256 lines
20 KiB
XML
256 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.93.10039" ID-GBIF-Dataset="ecfecb5a-ab9b-4ad0-931e-8407c681fd11" ID-GBIF-Taxon="182236851" ID-Pensoft-Pub="1860-0743-1-45" ID-Pensoft-UUID="8BF19C7ACDA45671A0A8BCF324660CD0" ID-Zenodo-Dep="250941" ID-ZooBank="0BA1B8432BD449FC8FDAF68041A5D167" ModsDocID="1860-0743-93-1-45" checkinTime="1621423694470" checkinUser="pensoft" docAuthor="Albano, Paolo G., Bakker, Piet A. J., Janssen, Ronald & Eschner, Anita" docDate="2017" docId="1C3C5364B065EB356106DB2E0357CF70" docLanguage="en" docName="ZoosystEvol 93(1): 45-94" docOrigin="Zoosystematics and Evolution 93 (1)" docPubDate="2017-01-18" docSource="http://dx.doi.org/10.3897/zse.93.10039" docTitle="Emarginula harmilensis Sturany 1903" docType="treatment" docVersion="3" id="8BF19C7ACDA45671A0A8BCF324660CD0" lastPageId="2" lastPageNumber="47" masterDocId="8BF19C7ACDA45671A0A8BCF324660CD0" masterDocTitle="An illustrated catalogue of Rudolf Sturany's type specimens in the Naturhistorisches Museum Wien, Austria (NHMW): Red Sea gastropods" masterLastPageNumber="94" masterPageNumber="45" pageId="1" pageNumber="46" updateTime="1643477094350" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>An illustrated catalogue of Rudolf Sturany's type specimens in the Naturhistorisches Museum Wien, Austria (NHMW): Red Sea gastropods</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Albano, Paolo G.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9876-1024</mods:nameIdentifier>
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<mods:affiliation>Department of Palaeontology, University of Vienna, Althanstrasse 14, 1090 Vienna, Austria</mods:affiliation>
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<mods:nameIdentifier type="email">pgalbano@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bakker, Piet A. J.</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Janssen, Ronald</mods:namePart>
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<mods:affiliation>Malacology Section, Senckenberg Research Institute and Natural History Museum, Senckenberganlage 25, 60325 Frankfurt am Main, Germany,</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Eschner, Anita</mods:namePart>
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<mods:affiliation>Naturhistorisches Museum Wien, 3. Zoologische Abteilung, Burgring 7, 1010 Wien</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2017-01-18</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>93</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>45</mods:start>
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<mods:end>94</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zse.93.10039</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.93.10039</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-1-45</mods:identifier>
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<mods:identifier type="Zenodo-Dep">250941</mods:identifier>
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<treatment ID-GBIF-Taxon="182236851" LSID="urn:lsid:plazi:treatment:1C3C5364B065EB356106DB2E0357CF70" httpUri="http://treatment.plazi.org/id/1C3C5364B065EB356106DB2E0357CF70" lastPageId="2" lastPageNumber="47" pageId="1" pageNumber="46">
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<subSubSection pageId="1" pageNumber="46" type="nomenclature">
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<paragraph pageId="1" pageNumber="46">
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<taxonomicName authority="Sturany, 1903" authorityName="Sturany" authorityYear="1903" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Emarginula harmilensis" order="Archaeogastropoda" pageId="1" pageNumber="46" phylum="Mollusca" rank="species" species="harmilensis">Emarginula harmilensis Sturany, 1903</taxonomicName>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Emarginula harmilensis Sturany, 1903, Station 143 (Harmil Island, Dahlak Archipelago, Eritrea, Red Sea). A-F, H. Holotype, NHMW 84290: top (A-B), aperture (C), left side (D-E), protoconch (F), microsculpture (H). G. Original figure by Sturany (1903). Scale bars: A-E: 1 mm, F: 0.1 mm, H: 0.5 mm." figureDoi="10.3897/zse.93.10039.figure1" httpUri="https://binary.pensoft.net/fig/118636" pageId="1" pageNumber="46">Figure 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="46" type="reference_group">
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<paragraph pageId="1" pageNumber="46">
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<taxonomicName authorityName="Sturany" authorityYear="1903" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Emarginula harmilensis" order="Archaeogastropoda" pageId="1" pageNumber="46" phylum="Mollusca" rank="species" species="harmilensis">Emarginula harmilensis</taxonomicName>
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Sturany, 1903: 235, plate V, figures 12a-b.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="46" type="type locality">
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<paragraph pageId="1" pageNumber="46">Type locality.</paragraph>
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<paragraph pageId="1" pageNumber="46">
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Station 143,
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<normalizedToken originalValue="“nächst">"naechst</normalizedToken>
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der Insel Harmil" [near Harmil Island, Dahlak Archipelago, Eritrea]
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<geoCoordinate degrees="17" direction="north" minutes="7" orientation="latitude" precision="925" value="17.116667">17°7'N</geoCoordinate>
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,
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<geoCoordinate degrees="39" direction="east" minutes="55" orientation="longitude" precision="925" value="39.916668">39°55'E</geoCoordinate>
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, 212 m.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="46" type="type material">
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<paragraph pageId="1" pageNumber="46">Type material.</paragraph>
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<paragraph pageId="1" pageNumber="46">Holotype: NHMW 84290, length 6.8 mm.</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="46" type="description">
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<paragraph pageId="1" pageNumber="46">Original description.</paragraph>
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<paragraph pageId="1" pageNumber="46">
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<emphasis italics="true" pageId="1" pageNumber="46">Von der Station 143 (212 m); 1 Exemplar.</emphasis>
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</paragraph>
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<paragraph pageId="1" pageNumber="46">
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<emphasis italics="true" pageId="1" pageNumber="46">
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Die Schale ist 7
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<normalizedToken originalValue="½">1/2</normalizedToken>
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mm lang, 4 mm hoch, 5
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<normalizedToken originalValue="¼">1/4</normalizedToken>
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mm breit. Der stark nach
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<normalizedToken originalValue="rückwärts">rueckwaerts</normalizedToken>
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und etwas nach unten gekehrte Apex
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<normalizedToken originalValue="fällt">faellt</normalizedToken>
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fast mit dem Hinterende der Schale zusammen: die absolute Distanz des Wirbelendes zum Schalenende
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<normalizedToken originalValue="beträgt">betraegt</normalizedToken>
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2 mm, die relative (bei Projection derselben zur Basis) nur
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<normalizedToken originalValue="½">1/2</normalizedToken>
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mm. Die
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<normalizedToken originalValue="schmutzigweiße">schmutzigweisse</normalizedToken>
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bis gelbe Grundfarbe des
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<normalizedToken originalValue="Gehäuses">Gehaeuses</normalizedToken>
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<normalizedToken originalValue="erhält">erhaelt</normalizedToken>
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durch
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<normalizedToken originalValue="radiär">radiaer</normalizedToken>
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angeordnete Fleckchen, Linien und Punkte von brauner bis
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<normalizedToken originalValue="grünlicher">gruenlicher</normalizedToken>
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Farbe ein gesprenkeltes Aussehen.
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<normalizedToken originalValue="Milchweiß">Milchweiss</normalizedToken>
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<normalizedToken originalValue="gefärbt">gefaerbt</normalizedToken>
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sind die zahlreichen
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<normalizedToken originalValue="Hauptradiärrippen">Hauptradiaerrippen</normalizedToken>
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, zwischen denen zartere Rippen liegen, die mitunter dunkler erscheinen (zwischen je 2 Hauptrippen liegt eine Nebenrippe). Die
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<normalizedToken originalValue="erwähnten">erwaehnten</normalizedToken>
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Rippen werden von vielen zarten Querlinien gekreuzt, an den Kreuzungsstellen der Hauptrippen sind
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<normalizedToken originalValue="perlenförmige">perlenfoermige</normalizedToken>
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Verdickungen ausgebildet. Der Einschnitt der Schale ist etwas
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<normalizedToken originalValue="länger">laenger</normalizedToken>
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als 3 mm: die
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<normalizedToken originalValue="Ränder">Raender</normalizedToken>
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derselben sind gerade und innen mit einem verdickten Belage versehen, der sich in Form einer immer
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<normalizedToken originalValue="stärker">staerker</normalizedToken>
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werdenden
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<normalizedToken originalValue="weißen">weissen</normalizedToken>
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Schwiele bis in die Wirbelgegend fortsetzt. Die Rinne zwischen der
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<normalizedToken originalValue="Wirbelhöhe">Wirbelhoehe</normalizedToken>
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und dem blinden Ende des Einschnittes ist stark vertieft,
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<normalizedToken originalValue="weiß">weiss</normalizedToken>
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<normalizedToken originalValue="gefärbt">gefaerbt</normalizedToken>
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und quer gestreift.
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</emphasis>
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</paragraph>
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<paragraph pageId="1" pageNumber="46">
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<emphasis italics="true" pageId="1" pageNumber="46">Die neue Art ist mit</emphasis>
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<taxonomicName authorityName="A.Adams" authorityYear="1852" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Emarginula bellula" order="Archaeogastropoda" pageId="1" pageNumber="46" phylum="Mollusca" rank="species" species="bellula">Emarginula bellula</taxonomicName>
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<emphasis italics="true" pageId="1" pageNumber="46">A. Ad. von den Philippinen verwandt.</emphasis>
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</paragraph>
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<caption ID-Zenodo-Dep="985077" doi="10.3897/zse.93.10039.figure1" httpUri="https://binary.pensoft.net/fig/118636" pageId="1" pageNumber="46" start="Figure 1" startId="F1">
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<paragraph pageId="1" pageNumber="46">
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<emphasis bold="true" pageId="1" pageNumber="46">Figure 1.</emphasis>
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<taxonomicName authorityName="Sturany" authorityYear="1903" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Emarginula harmilensis" order="Archaeogastropoda" pageId="1" pageNumber="46" phylum="Mollusca" rank="species" species="harmilensis">
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<emphasis italics="true" pageId="1" pageNumber="46">Emarginula harmilensis</emphasis>
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</taxonomicName>
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Sturany, 1903, Station 143 (Harmil Island, Dahlak Archipelago, Eritrea, Red Sea).
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<emphasis bold="true" pageId="1" pageNumber="46">A-F, H</emphasis>
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. Holotype, NHMW 84290: top (
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<emphasis bold="true" pageId="1" pageNumber="46">A-B</emphasis>
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), aperture (
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<emphasis bold="true" pageId="1" pageNumber="46">C</emphasis>
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), left side (
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<emphasis bold="true" pageId="1" pageNumber="46">D-E</emphasis>
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), protoconch (
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<emphasis bold="true" pageId="1" pageNumber="46">F</emphasis>
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), microsculpture (
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<emphasis bold="true" pageId="1" pageNumber="46">H</emphasis>
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).
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<emphasis bold="true" pageId="1" pageNumber="46">G</emphasis>
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. Original figure by
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<bibRefCitation author="Sturany, R" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe" pageId="40" pageNumber="85" pagination="210 - 283" refId="B30" refString="Sturany, R, 1903. Expeditionen S.M. Schiff "Pola" in das Rothe Meer, noerdliche und suedliche Haelfte, 1895/96 - 1897/98. Zoologische Ergebnisse XXIII. Gastropoden des Rothen Meeres. Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe 74: 210 - 283" title="Expeditionen S. M. Schiff " Pola " in das Rothe Meer, noerdliche und suedliche Haelfte, 1895 / 96 - 1897 / 98. Zoologische Ergebnisse XXIII. Gastropoden des Rothen Meeres." volume="74" year="1903">Sturany (1903)</bibRefCitation>
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. Scale bars:
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<emphasis bold="true" pageId="1" pageNumber="46">A-E</emphasis>
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: 1 mm,
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<emphasis bold="true" pageId="1" pageNumber="46">F</emphasis>
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: 0.1 mm,
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<emphasis bold="true" pageId="1" pageNumber="46">H</emphasis>
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: 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="47" type="translation">
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<paragraph pageId="2" pageNumber="47">
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<pageBreakToken pageId="2" pageNumber="47" start="start">Translation</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="2" pageNumber="47">From station 143 (212 m); one specimen.</paragraph>
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<paragraph pageId="2" pageNumber="47">The shell is 7.5 mm long, 4 mm high, 5.25 mm wide. Apex positioned strongly backward and downward almost coinciding with the end of the shell: the absolute distance from the apex to the shell margin is 2 mm, the distance between the apex projection on the base and the posterior margin is only 0.5 mm. The whitish to yellow background shell colour has a spotted appearance because of the radially arranged blotches, lines and dots of brown to greenish colour. The numerous main radial ribs are milky white, whereas the finer ribs in between are darker (between two main ribs there is one finer rib). The ribs are crossed by many delicate transverse lines, the intersections form pearl-shaped tubercles. The slit is slightly longer than 3 mm: its edges are straight showing internally a thickened coating, continuing as white callus in the spire. The groove between the apex and the slit [anal fasciole] is very deep, white and transversely ribbed.</paragraph>
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<paragraph pageId="2" pageNumber="47">
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The new species is related to
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<taxonomicName authorityName="A.Adams" authorityYear="1852" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Emarginula bellula" order="Archaeogastropoda" pageId="2" pageNumber="47" phylum="Mollusca" rank="species" species="bellula">
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<emphasis italics="true" pageId="2" pageNumber="47">Emarginula bellula</emphasis>
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</taxonomicName>
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A. Adams from the Philippines.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="47" type="comments">
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<paragraph pageId="2" pageNumber="47">Comments.</paragraph>
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<paragraph pageId="2" pageNumber="47">
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[text by D. Geiger] The species is clearly a member of
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<taxonomicName authorityName="Lamarck" authorityYear="1801" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Emarginula" order="Archaeogastropoda" pageId="2" pageNumber="47" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="2" pageNumber="47">Emarginula</emphasis>
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</taxonomicName>
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given the overall shell outline, the height of the shell as well as the slit with parallel margins. It is not a juvenile
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<taxonomicName authorityName="J.Fleming" authorityYear="1822" lsidName="" pageId="2" pageNumber="47" rank="subFamily" subFamily="Fissurellinae">Fissurellinae</taxonomicName>
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or
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<taxonomicName lsidName="" pageId="2" pageNumber="47" rank="subFamily" subFamily="Diodorinae">Diodorinae</taxonomicName>
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, because at the size of
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<normalizedToken originalValue="Sturany’s">Sturany's</normalizedToken>
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specimen a hole would have formed. Note that some juvenile
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<taxonomicName authorityName="J.E.Gray" authorityYear="1821" class="Gastropoda" family="Fissurellidae" genus="Diodora" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Diodora" order="Archaeogastropoda" pageId="2" pageNumber="47" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="2" pageNumber="47">Diodora</emphasis>
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</taxonomicName>
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have been described as distinct species, such as
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<taxonomicName authorityName="Palazzi & Villari" authorityYear="2001" class="Gastropoda" family="Fissurellidae" genus="Puncturella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Puncturella piccirida" order="Archaeogastropoda" pageId="2" pageNumber="47" phylum="Mollusca" rank="species" species="piccirida">
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<emphasis italics="true" pageId="2" pageNumber="47">Puncturella piccirida</emphasis>
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</taxonomicName>
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Palazzi & Villari, 2001 from the Mediterranean. However it lacks the distinctive internal septum of
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<taxonomicName authorityName="R.T.Lowe" authorityYear="1827" class="Gastropoda" family="Fissurellidae" genus="Puncturella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Puncturella" order="Archaeogastropoda" pageId="2" pageNumber="47" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="2" pageNumber="47">Puncturella</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="2" pageNumber="47">
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<normalizedToken originalValue="Sturany’s">Sturany's</normalizedToken>
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<taxonomicName authorityName="Sturany" authorityYear="1903" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Emarginula harmilensis" order="Archaeogastropoda" pageId="2" pageNumber="47" phylum="Mollusca" rank="species" species="harmilensis">
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<emphasis italics="true" pageId="2" pageNumber="47">Emarginula harmilensis</emphasis>
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</taxonomicName>
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is characterized by a rather uncommon color pattern of tan spiral bands. Those are also found in
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<taxonomicName authorityName="Deshayes" authorityYear="1863" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Emarginula costulata" order="Archaeogastropoda" pageId="2" pageNumber="47" phylum="Mollusca" rank="species" species="costulata">
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<emphasis italics="true" pageId="2" pageNumber="47">Emarginula costulata</emphasis>
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</taxonomicName>
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Deshayes, 1863, described from Reunion Island.
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<bibRefCitation DOI="https://doi.org/10.1080/02541858.1987.11448013" author="Herbert, DG" journalOrPublisher="South African Journal of Zoology" pageId="39" pageNumber="84" pagination="1 - 13" refId="B16" refString="Herbert, DG, 1987. Taxonomic studies on the Emarginulinae (Mollusca: Gastropoda: Fissurellidae) of southern Africa and Mozambique. Hemitoma, Clypidina, Tugali, Scutus, Zeidora and two species of Emarginula. South African Journal of Zoology 22 (1): 1 - 13, DOI: https://doi.org/10.1080/02541858.1987.11448013" title="Taxonomic studies on the Emarginulinae (Mollusca: Gastropoda: Fissurellidae) of southern Africa and Mozambique. Hemitoma, Clypidina, Tugali, Scutus, Zeidora and two species of Emarginula." url="https://doi.org/10.1080/02541858.1987.11448013" volume="22" year="1987">Herbert (1987</bibRefCitation>
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: figs 39-40) illustrated the by him designated lectotype, and a more typical, fully-grown specimen (figs 41-42), and discussed previous misidentifications of the species as
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<taxonomicName authorityName="G.B.Sowerby II" authorityYear="1863" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Emarginula tenuicostata" order="Archaeogastropoda" pageId="2" pageNumber="47" phylum="Mollusca" rank="species" species="tenuicostata">
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<emphasis italics="true" pageId="2" pageNumber="47">Emarginula tenuicostata</emphasis>
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</taxonomicName>
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Adams & Sowerby, 1863. The shared characters include overall shape, placement of the spire, length of the slit, color pattern, and number and strength of the ribs. Many species found along the coast of East Africa are also encountered in the Red Sea. Accordingly,
|
||
<taxonomicName authorityName="Sturany" authorityYear="1903" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Emarginula harmilensis" order="Archaeogastropoda" pageId="2" pageNumber="47" phylum="Mollusca" rank="species" species="harmilensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="47">Emarginula harmilensis</emphasis>
|
||
</taxonomicName>
|
||
Sturany, 1903 is a junior synonym of
|
||
<taxonomicName authorityName="Deshayes" authorityYear="1863" class="Gastropoda" family="Fissurellidae" genus="Emarginula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Emarginula costulata" order="Archaeogastropoda" pageId="2" pageNumber="47" phylum="Mollusca" rank="species" species="costulata">
|
||
<emphasis italics="true" pageId="2" pageNumber="47">Emarginula costulata</emphasis>
|
||
</taxonomicName>
|
||
Deshayes, 1963.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |