae Framenau, 2007. A, Tetralycosa oraria (L. Koch, 1876), male pedipalp ventral; B, T. alteripa (McKay, 1976), epigyne ventral; C, T. eyrei (Hickman, 1944), epigyne ventral; D, T. arabanae Framenau, Gotch & Austin, 2006, epigyne ventral; E, T. oraria, epigyne ventral; F, G, generic frontal views of Lycosidae; H, Kochosa australia sp. nov., male dorsal view; I, K. timwintoni sp. nov., male dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7634803" httpUri="https://zenodo.org/record/7634803/files/figure.png" pageId="6" pageNumber="307">2H, I</figureCitation>
|
||
)....
|
||
<taxonomicName authority="Framenau & Castanheira & Yoo, 2023" authorityName="Framenau & Castanheira & Yoo" authorityYear="2023" box="[1258,1343,844,866]" class="Arachnida" family="Lycosidae" genus="Kochosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis bold="true" box="[1258,1343,844,866]" italics="true" pageId="6" pageNumber="307">Kochosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1349,1437,845,866]" pageId="6" pageNumber="307" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,580,1189]" box="[151,1436,873,895]" pageId="6" pageNumber="307">
|
||
<keyLead box="[151,1436,873,895]" pageId="6" pageNumber="307">- Light cardiac mark of abdomen not as distinct and not surrounded by dark margin/long setae posteriorly................ 4</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="307">
|
||
<paragraph blockId="6.[151,1437,580,1189]" pageId="6" pageNumber="307">
|
||
<keyLead pageId="6" pageNumber="307">
|
||
4. Carapace with distinct light lateral bands that reach to the carapace margin; abdomen often with light cardiac mark that separates a central, generally diamond-shaped black spot into two triangles and a posterior rectangular or subtriangular pale patch (
|
||
<figureCitation captionStart="FIGURE 1" captionStartId="4.[151,250,1820,1844]" captionTargetBox="[183,1405,179,1794]" captionTargetId="figure-24@4.[183,1405,179,1794]" captionTargetPageId="4" captionText="FIGURE 1A–H. Life images of members of the subfamily Artoriinae Framenau, 2007. A, Artoria cingulipes Simon, 1909, male (Western Australia); B, Artoriopsis expolita (L. Koch, 1877), female (Western Australia); C, Diahogna martensii (Karsch, 1878), female (Tasmania); D, Kangarosa tristicula (L. Koch, 1877), female (no live image for genus available); E, Tetralycosa oraria (L. Koch, 1876), male (Western Australia); F, Tetralycosa baudinettei Framenau & Hudson, 2017, male (Western Australia); G, Kochosa australia sp. nov., female (Victoria); H, same species, male (Victoria) (both MV K-14336). All images © V.W. Framenau, except C, John Douglas; G, H, Joseph Schubert." figureDoi="http://doi.org/10.5281/zenodo.7634799" httpUri="https://zenodo.org/record/7634799/files/figure.png" pageId="6" pageNumber="307">Figs 1B</figureCitation>
|
||
,
|
||
<figureCitation box="[266,297,962,983]" captionStart="FIGURE 3" captionStartId="8.[152,255,1806,1830]" captionTargetBox="[292,1310,189,1765]" captionTargetId="figure-24@8.[278,1310,181,1782]" captionTargetPageId="8" captionText="FIGURE 3A–L. Key to the genera of Australian Artoriinae Framenau, 2007. A, Artoriopsis expolita, male, dorsal; B, A. bogabilla Framenau & Douglas, 2021, male, dorsal; Artoria cingulipes Simon, 1909, male, dorsal; D, Kangarosa tristicula (L. Koch, 1877), male pedipalp, ventral; E, Diahogna exculta (L. Koch, 1876), male pedipalp, ventral; F, D. martensii (Karsch, 1878), male pedipalp, ventral;G, K. tristicula, epigyne ventral;H, K. ludwigi Framenau, 2010; epigyne ventral; I, K. alboguttulata (L. Koch, 1878), epigyne ventral; J, D. exculta, epigyne ventral; D. hildegardae Framenau, 2006, epigyne ventral; D. martensii, epigyne ventral." figureDoi="http://doi.org/10.5281/zenodo.7634805" httpUri="https://zenodo.org/record/7634805/files/figure.png" pageId="6" pageNumber="307">3A</figureCitation>
|
||
), or abdomen uniformly dark olive-grey with an irregular light median band along its whole length (
|
||
<figureCitation box="[1262,1335,961,983]" captionStart="FIGURE 3" captionStartId="8.[152,255,1806,1830]" captionTargetBox="[292,1310,189,1765]" captionTargetId="figure-24@8.[278,1310,181,1782]" captionTargetPageId="8" captionText="FIGURE 3A–L. Key to the genera of Australian Artoriinae Framenau, 2007. A, Artoriopsis expolita, male, dorsal; B, A. bogabilla Framenau & Douglas, 2021, male, dorsal; Artoria cingulipes Simon, 1909, male, dorsal; D, Kangarosa tristicula (L. Koch, 1877), male pedipalp, ventral; E, Diahogna exculta (L. Koch, 1876), male pedipalp, ventral; F, D. martensii (Karsch, 1878), male pedipalp, ventral;G, K. tristicula, epigyne ventral;H, K. ludwigi Framenau, 2010; epigyne ventral; I, K. alboguttulata (L. Koch, 1878), epigyne ventral; J, D. exculta, epigyne ventral; D. hildegardae Framenau, 2006, epigyne ventral; D. martensii, epigyne ventral." figureDoi="http://doi.org/10.5281/zenodo.7634805" httpUri="https://zenodo.org/record/7634805/files/figure.png" pageId="6" pageNumber="307">Fig. 3B</figureCitation>
|
||
)..................................................................................................
