536 lines
57 KiB
XML
536 lines
57 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1077.67081" ID-GBIF-Dataset="66b827c0-65ac-44a6-90e0-b78cd9ef81c2" ID-PMC="PMC8692307" ID-Pensoft-Pub="1313-2970-1077-1" ID-Pensoft-UUID="734088641608531C8E2CC69397B000ED" ID-PubMed="35027858" ID-ZooBank="C28AD65A76F242CFBED7DFB3702CABCE" ModsDocID="1313-2970-1077-1" checkinTime="1639570129388" checkinUser="pensoft" docAuthor="De Mattia, Willy, Reier, Susanne & Haring, Elisabeth" docDate="2021" docId="BD9C4051F7C05EBA890AA01BD7B8CEBC" docLanguage="en" docName="ZooKeys 1077: 1-175" docOrigin="ZooKeys 1077" docPubDate="2021-12-14" docSource="http://dx.doi.org/10.3897/zookeys.1077.67081" docTitle="Sicania nobilis subsp. spezialensis , stat. nov. 2021, comb. nov." docType="treatment" docVersion="4" id="734088641608531C8E2CC69397B000ED" lastPageNumber="1" masterDocId="734088641608531C8E2CC69397B000ED" masterDocTitle="Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="1" updateTime="1668151235153" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>De Mattia, Willy</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0056-467X</mods:nameIdentifier>
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<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:nameIdentifier type="email">willy.demattia@icgeb.org</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reier, Susanne</mods:namePart>
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<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Haring, Elisabeth</mods:namePart>
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<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-12-14</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1077</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>175</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1077.67081</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1077.67081</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1077-1</mods:identifier>
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<mods:identifier type="ZooBank">C28AD65A76F242CFBED7DFB3702CABCE</mods:identifier>
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<mods:identifier type="Pensoft-UUID">734088641608531C8E2CC69397B000ED</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="191645930" LSID="urn:lsid:plazi:treatment:BD9C4051F7C05EBA890AA01BD7B8CEBC" httpUri="http://treatment.plazi.org/id/BD9C4051F7C05EBA890AA01BD7B8CEBC" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="BD9C4051-F7C0-5EBA-890A-A01BD7B8CEBC" authority="(Nordsieck, 1984), stat. nov." authorityName=", stat. nov." authorityYear="2021" baseAuthorityName="Nordsieck" baseAuthorityYear="1984" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis subsp. spezialensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="nobilis" status="comb. nov." subSpecies="spezialensis">Sicania nobilis spezialensis (Nordsieck, 1984), stat. nov.</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">comb. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Wagner's (1913, 1925) and Brandt's (1961) Siciliaria genital drawings A Siciliaria septemplicata, Wagner, 1913: plate 572, fig. 14 B Siciliaria grohmanniana grohmanniana, Wagner, 1925: plate 1, fig. 8 C Siciliaria calcarae calcarae, 1925: plate 3, fig. 25 D Siciliaria ferrox, Brandt, 1961: fig. 1 E overall Siciliaria / Sicania and Sicilian Mauritanica distribution F Siciliaria / Sicania and Sicilian Mauritanica distribution of taxa." figureDoi="10.3897/zookeys.1077.67081.figure1" httpUri="https://binary.pensoft.net/fig/622793" pageId="0" pageNumber="1">Figs 1.F, 35.1-35.7, 36.14-36.16</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria (Siciliaria) spezialensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="spezialensis" status="comb. nov." subGenus="Siciliaria">Siciliaria (Siciliaria) spezialensis</taxonomicName>
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<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" refId="B61" refString="Nordsieck, H, 1984. Neue Taxa rezenter europaeischer Clausilien, mit Bemerkungen zur Bastardierung bei Clausilien (Gastropoda: Clausiliidae). Archiv fuer Molluskenkunde 114(4/6): 189-211." title="Neue Taxa rezenter europaeischer Clausilien, mit Bemerkungen zur Bastardierung bei Clausilien (Gastropoda: Clausiliidae). Archiv fuer Molluskenkunde 114 (4 / 6): 189 - 211." year="1984">Nordsieck 1984</bibRefCitation>
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: 202.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria (Siciliaria) spezialensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="spezialensis" status="comb. nov." subGenus="Siciliaria">Siciliaria (Siciliaria) spezialensis</taxonomicName>
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-
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<bibRefCitation author="Manganelli, G" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" refId="B43" refString="Manganelli, G, Bodon, M, Favilli, L, Giusti, F, 1995. Fascicolo 16, Gastropoda Pulmonta. In: Minelli A, Ruffo S, La Posta S (Eds) Checklist delle specie della fauna italiana 16: 1-60. Calderini, Bologna." title="Fascicolo 16, Gastropoda Pulmonta. In: Minelli A, Ruffo S, La Posta S (Eds) Checklist delle specie della fauna italiana 16: 1 - 60. Calderini, Bologna." year="1995">Manganelli et al. 1995</bibRefCitation>
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: 25.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria spezialensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="spezialensis" status="comb. nov.">Charpentieria spezialensis</taxonomicName>
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-
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<bibRefCitation DOI="https://doi.org/10.1127/arch.moll/133/2004/185" author="Beckmann, KH" journalOrPublisher="Basteria" pageId="0" pageNumber="1" refId="B4" refString="Beckmann, KH, 2004. Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133(1/2): 185-191. https://doi.org/10.1127/arch.moll/133/2004/185" title="Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133 (1 / 2): 185 - 191." url="https://doi.org/10.1127/arch.moll/133/2004/185" year="2004">Beckmann 2004</bibRefCitation>
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: 188.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria spezialensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="spezialensis" status="comb. nov.">Siciliaria spezialensis</taxonomicName>
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-
