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<document id="04DDE03D3CEEE2AB51FAB7C5D52194BC" ID-DOI="10.11646/zootaxa.5293.2.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7960070" ID-ZooBank="47EF72B0-A13B-48FD-AF07-72102457E518" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1684797818070" checkinUser="plazi" docAuthor="Michelsen, Verner" docDate="2023" docId="B41A7D60FFADFFF400AD17B990CDFE8F" docLanguage="en" docName="zootaxa.5293.2.3.pdf" docOrigin="Zootaxa 5293 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5293.2.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helina atlantica" docType="treatment" docVersion="2" lastPageNumber="273" masterDocId="48230518FFBEFFE2003A11739121FF92" masterDocTitle="Macaronesian Muscidae (Diptera). IV. The genus Helina Robineau-Desvoidy, and description of the male of Spilogona maderensis Michelsen" masterLastPageNumber="276" masterPageNumber="251" pageNumber="270" updateTime="1684799522895" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="A9E5FEDA6F52E0FC16E32A71472CAE4A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="75F290BF0C9F1F489AB2D5BDB48D1440">Macaronesian Muscidae (Diptera). IV. The genus Helina Robineau-Desvoidy, and description of the male of Spilogona maderensis Michelsen</mods:title>
</mods:titleInfo>
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<mods:namePart id="CDD6E89F06CE07995547DEBEC56F7DE5">Michelsen, Verner</mods:namePart>
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<mods:title id="E8E94CD3CF08A1D18025CC3E6C442C8E">Zootaxa</mods:title>
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<mods:date id="86F23E5FCD10DDCBA73841958E7B290C">2023</mods:date>
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<mods:number id="7F4168FB0A84B9792B903907D5C18879">2023-05-22</mods:number>
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<mods:url id="0EB53503837FB82EBCB9229ED4E69E6B">http://dx.doi.org/10.11646/zootaxa.5293.2.3</mods:url>
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<mods:classification id="35D67AFE2DBB02D93E9D4CEE8A61AE2E">journal article</mods:classification>
<mods:identifier id="9C966811346E16F9696B75787043F2C3" type="DOI">10.11646/zootaxa.5293.2.3</mods:identifier>
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</mods:mods>
<treatment id="B41A7D60FFADFFF400AD17B990CDFE8F" LSID="urn:lsid:plazi:treatment:B41A7D60FFADFFF400AD17B990CDFE8F" httpUri="http://treatment.plazi.org/id/B41A7D60FFADFFF400AD17B990CDFE8F" lastPageId="22" lastPageNumber="273" pageId="19" pageNumber="270">
<subSubSection id="74A99FFDFFADFFF100AD17B99326F976" box="[151,519,1738,1765]" pageId="19" pageNumber="270" type="nomenclature">
<paragraph id="3C0CCC76FFADFFF100AD17B99326F976" blockId="19.[151,519,1738,1801]" box="[151,519,1738,1765]" pageId="19" pageNumber="270">
<heading id="67447B1AFFADFFF100AD17B99326F976" bold="true" box="[151,519,1738,1765]" fontSize="11" level="1" pageId="19" pageNumber="270" reason="1">
<emphasis id="0EC71064FFADFFF100AD17B99326F976" bold="true" box="[151,519,1738,1765]" pageId="19" pageNumber="270">
13.
<taxonomicName id="FBB3B7F5FFADFFF100F917B99326F976" ID-CoL="6LJZ8" authority="(Tiensuu)" baseAuthorityName="Tiensuu" baseAuthorityYear="1939" box="[195,519,1738,1765]" class="Insecta" family="Muscidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="270" phylum="Arthropoda" rank="species" species="atlantica">
<emphasis id="0EC71064FFADFFF100F917B990A9F977" bold="true" box="[195,392,1738,1765]" italics="true" pageId="19" pageNumber="270">Helina atlantica</emphasis>
(Tiensuu)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="74A99FFDFFADFFF100AD179C9011F89B" box="[151,304,1775,1801]" pageId="19" pageNumber="270" type="description">
<paragraph id="3C0CCC76FFADFFF100AD179C9011F89B" blockId="19.[151,519,1738,1801]" box="[151,304,1775,1801]" pageId="19" pageNumber="270">
(
<figureCitation id="A488D0F3FFADFFF100A5179C9009F89B" box="[159,296,1775,1801]" captionStart="FIGURE 13" captionStartId="21.[151,250,1856,1880]" captionTargetBox="[160,1396,593,1824]" captionTargetId="figure-220@21.[151,1436,561,1833]" captionTargetPageId="21" captionText="FIGURE 13. Helina atlantica (Tiensuu), male. Head, dorso-lateral (A); abdomen (B); sternite V (C); hypopygium, caudal (D) and lateral (E). Same scale for C–E." figureDoi="http://doi.org/10.5281/zenodo.7960110" httpUri="https://zenodo.org/record/7960110/files/figure.png" pageId="19" pageNumber="270">Figs 13A–E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="74A99FFDFFADFFF100AD164092D3F8F8" pageId="19" pageNumber="270" type="reference_group">
<paragraph id="3C0CCC76FFADFFF100AD16409538F8D9" blockId="19.[151,1049,1843,1898]" box="[151,1049,1843,1868]" pageId="19" pageNumber="270">
<treatmentCitationGroup id="1CA3EB58FFADFFF100AD16409538F8D9" box="[151,1049,1843,1868]" pageId="19" pageNumber="270">
<taxonomicName id="FBB3B7F5FFADFFF100AD164090FBF8D9" ID-CoL="39WSC" authority="Tiensuu" authorityName="Tiensuu" authorityYear="1938" box="[151,474,1843,1867]" class="Insecta" family="Muscidae" genus="Enoplopteryx" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="270" phylum="Arthropoda" rank="species" species="atlantica">
<emphasis id="0EC71064FFADFFF100AD164090A1F8D9" box="[151,384,1843,1867]" italics="true" pageId="19" pageNumber="270">Enoplopteryx atlantica</emphasis>
Tiensuu
</taxonomicName>
;
<treatmentCitation id="BD12EA67FFADFFF101DE164093A7F8D9" author="Tiensuu, L." box="[484,646,1843,1867]" page="5" pageId="19" pageNumber="270" year="1938">
<bibRefCitation id="5822B187FFADFFF101DE164093A7F8D9" author="Tiensuu, L." box="[484,646,1843,1867]" pageId="19" pageNumber="270" pagination="1 - 11" refId="ref16706" refString="Tiensuu, L. (1938) Die Arthropodenfauna von Madeira nach den Ergebnissen der Reise von Prof. Dr. O. Lundblad Juli - August 1935. XII. Diptera: Muscidae und Tachinidae. Arkiv f ˆ r Zoologi, 30 A, 22, 1 - 11." type="journal article" year="1938">Tiensuu 1938: 5</bibRefCitation>
</treatmentCitation>
ff. (Madeira);
<treatmentCitation id="BD12EA67FFADFFF1032216479284F8D9" author="Frey, R." box="[792,933,1843,1868]" page="43" pageId="19" pageNumber="270" year="1949">
<bibRefCitation id="5822B187FFADFFF1032216479284F8D9" author="Frey, R." box="[792,933,1843,1868]" pageId="19" pageNumber="270" pagination="1 - 47" refId="ref15873" refString="Frey, R. (1949) Die Dipterenfauna der Insel Madeira. Commentationes biologicae, 8 (16), 1 - 47." type="journal article" year="1949">Frey 1949: 43</bibRefCitation>
</treatmentCitation>
(Madeira).
</treatmentCitationGroup>
</paragraph>
<paragraph id="3C0CCC76FFADFFF100AD162192D3F8F8" blockId="19.[151,1049,1843,1898]" box="[151,1010,1874,1898]" pageId="19" pageNumber="270">
<treatmentCitationGroup id="1CA3EB58FFADFFF100AD162192D3F8F8" box="[151,1010,1874,1898]" pageId="19" pageNumber="270">
<taxonomicName id="FBB3B7F5FFADFFF100AD16219089F8F8" ID-CoL="6LJZ8" authority="(Tiensuu)" baseAuthorityName="Tiensuu" baseAuthorityYear="1939" box="[151,424,1874,1898]" class="Insecta" family="Muscidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="270" phylum="Arthropoda" rank="species" species="atlantica">
<emphasis id="0EC71064FFADFFF100AD1621901DF8F8" box="[151,316,1874,1898]" italics="true" pageId="19" pageNumber="270">Helina atlantica</emphasis>
(Tiensuu)
</taxonomicName>
;
<treatmentCitation id="BD12EA67FFADFFF1018816219348F8F8" author="Hennig, W. &amp; Lindner, E. &amp; Die Fliegen" box="[434,617,1874,1898]" page="161" pageId="19" pageNumber="270" year="1957">
<bibRefCitation id="5822B187FFADFFF1018816219348F8F8" author="Hennig, W. &amp; Lindner, E. &amp; Die Fliegen" box="[434,617,1874,1898]" pageId="19" pageNumber="270" pagination="145 - 192" refId="ref15952" refString="Hennig, W. (1957 - 1958) 63 b. Muscidae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region. 7 (2). Lief. 197 (1957) &amp; Lief. 199 (1958). Schweizerbart´sche Verlagsbuchhandlung, Stuttgart, pp. 145 - 192 &amp; 193 - 232." type="journal article" year="1957">Hennig 1957: 161</bibRefCitation>
</treatmentCitation>
(Madeira);
<treatmentCitation id="BD12EA67FFADFFF102D8162192A1F8F8" author="Hennig, W." box="[738,896,1874,1898]" page="4" pageId="19" pageNumber="270" year="1978">
<bibRefCitation id="5822B187FFADFFF102D8162192A1F8F8" author="Hennig, W." box="[738,896,1874,1898]" pageId="19" pageNumber="270" pagination="1 - 12" refId="ref16017" refString="Hennig, W. (1978) Uber Muscidae (Diptera) der Kanarischen Inseln. Stuttgarter Beitrage zur Naturkunde, A (Biologie), 311, 1 - 12. [Posthumous paper.]" type="journal article" year="1978">Hennig 1978: 4</bibRefCitation>
</treatmentCitation>
(Madeira).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="74A99FFDFFADFFF100AD16EB93ADF868" pageId="19" pageNumber="270" type="materials_examined">
<paragraph id="3C0CCC76FFADFFF100AD16EB93ADF868" blockId="19.[151,1436,1944,2043]" pageId="19" pageNumber="270">
<emphasis id="0EC71064FFADFFF100AD16EB9303F820" bold="true" box="[151,546,1944,1970]" pageId="19" pageNumber="270">Material examined. MADEIRA:</emphasis>
[
<materialsCitation id="8CDBC62BFFADFFF1020E16EA93A8F868" collectingDate="1935-08-06" collectingDateMax="1957-06-19" collectingDateMin="1935-08-06" collectionCode="MZH, R, V" collectorName="O. Lundblad &amp; Calheta &amp; Frey &amp; Ribeira Frio &amp; Frey, J. &amp; Stora &amp; Rabacal &amp; Poiso &amp; Arieira &amp; H. Lindberg" elevation="1250" pageId="19" pageNumber="270" specimenCount="2" specimenCount-female="2" typeStatus="syntype">
<collectionCode id="5AA254B3FFADFFF1020E16EA935AF821" box="[564,635,1945,1971]" country="Finland" httpUri="http://grbio.org/cool/2vmj-fp0v" name="Finnish Museum of Natural History" pageId="19" pageNumber="270" type="Museum">MZH</collectionCode>
]. Caramujo,
<quantity id="FB4B6193FFADFFF1032B16EB9240F821" box="[785,865,1944,1971]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="19" pageNumber="270" unit="m" value="1250.0">
<elevation id="B79E2B45FFADFFF1032B16EB9240F821" box="[785,865,1944,1971]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="19" pageNumber="270" unit="m" value="1250.0">1250m</elevation>
</quantity>
,
<typeStatus id="E30872D4FFADFFF1034B16EA92F2F821" box="[881,979,1945,1971]" pageId="19" pageNumber="270">syntypes</typeStatus>
1♁,
<specimenCount id="2AB507FFFFADFFF1042916EB9517F820" box="[1043,1078,1944,1970]" pageId="19" pageNumber="270" type="female">1♀</specimenCount>
<date id="480DEAB6FFADFFF1047816EB95C4F821" box="[1090,1253,1944,1971]" pageId="19" pageNumber="270" value="1935-08-06" valueMax="1935-08-14" valueMin="1935-08-06">
<collectingDate id="5849135EFFADFFF1047816EB95C4F821" box="[1090,1253,1944,1971]" pageId="19" pageNumber="270" value="1935-08-06" valueMax="1935-08-14" valueMin="1935-08-06">6–14.viii.1935</collectingDate>
</date>
(
<collectorName id="9146A9A0FFADFFF104C316EB94B0F820" box="[1273,1425,1944,1970]" pageId="19" pageNumber="270">O. Lundblad</collectorName>
);
<collectorName id="9146A9A0FFADFFF100AD16CF91CEF844" box="[151,239,1980,2006]" pageId="19" pageNumber="270">Calheta</collectorName>
, 1♁
<date id="480DEAB6FFADFFF1011F16CF90A4F844" box="[293,389,1980,2006]" pageId="19" pageNumber="270" value="1938-05-07">
<collectingDate id="5849135EFFADFFF1011F16CF90A4F844" box="[293,389,1980,2006]" pageId="19" pageNumber="270" value="1938-05-07">7.v.1938</collectingDate>
</date>
(
<collectionCode id="5AA254B3FFADFFF101AF16CF908AF844" box="[405,427,1980,2006]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="270">R</collectionCode>
.
<collectorName id="9146A9A0FFADFFF1018C16CF90CFF844" box="[438,494,1980,2006]" pageId="19" pageNumber="270">Frey</collectorName>
);
<collectorName id="9146A9A0FFADFFF1023816CF93ACF844" box="[514,653,1980,2006]" pageId="19" pageNumber="270">Ribeira Frio</collectorName>
, 2♁
<date id="480DEAB6FFADFFF102F916CF9202F844" box="[707,803,1980,2006]" pageId="19" pageNumber="270" value="1938-05-03">
<collectingDate id="5849135EFFADFFF102F916CF9202F844" box="[707,803,1980,2006]" pageId="19" pageNumber="270" value="1938-05-03">3.v.1938</collectingDate>
</date>
(
<collectionCode id="5AA254B3FFADFFF1030916CF9268F844" box="[819,841,1980,2006]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="270">R</collectionCode>
.
<collectorName id="9146A9A0FFADFFF1036E16CE9285F845" box="[852,932,1981,2007]" pageId="19" pageNumber="270">Frey, J.</collectorName>
<collectionCode id="5AA254B3FFADFFF1039916CE9298F845" box="[931,953,1981,2007]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="19" pageNumber="270" type="Museum">V</collectionCode>
.
<collectorName id="9146A9A0FFADFFF103FE16CF9525F845" box="[964,1028,1980,2007]" pageId="19" pageNumber="270">Storå</collectorName>
);
<collectorName id="9146A9A0FFADFFF1042216CF9554F844" box="[1048,1141,1980,2006]" pageId="19" pageNumber="270">Rabaçal</collectorName>
, 1♁
<date id="480DEAB6FFADFFF1049016CF9406F844" box="[1194,1319,1980,2006]" pageId="19" pageNumber="270" value="1938-05-06" valueMax="1938-05-07" valueMin="1938-05-06">
<collectingDate id="5849135EFFADFFF1049016CF9406F844" box="[1194,1319,1980,2006]" pageId="19" pageNumber="270" value="1938-05-06" valueMax="1938-05-07" valueMin="1938-05-06">6–7.v.1938</collectingDate>
</date>
(
<collectionCode id="5AA254B3FFADFFF1050C16CF946DF844" box="[1334,1356,1980,2006]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="270">R</collectionCode>
.
<collectorName id="9146A9A0FFADFFF1056216CF94B1F844" box="[1368,1424,1980,2006]" pageId="19" pageNumber="270">Frey</collectorName>
);
<collectorName id="9146A9A0FFADFFF100AD169391F4F868" box="[151,213,2016,2042]" pageId="19" pageNumber="270">Poiso</collectorName>
‒
<collectorName id="9146A9A0FFADFFF100DE16939017F868" box="[228,310,2016,2042]" pageId="19" pageNumber="270">Arieira</collectorName>
,
<specimenCount id="2AB507FFFFADFFF1017B16939045F868" box="[321,356,2016,2042]" pageId="19" pageNumber="270" type="female">1♀</specimenCount>
<date id="480DEAB6FFADFFF10151169390C2F869" box="[363,483,2016,2043]" pageId="19" pageNumber="270" value="1957-06-19">
<collectingDate id="5849135EFFADFFF10151169390C2F869" box="[363,483,2016,2043]" pageId="19" pageNumber="270" value="1957-06-19">19.vi.1957</collectingDate>
</date>
(
<collectorName id="9146A9A0FFADFFF101C8169393A0F868" box="[498,641,2016,2042]" pageId="19" pageNumber="270">H. Lindberg</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="68CC9CFEFFAAFFF600AD179E943BF897" ID-DOI="http://doi.org/10.5281/zenodo.7960108" ID-Zenodo-Dep="7960108" box="[151,1306,1773,1797]" httpUri="https://zenodo.org/record/7960108/files/figure.png" pageId="20" pageNumber="271" startId="20.[151,250,1773,1797]" targetBox="[247,1382,208,1710]" targetPageId="20" targetType="figure">
<paragraph id="3C0CCC76FFAAFFF600AD179E943BF897" blockId="20.[151,1306,1773,1797]" box="[151,1306,1773,1797]" pageId="20" pageNumber="271">
<emphasis id="0EC71064FFAAFFF600AD179E9001F897" bold="true" box="[151,288,1773,1797]" pageId="20" pageNumber="271">FIGURE 12.</emphasis>
<taxonomicName id="FBB3B7F5FFAAFFF6011C179E933EF897" authority="Hennig" authorityName="Hennig" box="[294,543,1773,1797]" class="Insecta" family="Muscidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="vilissima">
<emphasis id="0EC71064FFAAFFF6011C179E90EAF897" box="[294,459,1773,1797]" italics="true" pageId="20" pageNumber="271">Helina vilissima</emphasis>
Hennig
</taxonomicName>
, male.
<emphasis id="0EC71064FFAAFFF6025D179E9354F897" bold="true" box="[615,629,1773,1797]" pageId="20" pageNumber="271">S</emphasis>
ternite V (
<emphasis id="0EC71064FFAAFFF602E6179E93CFF897" bold="true" box="[732,750,1773,1797]" pageId="20" pageNumber="271">A</emphasis>
); hypopygium, caudal (
<emphasis id="0EC71064FFAAFFF603D8179E92D2F897" bold="true" box="[994,1011,1773,1797]" pageId="20" pageNumber="271">B</emphasis>
) and lateral (
<emphasis id="0EC71064FFAAFFF60441179E95ACF897" bold="true" box="[1147,1165,1773,1797]" pageId="20" pageNumber="271">C</emphasis>
). Same scale.
</paragraph>
</caption>
<subSubSection id="74A99FFDFFAAFFF700FD16409361FD8B" lastPageId="21" lastPageNumber="272" pageId="20" pageNumber="271" type="description">
<paragraph id="3C0CCC76FFAAFFF700FD16409361FD8B" blockId="20.[151,1437,1843,2014]" lastBlockId="21.[151,1437,150,537]" lastPageId="21" lastPageNumber="272" pageId="20" pageNumber="271">
<emphasis id="0EC71064FFAAFFF600FD16409078F8DF" bold="true" box="[199,345,1843,1869]" pageId="20" pageNumber="271">Description.</emphasis>
Small to medium sized, wing length
<quantity id="FB4B6193FFAAFFF6033B164092A8F8DC" box="[769,905,1843,1870]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.7" metricValueMin="4.3" pageId="20" pageNumber="271" unit="mm" value="5.0" valueMax="5.7" valueMin="4.3">4.3–5.7 mm</quantity>
(n = 7).
<emphasis id="0EC71064FFAAFFF603D716409511F8DF" box="[1005,1072,1843,1869]" italics="true" pageId="20" pageNumber="271">Male:</emphasis>
Ground colour dark except legs with ochre yellow femoral apices and tibiae, most parts densely greyish pruinose. Head fully feminized (
<figureCitation id="A488D0F3FFAAFFF6051D162494AEF8E3" box="[1319,1423,1879,1905]" captionStart="FIGURE 13" captionStartId="21.[151,250,1856,1880]" captionTargetBox="[160,1396,593,1824]" captionTargetId="figure-220@21.[151,1436,561,1833]" captionTargetPageId="21" captionText="FIGURE 13. Helina atlantica (Tiensuu), male. Head, dorso-lateral (A); abdomen (B); sternite V (C); hypopygium, caudal (D) and lateral (E). Same scale for C–E." figureDoi="http://doi.org/10.5281/zenodo.7960110" httpUri="https://zenodo.org/record/7960110/files/figure.png" pageId="20" pageNumber="271">Fig. 13A</figureCitation>
); broad frontal vitta extensively brownish black, as grey frontal triangle reduced, only extended a little beyond ocellar tubercle; mesonotum with usual pattern of four dark stripes on mesonotum visible in posterior view; wing with both cross-veins narrowly shadowed; abdomen (
<figureCitation id="A488D0F3FFAAFFF602FF16B0920CF84F" box="[709,813,1987,2013]" captionStart="FIGURE 13" captionStartId="21.[151,250,1856,1880]" captionTargetBox="[160,1396,593,1824]" captionTargetId="figure-220@21.[151,1436,561,1833]" captionTargetPageId="21" captionText="FIGURE 13. Helina atlantica (Tiensuu), male. Head, dorso-lateral (A); abdomen (B); sternite V (C); hypopygium, caudal (D) and lateral (E). Same scale for C–E." figureDoi="http://doi.org/10.5281/zenodo.7960110" httpUri="https://zenodo.org/record/7960110/files/figure.png" pageId="20" pageNumber="271">Fig. 13B</figureCitation>
) with dark brown pattern on tergites III–V consisting of an oblong median mark flanked by subtriangular marks. Eyes bare. Aristal branches very long, up to 1.5 times as long as width of postpedicel. Frons (
<figureCitation id="A488D0F3FFABFFF7026911C89398FF47" box="[595,697,187,213]" captionStart="FIGURE 13" captionStartId="21.[151,250,1856,1880]" captionTargetBox="[160,1396,593,1824]" captionTargetId="figure-220@21.[151,1436,561,1833]" captionTargetPageId="21" captionText="FIGURE 13. Helina atlantica (Tiensuu), male. Head, dorso-lateral (A); abdomen (B); sternite V (C); hypopygium, caudal (D) and lateral (E). Same scale for C–E." figureDoi="http://doi.org/10.5281/zenodo.7960110" httpUri="https://zenodo.org/record/7960110/files/figure.png" pageId="21" pageNumber="272">Fig. 13A</figureCitation>
) formed by a broad, flat frontal vitta separating relatively narrow fronto-orbital plates; these bearing 2 strong, reclinate orbital setae on posterior part followed by 3–4 frontal setae of which the anterior one is strikingly enlarged. Dorsocentral setae 2 + 3; presutural acrostical field with setulae only arranged in 2 narrow, irregular rows; prealar seta very short, but slightly stronger than surrounding setulae; katepisternal setae 2 + 2. Vein C without dorsal setulae. Fore tibia with 1
<emphasis id="0EC71064FFABFFF703E3103892D4FEF7" box="[985,1013,331,357]" italics="true" pageId="21" pageNumber="272">ad</emphasis>
(short) and 1
<emphasis id="0EC71064FFABFFF704AC10389585FEF6" box="[1174,1188,331,356]" italics="true" pageId="21" pageNumber="272">p</emphasis>
seta; mid tibia with 0
<emphasis id="0EC71064FFABFFF700AD101C9192FE1B" box="[151,179,367,393]" italics="true" pageId="21" pageNumber="272">ad</emphasis>
and 2–3
<emphasis id="0EC71064FFABFFF70121101C9016FE1B" box="[283,311,367,393]" italics="true" pageId="21" pageNumber="272">pd</emphasis>
setae; hind femur with 1–3
<emphasis id="0EC71064FFABFFF7024B101C93AAFE1A" box="[625,651,367,392]" italics="true" pageId="21" pageNumber="272">pv</emphasis>
setae on middle third; hind tibia with 1
<emphasis id="0EC71064FFABFFF70477101C9546FE1A" box="[1101,1127,367,392]" italics="true" pageId="21" pageNumber="272">av</emphasis>
, 3–4
<emphasis id="0EC71064FFABFFF7049D101C95E2FE1B" box="[1191,1219,367,393]" italics="true" pageId="21" pageNumber="272">ad</emphasis>
setae and 1
<emphasis id="0EC71064FFABFFF70574101C944BFE1B" box="[1358,1386,367,393]" italics="true" pageId="21" pageNumber="272">pd</emphasis>
seta on basal third. Slender ovoid abdomen (
<figureCitation id="A488D0F3FFABFFF7026F10E09399FE3F" box="[597,696,403,429]" captionStart="FIGURE 13" captionStartId="21.[151,250,1856,1880]" captionTargetBox="[160,1396,593,1824]" captionTargetId="figure-220@21.[151,1436,561,1833]" captionTargetPageId="21" captionText="FIGURE 13. Helina atlantica (Tiensuu), male. Head, dorso-lateral (A); abdomen (B); sternite V (C); hypopygium, caudal (D) and lateral (E). Same scale for C–E." figureDoi="http://doi.org/10.5281/zenodo.7960110" httpUri="https://zenodo.org/record/7960110/files/figure.png" pageId="21" pageNumber="272">Fig. 13B</figureCitation>
) with marginal and discal setae on tergites IV and V abundant and strong; sternite V and hypopygium as in
<figureCitation id="A488D0F3FFABFFF7025810C493CAFE43" box="[610,747,439,465]" captionStart="FIGURE 13" captionStartId="21.[151,250,1856,1880]" captionTargetBox="[160,1396,593,1824]" captionTargetId="figure-220@21.[151,1436,561,1833]" captionTargetPageId="21" captionText="FIGURE 13. Helina atlantica (Tiensuu), male. Head, dorso-lateral (A); abdomen (B); sternite V (C); hypopygium, caudal (D) and lateral (E). Same scale for C–E." figureDoi="http://doi.org/10.5281/zenodo.7960110" httpUri="https://zenodo.org/record/7960110/files/figure.png" pageId="21" pageNumber="272">Figs 13C–E</figureCitation>
.
<emphasis id="0EC71064FFABFFF702CC10C49272FE43" box="[758,851,439,465]" italics="true" pageId="21" pageNumber="272">Female:</emphasis>
apart from usual sexual differences of the abdomen differing from the male by the presence of a greyish frontal triangle extended almost to anterior margin of frontal vitta and hind femur without
<emphasis id="0EC71064FFABFFF701E5108C90D8FD8A" box="[479,505,511,536]" italics="true" pageId="21" pageNumber="272">pv</emphasis>
setae.
</paragraph>
</subSubSection>
<caption id="68CC9CFEFFABFFF700AD16339327F8EF" ID-DOI="http://doi.org/10.5281/zenodo.7960110" ID-Zenodo-Dep="7960110" httpUri="https://zenodo.org/record/7960110/files/figure.png" pageId="21" pageNumber="272" startId="21.[151,250,1856,1880]" targetBox="[160,1396,593,1824]" targetPageId="21" targetType="figure">
<paragraph id="3C0CCC76FFABFFF700AD16339327F8EF" blockId="21.[151,1436,1856,1917]" pageId="21" pageNumber="272">
<emphasis id="0EC71064FFABFFF700AD16339001F8CA" bold="true" box="[151,288,1856,1880]" pageId="21" pageNumber="272">FIGURE 13.</emphasis>
<taxonomicName id="FBB3B7F5FFABFFF7011C16329317F8CB" authority="(Tiensuu)" baseAuthorityName="Tiensuu" baseAuthorityYear="1939" box="[294,566,1857,1881]" class="Insecta" family="Muscidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="272" phylum="Arthropoda" rank="species" species="atlantica">
<emphasis id="0EC71064FFABFFF7011C163290EDF8CB" box="[294,460,1857,1881]" italics="true" pageId="21" pageNumber="272">Helina atlantica</emphasis>
(Tiensuu)
</taxonomicName>
, male. Head, dorso-lateral (
<emphasis id="0EC71064FFABFFF7036A16339243F8CA" bold="true" box="[848,866,1856,1880]" pageId="21" pageNumber="272">A</emphasis>
); abdomen (
<emphasis id="0EC71064FFABFFF703D9163292D5F8CB" bold="true" box="[995,1012,1857,1881]" pageId="21" pageNumber="272">B</emphasis>
); sternite V (
<emphasis id="0EC71064FFABFFF70441163395ACF8CA" bold="true" box="[1147,1165,1856,1880]" pageId="21" pageNumber="272">C</emphasis>
); hypopygium, caudal (
<emphasis id="0EC71064FFABFFF705BB163294B2F8CB" bold="true" box="[1409,1427,1857,1881]" pageId="21" pageNumber="272">D</emphasis>
) and lateral (
<emphasis id="0EC71064FFABFFF7012A16169000F8EF" bold="true" box="[272,289,1893,1917]" pageId="21" pageNumber="272">E</emphasis>
). Same scale for C–E.
</paragraph>
</caption>
<subSubSection id="74A99FFDFFA8FFF400FD11E590CDFE8F" pageId="22" pageNumber="273" type="distribution">
<paragraph id="3C0CCC76FFA8FFF400FD11E59375FF23" blockId="22.[151,1436,150,285]" box="[199,596,150,177]" pageId="22" pageNumber="273">
<emphasis id="0EC71064FFA8FFF400FD11E59040FF22" bold="true" box="[199,353,150,176]" pageId="22" pageNumber="273">Distribution.</emphasis>
Endemic to Madeira.
</paragraph>
<paragraph id="3C0CCC76FFA8FFF400FD11C990CDFE8F" blockId="22.[151,1436,150,285]" pageId="22" pageNumber="273">
<emphasis id="0EC71064FFA8FFF400FD11C990BFFF47" bold="true" box="[199,414,186,213]" pageId="22" pageNumber="273">Taxonomic notes.</emphasis>
The enlarged setae on sternite V and elongated cerci and surstyli are obvious male synapomorphies that
<taxonomicName id="FBB3B7F5FFA8FFF401B711AC9366FF6B" baseAuthorityName="Tiensuu" baseAuthorityYear="1939" box="[397,583,223,249]" class="Insecta" family="Muscidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="atlantica">
<emphasis id="0EC71064FFA8FFF401B711AC9366FF6B" box="[397,583,223,249]" italics="true" pageId="22" pageNumber="273">Helina atlantica</emphasis>
</taxonomicName>
shares with
<taxonomicName id="FBB3B7F5FFA8FFF402E611AC9244FF6B" box="[732,869,223,249]" class="Insecta" family="Muscidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="vilissima">
<emphasis id="0EC71064FFA8FFF402E611AC9244FF6B" box="[732,869,223,249]" italics="true" pageId="22" pageNumber="273">H. vilissima</emphasis>
</taxonomicName>
, also endemic to Madeira, and
<taxonomicName id="FBB3B7F5FFA8FFF404EC11AC945EFF6B" baseAuthorityName="Stein" baseAuthorityYear="1907" box="[1238,1407,223,249]" class="Insecta" family="Muscidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="parcepilosa">
<emphasis id="0EC71064FFA8FFF404EC11AC945EFF6B" box="[1238,1407,223,249]" italics="true" pageId="22" pageNumber="273">H. parcepilosa</emphasis>
</taxonomicName>
, a widespread Palearctic species.
</paragraph>
</subSubSection>
</treatment>
</document>
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