treatments-xml/data/3E/38/0C/3E380C57FFC44A7627ACB07DFB40E0DC.xml

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<mods:title id="3F268F5CEDF3FD484E8EBCEE67099864">Scydmaenus Latreille of Australia: Revision of species in subgenera Choleropsis Franz, Kingius Franz, Mascarensia Franz, Parallomicrus Franz, Scottiscydmaenus Franz, and description of Ascydmaenus subgen. n. (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<paragraph id="B62EBD41FFC44A7227ACB07DFD24E47E" blockId="49.[151,763,529,556]" box="[151,763,529,556]" pageId="49" pageNumber="50">
<heading id="ED660A2DFFC44A7227ACB07DFD24E47E" bold="true" box="[151,763,529,556]" fontSize="11" level="2" pageId="49" pageNumber="50" reason="4">
<taxonomicName id="7191C6C2FFC44A7227ACB07DFD24E47E" ID-CoL="4W3XD" authority="Franz" authorityName="Franz" box="[151,763,529,556]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="clarkianus" subGenus="Scottiscydmaenus">
<emphasis id="84E56153FFC44A7227ACB07DFD24E47E" bold="true" box="[151,763,529,556]" pageId="49" pageNumber="50">
<emphasis id="84E56153FFC44A7227ACB07DFEF0E47E" bold="true" box="[151,303,529,556]" italics="true" pageId="49" pageNumber="50">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFC44A72267BB07DFDC8E47E" bold="true" box="[320,535,529,556]" italics="true" pageId="49" pageNumber="50">Scottiscydmaenus</emphasis>
) 
<emphasis id="84E56153FFC44A722513B07DFD76E47E" bold="true" box="[552,681,529,556]" italics="true" pageId="49" pageNumber="50">clarkianus</emphasis>
Franz
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFC44A7227ACB03AFE34E4C1" pageId="49" pageNumber="50" type="reference_group">
<paragraph id="B62EBD41FFC44A7227ACB03AFCD4E43C" blockId="49.[151,779,598,659]" box="[151,779,598,622]" pageId="49" pageNumber="50">
<treatmentCitationGroup id="96819A6FFFC44A7227ACB03AFCD4E43C" box="[151,779,598,622]" pageId="49" pageNumber="50">
<treatmentCitation id="37309B50FFC44A7227ACB03AFCD8E43C" author="Franz, H." box="[151,775,598,622]" page="278" pageId="49" pageNumber="50" year="1975">
<taxonomicName id="7191C6C2FFC44A7227ACB03AFCD8E43C" ID-CoL="4W3XD" authority="Franz, 1975: 278" authorityName="Franz" authorityPageNumber="278" authorityYear="1975" box="[151,775,598,622]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="clarkianus" subGenus="Scottiscydmaenus">
<emphasis id="84E56153FFC44A7227ACB03AFECBE43C" box="[151,276,598,622]" italics="true" pageId="49" pageNumber="50">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFC44A722618B03AFE08E43C" box="[291,471,598,622]" italics="true" pageId="49" pageNumber="50">Scottiscydmaenus</emphasis>
) 
<emphasis id="84E56153FFC44A7226DDB03AFD8EE43C" box="[486,593,598,622]" italics="true" pageId="49" pageNumber="50">clarkianus</emphasis>
<bibRefCitation id="D200C0B0FFC44A72256CB03BFCD8E43C" author="Franz, H." box="[599,775,598,622]" pageId="49" pageNumber="50" pagination="1 - 312" refId="ref50046" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">Franz, 1975: 278</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="B62EBD41FFC44A7227ACB014FE34E4C1" blockId="49.[151,779,598,659]" box="[151,491,632,659]" pageId="49" pageNumber="50">
(
<figureCitation id="2EAAA1C4FFC44A7227A4B014FEE3E4C1" box="[159,316,632,659]" captionStart-0="FIGURES 125–126" captionStart-1="FIGURES 127–129" captionStart-2="FIGURES 130–134" captionStartId-0="50.[151,264,1744,1768]" captionStartId-1="51.[151,264,1361,1385]" captionStartId-2="52.[151,264,1084,1108]" captionTargetBox-0="[151,1436,181,1721]" captionTargetBox-1="[151,1436,181,1337]" captionTargetBox-2="[157,1430,187,1054]" captionTargetId-0="figure-21@50.[151,1436,181,1721]" captionTargetId-1="figure-15@51.[151,1436,181,1337]" captionTargetId-2="figure-21@52.[151,1436,181,1060]" captionTargetPageId-0="50" captionTargetPageId-1="51" captionTargetPageId-2="52" captionText-0="FIGURES 125–126. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Head, pronotum and elytral base in dorsal view (125); head and prothorax in ventral view (126). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." captionText-1="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." captionText-2="FIGURES 130–134. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Dorsal habitus (130); aedeagus in dorsal (131, 133) and lateral (132, 134) views." figureDoi-0="http://doi.org/10.5281/zenodo.10144699" figureDoi-1="http://doi.org/10.5281/zenodo.10144701" figureDoi-2="http://doi.org/10.5281/zenodo.10144703" httpUri-0="https://zenodo.org/record/10144699/files/figure.png" httpUri-1="https://zenodo.org/record/10144701/files/figure.png" httpUri-2="https://zenodo.org/record/10144703/files/figure.png" pageId="49" pageNumber="50">Figs 125–134</figureCitation>
, 
<figureCitation id="2EAAA1C4FFC44A722673B014FE73E4C1" box="[328,428,632,659]" captionStart="FIGURES 155–162" captionStartId="60.[151,264,1338,1362]" captionTargetBox="[151,1435,181,1302]" captionTargetId="figure-21@60.[151,1435,181,1314]" captionTargetPageId="60" captionText="FIGURES 155–162. Scottiscydmaenus, heads in dorsal view (155–158) and profemora of males (159–162). Scydmaenus clar- kianus Franz (155, 159); S. optatus Sharp (156, 160); S. swanensis Franz (157, 161); S. brachyponerophilus sp. n. (158, 162)." figureDoi="http://doi.org/10.5281/zenodo.10144711" httpUri="https://zenodo.org/record/10144711/files/figure.png" pageId="49" pageNumber="50">155, 159</figureCitation>
, 
<figureCitation id="2EAAA1C4FFC44A722683B015FE3BE4C1" box="[440,484,633,659]" captionStart="FIGURES 189–205" captionStartId="71.[151,264,1644,1668]" captionTargetBox="[157,1429,187,1614]" captionTargetId="figure-15@71.[151,1436,181,1620]" captionTargetPageId="71" captionText="FIGURES 189–205. Historical labels of primary type specimens. Lectotype of Heterognathus gracilis King (189); holotype of Scydmaenus beechmonti Franz (190); lectotype of Heterognathus geniculatus King (191); holotype of Scydmaenus kurandae Franz (192); lectotype of Heterognathus princeps King (193); holotype of Scydmaenus marwillumbali Franz [© Lisha Jasper, (BM)] (194); lectotype of Heterognathus carinatus King (195); holotype of Scydmaenus formicarum Franz (196); holotype of Scydmaenus australiensis Franz (197); lectotype of Heterognathus myrmecophilus Lea (198); holotype of Scydmaenus inflatitibia Franz (199); holotype of Scydmaenus clarkianus Franz (200); lectotype of Scydmaenus optatus Sharp (201); holotype of Scydmaenus scotti Franz (202); lectotype of Scydmaenus swanensis Franz (203); holotype of Scydmaenus brachyponerophilus sp. n. (204); syntype of Heterognathus armitagei King [© Natalie Tees (AMS)] (205)." figureDoi="http://doi.org/10.5281/zenodo.10144723" httpUri="https://zenodo.org/record/10144723/files/figure.png" pageId="49" pageNumber="50">200</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFC44A7727ACB0ADFEADE37E" lastPageId="52" lastPageNumber="53" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="B62EBD41FFC44A7227ACB0ADFB31E515" blockId="49.[151,1437,704,2027]" pageId="49" pageNumber="50">
<emphasis id="84E56153FFC44A7227ACB0ADFE77E488" bold="true" box="[151,424,704,731]" pageId="49" pageNumber="50">Type material studied.</emphasis>
<materialsCitation id="06F9B71CFFC44A72268AB0ADFB31E515" collectingDate="2013-08-21" collectionCode="SAMA" collectorName="J. Clark" country="Australia" determinerName="H. Franz" location="WESTERN AUSTRALIA" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="84E56153FFC44A72268AB0ADFBECE488" bold="true" box="[433,1075,704,731]" pageId="49" pageNumber="50">
<typeStatus id="692A03E3FFC44A72268AB0ADFDC1E489" box="[433,542,705,731]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="CE86FDD1FFC44A722514B0ACFD05E488" box="[559,730,704,730]" name="Australia" pageId="49" pageNumber="50">AUSTRALIA</collectingCountry>
: 
<collectingRegion id="745573A3FFC44A7225DCB0ACFBF9E488" box="[743,1062,704,730]" country="Australia" name="Western Australia" pageId="49" pageNumber="50">WESTERN AUSTRALIA</collectingRegion>
):
</emphasis>
♂ (
<figureCitation id="2EAAA1C4FFC44A722358B0ACFB19E489" box="[1123,1222,704,731]" captionStart="FIGURES 130–134" captionStartId="52.[151,264,1084,1108]" captionTargetBox="[157,1430,187,1054]" captionTargetId="figure-21@52.[151,1436,181,1060]" captionTargetPageId="52" captionText="FIGURES 130–134. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Dorsal habitus (130); aedeagus in dorsal (131, 133) and lateral (132, 134) views." figureDoi="http://doi.org/10.5281/zenodo.10144703" httpUri="https://zenodo.org/record/10144703/files/figure.png" pageId="49" pageNumber="50">Fig. 130</figureCitation>
), with labels illustrated in 
<figureCitation id="2EAAA1C4FFC44A7227C5B088FEBEE4AD" box="[254,353,740,767]" captionStart="FIGURES 189–205" captionStartId="71.[151,264,1644,1668]" captionTargetBox="[157,1429,187,1614]" captionTargetId="figure-15@71.[151,1436,181,1620]" captionTargetPageId="71" captionText="FIGURES 189–205. Historical labels of primary type specimens. Lectotype of Heterognathus gracilis King (189); holotype of Scydmaenus beechmonti Franz (190); lectotype of Heterognathus geniculatus King (191); holotype of Scydmaenus kurandae Franz (192); lectotype of Heterognathus princeps King (193); holotype of Scydmaenus marwillumbali Franz [© Lisha Jasper, (BM)] (194); lectotype of Heterognathus carinatus King (195); holotype of Scydmaenus formicarum Franz (196); holotype of Scydmaenus australiensis Franz (197); lectotype of Heterognathus myrmecophilus Lea (198); holotype of Scydmaenus inflatitibia Franz (199); holotype of Scydmaenus clarkianus Franz (200); lectotype of Scydmaenus optatus Sharp (201); holotype of Scydmaenus scotti Franz (202); lectotype of Scydmaenus swanensis Franz (203); holotype of Scydmaenus brachyponerophilus sp. n. (204); syntype of Heterognathus armitagei King [© Natalie Tees (AMS)] (205)." figureDoi="http://doi.org/10.5281/zenodo.10144723" httpUri="https://zenodo.org/record/10144723/files/figure.png" pageId="49" pageNumber="50">Fig. 200</figureCitation>
: “Perth / 
<collectorName id="1B64D897FFC44A7226F5B089FDF6E4AC" box="[462,553,740,767]" pageId="49" pageNumber="50">J. Clark</collectorName>
” [brownish, handwritten], “ 
<taxonomicName id="7191C6C2FFC44A72245EB088FC2AE4AC" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[869,1013,740,766]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (
<taxonomicName id="7191C6C2FFC44A72232EB088FB3CE4AC" box="[1045,1251,740,766]" pageId="49" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">Scottiscydmaenus</taxonomicName>
) / clarkianus m. / det. 
<determinerName id="43F25185FFC44A7227EBB164FEF1E570" box="[208,302,776,802]" pageId="49" pageNumber="50">H.Franz</determinerName>
” [white, handwritten and printed], “Typus” [red, handwritten], “SAMA Database / No. 25-037033” [white, printed], “SAMA Digital Image / 
<date id="C22F9B81FFC44A72255CB141FCDFE515" box="[615,768,813,839]" pageId="49" pageNumber="50" value="2013-08-21">
<collectingDate id="D26B6269FFC44A72255CB141FCDFE515" box="[615,768,813,839]" pageId="49" pageNumber="50" value="2013-08-21">21. Aug 2013</collectingDate>
</date>
” [green, printed and handwritten] (
<collectionCode id="D0802584FFC44A7223B0B141FB3DE515" box="[1163,1250,813,839]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="49" pageNumber="50" type="Museum">SAMA</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="B62EBD41FFC44A7227FCB13DFC53E1B9" blockId="49.[151,1437,704,2027]" pageId="49" pageNumber="50">
<emphasis id="84E56153FFC44A7227FCB13DFD21E538" bold="true" box="[199,766,848,875]" pageId="49" pageNumber="50">
<typeStatus id="692A03E3FFC44A7227FCB13DFEE0E539" box="[199,319,849,875]" pageId="49" pageNumber="50" type="paratype">Paratypes</typeStatus>
(39 exx.). 
<collectingRegion id="745573A3FFC44A722686B13CFD26E538" box="[445,761,848,874]" country="Australia" name="Western Australia" pageId="49" pageNumber="50">WESTERN AUSTRALIA</collectingRegion>
:
</emphasis>
21 exx. mounted on 3 large cardboards, 7 on each, on one pin, with additional card with 4 ants (most likely 
<taxonomicName id="7191C6C2FFC44A7225E8B118FC6AE5DC" box="[723,949,884,910]" class="Insecta" family="Formicidae" genus="Brachyponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="84E56153FFC44A7225E8B118FC6AE5DC" box="[723,949,884,910]" italics="true" pageId="49" pageNumber="50">Brachyponera lutea</emphasis>
</taxonomicName>
), “Swan R. / J. S. Clark” [white, printed], “ 
<taxonomicName id="7191C6C2FFC44A72279AB1F4FEEEE5E0" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[161,305,920,946]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (
<taxonomicName id="7191C6C2FFC44A72266BB1F4FDC1E5E0" box="[336,542,920,946]" pageId="49" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">Scottiscydmaenus</taxonomicName>
) / clarki {sic!} m. / 
<typeStatus id="692A03E3FFC44A72243DB1F5FC78E5E1" box="[774,935,921,947]" pageId="49" pageNumber="50" type="paratype">PARATYPUS</typeStatus>
” [yellow, handwritten and printed], “SAMA 25- 43072” [white, printed] (
<collectionCode id="D0802584FFC44A7226D2B1D1FDE0E585" box="[489,575,957,983]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="49" pageNumber="50" type="Museum">SAMA</collectionCode>
; another mounting card with 
<specimenCount id="A09776C8FFC44A7224A1B1D1FBF7E584" box="[922,1064,956,983]" count="4" pageId="49" pageNumber="50" type="generic">4 specimens</specimenCount>
originally on the same pin was pinned separately during the current study and labelled “Swan R. / J. S. Clark” [white, printed], “part of / SAMA 25- 43072” [white, printed]— 
<specimenCount id="A09776C8FFC44A722687B668FD9CE24C" box="[444,579,1028,1054]" count="3" pageId="49" pageNumber="50" type="generic">3 specimens</specimenCount>
on the left are 
<taxonomicName id="7191C6C2FFC44A7225D2B668FC45E24D" box="[745,922,1028,1055]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="84E56153FFC44A7225D2B668FCACE24C" box="[745,883,1028,1054]" italics="true" pageId="49" pageNumber="50">Scydmaenus</emphasis>
sp.
</taxonomicName>
[not 
<taxonomicName id="7191C6C2FFC44A7224EFB669FBB9E24C" box="[980,1126,1028,1055]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="clarkianus">
<emphasis id="84E56153FFC44A7224EFB669FBB9E24C" box="[980,1126,1028,1055]" italics="true" pageId="49" pageNumber="50">S. clarkianus</emphasis>
</taxonomicName>
], 
<specimenCount id="A09776C8FFC44A722346B669FAC9E24C" box="[1149,1302,1028,1055]" count="1" pageId="49" pageNumber="50" type="generic">one specimen</specimenCount>
on the right is 
<taxonomicName id="7191C6C2FFC44A722788B644FE0CE211" authority="Franz" authorityName="Franz. Despite" authorityYear="1975" box="[179,467,1064,1091]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="mirus">
<emphasis id="84E56153FFC44A722788B644FE59E210" box="[179,390,1064,1090]" italics="true" pageId="49" pageNumber="50">Scydmaenus mirus</emphasis>
Franz
</taxonomicName>
); 
<specimenCount id="A09776C8FFC44A7226D3B645FDCBE211" box="[488,532,1065,1091]" count="1" pageId="49" pageNumber="50" type="female">1 ♀</specimenCount>
, “Swan R. / W Australia / J. Clark” [white, handwritten], “ 
<specimenCount id="A09776C8FFC44A722386B645FB10E211" box="[1213,1231,1065,1091]" count="1" pageId="49" pageNumber="50" type="female">♀</specimenCount>
” [white, printed], “ 
<taxonomicName id="7191C6C2FFC44A72279AB620FEEEE234" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[161,305,1100,1126]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (
<taxonomicName id="7191C6C2FFC44A722676B620FDC4E234" box="[333,539,1100,1126]" pageId="49" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">Scottiscydmaenus</taxonomicName>
) / clarkianus m. / 
<typeStatus id="692A03E3FFC44A7225D9B621FC5CE235" box="[738,899,1101,1127]" pageId="49" pageNumber="50" type="paratype">PARATYPUS</typeStatus>
” [yellow, handwritten and printed], “SAMA 25- 43082” [white, printed] (
<collectionCode id="D0802584FFC44A722691B61DFDDEE2D9" box="[426,513,1137,1163]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="49" pageNumber="50" type="Museum">SAMA</collectionCode>
); 
<specimenCount id="A09776C8FFC44A722528B61DFDE3E2D9" box="[531,572,1137,1163]" count="1" pageId="49" pageNumber="50" type="female">1 ♀</specimenCount>
, with 
<taxonomicName id="7191C6C2FFC44A7225BBB61CFC81E2D8" box="[640,862,1136,1162]" class="Insecta" family="Formicidae" genus="Brachyponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="84E56153FFC44A7225BBB61CFC81E2D8" box="[640,862,1136,1162]" italics="true" pageId="49" pageNumber="50">Brachyponera lutea</emphasis>
</taxonomicName>
, “Swan R. / W Australia / J. Clark” [white, printed], “ 
<taxonomicName id="7191C6C2FFC44A72279AB6F8FEEEE2FC" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[161,305,1172,1198]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ Pubesc much shorter / than in optatus / not-microps and has / 4jnt club” [brownish, handwritten], “ 
<taxonomicName id="7191C6C2FFC44A72279AB6D4FEEEE280" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[161,305,1208,1234]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (
<taxonomicName id="7191C6C2FFC44A722676B6D4FDC4E280" box="[333,539,1208,1234]" pageId="49" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">Scottiscydmaenus</taxonomicName>
) / clarkianus m. / 
<typeStatus id="692A03E3FFC44A7225D9B6D5FC5CE281" box="[738,899,1209,1235]" pageId="49" pageNumber="50" type="paratype">PARATYPUS</typeStatus>
” [yellow, handwritten and printed], “SAMA 25- 43075” [white, printed] (
<collectionCode id="D0802584FFC44A722695B6B1FDDAE2A5" box="[430,517,1245,1271]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="49" pageNumber="50" type="Museum">SAMA</collectionCode>
); 
<specimenCount id="A09776C8FFC44A722523B6B1FD9DE2A5" box="[536,578,1245,1271]" count="1" pageId="49" pageNumber="50" type="female">1 ♀</specimenCount>
, with unidentified ants,, “Swan R. / W Australia / J. Clark” [white, printed], 
<taxonomicName id="7191C6C2FFC44A7227ACB76CFEFAE348" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[151,293,1280,1306]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (
<taxonomicName id="7191C6C2FFC44A722671B76CFDC7E348" box="[330,536,1280,1306]" pageId="49" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">Scottiscydmaenus</taxonomicName>
) / clarkianus m. / 
<typeStatus id="692A03E3FFC44A7225CDB76DFC48E349" box="[758,919,1281,1307]" pageId="49" pageNumber="50" type="paratype">PARATYPUS</typeStatus>
” [yellow, handwritten and printed], “SAMA 25- 43080” [white, printed] (
<collectionCode id="D0802584FFC44A7226D9B749FDE6E36D" box="[482,569,1317,1343]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="49" pageNumber="50" type="Museum">SAMA</collectionCode>
); 
<specimenCount id="A09776C8FFC44A722575B749FDA5E36D" box="[590,634,1317,1343]" count="1" pageId="49" pageNumber="50" type="female">1 ♀</specimenCount>
, “Swan R. / W Australia / J. Clark” [white, handwritten], “ 
<specimenCount id="A09776C8FFC44A72221EB749FAE8E36D" box="[1317,1335,1317,1343]" count="1" pageId="49" pageNumber="50" type="female">♀</specimenCount>
” [white, printed], “ 
<taxonomicName id="7191C6C2FFC44A722631B724FE45E330" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[266,410,1352,1378]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (
<taxonomicName id="7191C6C2FFC44A722686B724FD54E330" box="[445,651,1352,1378]" pageId="49" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">Scottiscydmaenus</taxonomicName>
) / clarkianus m. / 
<typeStatus id="692A03E3FFC44A722458B725FBDBE331" box="[867,1028,1353,1379]" pageId="49" pageNumber="50" type="paratype">PARATYPUS</typeStatus>
” [yellow, handwritten and printed], “SAMA 25- 43083” [white, printed] (
<collectionCode id="D0802584FFC44A722578B701FD45E3D5" box="[579,666,1389,1415]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="49" pageNumber="50" type="Museum">SAMA</collectionCode>
); 
<specimenCount id="A09776C8FFC44A722594B701FD05E3D5" box="[687,730,1389,1415]" count="1" pageId="49" pageNumber="50" type="female">1 ♀</specimenCount>
, with 
<taxonomicName id="7191C6C2FFC44A72241FB700FBDBE3D4" box="[804,1028,1388,1414]" class="Insecta" family="Formicidae" genus="Brachyponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="84E56153FFC44A72241FB700FBDBE3D4" box="[804,1028,1388,1414]" italics="true" pageId="49" pageNumber="50">Brachyponera lutea</emphasis>
</taxonomicName>
, “Swan R. / W Australia / J. Clark” [white, handwritten], “ 
<specimenCount id="A09776C8FFC44A7226A5B7FDFE6FE3F9" box="[414,432,1425,1451]" count="1" pageId="49" pageNumber="50" type="female">♀</specimenCount>
” [white, printed], “ 
<taxonomicName id="7191C6C2FFC44A7225AEB7FCFCFAE3F8" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[661,805,1424,1450]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (
<taxonomicName id="7191C6C2FFC44A722471B7FCFBC7E3F8" box="[842,1048,1424,1450]" pageId="49" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">Scottiscydmaenus</taxonomicName>
) / clarkianus m. / 
<typeStatus id="692A03E3FFC44A7223C9B7FDFA4CE3F9" box="[1266,1427,1425,1451]" pageId="49" pageNumber="50" type="paratype">PARATYPUS</typeStatus>
” [yellow, handwritten and printed], “SAMA 25- 43078” [white, printed] (
<collectionCode id="D0802584FFC44A7224E1B7D9FBEEE39D" box="[986,1073,1461,1487]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="49" pageNumber="50" type="Museum">SAMA</collectionCode>
); 6 exx. on 4 cards on one pin, with 
<taxonomicName id="7191C6C2FFC44A7227E9B7B4FE6DE3A0" box="[210,434,1496,1522]" class="Insecta" family="Formicidae" genus="Brachyponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="84E56153FFC44A7227E9B7B4FE6DE3A0" box="[210,434,1496,1522]" italics="true" pageId="49" pageNumber="50">Brachyponera lutea</emphasis>
</taxonomicName>
, “Swan R. / J. S. Clark” [white, printed], “SAMA 25- 43079” [white, printed] (
<collectionCode id="D0802584FFC44A722279B7B5FA47E3A1" box="[1346,1432,1497,1523]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="49" pageNumber="50" type="Museum">SAMA</collectionCode>
; another mounting card with 
<specimenCount id="A09776C8FFC44A7226EEB791FD80E044" box="[469,607,1532,1559]" count="3" pageId="49" pageNumber="50" type="generic">3 specimens</specimenCount>
originally on the same pin was pinned separately during the current study and labelled “Swan R. / J. S. Clark” [white, printed], “part of / SAMA 25-43079” [white, printed]—specimen on the left is 
<taxonomicName id="7191C6C2FFC44A7227E5B428FE50E00D" box="[222,399,1604,1631]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="84E56153FFC44A7227E5B428FEB7E00C" box="[222,360,1604,1630]" italics="true" pageId="49" pageNumber="50">Scydmaenus</emphasis>
sp.
</taxonomicName>
[not 
<taxonomicName id="7191C6C2FFC44A7226F0B429FD82E00C" box="[459,605,1604,1631]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="clarkianus">
<emphasis id="84E56153FFC44A7226F0B429FD82E00C" box="[459,605,1604,1631]" italics="true" pageId="49" pageNumber="50">S. clarkianus</emphasis>
</taxonomicName>
], specimen in middle and on the right are 
<taxonomicName id="7191C6C2FFC44A722301B428FA87E00D" authority="Franz" authorityName="Franz. Despite" authorityYear="1975" box="[1082,1368,1604,1631]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="mirus">
<emphasis id="84E56153FFC44A722301B428FAD3E00C" box="[1082,1292,1604,1630]" italics="true" pageId="49" pageNumber="50">Scydmaenus mirus</emphasis>
Franz
</taxonomicName>
); 
<specimenCount id="A09776C8FFC44A722250B429FA4AE00D" box="[1387,1429,1605,1631]" count="1" pageId="49" pageNumber="50" type="female">1 ♀</specimenCount>
, “Swan R. / W Australia / J. Clark” [white, handwritten], “ 
<specimenCount id="A09776C8FFC44A722425B405FCEFE0D1" box="[798,816,1641,1667]" count="1" pageId="49" pageNumber="50" type="female">♀</specimenCount>
” [white, printed], “ 
<taxonomicName id="7191C6C2FFC44A722330B404FB44E0D0" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[1035,1179,1640,1666]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (
<taxonomicName id="7191C6C2FFC44A722383B404FA59E0D0" box="[1208,1414,1640,1666]" pageId="49" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">Scottiscydmaenus</taxonomicName>
) / clarkianus m. / 
<typeStatus id="692A03E3FFC44A72267EB4E1FE39E0F5" box="[325,486,1677,1703]" pageId="49" pageNumber="50" type="paratype">PARATYPUS</typeStatus>
” [yellow, handwritten and printed], “SAMA 25- 43076” [white, printed] (
<collectionCode id="D0802584FFC44A722218B4E1FAA5E0F5" box="[1315,1402,1677,1703]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="49" pageNumber="50" type="Museum">SAMA</collectionCode>
); 
<specimenCount id="A09776C8FFC44A7222B5B4E1FF64E099" count="1" pageId="49" pageNumber="50" type="generic">1 ex.</specimenCount>
, with 
<taxonomicName id="7191C6C2FFC44A7227C4B4DCFE01E098" box="[255,478,1712,1738]" class="Insecta" family="Formicidae" genus="Brachyponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="84E56153FFC44A7227C4B4DCFE01E098" box="[255,478,1712,1738]" italics="true" pageId="49" pageNumber="50">Brachyponera lutea</emphasis>
</taxonomicName>
, “Swan R. / W Australia / J. Clark” [white, handwritten], “ 
<taxonomicName id="7191C6C2FFC44A722354B4DCFB20E098" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[1135,1279,1712,1738]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (
<taxonomicName id="7191C6C2FFC44A722227B4DCFF31E0BD" pageId="49" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">Scottiscydmaenus</taxonomicName>
) / clarkianus m. / 
<typeStatus id="692A03E3FFC44A7226FAB4B9FDBDE0BD" box="[449,610,1749,1775]" pageId="49" pageNumber="50" type="paratype">PARATYPUS</typeStatus>
” [yellow, handwritten and printed], “SAMA 25- 43081” [white, printed] (
<collectionCode id="D0802584FFC44A7227A4B495FF29E141" box="[159,246,1785,1811]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="49" pageNumber="50" type="Museum">SAMA</collectionCode>
); 
<specimenCount id="A09776C8FFC44A722630B495FEE9E141" box="[267,310,1785,1811]" count="1" pageId="49" pageNumber="50" type="female">1 ♀</specimenCount>
, with 
<taxonomicName id="7191C6C2FFC44A722644B494FD80E140" box="[383,607,1784,1810]" class="Insecta" family="Formicidae" genus="Brachyponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="84E56153FFC44A722644B494FD80E140" box="[383,607,1784,1810]" italics="true" pageId="49" pageNumber="50">Brachyponera lutea</emphasis>
</taxonomicName>
, “Swan R. / W Australia / J. Clark” [white, handwritten], “ 
<taxonomicName id="7191C6C2FFC44A7223C7B494FA53E140" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[1276,1420,1784,1810]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (
<taxonomicName id="7191C6C2FFC44A7227A4B570FEB2E164" box="[159,365,1820,1846]" pageId="49" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">Scottiscydmaenus</taxonomicName>
) / clarkianus m. / 
<typeStatus id="692A03E3FFC44A72250FB571FD0AE165" box="[564,725,1821,1847]" pageId="49" pageNumber="50" type="paratype">PARATYPUS</typeStatus>
” [yellow, handwritten and printed], “SAMA 25- 43084” [white, printed] (
<collectionCode id="D0802584FFC44A7227C4B52DFE89E109" box="[255,342,1857,1883]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="49" pageNumber="50" type="Museum">SAMA</collectionCode>
); 3 exx. mounted on one card, “Perth / J. Clark” [brownish, handwritten], “ 
<taxonomicName id="7191C6C2FFC44A7223A7B52CFAF3E108" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[1180,1324,1856,1882]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ microps Lea / Id. by A.M. Lea” [brownish, handwritten], “ 
<taxonomicName id="7191C6C2FFC44A7225EDB508FCB9E12C" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[726,870,1892,1918]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (
<taxonomicName id="7191C6C2FFC44A7224B3B508FB89E12C" box="[904,1110,1892,1918]" pageId="49" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">Scottiscydmaenus</taxonomicName>
) / clarkianus m. / PARATY- PUS” [yellow, handwritten and printed], “SAMA 25- 43070” [white, printed] (
<collectionCode id="D0802584FFC44A722333B5E5FB80E1F1" box="[1032,1119,1929,1955]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="49" pageNumber="50" type="Museum">SAMA</collectionCode>
); 
<specimenCount id="A09776C8FFC44A722348B5E5FB42E1F1" box="[1139,1181,1929,1955]" count="1" pageId="49" pageNumber="50" type="female">1 ♀</specimenCount>
, “Perth / W Australia / J. Clark” [white, handwritten], “ 
<taxonomicName id="7191C6C2FFC44A72253BB5C0FD4FE194" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[512,656,1964,1990]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (
<taxonomicName id="7191C6C2FFC44A722595B5C0FCA3E194" box="[686,892,1964,1990]" pageId="49" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">Scottiscydmaenus</taxonomicName>
) / clarkianus m. / 
<typeStatus id="692A03E3FFC44A722372B5C1FB35E195" box="[1097,1258,1965,1991]" pageId="49" pageNumber="50" type="paratype">PARATYPUS</typeStatus>
” [yellow, handwritten and printed], “SAMA 25- 43074” [white, printed] (
<collectionCode id="D0802584FFC44A722412B5BDFC5FE1B9" box="[809,896,2001,2027]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="49" pageNumber="50" type="Museum">SAMA</collectionCode>
).
</paragraph>
<caption id="E2EEEDC9FFC74A7127ACB4BCFADEE107" ID-DOI="http://doi.org/10.5281/zenodo.10144699" ID-Zenodo-Dep="10144699" httpUri="https://zenodo.org/record/10144699/files/figure.png" pageId="50" pageNumber="51" startId="50.[151,264,1744,1768]" targetBox="[151,1436,181,1721]" targetPageId="50" targetType="figure">
<paragraph id="B62EBD41FFC74A7127ACB4BCFADEE107" blockId="50.[151,1437,1744,1877]" pageId="50" pageNumber="51">
<emphasis id="84E56153FFC74A7127ACB4BCFEB3E0BA" bold="true" box="[151,364,1744,1768]" pageId="50" pageNumber="51">FIGURES 125–126.</emphasis>
<taxonomicName id="7191C6C2FFC74A71264AB4BDFCB5E0BA" authority="Franz" authorityName="Franz" box="[369,874,1745,1769]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="clarkianus" subGenus="Scottiscydmaenus">
<emphasis id="84E56153FFC74A71264AB4BDFE31E0BB" box="[369,494,1745,1769]" italics="true" pageId="50" pageNumber="51">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFC74A7126C7B4BDFD6FE0BB" box="[508,688,1745,1769]" italics="true" pageId="50" pageNumber="51">Scottiscydmaenus</emphasis>
) 
<emphasis id="84E56153FFC74A712585B4BDFCF6E0BB" box="[702,809,1745,1769]" italics="true" pageId="50" pageNumber="51">clarkianus</emphasis>
Franz
</taxonomicName>
, holotype male. Head, pronotum and elytral base in dorsal view (
<emphasis id="84E56153FFC74A7127C1B498FEFFE15E" bold="true" box="[250,288,1780,1804]" pageId="50" pageNumber="51">125</emphasis>
); head and prothorax in ventral view (
<emphasis id="84E56153FFC74A712593B498FD11E15E" bold="true" box="[680,718,1780,1804]" pageId="50" pageNumber="51">126</emphasis>
). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum.
</paragraph>
</caption>
<caption id="E2EEEDC9FFC64A7027ACB73DFCDCE386" ID-DOI="http://doi.org/10.5281/zenodo.10144701" ID-Zenodo-Dep="10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="51" pageNumber="52" startId="51.[151,264,1361,1385]" targetBox="[151,1436,181,1337]" targetPageId="51" targetType="figure">
<paragraph id="B62EBD41FFC64A7027ACB73DFCDCE386" blockId="51.[151,1436,1361,1493]" pageId="51" pageNumber="52">
<emphasis id="84E56153FFC64A7027ACB73DFEB2E33B" bold="true" box="[151,365,1361,1385]" pageId="51" pageNumber="52">FIGURES 127–129.</emphasis>
<taxonomicName id="7191C6C2FFC64A702648B73DFCB1E33A" authority="Franz" authorityName="Franz" box="[371,878,1361,1385]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="clarkianus" subGenus="Scottiscydmaenus">
<emphasis id="84E56153FFC64A702648B73DFE2FE33B" box="[371,496,1361,1385]" italics="true" pageId="51" pageNumber="52">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFC64A7026C4B73DFD6CE33B" box="[511,691,1361,1385]" italics="true" pageId="51" pageNumber="52">Scottiscydmaenus</emphasis>
) 
<emphasis id="84E56153FFC64A7025FAB73DFCF3E33B" box="[705,812,1361,1385]" italics="true" pageId="51" pageNumber="52">clarkianus</emphasis>
Franz
</taxonomicName>
, holotype male. Distal region of right antenna in dorsal view (
<emphasis id="84E56153FFC64A7027ECB719FF22E3DF" bold="true" box="[215,253,1397,1421]" pageId="51" pageNumber="52">127</emphasis>
); pterothorax and base of abdomen in ventral view (
<emphasis id="84E56153FFC64A70242FB719FCE5E3DF" bold="true" box="[788,826,1397,1421]" pageId="51" pageNumber="52">128</emphasis>
); distal region of right fore leg in dorsal view (
<emphasis id="84E56153FFC64A702220B719FA9EE3DF" bold="true" box="[1307,1345,1397,1421]" pageId="51" pageNumber="52">129</emphasis>
). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus.
</paragraph>
</caption>
<paragraph id="B62EBD41FFC64A7027FCB46FFD6DE0FF" blockId="51.[151,1437,1539,2034]" pageId="51" pageNumber="52">
<emphasis id="84E56153FFC64A7027FCB46FFCBFE04F" bold="true" box="[199,864,1539,1565]" pageId="51" pageNumber="52">
Additional material studied. 
<collectingRegion id="745573A3FFC64A702524B46FFC85E04F" box="[543,858,1539,1565]" country="Australia" name="Western Australia" pageId="51" pageNumber="52">WESTERN AUSTRALIA</collectingRegion>
:
</emphasis>
1 ♂, originally mounted together on one card with specimens identified here as 
<taxonomicName id="7191C6C2FFC64A7026E7B44BFD06E013" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[476,729,1575,1601]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="brachyponerophilus" status="sp. n.">
<emphasis id="84E56153FFC64A7026E7B44BFD06E013" box="[476,729,1575,1601]" italics="true" pageId="51" pageNumber="52">S. brachyponerophilus</emphasis>
</taxonomicName>
<emphasis id="84E56153FFC64A7025DBB44BFCFFE013" bold="true" box="[736,800,1575,1601]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="9FD6DC28FFC64A7025DBB44BFCFFE013" box="[736,800,1575,1601]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, during present study separately mounted and pinned on original short pin, with white printed copy of original locality label “Swan R. / W Australia / J. Clark”, and with one of two original labels “ 
<taxonomicName id="7191C6C2FFC64A7026A3B403FDF7E0DB" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[408,552,1647,1673]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ 
<taxonomicName id="7191C6C2FFC64A702505B41CFD08E0DB" authority="Sharp" authorityName="Sharp" box="[574,727,1647,1674]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="optatus">optatus Sharp</taxonomicName>
/ Id. by A. M. Lea” [white, handwritten and printed], and “part of SAMA 25- 43099” [white, printed] (
<collectionCode id="D0802584FFC64A702574B4FFFD79E0FF" box="[591,678,1683,1709]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="51" pageNumber="52" type="Museum">SAMA</collectionCode>
).
</paragraph>
<paragraph id="B62EBD41FFC64A7027FCB4DBFBE8E1A3" blockId="51.[151,1437,1539,2034]" pageId="51" pageNumber="52">
<emphasis id="84E56153FFC64A7027FCB4DBFC69E083" bold="true" box="[199,950,1719,1746]" pageId="51" pageNumber="52">
Specimens excluded from type series of 
<taxonomicName id="7191C6C2FFC64A70259EB4DBFCE0E080" box="[677,831,1719,1746]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="clarkianus">
<emphasis id="84E56153FFC64A70259EB4DBFCE0E080" bold="true" box="[677,831,1719,1746]" italics="true" pageId="51" pageNumber="52">S. clarkianus</emphasis>
</taxonomicName>
(13 exx.):
</emphasis>
4 exx. mounted on one card and originally pinned as a part of SAMA 25-43072 (see above)— 
<specimenCount id="A09776C8FFC64A7025E3B4B7FCBFE0A7" box="[728,864,1755,1781]" count="3" pageId="51" pageNumber="52" type="generic">3 specimens</specimenCount>
on the left are 
<taxonomicName id="7191C6C2FFC64A702330B4B7FB62E0A4" box="[1035,1213,1755,1782]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="84E56153FFC64A702330B4B7FB4AE0A7" box="[1035,1173,1755,1781]" italics="true" pageId="51" pageNumber="52">Scydmaenus</emphasis>
sp.
</taxonomicName>
[not 
<taxonomicName id="7191C6C2FFC64A7023C2B4B7FA53E0A7" box="[1273,1420,1755,1781]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="clarkianus">
<emphasis id="84E56153FFC64A7023C2B4B7FA53E0A7" box="[1273,1420,1755,1781]" italics="true" pageId="51" pageNumber="52">S. clarkianus</emphasis>
</taxonomicName>
], 
<specimenCount id="A09776C8FFC64A7027ACB56CFEEFE14B" box="[151,304,1791,1818]" count="1" pageId="51" pageNumber="52" type="generic">one specimen</specimenCount>
on the right is 
<taxonomicName id="7191C6C2FFC64A7026ECB493FD2EE14B" authority="Franz" authorityName="Franz. Despite" authorityYear="1975" box="[471,753,1791,1817]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="mirus">
<emphasis id="84E56153FFC64A7026ECB493FD76E14B" box="[471,681,1791,1817]" italics="true" pageId="51" pageNumber="52">Scydmaenus mirus</emphasis>
Franz
</taxonomicName>
) (
<collectionCode id="D0802584FFC64A70243CB493FC81E14B" box="[775,862,1791,1817]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="51" pageNumber="52" type="Museum">SAMA</collectionCode>
); 3 exx. mounted on one card and originally pinned as a part of SAMA 25-43079 (see above)—specimen on the left is 
<taxonomicName id="7191C6C2FFC64A7024BBB54FFBEDE16C" box="[896,1074,1827,1854]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="84E56153FFC64A7024BBB54FFBD5E16F" box="[896,1034,1827,1853]" italics="true" pageId="51" pageNumber="52">Scydmaenus</emphasis>
sp.
</taxonomicName>
[not 
<taxonomicName id="7191C6C2FFC64A702357B54FFB21E16F" box="[1132,1278,1827,1853]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="clarkianus">
<emphasis id="84E56153FFC64A702357B54FFB21E16F" box="[1132,1278,1827,1853]" italics="true" pageId="51" pageNumber="52">S. clarkianus</emphasis>
</taxonomicName>
], specimen in middle and on the right are 
<taxonomicName id="7191C6C2FFC64A7026EAB52BFD2FE133" authority="Franz" authorityName="Franz. Despite" authorityYear="1975" box="[465,752,1863,1889]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="mirus">
<emphasis id="84E56153FFC64A7026EAB52BFD7CE133" box="[465,675,1863,1889]" italics="true" pageId="51" pageNumber="52">Scydmaenus mirus</emphasis>
Franz
</taxonomicName>
); 3 exx. mounted on one card, SAMA 25-43085 (from Swan River), all 
<taxonomicName id="7191C6C2FFC64A702636B507FD96E1D7" authority="(SAMA)" baseAuthorityName="SAMA" box="[269,585,1899,1925]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="mirus">
<emphasis id="84E56153FFC64A702636B507FE02E1D7" box="[269,477,1899,1925]" italics="true" pageId="51" pageNumber="52">Scydmaenus mirus</emphasis>
(SAMA)
</taxonomicName>
; 
<specimenCount id="A09776C8FFC64A702568B507FD57E1D4" box="[595,648,1899,1926]" count="1" pageId="51" pageNumber="52" type="generic">1 ex.</specimenCount>
of 
<taxonomicName id="7191C6C2FFC64A702591B507FCA5E1D7" box="[682,890,1899,1925]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="mirus">
<emphasis id="84E56153FFC64A702591B507FCA5E1D7" box="[682,890,1899,1925]" italics="true" pageId="51" pageNumber="52">Scydmaenus mirus</emphasis>
</taxonomicName>
, with unidentified ants, from Swan River, SAMA 25-43077 (
<collectionCode id="D0802584FFC64A70262BB5E3FEB8E1FB" box="[272,359,1935,1961]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="51" pageNumber="52" type="Museum">SAMA</collectionCode>
); 
<specimenCount id="A09776C8FFC64A702642B5E3FE70E1F8" box="[377,431,1935,1962]" count="1" pageId="51" pageNumber="52" type="generic">1 ex.</specimenCount>
of 
<taxonomicName id="7191C6C2FFC64A7026EAB5E3FD7EE1FB" box="[465,673,1935,1961]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="mirus">
<emphasis id="84E56153FFC64A7026EAB5E3FD7EE1FB" box="[465,673,1935,1961]" italics="true" pageId="51" pageNumber="52">Scydmaenus mirus</emphasis>
</taxonomicName>
, with unidentified ants, from Perth, SAMA 25-43086 (
<collectionCode id="D0802584FFC64A7023C5B5E3FA8AE1FB" box="[1278,1365,1935,1961]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="51" pageNumber="52" type="Museum">SAMA</collectionCode>
); 
<specimenCount id="A09776C8FFC64A70225CB5E3FA42E1F8" box="[1383,1437,1935,1962]" count="1" pageId="51" pageNumber="52" type="generic">1 ex.</specimenCount>
of 
<taxonomicName id="7191C6C2FFC64A70278FB5DFFE5AE19F" box="[180,389,1971,1997]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="mirus">
<emphasis id="84E56153FFC64A70278FB5DFFE5AE19F" box="[180,389,1971,1997]" italics="true" pageId="51" pageNumber="52">Scydmaenus mirus</emphasis>
</taxonomicName>
, “with 
<taxonomicName id="7191C6C2FFC64A7026EDB5D8FD5DE19F" authorityName="auctt, non Patella lutea Linnaeus" authorityYear="1758" box="[470,642,1971,1997]" class="Insecta" family="Formicidae" genus="Euponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="84E56153FFC64A7026EDB5D8FD5DE19F" box="[470,642,1971,1997]" italics="true" pageId="51" pageNumber="52">Euponera lutea</emphasis>
</taxonomicName>
”, from Swan River, SAMA 25- 43071; 1 ♂ designated here as 
<typeStatus id="692A03E3FFC64A702201B5DFFA43E19F" box="[1338,1436,1971,1997]" pageId="51" pageNumber="52" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="7191C6C2FFC64A70278DB5BBFDF8E1A3" authorityName="Jałoszyński" authorityYear="2023" box="[182,551,2007,2033]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="brachyponerophilus">
<emphasis id="84E56153FFC64A70278DB5BBFDF8E1A3" box="[182,551,2007,2033]" italics="true" pageId="51" pageNumber="52">Scydmaenus brachyponerophilus</emphasis>
</taxonomicName>
, from Swan River, SAMA 25-43073 (
<collectionCode id="D0802584FFC64A7024EFB5BBFBF4E1A3" box="[980,1067,2007,2033]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="51" pageNumber="52" type="Museum">SAMA</collectionCode>
).
</paragraph>
<caption id="E2EEEDC9FFC14A7727ACB650FD93E22A" ID-DOI="http://doi.org/10.5281/zenodo.10144703" ID-Zenodo-Dep="10144703" httpUri="https://zenodo.org/record/10144703/files/figure.png" pageId="52" pageNumber="53" startId="52.[151,264,1084,1108]" targetBox="[157,1430,187,1054]" targetPageId="52" targetType="figure">
<paragraph id="B62EBD41FFC14A7727ACB650FD93E22A" blockId="52.[151,1436,1084,1144]" pageId="52" pageNumber="53">
<emphasis id="84E56153FFC14A7727ACB650FEB1E206" bold="true" box="[151,366,1084,1108]" pageId="52" pageNumber="53">FIGURES 130–134.</emphasis>
<taxonomicName id="7191C6C2FFC14A77264FB650FCAFE201" authority="Franz" authorityName="Franz" box="[372,880,1084,1108]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="clarkianus" subGenus="Scottiscydmaenus">
<emphasis id="84E56153FFC14A77264FB650FE2EE206" box="[372,497,1084,1108]" italics="true" pageId="52" pageNumber="53">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFC14A77253BB650FD6BE206" box="[512,692,1084,1108]" italics="true" pageId="52" pageNumber="53">Scottiscydmaenus</emphasis>
) 
<emphasis id="84E56153FFC14A7725F8B650FCF1E206" box="[707,814,1084,1108]" italics="true" pageId="52" pageNumber="53">clarkianus</emphasis>
Franz
</taxonomicName>
, holotype male. Dorsal habitus (
<emphasis id="84E56153FFC14A772382B650FB00E206" bold="true" box="[1209,1247,1084,1108]" pageId="52" pageNumber="53">130</emphasis>
); aedeagus in dorsal (
<emphasis id="84E56153FFC14A7727F9B60CFEC4E22A" bold="true" box="[194,283,1120,1144]" pageId="52" pageNumber="53">131, 133</emphasis>
) and lateral (
<emphasis id="84E56153FFC14A772699B60CFE24E22A" bold="true" box="[418,507,1120,1144]" pageId="52" pageNumber="53">132, 134</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="B62EBD41FFC14A7727FCB6CAFEADE37E" blockId="52.[151,1437,1190,2009]" pageId="52" pageNumber="53">
<emphasis id="84E56153FFC14A7727FCB6CAFDB7E292" bold="true" box="[199,616,1190,1216]" pageId="52" pageNumber="53">Revised diagnosis (based on males)</emphasis>
. Head clearly elongate (
<figureCitation id="2EAAA1C4FFC14A772442B6CAFC03E293" box="[889,988,1190,1217]" captionStart="FIGURES 155–162" captionStartId="60.[151,264,1338,1362]" captionTargetBox="[151,1435,181,1302]" captionTargetId="figure-21@60.[151,1435,181,1314]" captionTargetPageId="60" captionText="FIGURES 155–162. Scottiscydmaenus, heads in dorsal view (155–158) and profemora of males (159–162). Scydmaenus clar- kianus Franz (155, 159); S. optatus Sharp (156, 160); S. swanensis Franz (157, 161); S. brachyponerophilus sp. n. (158, 162)." figureDoi="http://doi.org/10.5281/zenodo.10144711" httpUri="https://zenodo.org/record/10144711/files/figure.png" pageId="52" pageNumber="53">Fig. 155</figureCitation>
); profemoral tooth at most right-angled, broader than long or as long as broad (
<figureCitation id="2EAAA1C4FFC14A77257AB6A6FD7CE2B7" box="[577,675,1226,1253]" captionStart="FIGURES 155–162" captionStartId="60.[151,264,1338,1362]" captionTargetBox="[151,1435,181,1302]" captionTargetId="figure-21@60.[151,1435,181,1314]" captionTargetPageId="60" captionText="FIGURES 155–162. Scottiscydmaenus, heads in dorsal view (155–158) and profemora of males (159–162). Scydmaenus clar- kianus Franz (155, 159); S. optatus Sharp (156, 160); S. swanensis Franz (157, 161); S. brachyponerophilus sp. n. (158, 162)." figureDoi="http://doi.org/10.5281/zenodo.10144711" httpUri="https://zenodo.org/record/10144711/files/figure.png" pageId="52" pageNumber="53">Fig. 159</figureCitation>
); length of temple in dorsal view about 4 × as long as eye (
<figureCitation id="2EAAA1C4FFC14A772215B6A6FA50E2B7" box="[1326,1423,1226,1253]" captionStart="FIGURES 155–162" captionStartId="60.[151,264,1338,1362]" captionTargetBox="[151,1435,181,1302]" captionTargetId="figure-21@60.[151,1435,181,1314]" captionTargetPageId="60" captionText="FIGURES 155–162. Scottiscydmaenus, heads in dorsal view (155–158) and profemora of males (159–162). Scydmaenus clar- kianus Franz (155, 159); S. optatus Sharp (156, 160); S. swanensis Franz (157, 161); S. brachyponerophilus sp. n. (158, 162)." figureDoi="http://doi.org/10.5281/zenodo.10144711" httpUri="https://zenodo.org/record/10144711/files/figure.png" pageId="52" pageNumber="53">Fig. 155</figureCitation>
); aedeagus in dorsal view (
<figureCitation id="2EAAA1C4FFC14A77268EB682FDCAE35A" box="[437,533,1262,1288]" captionStart="FIGURES 130–134" captionStartId="52.[151,264,1084,1108]" captionTargetBox="[157,1430,187,1054]" captionTargetId="figure-21@52.[151,1436,181,1060]" captionTargetPageId="52" captionText="FIGURES 130–134. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Dorsal habitus (130); aedeagus in dorsal (131, 133) and lateral (132, 134) views." figureDoi="http://doi.org/10.5281/zenodo.10144703" httpUri="https://zenodo.org/record/10144703/files/figure.png" pageId="52" pageNumber="53">Fig. 131</figureCitation>
) almost gradually narrowing from sub-basal region to truncate and weakly concave apical margin.
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFC14A7627FCB75AFEEFE3EA" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="description">
<paragraph id="B62EBD41FFC14A7727FCB75AFE29E3CA" blockId="52.[151,1437,1190,2009]" pageId="52" pageNumber="53">
<emphasis id="84E56153FFC14A7727FCB75AFEAAE302" bold="true" box="[199,373,1334,1360]" pageId="52" pageNumber="53">Redescription.</emphasis>
Body in male (
<figureCitation id="2EAAA1C4FFC14A77251EB75AFD5AE302" box="[549,645,1334,1360]" captionStart="FIGURES 130–134" captionStartId="52.[151,264,1084,1108]" captionTargetBox="[157,1430,187,1054]" captionTargetId="figure-21@52.[151,1436,181,1060]" captionTargetPageId="52" captionText="FIGURES 130–134. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Dorsal habitus (130); aedeagus in dorsal (131, 133) and lateral (132, 134) views." figureDoi="http://doi.org/10.5281/zenodo.10144703" httpUri="https://zenodo.org/record/10144703/files/figure.png" pageId="52" pageNumber="53">Fig. 130</figureCitation>
) strongly convex, elongate and slender, BL 
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; pigmentation uniformly light to dark brown (including appendages); cuticle moderately glossy, covered with vestiture of setae slightly lighter than body.
</paragraph>
<paragraph id="B62EBD41FFC14A7727FCB7CEFCE3E18A" blockId="52.[151,1437,1190,2009]" pageId="52" pageNumber="53">
Head (
<figureCitation id="2EAAA1C4FFC14A77262BB7CEFE72E3EF" box="[272,429,1442,1469]" captionStart="FIGURES 125–126" captionStartId="50.[151,264,1744,1768]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-21@50.[151,1436,181,1721]" captionTargetPageId="50" captionText="FIGURES 125–126. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Head, pronotum and elytral base in dorsal view (125); head and prothorax in ventral view (126). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10144699" httpUri="https://zenodo.org/record/10144699/files/figure.png" pageId="52" pageNumber="53">Figs 125–126</figureCitation>
, 
<figureCitation id="2EAAA1C4FFC14A77268CB7CEFE3DE3EF" box="[439,482,1442,1469]" captionStart="FIGURES 155–162" captionStartId="60.[151,264,1338,1362]" captionTargetBox="[151,1435,181,1302]" captionTargetId="figure-21@60.[151,1435,181,1314]" captionTargetPageId="60" captionText="FIGURES 155–162. Scottiscydmaenus, heads in dorsal view (155–158) and profemora of males (159–162). Scydmaenus clar- kianus Franz (155, 159); S. optatus Sharp (156, 160); S. swanensis Franz (157, 161); S. brachyponerophilus sp. n. (158, 162)." figureDoi="http://doi.org/10.5281/zenodo.10144711" httpUri="https://zenodo.org/record/10144711/files/figure.png" pageId="52" pageNumber="53">155</figureCitation>
) in dorsal view elongate and rounded, broadest at eyes, HL 
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, HW 
<quantity id="716910A4FFC14A772266B7CEFF20E3B3" metricMagnitude="-4" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="52" pageNumber="53" unit="mm" value="0.325" valueMax="0.34" valueMin="0.31">0.31– 0.34 mm</quantity>
; vertex and frons confluent and weakly convex, posterior margin of vertex arcuate and weakly anteriorly concave; tempora about 4 × as long as length of eye in dorsal view; supraantennal tubercles indistinct; frons over antennal fossae broadly subtriangular and with blunt, broadly subtriangular anterior margin. Eyes conspicuously small, almost circular, not emarginate posteriorly. Punctures on frons and vertex fine, inconspicuous; setae (including those on tempora) short, sparse, nearly recumbent. Genae (
<figureCitation id="2EAAA1C4FFC14A772454B43AFC0EE022" box="[879,977,1622,1648]" captionStart="FIGURES 125–126" captionStartId="50.[151,264,1744,1768]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-21@50.[151,1436,181,1721]" captionTargetPageId="50" captionText="FIGURES 125–126. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Head, pronotum and elytral base in dorsal view (125); head and prothorax in ventral view (126). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10144699" httpUri="https://zenodo.org/record/10144699/files/figure.png" pageId="52" pageNumber="53">Fig. 126</figureCitation>
) as sparsely setose as frons and vertex. Anterior (exposed) region of head capsule demarcated from neck region by short abrupt impression around occipital constriction, anterior margin of gular plate on neck region with indistinct, narrow anteriorly-directed projection. Submentum (
<figureCitation id="2EAAA1C4FFC14A77260AB4AEFE4CE08E" box="[305,403,1730,1756]" captionStart="FIGURES 125–126" captionStartId="50.[151,264,1744,1768]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-21@50.[151,1436,181,1721]" captionTargetPageId="50" captionText="FIGURES 125–126. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Head, pronotum and elytral base in dorsal view (125); head and prothorax in ventral view (126). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10144699" httpUri="https://zenodo.org/record/10144699/files/figure.png" pageId="52" pageNumber="53">Fig. 126</figureCitation>
) without submental lobes; hypostomal ridges (
<figureCitation id="2EAAA1C4FFC14A772488B4AEFBCAE08E" box="[947,1045,1730,1756]" captionStart="FIGURES 125–126" captionStartId="50.[151,264,1744,1768]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-21@50.[151,1436,181,1721]" captionTargetPageId="50" captionText="FIGURES 125–126. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Head, pronotum and elytral base in dorsal view (125); head and prothorax in ventral view (126). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10144699" httpUri="https://zenodo.org/record/10144699/files/figure.png" pageId="52" pageNumber="53">Fig. 126</figureCitation>
) extend mesally and anteriorly to connect at middle behind anterior margin of submentum where they form distinct transverse carina. Antennae (
<figureCitation id="2EAAA1C4FFC14A772251B48AFF1CE176" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="52" pageNumber="53">Figs 127</figureCitation>
, 
<figureCitation id="2EAAA1C4FFC14A7727EBB566FF24E176" box="[208,251,1802,1828]" captionStart="FIGURES 130–134" captionStartId="52.[151,264,1084,1108]" captionTargetBox="[157,1430,187,1054]" captionTargetId="figure-21@52.[151,1436,181,1060]" captionTargetPageId="52" captionText="FIGURES 130–134. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Dorsal habitus (130); aedeagus in dorsal (131, 133) and lateral (132, 134) views." figureDoi="http://doi.org/10.5281/zenodo.10144703" httpUri="https://zenodo.org/record/10144703/files/figure.png" pageId="52" pageNumber="53">130</figureCitation>
) long and slender, AnL 
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; three terminal antennomeres forming club, but antennomere 9 is so slender that club appears as dimerous; scape 3 × as long as broad, distinctly broadening distally; pedicel slightly more than twice as long as broad; antennomeres 3 and 4 each twice as long as broad, 5 2.5 × as long as broad, 6–8 each about twice as long as broad, 9 2.5 × as long as broad, gradually broadening distally, 10 nearly twice as long as broad and distinctly broadening distally but again narrowing in short distal region, 11 much shorter than 9 and 10 combined, about 2.2 × as long as broad, nearly symmetrical.
</paragraph>
<paragraph id="B62EBD41FFC04A7627FCB2FBFC19E7A0" blockId="53.[151,1437,150,1678]" pageId="53" pageNumber="54">
Pronotum in dorsal view (
<figureCitation id="2EAAA1C4FFC04A7626DFB2FBFD9DE6E3" box="[484,578,150,177]" captionStart="FIGURES 125–126" captionStartId="50.[151,264,1744,1768]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-21@50.[151,1436,181,1721]" captionTargetPageId="50" captionText="FIGURES 125–126. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Head, pronotum and elytral base in dorsal view (125); head and prothorax in ventral view (126). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10144699" httpUri="https://zenodo.org/record/10144699/files/figure.png" pageId="53" pageNumber="54">Fig. 125</figureCitation>
) elongate, broadest near anterior third, PL 
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, PW 
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; anterior margin nearly straight; anterior corners weakly marked, obtuse-angled and blunt; sides strongly rounded in anterior half; posterior corners barely marked; posterior margin weakly arcuate; posterior marginal carina obliterated. Pronotal disc covered with fine and shallow, unremarkable sparse punctures; setae similar to those on head, moderately dense and long, suberect. Ventrally (
<figureCitation id="2EAAA1C4FFC04A76258DB349FCCAE76D" box="[694,789,293,319]" captionStart="FIGURES 125–126" captionStartId="50.[151,264,1744,1768]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-21@50.[151,1436,181,1721]" captionTargetPageId="50" captionText="FIGURES 125–126. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Head, pronotum and elytral base in dorsal view (125); head and prothorax in ventral view (126). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10144699" httpUri="https://zenodo.org/record/10144699/files/figure.png" pageId="53" pageNumber="54">Fig. 126</figureCitation>
) prothorax with nearly asetose and impunctate hypomera and basisternal region only slightly longer than procoxal rests, sparsely covered with moderately long recumbent setae, with short and relatively indistinct anterior ‘collar’, and with distinct vestiges of notosternal sutures visible as notches on sides of anterior prothoracic margin; hypomeral ridges distinct and complete, demarcating narrow inner (adcoxal) region of each hypomeron, anteriorly running along procoxal rests and connecting at middle to form biarcuate anteprocoxal carina demarcating basisternal region posteriorly.
</paragraph>
<paragraph id="B62EBD41FFC04A7627FCB390FE94E4D3" blockId="53.[151,1437,150,1678]" pageId="53" pageNumber="54">
Elytra (
<figureCitation id="2EAAA1C4FFC04A762627B390FE5CE445" box="[284,387,508,535]" captionStart="FIGURES 125–126" captionStartId="50.[151,264,1744,1768]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-21@50.[151,1436,181,1721]" captionTargetPageId="50" captionText="FIGURES 125–126. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Head, pronotum and elytral base in dorsal view (125); head and prothorax in ventral view (126). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10144699" httpUri="https://zenodo.org/record/10144699/files/figure.png" pageId="53" pageNumber="54">Figs 125</figureCitation>
, 
<figureCitation id="2EAAA1C4FFC04A7626AAB390FE63E444" box="[401,444,508,534]" captionStart="FIGURES 130–134" captionStartId="52.[151,264,1084,1108]" captionTargetBox="[157,1430,187,1054]" captionTargetId="figure-21@52.[151,1436,181,1060]" captionTargetPageId="52" captionText="FIGURES 130–134. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Dorsal habitus (130); aedeagus in dorsal (131, 133) and lateral (132, 134) views." figureDoi="http://doi.org/10.5281/zenodo.10144703" httpUri="https://zenodo.org/record/10144703/files/figure.png" pageId="53" pageNumber="54">130</figureCitation>
) slightly rhomboidal, broadest at or slightly behind of middle, EL 
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, EW 
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, EI 1.43–1.58. Humeral calli, basal impressions and basal foveae lacking; apices separately rounded. Elytral punctures fine and inconspicuous; setae similar to those on pronotum, moderately dense. Hind wings completely reduced.
</paragraph>
<paragraph id="B62EBD41FFC04A7627FCB0E7FC03E5F2" blockId="53.[151,1437,150,1678]" pageId="53" pageNumber="54">
Mesoventrite (
<figureCitation id="2EAAA1C4FFC04A762655B0E7FE0EE4F7" box="[366,465,651,677]" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="53" pageNumber="54">Fig. 128</figureCitation>
) with carinate subrectangular mesoventral intermesocoxal process posteriorly fused with elongate and posteriorly broadening anterior metaventral process, fusion site indistinctly marked on surface by transverse ridge. Metanepisterna (
<figureCitation id="2EAAA1C4FFC04A76252DB0BEFDA9E4BE" box="[534,630,722,748]" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="53" pageNumber="54">Fig. 128</figureCitation>
) completely fused with metaventrite, but fusion site marked on surface as line along which sides of thorax are bent dorsally. Metaventral intermetacoxal process (
<figureCitation id="2EAAA1C4FFC04A762347B09AFB04E542" box="[1148,1243,758,784]" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="53" pageNumber="54">Fig. 128</figureCitation>
) broad and short, with distinctly concave posterior margin, its lateral portions form short subtriangular processes weakly projecting posteriorly; distance between metacoxae subequal to 1/3 width of metaventrite at its posterior margin (excluding metanepisterna) and slightly wider than one metacoxa. Metaventrite (
<figureCitation id="2EAAA1C4FFC04A7624A4B10DFC20E529" box="[927,1023,865,891]" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="53" pageNumber="54">Fig. 128</figureCitation>
) weakly convex, slightly flattened at middle, unmodified and covered with sparse, short, slightly suberect setae.
</paragraph>
<paragraph id="B62EBD41FFC04A7627FCB1C5FE8BE2EF" blockId="53.[151,1437,150,1678]" pageId="53" pageNumber="54">
Legs (
<figureCitation id="2EAAA1C4FFC04A762635B1C5FEABE591" box="[270,372,937,963]" captionStart="FIGURES 125–126" captionStartId="50.[151,264,1744,1768]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-21@50.[151,1436,181,1721]" captionTargetPageId="50" captionText="FIGURES 125–126. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Head, pronotum and elytral base in dorsal view (125); head and prothorax in ventral view (126). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10144699" httpUri="https://zenodo.org/record/10144699/files/figure.png" pageId="53" pageNumber="54">Figs 126</figureCitation>
, 
<figureCitation id="2EAAA1C4FFC04A7626B9B1C5FE38E591" box="[386,487,937,963]" captionStart-0="FIGURES 127–129" captionStart-1="FIGURES 130–134" captionStartId-0="51.[151,264,1361,1385]" captionStartId-1="52.[151,264,1084,1108]" captionTargetBox-0="[151,1436,181,1337]" captionTargetBox-1="[157,1430,187,1054]" captionTargetId-0="figure-15@51.[151,1436,181,1337]" captionTargetId-1="figure-21@52.[151,1436,181,1060]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionText-0="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." captionText-1="FIGURES 130–134. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Dorsal habitus (130); aedeagus in dorsal (131, 133) and lateral (132, 134) views." figureDoi-0="http://doi.org/10.5281/zenodo.10144701" figureDoi-1="http://doi.org/10.5281/zenodo.10144703" httpUri-0="https://zenodo.org/record/10144701/files/figure.png" httpUri-1="https://zenodo.org/record/10144703/files/figure.png" pageId="53" pageNumber="54">128–130</figureCitation>
, 
<figureCitation id="2EAAA1C4FFC04A7626CFB1C5FDC0E596" box="[500,543,937,964]" captionStart="FIGURES 155–162" captionStartId="60.[151,264,1338,1362]" captionTargetBox="[151,1435,181,1302]" captionTargetId="figure-21@60.[151,1435,181,1314]" captionTargetPageId="60" captionText="FIGURES 155–162. Scottiscydmaenus, heads in dorsal view (155–158) and profemora of males (159–162). Scydmaenus clar- kianus Franz (155, 159); S. optatus Sharp (156, 160); S. swanensis Franz (157, 161); S. brachyponerophilus sp. n. (158, 162)." figureDoi="http://doi.org/10.5281/zenodo.10144711" httpUri="https://zenodo.org/record/10144711/files/figure.png" pageId="53" pageNumber="54">159</figureCitation>
) long and slender; profemur with short, broad and relatively blunt distal tooth (
<figureCitation id="2EAAA1C4FFC04A7627A4B1A0FEDDE5B5" box="[159,258,972,999]" captionStart="FIGURES 155–162" captionStartId="60.[151,264,1338,1362]" captionTargetBox="[151,1435,181,1302]" captionTargetId="figure-21@60.[151,1435,181,1314]" captionTargetPageId="60" captionText="FIGURES 155–162. Scottiscydmaenus, heads in dorsal view (155–158) and profemora of males (159–162). Scydmaenus clar- kianus Franz (155, 159); S. optatus Sharp (156, 160); S. swanensis Franz (157, 161); S. brachyponerophilus sp. n. (158, 162)." figureDoi="http://doi.org/10.5281/zenodo.10144711" httpUri="https://zenodo.org/record/10144711/files/figure.png" pageId="53" pageNumber="54">Fig. 159</figureCitation>
) slightly variable within studied specimens, but never elongate; protibia (
<figureCitation id="2EAAA1C4FFC04A76236DB1A0FB67E5B5" box="[1110,1208,972,999]" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="53" pageNumber="54">Fig. 129</figureCitation>
) slightly broadened distally; protarsus with barely discernibly broadened proximal half and tarsomeres 1–3 covered ventrally with dense setae, of which only a few have spatulate adhesive tips. Protarsomere 1 nearly twice as long as broad, 2–4 each indistinctly elongate, 5 about 3 × as long as broad; mesotarsi longer than protarsi, mesotarsomere 1 about 3 × as long as broad, tarsomeres 2–4 each weakly elongate, tarsomere 5 about 2.5 × as long as broad; metatarsi slightly longer than mesotarsi, metatarsomere 1 3 × as long as broad, tarsomeres 2–4 each weakly elongate, tarsomere 5 about 3 × as long as broad.
</paragraph>
<paragraph id="B62EBD41FFC04A7627FCB6ABFE2FE31E" blockId="53.[151,1437,150,1678]" pageId="53" pageNumber="54">
Aedeagus (
<figureCitation id="2EAAA1C4FFC04A76267EB6ABFE3EE2B3" box="[325,481,1223,1249]" captionStart="FIGURES 130–134" captionStartId="52.[151,264,1084,1108]" captionTargetBox="[157,1430,187,1054]" captionTargetId="figure-21@52.[151,1436,181,1060]" captionTargetPageId="52" captionText="FIGURES 130–134. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Dorsal habitus (130); aedeagus in dorsal (131, 133) and lateral (132, 134) views." figureDoi="http://doi.org/10.5281/zenodo.10144703" httpUri="https://zenodo.org/record/10144703/files/figure.png" pageId="53" pageNumber="54">Figs 131–134</figureCitation>
) elongate and slender, AeL 
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, in dorsal view median lobe broadest near base, narrowing distally toward truncate apex, apical margin slightly concave, with lateral groups of minute setae; flagellum broadened in proximal region to form three consecutive symmetrical chambers; ostium situated in distal half of median lobe, far from its apex.
</paragraph>
<paragraph id="B62EBD41FFC04A7627FCB73AFEEFE3EA" blockId="53.[151,1437,150,1678]" pageId="53" pageNumber="54">
<emphasis id="84E56153FFC04A7627FCB73AFEFAE322" bold="true" box="[199,293,1366,1392]" pageId="53" pageNumber="54">Female.</emphasis>
Externally differs from male in profemora lacking teeth. BL 
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; HL 
<quantity id="716910A4FFC04A762387B739FABDE322" box="[1212,1378,1365,1392]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="53" pageNumber="54" unit="mm" value="0.365" valueMax="0.38" valueMin="0.35">0.35–0.38 mm</quantity>
, HW 
<quantity id="716910A4FFC04A7627ACB715FEE3E3C6" box="[151,316,1401,1428]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="53" pageNumber="54" unit="mm" value="0.32" valueMax="0.33" valueMin="0.31">0.31–0.33 mm</quantity>
, AnL 
<quantity id="716910A4FFC04A762644B715FDFAE3C6" box="[383,549,1401,1428]" metricMagnitude="-4" metricUnit="m" metricValue="9.9" metricValueMax="10.3" metricValueMin="9.5" pageId="53" pageNumber="54" unit="mm" value="0.99" valueMax="1.03" valueMin="0.95">0.95–1.03 mm</quantity>
; PL 
<quantity id="716910A4FFC04A76256CB715FD23E3C6" box="[599,764,1401,1428]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="53" pageNumber="54" unit="mm" value="0.525" valueMax="0.55" valueMin="0.5">0.50–0.55 mm</quantity>
, PW 
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; EL 
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, EW 
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, EI 1.37–1.45.
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFC04A7627FCB7ACFB75E389" box="[199,1194,1472,1499]" pageId="53" pageNumber="54" type="distribution">
<paragraph id="B62EBD41FFC04A7627FCB7ACFB75E389" blockId="53.[151,1437,150,1678]" box="[199,1194,1472,1499]" pageId="53" pageNumber="54">
<emphasis id="84E56153FFC04A7627FCB7ACFEBEE388" bold="true" box="[199,353,1472,1498]" pageId="53" pageNumber="54">Distribution.</emphasis>
Known only from SW 
<collectingCountry id="CE86FDD1FFC04A76255DB7ADFD0EE389" box="[614,721,1473,1499]" name="Australia" pageId="53" pageNumber="54">Australia</collectingCountry>
: SW 
<collectingRegion id="745573A3FFC04A762430B7ADFC0CE389" box="[779,979,1473,1499]" country="Australia" name="Western Australia" pageId="53" pageNumber="54">Western Australia</collectingRegion>
(vicinity of Perth).
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFC04A7627FCB789FB40E0DC" pageId="53" pageNumber="54" type="discussion">
<paragraph id="B62EBD41FFC04A7627FCB789FB40E0DC" blockId="53.[151,1437,150,1678]" pageId="53" pageNumber="54">
<emphasis id="84E56153FFC04A7627FCB789FEE4E3AD" bold="true" box="[199,315,1509,1535]" pageId="53" pageNumber="54">Remarks.</emphasis>
<taxonomicName id="7191C6C2FFC04A762679B788FD96E3AC" box="[322,585,1508,1534]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="clarkianus">
<emphasis id="84E56153FFC04A762679B788FD96E3AC" box="[322,585,1508,1534]" italics="true" pageId="53" pageNumber="54">Scydmaenus clarkianus</emphasis>
</taxonomicName>
is distinct in the slender body, elytra strongly narrowing anteriorly, conspicuously small eyes, and a broad and short profemoral tooth. The long 
<typeStatus id="692A03E3FFC04A7624A6B465FC12E071" box="[925,973,1545,1571]" pageId="53" pageNumber="54">type</typeStatus>
series was found to be composed of four species; in addition to 
<taxonomicName id="7191C6C2FFC04A7626A2B440FDF2E014" box="[409,557,1580,1606]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="clarkianus">
<emphasis id="84E56153FFC04A7626A2B440FDF2E014" box="[409,557,1580,1606]" italics="true" pageId="53" pageNumber="54">S. clarkianus</emphasis>
</taxonomicName>
it also contains specimens of 
<taxonomicName id="7191C6C2FFC04A7624BDB440FA43E015" authority="Franz, 1975" authorityName="Franz. Despite" authorityYear="1975" box="[902,1436,1580,1607]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="mirus" subGenus="Heteromicrus">
<emphasis id="84E56153FFC04A7624BDB440FBCFE014" box="[902,1040,1580,1606]" italics="true" pageId="53" pageNumber="54">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFC04A762319B441FB64E014" box="[1058,1211,1581,1606]" italics="true" pageId="53" pageNumber="54">Heteromicrus</emphasis>
) 
<emphasis id="84E56153FFC04A7623F6B441FAD2E014" box="[1229,1293,1581,1606]" italics="true" pageId="53" pageNumber="54">mirus</emphasis>
<bibRefCitation id="D200C0B0FFC04A76222EB441FA43E015" author="Franz, H." box="[1301,1436,1580,1607]" pageId="53" pageNumber="54" pagination="1 - 312" refId="ref50046" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">Franz, 1975</bibRefCitation>
</taxonomicName>
(in fact three times as many as the 
<typeStatus id="692A03E3FFC04A76251BB43CFD8FE038" box="[544,592,1616,1642]" pageId="53" pageNumber="54">type</typeStatus>
series of 
<taxonomicName id="7191C6C2FFC04A762585B43CFCC4E03B" authorityName="Franz. Despite" authorityYear="1975" box="[702,795,1616,1642]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="mirus">
<emphasis id="84E56153FFC04A762585B43CFCC4E03B" box="[702,795,1616,1642]" italics="true" pageId="53" pageNumber="54">S. mirus</emphasis>
</taxonomicName>
includes!), a new species of 
<taxonomicName id="7191C6C2FFC04A762359B43CFAF6E038" box="[1122,1321,1616,1642]" pageId="53" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">
<emphasis id="84E56153FFC04A762359B43CFAF6E038" box="[1122,1321,1616,1642]" italics="true" pageId="53" pageNumber="54">Scottiscydmaenus</emphasis>
</taxonomicName>
described below as 
<taxonomicName id="7191C6C2FFC04A76263AB418FE22E0DC" authorityName="Jałoszyński" authorityYear="2023" box="[257,509,1652,1678]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="brachyponerophilus">
<emphasis id="84E56153FFC04A76263AB418FE22E0DC" box="[257,509,1652,1678]" italics="true" pageId="53" pageNumber="54">S. brachyponerophilus</emphasis>
</taxonomicName>
, and specimens of an undetermined species of 
<taxonomicName id="7191C6C2FFC04A762336B418FB48E0DC" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[1037,1175,1652,1678]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="84E56153FFC04A762336B418FB48E0DC" box="[1037,1175,1652,1678]" italics="true" pageId="53" pageNumber="54">Scydmaenus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>