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<bibRefCitation author="Welter-Schultes, F" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="1" refId="B101" refString="Welter-Schultes, F, 2012. European non-marine molluscs, a quide for species identification. Planet Poster Editions, Goettingen" title="European non-marine molluscs, a quide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
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: 343.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria spezialensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="spezialensis" status="comb. nov.">Siciliaria spezialensis</taxonomicName>
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-
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<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck 2013b</bibRefCitation>
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: 9.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria spezialensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="spezialensis" status="comb. nov.">Charpentieria spezialensis</taxonomicName>
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-
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<bibRefCitation author="De Mattia, W" journalOrPublisher="Molecular Ecology Resources" pageId="0" pageNumber="1" publicationUrl="https://www.iucnredlist.org/species/170841/1317462" refId="B23" refString="De Mattia, W, 2017j. Charpentieria spezialensis. The IUCN Red List of Threatened Species. https://www.iucnredlist.org/species/170841/1317462" title="Charpentieria spezialensis. The IUCN Red List of Threatened Species." url="https://www.iucnredlist.org/species/170841/1317462" year="2017 j">De Mattia 2017j</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation accessionNumber="MW758910, MW757039, MW757040, MW758911" collectingDate="2017-04-14" collectorName="Macari, E, Lab ID, W. De Mattia, J. Macor" country="Italy" elevation="75" latitude="38.12827" location="Monte Speziale" longLatPrecision="1" longitude="12.7359" municipality="San Vito Lo Capo" specimenCount="1" stateProvince="Sicily">
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<collectingCountry name="Italy">Italy</collectingCountry>
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,
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<collectingRegion country="Italy" name="Sicilia">Sicily</collectingRegion>
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,
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<collectingMunicipality>San Vito Lo Capo</collectingMunicipality>
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,
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<collectorName>Macari, E</collectorName>
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side of
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<location LSID="urn:lsid:plazi:treatment:BD9C4051F7C05EBA890AA01BD7B8CEBC:71984345B38E6DDA2D4F0B1E467E32E1" country="Italy" latitude="38.12827" longLatPrecision="1" longitude="12.7359" municipality="San Vito Lo Capo" name="Monte Speziale" stateProvince="Sicily">Monte Speziale</location>
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,
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<elevation metricMagnitude="1" metricUnit="m" metricValue="7.5" unit="m" value="75.0">
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<quantity metricMagnitude="1" metricUnit="m" metricValue="7.5" unit="m" value="75.0">75 m</quantity>
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asl
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</elevation>
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,
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<geoCoordinate degrees="38" direction="north" minutes="07" orientation="latitude" precision="1" seconds="41.78" value="38.12827">38°07'41.78"N</geoCoordinate>
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,
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<geoCoordinate degrees="12" direction="east" minutes="44" orientation="longitude" precision="1" seconds="9.24" value="12.7359">12°44'9.24"E</geoCoordinate>
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, [
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<collectorName>Lab ID</collectorName>
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63_1, COI:
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW758910">MW758910</accessionNumber>
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, ITS2:
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<accessionNumber isEnumeration="true">MW757039, MW757040</accessionNumber>
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;
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<collectorName>Lab ID</collectorName>
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63_2, COI:
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW758911">MW758911</accessionNumber>
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],
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<collectorName>W. De Mattia</collectorName>
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and
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<collectorName>J. Macor</collectorName>
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leg.,
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<collectingDate value="2017-04-14">14.iv.2017</collectingDate>
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. 4 dissected spm
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="shell">
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<paragraph pageId="0" pageNumber="1">Shell</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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(Figs
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<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Sicania nobilis nobilis (A. Schmidt, 1868), comb. nov., Castelluzzo 36.1 shell 36.2 detail of the aperture 36.3 clausiliar plate double side. San Vito Lo capo, camping 36.4 shell 36.5 detail of the aperture 36.6 clausiliar plate double side. San Vito Lo Capo, boulders 36.7 shell 36.8 detail of the aperture 36.9 clausiliar plate double side. Torre delle Usciere (ex. forma Sicania nobilis nobilis episoma) 36.10 shell 36.11 detail of the aperture 36.12 clausiliar plate double side. Monte Cofano, Tonnara Cofano 36.13 shell. Sicania nobilis spezialensis (Nordsieck 1984), stat. nov., comb. nov. Macari 36.14 shell 36.15 detail of the aperture 36.16 clausiliar plate double side." figureDoi="10.3897/zookeys.1077.67081.figure36" httpUri="https://binary.pensoft.net/fig/622828" pageId="0" pageNumber="1">36</figureCitation>
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.14-36.16).
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</emphasis>
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Shell decollate; whorls densely ribbed, without or with very weak sutural papillae; dorsal keel absent; inferior lamella moderately high; anterior upper palatal plica absent, posterior upper palatal plica very short; ALPP very short to knob-like, posterior lower reduced; palatal edge of clausilium plate distally not receding, palatal edge distally not bent upwards (see Nordsieck, 1984: 202).
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</paragraph>
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<caption doi="10.3897/zookeys.1077.67081.figure34" httpUri="https://binary.pensoft.net/fig/622826" pageId="0" pageNumber="1" start="Figure 34" startId="F34">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 34.</emphasis>
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<taxonomicName authority="nobilis" authorityName="nobilis (L. Pfeiffer" authorityYear="1848" baseAuthorityName="nobilis (L. Pfeiffer" baseAuthorityYear="1848" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis subsp. nobilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="nobilis" subSpecies="nobilis">
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<emphasis italics="true" pageId="0" pageNumber="1">Sicania nobilis nobilis</emphasis>
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</taxonomicName>
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(A. Schmidt, 1868), comb. nov., Castelluzzo
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<emphasis bold="true" pageId="0" pageNumber="1">34.1</emphasis>
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whole distal genital organs
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<emphasis bold="true" pageId="0" pageNumber="1">34.2</emphasis>
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internal distal part of genital organs. San Vito Lo capo, camping
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<emphasis bold="true" pageId="0" pageNumber="1">34.3</emphasis>
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whole distal genital organs
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<emphasis bold="true" pageId="0" pageNumber="1">34.4</emphasis>
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internal distal part of genital organs. San Vito Lo Capo, boulders
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<emphasis bold="true" pageId="0" pageNumber="1">34.5</emphasis>
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whole distal genital organs
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<emphasis bold="true" pageId="0" pageNumber="1">34.6</emphasis>
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internal distal part of genital organs. Torre delle Usciere (ex. forma
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<taxonomicName class="Gastropoda" family="Clausiliidae" form="episoma" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis subsp. nobilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="form" species="nobilis" subSpecies="nobilis">Sicania nobilis nobilis episoma</taxonomicName>
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)
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<emphasis bold="true" pageId="0" pageNumber="1">34.7</emphasis>
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whole distal genital organs
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<emphasis bold="true" pageId="0" pageNumber="1">34.8</emphasis>
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internal distal part of genital organs.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Measurements</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">(n = 24, decollate).</emphasis>
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shell height 22.2
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<normalizedToken originalValue="±">+/-</normalizedToken>
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1.3, whorl width 5.1
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.2, aperture height 4.1
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.3, aperture width 2.8
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.3.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="external morphology of the genital organs">
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<paragraph pageId="0" pageNumber="1">External morphology of the genital organs</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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(Figs
|
||
<figureCitation captionStart="Figure" captionStartId="F35" captionText="Figure. 35. Sicania nobilis spezialensis (Nordsieck 1984), stat. nov., comb. nov. Macari 35.1 whole distal genital organs 35.2 internal distal part of genital organs 35.3 whole distal genital organs. 35.4 internal distal part of genital organs 35.5 cross section of the pseudopapilla 35.6 cross section of the epiphallus 35.7 detail of the pseudopapilla." figureDoi="10.3897/zookeys.1077.67081.figure" httpUri="https://binary.pensoft.net/fig/622827" pageId="0" pageNumber="1">35</figureCitation>
|
||
.1, 35.3).
|
||
</emphasis>
|
||
The FO is longer than the V (FO/V range 1.9-2.2). The VD is thin along its whole course. The FDBC is slightly longer than the BC+SDBC (FDBC/BC+SDBC range 1.1-1.2). The BC+SDBC is club-like and longer than the V (BC+SDBC/V range 1.7-1.8), with no clear distinction between the SDBC and the BC. The apex is big and rounded. The D is longer than the V (D/V range 2.1-2.3) and longer that the BC+SDBC (D/BC+SDBC range 1.3-1.7), thinner than the BC+SDBC and with a small and round apex. The V is cylindrical. The A is very large. The PC is longer than the V (P+E/V range 1.8-2.0). The PR is short and robust. The E is almost as long as the P (E/P range 0.8-0.9), gradually shrinking and turning into the VD.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="internal morphology of the genital organs">
|
||
<paragraph pageId="0" pageNumber="1">Internal morphology of the genital organs</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
(Figs
|
||
<figureCitation captionStart="Figure" captionStartId="F35" captionText="Figure. 35. Sicania nobilis spezialensis (Nordsieck 1984), stat. nov., comb. nov. Macari 35.1 whole distal genital organs 35.2 internal distal part of genital organs 35.3 whole distal genital organs. 35.4 internal distal part of genital organs 35.5 cross section of the pseudopapilla 35.6 cross section of the epiphallus 35.7 detail of the pseudopapilla." figureDoi="10.3897/zookeys.1077.67081.figure" httpUri="https://binary.pensoft.net/fig/622827" pageId="0" pageNumber="1">35</figureCitation>
|
||
.2, 35.4-35.7).
|
||
</emphasis>
|
||
The V distally presents a set of irregular, weak, and fleshy folds that proximally become a set of oblique smooth pleats. The A shows a set of irregular fleshy folds. The P presents 4-to 6 longitudinal fleshy smooth pleats, slightly depressed in the middle forming a sort of longitudinal hollow. These pleats distally become weak and irregular entering in the A. The fine structure of the wall is smooth. The PP is little, rhombus-shaped and smooth. The P-E transition presents a first distal ER, whereas the PP and the ELP originate from the second proximal ER. The epiphallar formula is: 1ER+2ER(PP+ELP). The E shows a pattern of three or four irregular longitudinal fringed pleats. The wall of the E is smooth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="ecology">
|
||
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName=", stat. nov." authorityYear="2021" baseAuthorityName="Nordsieck" baseAuthorityYear="1984" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis subsp. spezialensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="nobilis" subSpecies="spezialensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania nobilis spezialensis</emphasis>
|
||
</taxonomicName>
|
||
stat. nov., comb. nov. is a strictly limestone dweller and inhabits cliffs, hiding among rock cracks and crevices. As previously deemed as a species, according to
|
||
<bibRefCitation author="De Mattia, W" journalOrPublisher="Molecular Ecology Resources" pageId="0" pageNumber="1" publicationUrl="https://www.iucnredlist.org/species/170841/1317462" refId="B23" refString="De Mattia, W, 2017j. Charpentieria spezialensis. The IUCN Red List of Threatened Species. https://www.iucnredlist.org/species/170841/1317462" title="Charpentieria spezialensis. The IUCN Red List of Threatened Species." url="https://www.iucnredlist.org/species/170841/1317462" year="2017 j">De Mattia (2017j)</bibRefCitation>
|
||
the taxon is Critically Endangered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName=", stat. nov." authorityYear="2021" baseAuthorityName="Nordsieck" baseAuthorityYear="1984" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis subsp. spezialensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="nobilis" subSpecies="spezialensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania nobilis spezialensis</emphasis>
|
||
</taxonomicName>
|
||
stat. nov., comb. nov. is known only from the type locality: along the western cliffs of Monte Speziale in the surroundings of Macari (San Vito lo Capo). No overlapping zone with
|
||
<taxonomicName authority="nobilis" authorityName="nobilis (L. Pfeiffer" authorityYear="1848" baseAuthorityName="nobilis (L. Pfeiffer" baseAuthorityYear="1848" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis subsp. nobilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="nobilis" subSpecies="nobilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania nobilis nobilis</emphasis>
|
||
</taxonomicName>
|
||
stat. nov., comb. nov. is known until now. Although, in its original description, the taxon was reported for one locality only (
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" refId="B61" refString="Nordsieck, H, 1984. Neue Taxa rezenter europaeischer Clausilien, mit Bemerkungen zur Bastardierung bei Clausilien (Gastropoda: Clausiliidae). Archiv fuer Molluskenkunde 114(4/6): 189-211." title="Neue Taxa rezenter europaeischer Clausilien, mit Bemerkungen zur Bastardierung bei Clausilien (Gastropoda: Clausiliidae). Archiv fuer Molluskenkunde 114 (4 / 6): 189 - 211." year="1984">Nordsieck 1984</bibRefCitation>
|
||
: 202 "Monte Speziale bei San Vito"), the same author reported later and extended area of distribution (
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck 2013b</bibRefCitation>
|
||
: "from Macari and Castelluzzo to Monte Speziale and Monte Acci").
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName=", stat. nov." authorityYear="2021" baseAuthorityName="Nordsieck" baseAuthorityYear="1984" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis subsp. spezialensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="nobilis" subSpecies="spezialensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania nobilis spezialensis</emphasis>
|
||
</taxonomicName>
|
||
stat. nov., comb. nov. is now considered as a subspecies of
|
||
<taxonomicName lsidName="S. nobilis" pageId="0" pageNumber="1" rank="species" species="nobilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. nobilis</emphasis>
|
||
</taxonomicName>
|
||
. It is well embedded within the clade of
|
||
<taxonomicName authorityName=", s. l." authorityYear="2021" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1848" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="nobilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania nobilis</emphasis>
|
||
</taxonomicName>
|
||
in both COI and combined trees.
|
||
<taxonomicName authorityName=", stat. nov." authorityYear="2021" baseAuthorityName="Nordsieck" baseAuthorityYear="1984" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis subsp. spezialensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="nobilis" subSpecies="spezialensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania nobilis spezialensis</emphasis>
|
||
</taxonomicName>
|
||
stat. nov., comb. nov. is a local isolated, spatially restricted ribbed form of the more widespread
|
||
<taxonomicName authorityName=", s. l." authorityYear="2021" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1848" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="nobilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania nobilis</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. with few different features of the shell, viz. the lack of the anterior upper palatal plica and a clausilium plate that fits into a frame of the palatal plicae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="mauritanica taxonomic part">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="O. Boettger" authorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica</emphasis>
|
||
</taxonomicName>
|
||
taxonomic part
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The genus
|
||
<taxonomicName authorityName="O. Boettger" authorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica</emphasis>
|
||
</taxonomicName>
|
||
comprises four species that are distributed over northeastern Algeria and central-northern Tunisia (
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="27 - 39" refId="B64" refString="Nordsieck, H, 2002. Contributions to the knowldge of the Delimini (Gastropoda: Stylommatophora: Clausiliidae). Mitteilungen der deutschen malakozoologischen Gesellschaft 67: 27 - 39" title="Contributions to the knowldge of the Delimini (Gastropoda: Stylommatophora: Clausiliidae)." volume="67" year="2002">Nordsieck 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="311 - 346" refId="B91" refString="Sparacio, I, Liberto, F, La Mantia, T, 2020. The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia. Biodiversity Journal 11 (1): 311 - 346, DOI: https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" title="The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia." url="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" volume="11" year="2020">Sparacio et al. 2020</bibRefCitation>
|
||
). It is currently considered as a subgenus of
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
(MolluscaBase 2021). Only the Tunisian taxa were recently reviewed (
|
||
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="311 - 346" refId="B91" refString="Sparacio, I, Liberto, F, La Mantia, T, 2020. The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia. Biodiversity Journal 11 (1): 311 - 346, DOI: https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" title="The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia." url="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" volume="11" year="2020">Sparacio et al. 2020</bibRefCitation>
|
||
), mainly by means of shell morphology and a brief analysis of the anatomy of the genital organs, discussing the position of the genus and separating it from
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
without involving additional morphological and molecular genetic data.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1077.67081.figure" httpUri="https://binary.pensoft.net/fig/622827" pageId="0" pageNumber="1" start="Figure" startId="F35">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure.</emphasis>
|
||
<emphasis bold="true" pageId="0" pageNumber="1">35.</emphasis>
|
||
<taxonomicName authorityName=", stat. nov." authorityYear="2021" baseAuthorityName="Nordsieck" baseAuthorityYear="1984" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis subsp. spezialensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="nobilis" subSpecies="spezialensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania nobilis spezialensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" refId="B61" refString="Nordsieck, H, 1984. Neue Taxa rezenter europaeischer Clausilien, mit Bemerkungen zur Bastardierung bei Clausilien (Gastropoda: Clausiliidae). Archiv fuer Molluskenkunde 114(4/6): 189-211." title="Neue Taxa rezenter europaeischer Clausilien, mit Bemerkungen zur Bastardierung bei Clausilien (Gastropoda: Clausiliidae). Archiv fuer Molluskenkunde 114 (4 / 6): 189 - 211." year="1984">Nordsieck 1984</bibRefCitation>
|
||
), stat. nov., comb. nov. Macari
|
||
<emphasis bold="true" pageId="0" pageNumber="1">35.1</emphasis>
|
||
whole distal genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">35.2</emphasis>
|
||
internal distal part of genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">35.3</emphasis>
|
||
whole distal genital organs.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">35.4</emphasis>
|
||
internal distal part of genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">35.5</emphasis>
|
||
cross section of the pseudopapilla
|
||
<emphasis bold="true" pageId="0" pageNumber="1">35.6</emphasis>
|
||
cross section of the epiphallus
|
||
<emphasis bold="true" pageId="0" pageNumber="1">35.7</emphasis>
|
||
detail of the pseudopapilla.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
In our molecular genetic analysis (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. BI tree of based on the COI data set of Siciliaria, Sicania and related genera. BI posterior probability values (> 0.90) of major nodes are indicated." figureDoi="10.3897/zookeys.1077.67081.figure4" httpUri="https://binary.pensoft.net/fig/622796" pageId="0" pageNumber="1">4</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. BI tree of based on the concatenated COI and ITS 2 data set of Siciliaria, Sicania and related genera. Colours of Siciliaria and Sicania clades correspond to those in the distribution maps (Fig. 1 E, F). BI posterior probability values (> 0.90) of major nodes are indicated." figureDoi="10.3897/zookeys.1077.67081.figure6" httpUri="https://binary.pensoft.net/fig/622798" pageId="0" pageNumber="1">6</figureCitation>
|
||
) we only had
|
||
<taxonomicName baseAuthorityName="O. Boettger" baseAuthorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica perinni subsp. polygyra" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perinni" subSpecies="polygyra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica perinni polygyra</emphasis>
|
||
</taxonomicName>
|
||
available. In the COI, ITS2 and combined trees, this species falls outside the
|
||
<taxonomicName authorityName="Vest" authorityYear="1867" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName authorityName="Tomlin" authorityYear="1929" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania</emphasis>
|
||
</taxonomicName>
|
||
clades, forming a separate independent clade together with
|
||
<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria scarificata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scarificata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria scarificata</emphasis>
|
||
</taxonomicName>
|
||
, endemic to Marettimo. In the COI tree, this clade represents the
|
||
<normalizedToken originalValue="“sister”">"sister"</normalizedToken>
|
||
clade of
|
||
<taxonomicName authorityName="Tomlin" authorityYear="1929" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania</emphasis>
|
||
</taxonomicName>
|
||
(average p distance 17%) and the same is found in the ITS2 tree (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. BI tree of based on the ITS 2 data set of Siciliaria, Sicania and related genera. Colours of Siciliaria and Sicania clades correspond to those in the distribution maps (Figs 1 E, F). BI posterior probability values (> 0.90) of major nodes are indicated." figureDoi="10.3897/zookeys.1077.67081.figure5" httpUri="https://binary.pensoft.net/fig/622797" pageId="0" pageNumber="1">5</figureCitation>
|
||
), yet with lower support. Despite low sample number, no association with any of the other
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
lineages was observed.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1077.67081.figure36" httpUri="https://binary.pensoft.net/fig/622828" pageId="0" pageNumber="1" start="Figure 36" startId="F36">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 36.</emphasis>
|
||
<taxonomicName authority="nobilis" authorityName="nobilis (L. Pfeiffer" authorityYear="1848" baseAuthorityName="nobilis (L. Pfeiffer" baseAuthorityYear="1848" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis subsp. nobilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="nobilis" subSpecies="nobilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania nobilis nobilis</emphasis>
|
||
</taxonomicName>
|
||
(A. Schmidt, 1868), comb. nov., Castelluzzo
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.1</emphasis>
|
||
shell
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.2</emphasis>
|
||
detail of the aperture
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.3</emphasis>
|
||
clausiliar plate double side. San Vito Lo capo, camping
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.4</emphasis>
|
||
shell
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.5</emphasis>
|
||
detail of the aperture
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.6</emphasis>
|
||
clausiliar plate double side. San Vito Lo Capo, boulders
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.7</emphasis>
|
||
shell
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.8</emphasis>
|
||
detail of the aperture
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.9</emphasis>
|
||
clausiliar plate double side. Torre delle Usciere (ex. forma
|
||
<taxonomicName class="Gastropoda" family="Clausiliidae" form="episoma" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis subsp. nobilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="form" species="nobilis" subSpecies="nobilis">Sicania nobilis nobilis episoma</taxonomicName>
|
||
)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.10</emphasis>
|
||
shell
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.11</emphasis>
|
||
detail of the aperture
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.12</emphasis>
|
||
clausiliar plate double side. Monte Cofano, Tonnara Cofano
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.13</emphasis>
|
||
shell.
|
||
<taxonomicName authorityName=", stat. nov." authorityYear="2021" baseAuthorityName="Nordsieck" baseAuthorityYear="1984" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis subsp. spezialensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="nobilis" subSpecies="spezialensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania nobilis spezialensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" refId="B61" refString="Nordsieck, H, 1984. Neue Taxa rezenter europaeischer Clausilien, mit Bemerkungen zur Bastardierung bei Clausilien (Gastropoda: Clausiliidae). Archiv fuer Molluskenkunde 114(4/6): 189-211." title="Neue Taxa rezenter europaeischer Clausilien, mit Bemerkungen zur Bastardierung bei Clausilien (Gastropoda: Clausiliidae). Archiv fuer Molluskenkunde 114 (4 / 6): 189 - 211." year="1984">Nordsieck 1984</bibRefCitation>
|
||
), stat. nov., comb. nov. Macari
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.14</emphasis>
|
||
shell
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.15</emphasis>
|
||
detail of the aperture
|
||
<emphasis bold="true" pageId="0" pageNumber="1">36.16</emphasis>
|
||
clausiliar plate double side.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The separation of
|
||
<taxonomicName authorityName="O. Boettger" authorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<emphasis italics="true" pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Vest" authorityYear="1867" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Siciliaria</taxonomicName>
|
||
/
|
||
<taxonomicName authorityName="Tomlin" authorityYear="1929" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sicania</taxonomicName>
|
||
</emphasis>
|
||
is consistent through all the trees, thus we provisionally keep
|
||
<taxonomicName authorityName="O. Boettger" authorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica</emphasis>
|
||
</taxonomicName>
|
||
as a distinct genus, although the type species
|
||
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1860" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica tristrami" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tristrami">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica tristrami</emphasis>
|
||
</taxonomicName>
|
||
(L. Pfeiffer,1861) was not available to us. In order to fully understand the phylogenetic relationships of
|
||
<taxonomicName authorityName="O. Boettger" authorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName authorityName="Vest" authorityYear="1867" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName authorityName="Tomlin" authorityYear="1929" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania</emphasis>
|
||
</taxonomicName>
|
||
more genetic markers and specimens are essential.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Both
|
||
<taxonomicName lsidName="M. perinni subsp. polygyra" pageId="0" pageNumber="1" rank="subSpecies" species="perinni" subSpecies="polygyra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">M. perinni polygyra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="M. scarificata" pageId="0" pageNumber="1" rank="species" species="scarificata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">M. scarificata</emphasis>
|
||
</taxonomicName>
|
||
underwent anatomical investigation. A similar external genital morphology was detected. The only noticeable difference is the relative length of the FO, with a V/FO ratio of 1.1 for
|
||
<taxonomicName lsidName="M. scarificata" pageId="0" pageNumber="1" rank="species" species="scarificata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">M. scarificata</emphasis>
|
||
</taxonomicName>
|
||
and 0.6 for
|
||
<taxonomicName lsidName="M. perinni subsp. polygyra" pageId="0" pageNumber="1" rank="subSpecies" species="perinni" subSpecies="polygyra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">M. perinni polygyra</emphasis>
|
||
</taxonomicName>
|
||
. Remarkable differences were detected regarding the internal details of the penial complex.
|
||
<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1856" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica scarificata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scarificata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica scarificata</emphasis>
|
||
</taxonomicName>
|
||
presents 3 ERs with a very short and globose PP that makes it fall into the general
|
||
<taxonomicName authorityName="Vest" authorityYear="1867" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName authorityName="Tomlin" authorityYear="1929" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania</emphasis>
|
||
</taxonomicName>
|
||
genital anatomical arrangement.
|
||
<taxonomicName baseAuthorityName="O. Boettger" baseAuthorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica perinni subsp. polygyra" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perinni" subSpecies="polygyra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica perinni polygyra</emphasis>
|
||
</taxonomicName>
|
||
lacks any ER and the PP is irregular with a convolute surface pattern. It is directly connected to the OELP. Unfortunately, the genital descriptions and the pictures provided by
|
||
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="311 - 346" refId="B91" refString="Sparacio, I, Liberto, F, La Mantia, T, 2020. The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia. Biodiversity Journal 11 (1): 311 - 346, DOI: https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" title="The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia." url="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" volume="11" year="2020">Sparacio et al. (2020)</bibRefCitation>
|
||
do not include fine details of the internal genital organs (ER, PP structure and origin, ELP etc) so that any comparison to our results is impossible.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Following our genital anatomical investigations, the common genital anatomical traits deemed by
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" refId="B68" refString="Nordsieck, H, 2013a. Delimini (Gastropoda, Pulmonata, Clausiliidae) from Apennine Italy, with the description of three new subspecies from Calabria. Conchylia 44(1/2): 3-14." title="Delimini (Gastropoda, Pulmonata, Clausiliidae) from Apennine Italy, with the description of three new subspecies from Calabria. Conchylia 44 (1 / 2): 3 - 14." year="2013 a">Nordsieck (2013a</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">2013b</bibRefCitation>
|
||
) to justify his proposal to consider
|
||
<taxonomicName authorityName="O. Boettger" authorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica</emphasis>
|
||
</taxonomicName>
|
||
as a subgenus of
|
||
<taxonomicName authorityName="Vest" authorityYear="1867" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria</emphasis>
|
||
</taxonomicName>
|
||
, namely, "the presence of a thickening in the passage from the penis to the epiphallus and the indistinct delimitation between proximal and distal penis" revealed that it is probably not a reliable feature. The thickening was not found in
|
||
<taxonomicName baseAuthorityName="O. Boettger" baseAuthorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica perinni subsp. polygyra" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perinni" subSpecies="polygyra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica perinni polygyra</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Mauritanica scarificata (L. Pfeiffer, 1856), comb. nov., Marettimo 37.1 whole distal genital organs 37.2 internal distal part of genital organs 37.3 whole distal genital organs 37.4 internal distal part of genital organs 37.5 detail of the pseudopapilla. Mauritanica perinni polygyra (O. Boettger, 1879), Tunisia, Zaghouan 37.6 whole distal genital organs 37.7 internal distal part of genital organs 37.8 - 37.9 spermatophore 37.10 cross section of the spermatophore." figureDoi="10.3897/zookeys.1077.67081.figure37" httpUri="https://binary.pensoft.net/fig/622829" pageId="0" pageNumber="1">37</figureCitation>
|
||
). Moreover, the delimitation between proximal and distal penis revealed to be an extremely subtle and variable character also in
|
||
<taxonomicName authorityName="Vest" authorityYear="1867" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="O.Boettger" authorityYear="1877" class="Gastropoda" family="Clausiliidae" genus="Stigmatica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stigmatica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Stigmatica</emphasis>
|
||
</taxonomicName>
|
||
. Only few taxa of
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
show a detectable, constant and distinct delimitation between proximal and distal penis [e.g.,
|
||
<taxonomicName baseAuthorityName="Gredler" baseAuthorityYear="1869" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. lorinae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="lorinae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala lorinae</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Charpentieria itala clavata (Rossmaessler, 1836), Ballabio, Lecco 46.1 whole distal genital organs 46.2 internal distal part of genital organs 46.3 cross section of the pseudopapilla 46.3 spermatophore. Charpentieria itala latestriata (Kuester, 1850), Bedulita, Bergamo 46.5 whole distal genital organs 46.6 internal distal part of genital organs. Charpentieria itala lorinae (Gredler, 1869), Val Lorina, Storo 46.7 whole distal genital organs 46.8 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure46" httpUri="https://binary.pensoft.net/fig/622838" pageId="0" pageNumber="1">46</figureCitation>
|
||
.7) and
|
||
<taxonomicName baseAuthorityName="Kaeufel" baseAuthorityYear="1928" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. trepida" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="trepida">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala trepida</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Charpentieria itala triumplinae Nardi, 2011, Lavenone, Brescia 49.1 whole distal genital organs 49.2 internal distal part of genital organs. Charpentieria itala variscoi (Pini, 1883), Val Taleggio, Bergamo 49.3 whole distal genital organs 49.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure49" httpUri="https://binary.pensoft.net/fig/622841" pageId="0" pageNumber="1">49</figureCitation>
|
||
.1)].
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |