treatments-xml/data/21/C1/FB/21C1FBCEA8B652A6A7D42869B54D2625.xml
2024-06-21 12:31:17 +02:00

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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.12.e115556" ID-Pensoft-Pub="1314-2828-12-e115556" ID-Pensoft-UUID="58CD7053908A596FBFE9C5B82A839FCC" ID-ZooBank="C0916935E263450FAB28ABB7E82FB4F9" ModsDocID="1314-2828-12-e115556" checkinTime="1708088559296" checkinUser="pensoft" docAuthor="Lee, Jie Ying, Nasir, Nurul Syafiqah, Marzuki, Mohammad Effendi, Vermeulen, Jaap J. &amp; Khalik, Mohd Zacaery" docDate="2024" docId="21C1FBCEA8B652A6A7D42869B54D2625" docLanguage="en" docName="BiodivDatJour 12: e115556" docOrigin="Biodiversity Data Journal 12" docPubDate="2024-02-16" docSource="http://dx.doi.org/10.3897/BDJ.12.e115556" docTitle="Helicarion dyakanum" docType="treatment" docVersion="1" id="58CD7053908A596FBFE9C5B82A839FCC" lastPageNumber="115556" masterDocId="58CD7053908A596FBFE9C5B82A839FCC" masterDocTitle="The land snail fauna of Batu Kudik, isolated limestone outcrop near Simunjan, Sarawak, Malaysian Borneo (Mollusca, Gastropoda)" masterLastPageNumber="115556" masterPageNumber="115556" pageNumber="115556" updateTime="1708088559296" updateUser="pensoft">
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<mods:title id="768CF0CDF88929CBF9541B9B52C0E151">The land snail fauna of Batu Kudik, isolated limestone outcrop near Simunjan, Sarawak, Malaysian Borneo (Mollusca, Gastropoda)</mods:title>
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<mods:name id="4135437ACABBF6BF84D42F6553530348" type="personal">
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<mods:namePart id="7388AA1FFF2802F8A8F24A2B077990D9">Lee, Jie Ying</mods:namePart>
<mods:nameIdentifier id="43158B60168BCE511D77AC83C8C8B5B2" type="ORCID">https://orcid.org/0009-0002-6223-3814</mods:nameIdentifier>
<mods:affiliation id="C1784610D10AE67E0AD1172D4E196B8B">Faculty of Resource Science and Technology, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia &amp; Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia</mods:affiliation>
<mods:nameIdentifier id="AA05832C3B6DC93C2BCF55693202EDDF" type="email">winglee99@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="B5CD5F50363908AAF8B309F266C79F73">Nasir, Nurul Syafiqah</mods:namePart>
<mods:affiliation id="624B72FD03BB526AF96F26D2911C61F5">Faculty of Resource Science and Technology, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia &amp; Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia</mods:affiliation>
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<mods:namePart id="6CAC81E881B6C617AF09952DE7BE90F3">Marzuki, Mohammad Effendi</mods:namePart>
<mods:affiliation id="D23B2430AD16F9BCD0951396157DF6C6">Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia</mods:affiliation>
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<mods:namePart id="0B79C084FCA9AD9063F06D3D9D94BA89">Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation id="FF4FA8B6D168DEECFEBAEA268E384956">JK art and science, Lauwerbes 8 2318 AT, Leiden, Netherlands</mods:affiliation>
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<mods:namePart id="E7952BC5306632AC75D33906BCF47332">Khalik, Mohd Zacaery</mods:namePart>
<mods:nameIdentifier id="82A1910D679F6F4659C282487FD97D06" type="ORCID">https://orcid.org/0000-0001-7098-0743</mods:nameIdentifier>
<mods:affiliation id="047CCF1458AB09B1B61CCB08D92E6C98">Faculty of Resource Science and Technology, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia &amp; Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia</mods:affiliation>
<mods:nameIdentifier id="87D41A2ED2B75F05E96B10D6ECB4666B" type="email">kmzacaery@unimas.my</mods:nameIdentifier>
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<mods:typeOfResource id="EE381E8F43EC51E9CBAF4B7338535BC9">text</mods:typeOfResource>
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<mods:title id="5FEA524F4A7607398507A1047C30A701">Biodiversity Data Journal</mods:title>
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<mods:date id="10A64213B622C4D6CDD3F48B6E0F7B6C">2024</mods:date>
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<mods:number id="B93FEF7DFCD779039EF7D92FD50C124D">2024-02-16</mods:number>
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<mods:number id="A3FC0E508536745312285567763887CC">12</mods:number>
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<treatment id="21C1FBCEA8B652A6A7D42869B54D2625" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625" httpUri="http://treatment.plazi.org/id/21C1FBCEA8B652A6A7D42869B54D2625" lastPageNumber="115556" pageId="0" pageNumber="115556">
<subSubSection id="AF4AAD484F23B2A162436BC75D2958B7" pageId="0" pageNumber="115556" type="nomenclature">
<paragraph id="F7BFC0D8402BB1C44971BF089E5E4D9F" pageId="0" pageNumber="115556">
<taxonomicName id="8E1148E8415EC06B3A7D4192CE5B5925" LSID="21C1FBCE-A8B6-52A6-A7D4-2869B54D2625" authority="(Godwin-Austen, 1891)" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1891" class="Gastropoda" family="Helicarionidae" genus="Helicarion" kingdom="Animalia" lsidName="Helicarion dyakanum" order="Stylommatophora" pageId="0" pageNumber="115556" phylum="Mollusca" rank="species" species="dyakanum">Helicarion dyakanum (Godwin-Austen, 1891)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="5A5DA9AC20DF54ABA013E8D21EE5549B" pageId="0" pageNumber="115556" type="materials_examined">
<paragraph id="6C72A75CDC288974ACB37A41263CDC44" pageId="0" pageNumber="115556">Materials</paragraph>
<paragraph id="CEB4D45C9AB7C11CA08C56A211A6D093" pageId="0" pageNumber="115556">
<materialsCitation id="8489BBEF8646B2178FBD4F523BBD5B1F" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectionCode="MZU" country="Sarawak" location="Taxon" municipality="Simunjan" pageId="0" pageNumber="115556" specimenCode="MZU. MOL. 22.452" specimenCount="1" typeStatus="Other material">
<emphasis id="460D6DFA5AD72A55F4FC6CAD55553773" bold="true" pageId="0" pageNumber="115556">Type status:</emphasis>
<materialsCitation id="A84DEF803DA23B22320FA2674123F58C" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectionCode="MZU" country="Sarawak" county="Type" location="Taxon" municipality="Simunjan" specimenCode="MZU. MOL. 22.452" specimenCount="1" typeStatus="Other material">
<typeStatus id="9BBCC4F3B25A43CEEE51F51F8353F3AF" pageId="0" pageNumber="115556">Other material</typeStatus>
.
<emphasis id="489859A3209379256F12228135CA2CB5" bold="true" pageId="0" pageNumber="115556">Occurrence:</emphasis>
catalogNumber:
<specimenCode id="4504A6DCE059FB084B9CCF66388A4717" pageId="0" pageNumber="115556">MZU.MOL.22.452</specimenCode>
; individualCount:
<specimenCount id="D2DD4889237452346D5E236491954560" count="1" pageId="0" pageNumber="115556" type="generic">1</specimenCount>
; occurrenceID:
<uuid id="23C0416B73A9909BF1C08C5F7480BE67">9501608E-3784-54A9-8992-A699C780D0B6</uuid>
;
<emphasis id="018792972EB32D53F0126A8A1E2B2377" bold="true" pageId="0" pageNumber="115556">
<location id="1067D90AB5402F16154BE8AFB6ED509A" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:1067D90AB5402F16154BE8AFB6ED509A" country="Sarawak" county="Type" municipality="Simunjan" name="Taxon">Taxon</location>
:
</emphasis>
family:
<location id="2C43CAA88712154710FFB0BF262B2A9F" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:2C43CAA88712154710FFB0BF262B2A9F" country="Sarawak" county="Type" municipality="Simunjan" name="Helicarionidae">Helicarionidae</location>
; genus:
<location id="E7A25E6D9F884D8E4E46213CB3E2FBD3" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:E7A25E6D9F884D8E4E46213CB3E2FBD3" country="Sarawak" county="Type" municipality="Simunjan" name="Helicarion">Helicarion</location>
; scientificNameAuthorship: (Godwin-Austen, 1891);
<emphasis id="4F06F5B44C0F350B6357D17783668E33" bold="true" pageId="0" pageNumber="115556">
<location id="29647CE16BBE5BE94AB57892B9ECF955" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:29647CE16BBE5BE94AB57892B9ECF955" country="Sarawak" county="Type" municipality="Simunjan" name="Location">Location</location>
:
</emphasis>
locality:
<location id="F14DC8D844E3E3FCEDFCB4E0F389E554" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:F14DC8D844E3E3FCEDFCB4E0F389E554" country="Sarawak" county="Type" municipality="Simunjan" name="Batu Kudik, Simunjan, Sarawak" pageId="0" pageNumber="115556">
<location id="7694B27F7CBE2987D220404187367C99" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:7694B27F7CBE2987D220404187367C99" country="Sarawak" county="Type" municipality="Simunjan" name="Batu Kudik">Batu Kudik</location>
,
<collectingMunicipality id="624A49616A55A0C8143BE079BEA72647">Simunjan</collectingMunicipality>
,
<collectingCountry id="CC7C22BBE0631CD84B46CCC0C6959F0A" name="Malaysia">Sarawak</collectingCountry>
</location>
<materialsCitation id="B95318178698AB8A9AB290667D833588" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectionCode="ME" country="Sarawak" county="Type" location="Taxon" municipality="Simunjan" pageId="0" pageNumber="115556" specimenCode="ME 13367" specimenCount="5" typeStatus="Other material">
<emphasis id="6FAB0A394C4921A1A7ED5E0C18523F2D" bold="true" pageId="0" pageNumber="115556">
<collectingCounty id="4804299BF6FADEAA29AF6E71DB3A5EA5">Type</collectingCounty>
status:
</emphasis>
<materialsCitation id="3CDF8430C30E00D5EB33264B26C3BB2A" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectionCode="ME" country="Sarawak" county="Type" location="Taxon" municipality="Simunjan" specimenCode="ME 13367" specimenCount="5" typeStatus="Other material">
<typeStatus id="00A947FF283E8EDE4F3064B56035ADE6" pageId="0" pageNumber="115556">Other material</typeStatus>
.
<emphasis id="CC4391321047F7C11A16097307E55C4E" bold="true" pageId="0" pageNumber="115556">Occurrence:</emphasis>
catalogNumber:
<specimenCode id="F44D54CDA04AA6DF85C134985444DE4B" pageId="0" pageNumber="115556">ME 13367</specimenCode>
; individualCount:
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; occurrenceID:
<uuid id="C71F18536FF26B5D1702646C06B0E1DE">FF103876-92B6-5BC6-B205-3DFBC37D5D70</uuid>
;
<emphasis id="4FC1C80CEDFF28ED4013376272713526" bold="true" pageId="0" pageNumber="115556">
<location id="175F1D72797D44A053475435B55B0652" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:175F1D72797D44A053475435B55B0652" country="Sarawak" county="Type" municipality="Simunjan" name="Taxon">Taxon</location>
:
</emphasis>
family:
<location id="EE1FBEB65CB1E6D70B597C0ED99C78B2" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:EE1FBEB65CB1E6D70B597C0ED99C78B2" country="Sarawak" county="Type" municipality="Simunjan" name="Helicarionidae">Helicarionidae</location>
; genus:
<location id="DFF6D87EBFA940F916727584DCEF5F87" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:DFF6D87EBFA940F916727584DCEF5F87" country="Sarawak" county="Type" municipality="Simunjan" name="Helicarion">Helicarion</location>
; scientificNameAuthorship: (Godwin-Austen, 1891);
<emphasis id="0AD683A2011D96976C46537F100020D6" bold="true" pageId="0" pageNumber="115556">
<location id="35AFE680FCFE7B43677BC1EC8095ECBD" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:35AFE680FCFE7B43677BC1EC8095ECBD" country="Sarawak" county="Type" municipality="Simunjan" name="Location">Location</location>
:
</emphasis>
locality:
<location id="B5C092D69444DB34ED86608E80F5B30B" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:B5C092D69444DB34ED86608E80F5B30B" country="Sarawak" county="Type" municipality="Simunjan" name="Batu Kudik, Simunjan, Sarawak" pageId="0" pageNumber="115556">
<location id="D0A314DDC5BEF142C2A9F18F2005C45B" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:D0A314DDC5BEF142C2A9F18F2005C45B" country="Sarawak" county="Type" municipality="Simunjan" name="Batu Kudik">Batu Kudik</location>
,
<collectingMunicipality id="4C320443BFDDA0FB1B38903C17AD1CF3">Simunjan</collectingMunicipality>
,
<collectingCountry id="FE311111D31FE0AB9DA31FED578BD049" name="Malaysia">Sarawak</collectingCountry>
</location>
<materialsCitation id="F7CE1C89E97A91C5555A90C8BCF794E8" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectionCode="ME" country="Sarawak" county="Type" location="Taxon" municipality="Batu Kudik" pageId="0" pageNumber="115556" specimenCode="ME 13899" specimenCount="2" typeStatus="Other material">
<emphasis id="AB682292040DE1723A7AD35250064214" bold="true" pageId="0" pageNumber="115556">
<collectingCounty id="B1B16318C42104B4AC66739FB913B989">Type</collectingCounty>
status:
</emphasis>
<materialsCitation id="E31C5AE0280D16CB8DE58B600614C6C4" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectionCode="ME" country="Sarawak" county="Simunjan" location="Taxon" municipality="Batu Kudik" specimenCode="ME 13899" specimenCount="2" typeStatus="Other material">
<typeStatus id="A3845E9E688C351A7FAC880648DFF9A1" pageId="0" pageNumber="115556">Other material</typeStatus>
.
<emphasis id="E47B59FEB68978259F11ABCD7E5EEEF9" bold="true" pageId="0" pageNumber="115556">Occurrence:</emphasis>
catalogNumber:
<specimenCode id="CAED50FB09B22BCC1ED43DC7209FD67A" pageId="0" pageNumber="115556">ME 13899</specimenCode>
; individualCount:
<specimenCount id="71D22B94701423374E44AAC3AEC6D8E2" count="2" pageId="0" pageNumber="115556" type="generic">2</specimenCount>
; occurrenceID:
<uuid id="7733E30470C20DDFA1E3CD286952D24D">8E622615-3086-5C96-B118-8EC3F0636AD5</uuid>
;
<emphasis id="06E476B6EB91582A6CCB81ABD6A28FD0" bold="true" pageId="0" pageNumber="115556">
<location id="2F0CE63C592291FDF0E2B48218FD4A1A" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:2F0CE63C592291FDF0E2B48218FD4A1A" country="Sarawak" county="Simunjan" municipality="Batu Kudik" name="Taxon">Taxon</location>
:
</emphasis>
family:
<location id="75F5ED5389F67DF4986C2782BC6D7F0F" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:75F5ED5389F67DF4986C2782BC6D7F0F" country="Sarawak" county="Simunjan" municipality="Batu Kudik" name="Helicarionidae">Helicarionidae</location>
; genus:
<location id="FEABE532D5878BA405AABF68EAAB96A5" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:FEABE532D5878BA405AABF68EAAB96A5" country="Sarawak" county="Simunjan" municipality="Batu Kudik" name="Helicarion">Helicarion</location>
; scientificNameAuthorship: (Godwin-Austen, 1891);
<emphasis id="A2E9D24564F11517666890F977A5ECDE" bold="true" pageId="0" pageNumber="115556">
<location id="4774E6D050DD2ED2573F97E16FC50DBB" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:4774E6D050DD2ED2573F97E16FC50DBB" country="Sarawak" county="Simunjan" municipality="Batu Kudik" name="Location">Location</location>
:
</emphasis>
locality:
<location id="A9ADB9A4AF04A4D2637AD25DFECED421" LSID="urn:lsid:plazi:treatment:21C1FBCEA8B652A6A7D42869B54D2625:A9ADB9A4AF04A4D2637AD25DFECED421" country="Sarawak" county="Simunjan" municipality="Batu Kudik" name="Batu Kudik, Simunjan, Sarawak" pageId="0" pageNumber="115556">
<collectingMunicipality id="D229CCB31E6E6A3466874346C1DC4739">Batu Kudik</collectingMunicipality>
,
<collectingCounty id="21F60CFB4B8E5337023A50CA8C5C2B9D">Simunjan</collectingCounty>
,
<collectingCountry id="0F8B664AC4D110C09E2665DD36B31DE5" name="Malaysia">Sarawak</collectingCountry>
</location>
</materialsCitation>
</materialsCitation>
</materialsCitation>
</materialsCitation>
</materialsCitation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="AB93E57C38C0D347841537149596C220" pageId="0" pageNumber="115556" type="distribution">
<paragraph id="411CC6ED555CCDD941327CA829F30C7A" pageId="0" pageNumber="115556">Distribution</paragraph>
<paragraph id="5ED3C58111C15D611869AF1A29FC2BE1" pageId="0" pageNumber="115556">
Sarawak: Kuching, Serian, Samarahan and Miri Divisions (
<bibRefCitation id="2E61FA7466E306E85909E29EE9CC0516" author="Godwin-Austen, H. H." journalOrPublisher="Proceedings of the Zo-ological Society of London" pageId="0" pageNumber="115556" pagination="22 - 47" refId="B10565240" refString="Godwin-Austen, H. H., 1891. On a collection of land-shells made in Borneo by Mr. A. Everett, with descriptions of supposed new species, part II. Zonitidae and Helicidae. Proceedings of the Zo-ological Society of London 1891: 22 - 47" title="On a collection of land-shells made in Borneo by Mr. A. Everett, with descriptions of supposed new species, part II. Zonitidae and Helicidae." volume="1891" year="1891">Godwin-Austen 1891</bibRefCitation>
;
<bibRefCitation id="8FD91D2F38D35B9023BCFC800EA35603" author="Marzuki, M. E." journalOrPublisher="ZooKeys" pageId="0" pageNumber="115556" pagination="13 - 113" refId="B10565373" refString="Marzuki, M. E., Liew, T. S., Jayasilan., Mohd-Azlan, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 13 - 113" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
). Sabah: West Coast Division (
<bibRefCitation id="046F3614FA9BA350F01617CC06774339" author="Vermeulen, J. J." journalOrPublisher="Institude for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu" pageId="0" pageNumber="115556" refId="B10565564" refString="Vermeulen, J. J., Liew, T. S., 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institude for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu" title="Land snails and slugs of Sabah and Labuan (Malaysia)." year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
<emphasis id="EA2175F3FF3B197735AC2FCF87598159" bold="true" pageId="0" pageNumber="115556">Distribution elsewhere</emphasis>
: Indonesia: Lombok (
<bibRefCitation id="9E1641B4784C250D28993F904E78BD35" author="Smith, E. A." journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="115556" pagination="26 - 32" refId="B10565485" refString="Smith, E. A., 1899. A list of the land-shells of the island of Lombock, with descriptions of new species. Proceedings of the Malacological Society of London 3: 26 - 32" title="A list of the land-shells of the island of Lombock, with descriptions of new species." volume="3" year="1899">Smith 1899</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="51B71B2DE3315295D394F2D9DA97B4B2" pageId="0" pageNumber="115556" type="notes">
<paragraph id="1CB691252DCE57BF3BFA5A72AA41E9B0" pageId="0" pageNumber="115556">Notes</paragraph>
<paragraph id="1C9EC9E02C2D1513A665D28A9A23F78E" pageId="0" pageNumber="115556">
Living snails were observed within the arboreal area, on leaves of palms or trees at the base of the limestone cliff. The representative is shown in Fig.
<figureCitation id="40958CE210985C9984BE6FA15C385B0C" captionStart="Figure 20" captionStartId="F10617696" captionText="Figure 20. Apertural, posterial, umbilical and apical views of Helicarion dyakanum (ME 13367). Scale = 2.5 mm." figureDoi="10.3897/BDJ.12.e115556.figure20" httpUri="https://binary.pensoft.net/fig/949252" pageId="0" pageNumber="115556">20</figureCitation>
. The identification of the species was based on the original description by
<bibRefCitation id="B660E9AC77F9401110E3D343A4C3EB8D" author="Godwin-Austen, H. H." journalOrPublisher="Proceedings of the Zo-ological Society of London" pageId="0" pageNumber="115556" pagination="22 - 47" refId="B10565240" refString="Godwin-Austen, H. H., 1891. On a collection of land-shells made in Borneo by Mr. A. Everett, with descriptions of supposed new species, part II. Zonitidae and Helicidae. Proceedings of the Zo-ological Society of London 1891: 22 - 47" title="On a collection of land-shells made in Borneo by Mr. A. Everett, with descriptions of supposed new species, part II. Zonitidae and Helicidae." volume="1891" year="1891">Godwin-Austen (1891)</bibRefCitation>
and species description by
<bibRefCitation id="CBE5BA984498CC42194D3F7996465C18" author="Marzuki, M. E." journalOrPublisher="ZooKeys" pageId="0" pageNumber="115556" pagination="13 - 113" refId="B10565373" refString="Marzuki, M. E., Liew, T. S., Jayasilan., Mohd-Azlan, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 13 - 113" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." volume="1035" year="2021">Marzuki et al. (2021)</bibRefCitation>
. Holotype, not seen (NHMUK 91.3.9.4).
</paragraph>
</subSubSection>
</treatment>
</document>