|
||
<taxonomicName authorityName="Framenau" authorityYear="2007" box="[1327,1436,991,1012]" class="Arachnida" family="Lycosidae" genus="Artoriopsis" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis bold="true" box="[1327,1436,991,1012]" italics="true" pageId="6" pageNumber="307">Artoriopsis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,580,1189]" pageId="6" pageNumber="307">
|
||
<keyLead pageId="6" pageNumber="307">
|
||
- Carapace without lateral light bands, but generally with an irregular light submarginal band (i.e., there is a dark edge along the carapace margin) (
|
||
<figureCitation box="[403,478,1049,1071]" captionStart="FIGURE 3" captionStartId="8.[152,255,1806,1830]" captionTargetBox="[292,1310,189,1765]" captionTargetId="figure-24@8.[278,1310,181,1782]" captionTargetPageId="8" captionText="FIGURE 3A–L. Key to the genera of Australian Artoriinae Framenau, 2007. A, Artoriopsis expolita, male, dorsal; B, A. bogabilla Framenau & Douglas, 2021, male, dorsal; Artoria cingulipes Simon, 1909, male, dorsal; D, Kangarosa tristicula (L. Koch, 1877), male pedipalp, ventral; E, Diahogna exculta (L. Koch, 1876), male pedipalp, ventral; F, D. martensii (Karsch, 1878), male pedipalp, ventral;G, K. tristicula, epigyne ventral;H, K. ludwigi Framenau, 2010; epigyne ventral; I, K. alboguttulata (L. Koch, 1878), epigyne ventral; J, D. exculta, epigyne ventral; D. hildegardae Framenau, 2006, epigyne ventral; D. martensii, epigyne ventral." figureDoi="http://doi.org/10.5281/zenodo.7634805" httpUri="https://zenodo.org/record/7634805/files/figure.png" pageId="6" pageNumber="307">Fig. 3C</figureCitation>
|
||
); abdomen with a yellow to yellow-brown cardiac mark, sometimes indistinct, but never with central diamond shaped spot or without irregular light median band along its whole length (
|
||
<figureCitation box="[1033,1113,1079,1101]" captionStart="FIGURE 1" captionStartId="4.[151,250,1820,1844]" captionTargetBox="[183,1405,179,1794]" captionTargetId="figure-24@4.[183,1405,179,1794]" captionTargetPageId="4" captionText="FIGURE 1A–H. Life images of members of the subfamily Artoriinae Framenau, 2007. A, Artoria cingulipes Simon, 1909, male (Western Australia); B, Artoriopsis expolita (L. Koch, 1877), female (Western Australia); C, Diahogna martensii (Karsch, 1878), female (Tasmania); D, Kangarosa tristicula (L. Koch, 1877), female (no live image for genus available); E, Tetralycosa oraria (L. Koch, 1876), male (Western Australia); F, Tetralycosa baudinettei Framenau & Hudson, 2017, male (Western Australia); G, Kochosa australia sp. nov., female (Victoria); H, same species, male (Victoria) (both MV K-14336). All images © V.W. Framenau, except C, John Douglas; G, H, Joseph Schubert." figureDoi="http://doi.org/10.5281/zenodo.7634799" httpUri="https://zenodo.org/record/7634799/files/figure.png" pageId="6" pageNumber="307">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation box="[1123,1151,1079,1100]" captionStart="FIGURE 3" captionStartId="8.[152,255,1806,1830]" captionTargetBox="[292,1310,189,1765]" captionTargetId="figure-24@8.[278,1310,181,1782]" captionTargetPageId="8" captionText="FIGURE 3A–L. Key to the genera of Australian Artoriinae Framenau, 2007. A, Artoriopsis expolita, male, dorsal; B, A. bogabilla Framenau & Douglas, 2021, male, dorsal; Artoria cingulipes Simon, 1909, male, dorsal; D, Kangarosa tristicula (L. Koch, 1877), male pedipalp, ventral; E, Diahogna exculta (L. Koch, 1876), male pedipalp, ventral; F, D. martensii (Karsch, 1878), male pedipalp, ventral;G, K. tristicula, epigyne ventral;H, K. ludwigi Framenau, 2010; epigyne ventral; I, K. alboguttulata (L. Koch, 1878), epigyne ventral; J, D. exculta, epigyne ventral; D. hildegardae Framenau, 2006, epigyne ventral; D. martensii, epigyne ventral." figureDoi="http://doi.org/10.5281/zenodo.7634805" httpUri="https://zenodo.org/record/7634805/files/figure.png" pageId="6" pageNumber="307">3C</figureCitation>
|
||
)................
|
||
<taxonomicName authorityName="Thorell" authorityYear="1877" box="[1364,1436,1079,1100]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis bold="true" box="[1364,1436,1079,1100]" italics="true" pageId="6" pageNumber="307">Artoria</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="307">
|
||
<paragraph blockId="6.[151,1437,580,1189]" pageId="6" pageNumber="307">
|
||
<keyLead pageId="6" pageNumber="307">
|
||
5. Tegular apophysis a long hook (
|
||
<figureCitation box="[532,608,1108,1130]" captionStart="FIGURE 3" captionStartId="8.[152,255,1806,1830]" captionTargetBox="[292,1310,189,1765]" captionTargetId="figure-24@8.[278,1310,181,1782]" captionTargetPageId="8" captionText="FIGURE 3A–L. Key to the genera of Australian Artoriinae Framenau, 2007. A, Artoriopsis expolita, male, dorsal; B, A. bogabilla Framenau & Douglas, 2021, male, dorsal; Artoria cingulipes Simon, 1909, male, dorsal; D, Kangarosa tristicula (L. Koch, 1877), male pedipalp, ventral; E, Diahogna exculta (L. Koch, 1876), male pedipalp, ventral; F, D. martensii (Karsch, 1878), male pedipalp, ventral;G, K. tristicula, epigyne ventral;H, K. ludwigi Framenau, 2010; epigyne ventral; I, K. alboguttulata (L. Koch, 1878), epigyne ventral; J, D. exculta, epigyne ventral; D. hildegardae Framenau, 2006, epigyne ventral; D. martensii, epigyne ventral." figureDoi="http://doi.org/10.5281/zenodo.7634805" httpUri="https://zenodo.org/record/7634805/files/figure.png" pageId="6" pageNumber="307">Fig. 3D</figureCitation>
|
||
); atrium of female epigyne broader anteriorly than posteriorly, without median septum (
|
||
<figureCitation box="[233,337,1137,1159]" captionStart="FIGURE 3" captionStartId="8.[152,255,1806,1830]" captionTargetBox="[292,1310,189,1765]" captionTargetId="figure-24@8.[278,1310,181,1782]" captionTargetPageId="8" captionText="FIGURE 3A–L. Key to the genera of Australian Artoriinae Framenau, 2007. A, Artoriopsis expolita, male, dorsal; B, A. bogabilla Framenau & Douglas, 2021, male, dorsal; Artoria cingulipes Simon, 1909, male, dorsal; D, Kangarosa tristicula (L. Koch, 1877), male pedipalp, ventral; E, Diahogna exculta (L. Koch, 1876), male pedipalp, ventral; F, D. martensii (Karsch, 1878), male pedipalp, ventral;G, K. tristicula, epigyne ventral;H, K. ludwigi Framenau, 2010; epigyne ventral; I, K. alboguttulata (L. Koch, 1878), epigyne ventral; J, D. exculta, epigyne ventral; D. hildegardae Framenau, 2006, epigyne ventral; D. martensii, epigyne ventral." figureDoi="http://doi.org/10.5281/zenodo.7634805" httpUri="https://zenodo.org/record/7634805/files/figure.png" pageId="6" pageNumber="307">Fig. 3G, H</figureCitation>
|
||
) or as in
|
||
<figureCitation box="[427,492,1137,1159]" captionStart="FIGURE 3" captionStartId="8.[152,255,1806,1830]" captionTargetBox="[292,1310,189,1765]" captionTargetId="figure-24@8.[278,1310,181,1782]" captionTargetPageId="8" captionText="FIGURE 3A–L. Key to the genera of Australian Artoriinae Framenau, 2007. A, Artoriopsis expolita, male, dorsal; B, A. bogabilla Framenau & Douglas, 2021, male, dorsal; Artoria cingulipes Simon, 1909, male, dorsal; D, Kangarosa tristicula (L. Koch, 1877), male pedipalp, ventral; E, Diahogna exculta (L. Koch, 1876), male pedipalp, ventral; F, D. martensii (Karsch, 1878), male pedipalp, ventral;G, K. tristicula, epigyne ventral;H, K. ludwigi Framenau, 2010; epigyne ventral; I, K. alboguttulata (L. Koch, 1878), epigyne ventral; J, D. exculta, epigyne ventral; D. hildegardae Framenau, 2006, epigyne ventral; D. martensii, epigyne ventral." figureDoi="http://doi.org/10.5281/zenodo.7634805" httpUri="https://zenodo.org/record/7634805/files/figure.png" pageId="6" pageNumber="307">Fig. 3I</figureCitation>
|
||
.....................................................................
|
||
<taxonomicName authorityName="Framenau" authorityYear="2010" box="[1328,1436,1138,1159]" class="Arachnida" family="Lycosidae" genus="Kangarosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis bold="true" box="[1328,1436,1138,1159]" italics="true" pageId="6" pageNumber="307">Kangarosa</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,580,1189]" box="[151,1437,1166,1189]" pageId="6" pageNumber="307">
|
||
<keyLead box="[151,1437,1166,1189]" pageId="6" pageNumber="307">
|
||
- Tegular apophysis not hook-shaped (
|
||
<figureCitation box="[579,679,1167,1189]" captionStart="FIGURE 3" captionStartId="8.[152,255,1806,1830]" captionTargetBox="[292,1310,189,1765]" captionTargetId="figure-24@8.[278,1310,181,1782]" captionTargetPageId="8" captionText="FIGURE 3A–L. Key to the genera of Australian Artoriinae Framenau, 2007. A, Artoriopsis expolita, male, dorsal; B, A. bogabilla Framenau & Douglas, 2021, male, dorsal; Artoria cingulipes Simon, 1909, male, dorsal; D, Kangarosa tristicula (L. Koch, 1877), male pedipalp, ventral; E, Diahogna exculta (L. Koch, 1876), male pedipalp, ventral; F, D. martensii (Karsch, 1878), male pedipalp, ventral;G, K. tristicula, epigyne ventral;H, K. ludwigi Framenau, 2010; epigyne ventral; I, K. alboguttulata (L. Koch, 1878), epigyne ventral; J, D. exculta, epigyne ventral; D. hildegardae Framenau, 2006, epigyne ventral; D. martensii, epigyne ventral." figureDoi="http://doi.org/10.5281/zenodo.7634805" httpUri="https://zenodo.org/record/7634805/files/figure.png" pageId="6" pageNumber="307">Fig. 3E, F</figureCitation>
|
||
); female epigyne as in
|
||
<figureCitation box="[901,996,1167,1189]" captionStart="FIGURE 3" captionStartId="8.[152,255,1806,1830]" captionTargetBox="[292,1310,189,1765]" captionTargetId="figure-24@8.[278,1310,181,1782]" captionTargetPageId="8" captionText="FIGURE 3A–L. Key to the genera of Australian Artoriinae Framenau, 2007. A, Artoriopsis expolita, male, dorsal; B, A. bogabilla Framenau & Douglas, 2021, male, dorsal; Artoria cingulipes Simon, 1909, male, dorsal; D, Kangarosa tristicula (L. Koch, 1877), male pedipalp, ventral; E, Diahogna exculta (L. Koch, 1876), male pedipalp, ventral; F, D. martensii (Karsch, 1878), male pedipalp, ventral;G, K. tristicula, epigyne ventral;H, K. ludwigi Framenau, 2010; epigyne ventral; I, K. alboguttulata (L. Koch, 1878), epigyne ventral; J, D. exculta, epigyne ventral; D. hildegardae Framenau, 2006, epigyne ventral; D. martensii, epigyne ventral." figureDoi="http://doi.org/10.5281/zenodo.7634805" httpUri="https://zenodo.org/record/7634805/files/figure.png" pageId="6" pageNumber="307">Figs 3J–L</figureCitation>
|
||
............................
|
||
<taxonomicName authorityName="Roewer" authorityYear="1960" box="[1338,1437,1166,1188]" class="Arachnida" family="Lycosidae" genus="Diahogna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis bold="true" box="[1338,1437,1166,1188]" italics="true" pageId="6" pageNumber="307">Diahogna</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |