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<document ID-DOI="http://doi.org/10.5281/zenodo.6485240" ID-GBIF-Dataset="3dfc64e7-8c23-456d-9678-1bb0858ac470" ID-GBIF-Taxon="150645365" ID-Zenodo-Dep="6485240" checkinTime="1545431109525" checkinUser="PlaziZenodoSync" docAuthor="Jan van Tol &amp; André Günther" docDate="2018" docId="0395A91CAC4FFFC595D1FF52FDAFFE11" docLanguage="en" docName="10.5281zenodo.1481114_vanTol_Guenther_2018.pdf" docOrigin="Odonatologica 47 (3)" docStyle="DocumentStyle{}" docTitle="Rhinocypha sangihensis Jan van Tol &amp; André Günther 2018, sp. nov." docType="treatment" docVersion="4" lastPageId="67" lastPageNumber="366" masterDocId="FFACD164AC71FF869443FFAFFF05FFBC" masterDocTitle="The Odonata of Sulawesi and adjacent islands. Part 8. Revision of the genus Rhinocypha Rambur, 1842 (Chlorocyphidae)" masterLastPageNumber="386" masterPageNumber="299" originalUpdateDomain="Boston" originalUpdateTime="1544520335069" originalUpdateUser="PlaziZenodoSync" pageId="62" pageNumber="361" updateTime="1650928538772" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Odonata of Sulawesi and adjacent islands. Part 8. Revision of the genus Rhinocypha Rambur, 1842 (Chlorocyphidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jan van Tol</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>André Günther</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Odonatologica</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-12-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>47</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>299</mods:start>
<mods:end>386</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.1481114</mods:identifier>
<mods:identifier type="GBIF-Dataset">3dfc64e7-8c23-456d-9678-1bb0858ac470</mods:identifier>
<mods:identifier type="Zenodo-Dep">1481114</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6485240" ID-GBIF-Taxon="150645365" ID-Zenodo-Dep="6485240" LSID="urn:lsid:plazi:treatment:0395A91CAC4FFFC595D1FF52FDAFFE11" httpUri="http://treatment.plazi.org/id/0395A91CAC4FFFC595D1FF52FDAFFE11" lastPageId="67" lastPageNumber="366" pageId="62" pageNumber="361">
<subSubSection box="[402,792,253,281]" pageId="62" pageNumber="361" type="nomenclature">
<paragraph blockId="62.[159,1037,253,662]" box="[402,792,253,281]" pageId="62" pageNumber="361">
<heading bold="true" box="[402,792,253,281]" centered="true" fontSize="11" level="2" pageId="62" pageNumber="361" reason="4">
<emphasis bold="true" box="[402,792,253,281]" pageId="62" pageNumber="361">
<taxonomicName authority="Jan van Tol &amp; André Günther, 2018" authorityName="Jan van Tol &amp; André Günther" authorityYear="2018" box="[402,692,253,281]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="62" pageNumber="361" phylum="Arthropoda" rank="species" species="sangihensis" status="sp. nov.">
<emphasis bold="true" box="[402,692,253,281]" italics="true" pageId="62" pageNumber="361">Rhinocypha sangihensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[699,792,253,281]" pageId="62" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[525,669,287,315]" pageId="62" pageNumber="361" type="description">
<paragraph blockId="62.[159,1037,253,662]" box="[525,669,287,315]" pageId="62" pageNumber="361">
<heading box="[525,669,287,315]" centered="true" fontSize="11" level="4" pageId="62" pageNumber="361" reason="4">
(
<figureCitation box="[531,662,287,315]" captionStart-0="Figure 66" captionStart-1="Figures 6770" captionStartId-0="63.[159,227,1496,1521]" captionStartId-1="65.[159,238,1377,1402]" captionTargetBox-0="[249,946,898,1454]" captionTargetBox-1="[178,1025,644,1347]" captionTargetId-0="figure@63.[249,946,898,1454]" captionTargetId-1="figure@65.[251,926,944,1030]" captionTargetPageId-0="63" captionTargetPageId-1="65" captionText-0="Figure 66. Rhinocypha sangihensis sp. nov. Male, holotype. Sulawesi Utara. Sangihe Is. Manganitu, 1025-v-1985 [JvT 05310]. Photo: JvT" captionText-1="Figures 6770. Rhinocypha sangihensis sp. nov. All from Sulawesi Utara, Sangihe Islands. 67 Male, head, frontal view. Sangihe, Gunung Sahendaruman, 1219-v- ­1985 [JvT 05309, reconstructed]; 68 Male, pronotum, lef lateral view. Sangihe, Manganitu, 1025­v­1985 [JvT 33005]; 69 Male, synthorax, lef lateral view. Lo- cality as in Fig. 67 [JvT 05309]; 70 Male, abdomen, lef lateral view. Locality as in Fig. 68 [JvT 05310]. Artwork by JvT" httpUri-0="https://zenodo.org/record/2200523/files/figure.png" httpUri-1="https://zenodo.org/record/2200529/files/figure.png" pageId="62" pageNumber="361">Figs 6670</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="62" pageNumber="361" type="materials_examined">
<paragraph blockId="62.[159,1037,253,662]" box="[159,360,333,359]" pageId="62" pageNumber="361">
<heading bold="true" box="[159,360,333,359]" fontSize="11" level="3" pageId="62" pageNumber="361" reason="6">
<emphasis bold="true" box="[159,360,333,359]" pageId="62" pageNumber="361">Material studied</emphasis>
</heading>
</paragraph>
<paragraph blockId="62.[159,1037,253,662]" pageId="62" pageNumber="361">
<materialsCitation ID-GBIF-Occurrence="1949138301" collectingDate="1985-05-25" country="Indonesia" latitude="3.5833333" location="Manganitu. Riverine gar- dens, coconut plantations, secondary forest" longLatPrecision="1307" longitude="125.53333" municipality="Sangihe Islands" pageId="62" pageNumber="361" specimenCode="JvT 5310" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[159,273,371,397]" pageId="62" pageNumber="361">
<emphasis bold="true" box="[159,273,371,397]" pageId="62" pageNumber="361">Holotype</emphasis>
</typeStatus>
<specimenCount box="[284,306,371,397]" pageId="62" pageNumber="361" type="male"></specimenCount>
. »
<collectingCountry box="[337,495,371,397]" name="Indonesia" pageId="62" pageNumber="361">INDONESIA</collectingCountry>
/
<collectingMunicipality box="[527,718,371,397]" pageId="62" pageNumber="361">Sangihe Islands</collectingMunicipality>
:
<location LSID="urn:lsid:plazi:treatment:0395A91CAC4FFFC595D1FF52FDAFFE11:8EE34ED1AC4FFFB8969FFEDCFD9DFE0E" country="Indonesia" latitude="3.5833333" longLatPrecision="1307" longitude="125.53333" municipality="Sangihe Islands" name="Manganitu. Riverine gar- dens, coconut plantations, secondary forest" pageId="62" pageNumber="361">Manganitu. Riverine gar- dens, coconut plantations, secondary forest</location>
.
<collectingDate box="[672,822,408,435]" pageId="62" pageNumber="361" value="1985-05-25">
<date box="[672,822,408,435]" pageId="62" pageNumber="361" value="1985-05-25">25 May 1985</date>
</collectingDate>
.
<geoCoordinate box="[832,915,408,435]" direction="north" orientation="latitude" pageId="62" pageNumber="361" precision="925" value="3.5833333">3°35N</geoCoordinate>
,
<geoCoordinate box="[925,1032,408,435]" direction="east" orientation="longitude" pageId="62" pageNumber="361" precision="925" value="125.53333">125°32E</geoCoordinate>
.
<location LSID="urn:lsid:plazi:treatment:0395A91CAC4FFFC595D1FF52FDAFFE11:8EE34ED1AC4FFFB894DCFE12FE8EFE6B" box="[159,395,445,471]" country="Indonesia" latitude="3.5833333" longLatPrecision="1307" longitude="125.53333" municipality="Sangihe Islands" name="Leg. F. G. Rozendaal" pageId="62" pageNumber="361">Leg. F.G. Rozendaal</location>
« (
<specimenCode box="[418,527,445,472]" pageId="62" pageNumber="361">JvT 5310</specimenCode>
, in RMNH;
<figureCitation box="[670,753,445,472]" captionStart="Figure 66" captionStartId="63.[159,227,1496,1521]" captionTargetBox="[249,946,898,1454]" captionTargetId="figure@63.[249,946,898,1454]" captionTargetPageId="63" captionText="Figure 66. Rhinocypha sangihensis sp. nov. Male, holotype. Sulawesi Utara. Sangihe Is. Manganitu, 1025-v-1985 [JvT 05310]. Photo: JvT" httpUri="https://zenodo.org/record/2200523/files/figure.png" pageId="62" pageNumber="361">Fig. 66</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph blockId="62.[159,1037,253,662]" box="[159,693,482,509]" pageId="62" pageNumber="361">
<emphasis bold="true" box="[159,279,483,509]" pageId="62" pageNumber="361">Paratypes</emphasis>
(5♂ 1♀) [Sulawesi Utara province]
</paragraph>
<paragraph blockId="62.[159,1037,253,662]" pageId="62" pageNumber="361">2♂ 1♀, »Sangihe Islands. Manganitu. 3°35N, 125°32E. Riverine gardens, coconut plantations, secondary forest. 1025 May 1985. Leg. F.G. Rozendaal«; 3♂, »Sangihe Islands. SSW of Liwung. NW slope of Gn Sahendaruman. [Alt.] 600650 m a.s.l. 3°32N, 125°32E. Primary forest and forest edge, water trickle and small stream. 1219 May 1985. Leg. F.G. Rozendaal«.</paragraph>
</subSubSection>
<subSubSection box="[159,855,711,774]" pageId="62" pageNumber="361" type="etymology">
<paragraph blockId="62.[159,855,711,774]" box="[159,292,711,737]" pageId="62" pageNumber="361">
<heading bold="true" box="[159,292,711,737]" fontSize="11" level="3" pageId="62" pageNumber="361" reason="6">
<emphasis bold="true" box="[159,292,711,737]" pageId="62" pageNumber="361">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="62.[159,855,711,774]" box="[159,855,748,774]" pageId="62" pageNumber="361">
<emphasis box="[159,292,748,774]" italics="true" pageId="62" pageNumber="361">Sangihensis</emphasis>
, an adjective based on the type locality Sangihe.
</paragraph>
</subSubSection>
<subSubSection lastPageId="63" lastPageNumber="362" pageId="62" pageNumber="361" type="diagnosis">
<paragraph blockId="62.[159,1036,822,1484]" box="[159,281,822,848]" pageId="62" pageNumber="361">
<heading bold="true" box="[159,281,822,848]" fontSize="11" level="3" pageId="62" pageNumber="361" reason="6">
<emphasis bold="true" box="[159,281,822,848]" pageId="62" pageNumber="361">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="62.[159,1036,822,1484]" lastBlockId="63.[159,1036,253,317]" lastPageId="63" lastPageNumber="362" pageId="62" pageNumber="361">
Small and slender species with narrow wings. Only
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[749,886,860,886]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="62" pageNumber="361" phylum="Arthropoda" rank="genus">
<emphasis box="[749,886,860,886]" italics="true" pageId="62" pageNumber="361">Rhinocypha</emphasis>
</taxonomicName>
known from the Sangihe Islands. The males share the following characters with
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="62" pageNumber="361" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis italics="true" pageId="62" pageNumber="361">R. frontalis</emphasis>
</taxonomicName>
from northern Sulawesi: head with paired spot on frons (but crescent- shaped and very small in
<taxonomicName authorityName="Jan van Tol &amp; André Günther" authorityYear="2018" box="[459,620,972,998]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="62" pageNumber="361" phylum="Arthropoda" rank="species" species="sangihensis">
<emphasis box="[459,620,972,998]" italics="true" pageId="62" pageNumber="361">R. sangihensis</emphasis>
</taxonomicName>
), a paired pale spot on the anterior lobe of the pronotum, extensive and multiple spots on lateral lobe of pro- notum, synthorax with both ante-humeral stripe on mesepisternum and a short stripe posteriorly on mesepimeron, band on synthorax with dorsoposterior projection, which is, however, distinctly shorter in
<taxonomicName authorityName="Jan van Tol &amp; André Günther" authorityYear="2018" box="[868,1029,1121,1147]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="62" pageNumber="361" phylum="Arthropoda" rank="species" species="sangihensis">
<emphasis box="[868,1029,1121,1147]" italics="true" pageId="62" pageNumber="361">R. sangihensis</emphasis>
</taxonomicName>
; white areas on tibiae of mid and hind legs short, approximately ⅔ length of tibiae; abdomen with large blue marking on S9.
<taxonomicName authorityName="Jan van Tol &amp; André Günther" authorityYear="2018" box="[761,1035,1196,1222]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="62" pageNumber="361" phylum="Arthropoda" rank="species" species="sangihensis">
<emphasis box="[761,1035,1196,1222]" italics="true" pageId="62" pageNumber="361">Rhinocypha sangihensis</emphasis>
</taxonomicName>
males differ from
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[366,491,1233,1259]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="62" pageNumber="361" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[366,491,1233,1259]" italics="true" pageId="62" pageNumber="361">R. frontalis</emphasis>
</taxonomicName>
by the following characters: smaller size, white base of labium, mid lobe of pronotum with small triangular spot in latero- anterior corner (but no pale marking in lateral corner of anterior lobe), blue markings of S4S8 extended dorso-anteriorly and running along anterior margin to dorsum, wings with narrower opaque patches, although there is much variation in
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[379,507,1420,1446]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="62" pageNumber="361" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[379,507,1420,1446]" italics="true" pageId="62" pageNumber="361">R. frontalis</emphasis>
</taxonomicName>
. Diagnostic characters of females difficult to define due to insufficient material, but
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[629,759,1457,1483]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="62" pageNumber="361" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[629,759,1457,1483]" italics="true" pageId="62" pageNumber="361">R. frontalis</emphasis>
</taxonomicName>
seems to have a larger opaque brown band on the wings, frequently have no pale markings on the frons, or on mid lobe of the pronotum.
</paragraph>
</subSubSection>
<subSubSection lastPageId="66" lastPageNumber="365" pageId="63" pageNumber="362" type="description">
<paragraph blockId="63.[159,1036,365,877]" box="[159,305,365,391]" pageId="63" pageNumber="362">
<heading bold="true" box="[159,305,365,391]" fontSize="11" level="3" pageId="63" pageNumber="362" reason="6">
<emphasis bold="true" box="[159,305,365,391]" pageId="63" pageNumber="362">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="63.[159,1036,365,877]" box="[159,882,403,430]" pageId="63" pageNumber="362">
<emphasis bold="true" box="[159,220,403,429]" pageId="63" pageNumber="362">Male</emphasis>
(holotype; JvT 5310; head separate and crushed;
<figureCitation box="[793,873,403,430]" captionStart="Figure 66" captionStartId="63.[159,227,1496,1521]" captionTargetBox="[249,946,898,1454]" captionTargetId="figure@63.[249,946,898,1454]" captionTargetPageId="63" captionText="Figure 66. Rhinocypha sangihensis sp. nov. Male, holotype. Sulawesi Utara. Sangihe Is. Manganitu, 1025-v-1985 [JvT 05310]. Photo: JvT" httpUri="https://zenodo.org/record/2200523/files/figure.png" pageId="63" pageNumber="362">Fig. 66</figureCitation>
)
</paragraph>
<paragraph blockId="63.[159,1036,365,877]" pageId="63" pageNumber="362">
Head (
<figureCitation box="[239,326,440,467]" captionStart="Figures 6770" captionStartId="65.[159,238,1377,1402]" captionTargetBox="[178,1025,644,1347]" captionTargetId="figure@65.[251,926,944,1030]" captionTargetPageId="65" captionText="Figures 6770. Rhinocypha sangihensis sp. nov. All from Sulawesi Utara, Sangihe Islands. 67 Male, head, frontal view. Sangihe, Gunung Sahendaruman, 1219-v- ­1985 [JvT 05309, reconstructed]; 68 Male, pronotum, lef lateral view. Sangihe, Manganitu, 1025­v­1985 [JvT 33005]; 69 Male, synthorax, lef lateral view. Lo- cality as in Fig. 67 [JvT 05309]; 70 Male, abdomen, lef lateral view. Locality as in Fig. 68 [JvT 05310]. Artwork by JvT" httpUri="https://zenodo.org/record/2200529/files/figure.png" pageId="63" pageNumber="362">Fig. 67</figureCitation>
) — Labium with extensive pale coloration; base and basal half of (divided) middle lobe creamy yellow; base of lateral lobes creamy yellow; remainder black. Labrum glossy brownish black. Mandibles with large bluish white spot in basal part, covering nearly entire base; anteclypeus brownish black, shiny; postclypeus, frons, vertex, occiput and post-ocular lobe velvet black with a paired, irregular spot at base of frons
<emphasis box="[858,883,628,653]" italics="true" pageId="63" pageNumber="362">ca</emphasis>
the diameter of distance between spot an antennal base (these markings much smaller than in
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[248,373,701,727]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="63" pageNumber="362" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[248,373,701,727]" italics="true" pageId="63" pageNumber="362">R. frontalis</emphasis>
</taxonomicName>
); spots behind lateral ocelli yellow, somewhat larger than ocelli, and a paired oval yellow spot on post-ocular lobe close to occiput; ge- nae with extensive bluish white markings, being almost completely covered at level of base of mandibles with separate oval spot just above it; marking on gena not extending dorsally along eye.
</paragraph>
<caption httpUri="https://zenodo.org/record/2200523/files/figure.png" pageId="63" pageNumber="362" startId="63.[159,227,1496,1521]" targetBox="[249,946,898,1454]" targetPageId="63">
<paragraph blockId="63.[159,1036,1496,1550]" pageId="63" pageNumber="362">
<emphasis bold="true" box="[159,267,1496,1521]" pageId="63" pageNumber="362">Figure 66.</emphasis>
<taxonomicName authority="Jan van Tol &amp; André Günther, 2018" authorityName="Jan van Tol &amp; André Günther" authorityYear="2018" box="[273,529,1496,1520]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="63" pageNumber="362" phylum="Arthropoda" rank="species" species="sangihensis" status="sp. nov.">
<emphasis box="[273,529,1496,1520]" italics="true" pageId="63" pageNumber="362">Rhinocypha sangihensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[535,617,1496,1521]" pageId="63" pageNumber="362" rank="species">sp. nov.</taxonomicNameLabel>
Male, holotype. Sulawesi Utara. Sangihe Is. Manganitu, 1025-v-1985 [JvT 05310]. Photo: JvT
</paragraph>
</caption>
<paragraph blockId="64.[159,1037,253,840]" pageId="64" pageNumber="363">
Thorax — Pronotum (
<figureCitation box="[417,503,253,280]" captionStart="Figures 6770" captionStartId="65.[159,238,1377,1402]" captionTargetBox="[178,1025,644,1347]" captionTargetId="figure@65.[251,926,944,1030]" captionTargetPageId="65" captionText="Figures 6770. Rhinocypha sangihensis sp. nov. All from Sulawesi Utara, Sangihe Islands. 67 Male, head, frontal view. Sangihe, Gunung Sahendaruman, 1219-v- ­1985 [JvT 05309, reconstructed]; 68 Male, pronotum, lef lateral view. Sangihe, Manganitu, 1025­v­1985 [JvT 33005]; 69 Male, synthorax, lef lateral view. Lo- cality as in Fig. 67 [JvT 05309]; 70 Male, abdomen, lef lateral view. Locality as in Fig. 68 [JvT 05310]. Artwork by JvT" httpUri="https://zenodo.org/record/2200529/files/figure.png" pageId="64" pageNumber="363">Fig. 68</figureCitation>
): anterior lobe with paired oval marking, well removed from lateral margin, median lobe with a small bluish-white mark- ing in latero-anterior corner, pointed posteriorly; lateral lobe with large creamy white spot covering most of this lobe, extending on ventral margin of pronotum as pale stripe.
</paragraph>
<paragraph blockId="64.[159,1037,253,840]" pageId="64" pageNumber="363">
Synthorax (
<figureCitation box="[323,417,440,467]" captionStart="Figures 6770" captionStartId="65.[159,238,1377,1402]" captionTargetBox="[178,1025,644,1347]" captionTargetId="figure@65.[251,926,944,1030]" captionTargetPageId="65" captionText="Figures 6770. Rhinocypha sangihensis sp. nov. All from Sulawesi Utara, Sangihe Islands. 67 Male, head, frontal view. Sangihe, Gunung Sahendaruman, 1219-v- ­1985 [JvT 05309, reconstructed]; 68 Male, pronotum, lef lateral view. Sangihe, Manganitu, 1025­v­1985 [JvT 33005]; 69 Male, synthorax, lef lateral view. Lo- cality as in Fig. 67 [JvT 05309]; 70 Male, abdomen, lef lateral view. Locality as in Fig. 68 [JvT 05310]. Artwork by JvT" httpUri="https://zenodo.org/record/2200529/files/figure.png" pageId="64" pageNumber="363">Fig. 69</figureCitation>
). Mesopreepisternum yellowish white. Black with bluish or creamy white markings. Mesepisternum anteriorly with triangu- lar marking, widest anteriorly; mesepimeron black with short pale stripe in dorso-posterior corner; wide blue stripe from mesokatepisternum over mesepimeron, metepisternum and metepimeron; stripe dorsally very ir- regular, ventrally more or less straight; mesokatepisternum with trian- gular spot in posterior corner against coxa, continuing on mesepimeron and metepisternum, widening dorsally at level of metathoracic spiracle, halfway the height of mesepimeron; posterior side on metepisternum with posterior projection, continuing posteriorly towards, but not touching hind margin.
</paragraph>
<paragraph blockId="64.[159,1037,888,989]" pageId="64" pageNumber="363">Legs — Hind margin of coxae of all legs with triangular pale blue markings. Legs black, but white innerside in distal ⁵/₇ of mid tibiae, and distal ³/₅ of hind leg; posterior side of coxae with sub-triangular spots.</paragraph>
<paragraph blockId="64.[159,1036,1037,1213]" pageId="64" pageNumber="363">
Wings — Dark opaque patch posterior to nodus or Px1, hind wing with indistinct pattern of metallic sheen on ventral side, purplish or posteriorly more brownish; base of wings transparent with brownish tinge; pterostigma of fore and hind wings brownish black, covering only five underlying cells (usually 7 in
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[310,436,1187,1213]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="64" pageNumber="363" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[310,436,1187,1213]" italics="true" pageId="64" pageNumber="363">
R.
<taxonomicName box="[341,436,1187,1213]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="64" pageNumber="363" phylum="Arthropoda" rank="species" species="frontalis">frontalis</taxonomicName>
</emphasis>
</taxonomicName>
, and 8 or more in
<taxonomicName box="[652,812,1187,1213]" pageId="64" pageNumber="363">
<emphasis box="[652,812,1187,1213]" italics="true" pageId="64" pageNumber="363">R. monochroa</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph blockId="64.[159,1037,1261,1549]" pageId="64" pageNumber="363">
Abdomen (
<figureCitation box="[293,380,1262,1288]" captionStart="Figures 6770" captionStartId="65.[159,238,1377,1402]" captionTargetBox="[178,1025,644,1347]" captionTargetId="figure@65.[251,926,944,1030]" captionTargetPageId="65" captionText="Figures 6770. Rhinocypha sangihensis sp. nov. All from Sulawesi Utara, Sangihe Islands. 67 Male, head, frontal view. Sangihe, Gunung Sahendaruman, 1219-v- ­1985 [JvT 05309, reconstructed]; 68 Male, pronotum, lef lateral view. Sangihe, Manganitu, 1025­v­1985 [JvT 33005]; 69 Male, synthorax, lef lateral view. Lo- cality as in Fig. 67 [JvT 05309]; 70 Male, abdomen, lef lateral view. Locality as in Fig. 68 [JvT 05310]. Artwork by JvT" httpUri="https://zenodo.org/record/2200529/files/figure.png" pageId="64" pageNumber="363">Fig. 70</figureCitation>
) — Dorsum black; side of tergites with extensive blue markings as follows: on segment 1 sub-triangular spot, anteriorly tapering; on S2 and S3 rectangular markings, S2 with distinct and S3 with small ven- tro-anterior emargination; S4S9 with progressively smaller spots, tapered posteriorly, especially on dorsal side, those on S6S9 not reaching posterior end of segment; blue markings on S4S8 dorso-anteriorly projected run- ning dorsad along the anterior border of segment; S10 and anal appendages black.
</paragraph>
<paragraph blockId="65.[159,1036,253,318]" pageId="65" pageNumber="364">Measurements [mm] —Hind wing length 21.0 mm, hind wing width 5.0 mm; abdomen (including appendages) 17.5.</paragraph>
<paragraph blockId="65.[159,1036,365,616]" box="[159,378,365,392]" pageId="65" pageNumber="364">
<emphasis bold="true" box="[159,247,366,392]" pageId="65" pageNumber="364">Female</emphasis>
[JvT 5353]
</paragraph>
<paragraph blockId="65.[159,1036,365,616]" box="[159,897,403,430]" pageId="65" pageNumber="364">A teneral, crushed and incomplete specimen (S5S10 missing).</paragraph>
<paragraph blockId="65.[159,1036,365,616]" pageId="65" pageNumber="364">Head — Dark, base of labium pale coloured; mandibles with large creamy spot, with a small triangular pale spot on lowest part of gena next to man- dible; another pale spot of irregular shape more dorsal on gena, with a ven- tral extension along the eye; a short and narrow stripe along the eye at level of antenna; frons between antennae with irregular creamy paired spot, ap-</paragraph>
<caption httpUri="https://zenodo.org/record/2200529/files/figure.png" pageId="65" pageNumber="364" startId="65.[159,238,1377,1402]" targetBox="[178,1025,644,1347]" targetPageId="65">
<paragraph blockId="65.[159,1037,1377,1549]" pageId="65" pageNumber="364">
<emphasis bold="true" box="[159,319,1377,1402]" pageId="65" pageNumber="364">Figures 6770.</emphasis>
<taxonomicName authority="Jan van Tol &amp; André Günther, 2018" authorityName="Jan van Tol &amp; André Günther" authorityYear="2018" box="[327,585,1377,1401]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="65" pageNumber="364" phylum="Arthropoda" rank="species" species="sangihensis" status="sp. nov.">
<emphasis box="[327,585,1377,1401]" italics="true" pageId="65" pageNumber="364">Rhinocypha sangihensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[591,675,1377,1402]" pageId="65" pageNumber="364" rank="species">sp. nov.</taxonomicNameLabel>
All from Sulawesi Utara, Sangihe Islands. 67 Male, head, frontal view. Sangihe, Gunung Sahendaruman, 1219-v- ­1985 [JvT 0 5309, reconstructed]; 68 Male, pronotum, lef lateral view. Sangihe, Manganitu, 1025­v­1985 [JvT 33005]; 69 Male, synthorax, lef lateral view. Lo- cality as in Fig. 67 [JvT 05309]; 70 Male, abdomen, lef lateral view. Locality as in Fig. 68 [JvT 05310]. Artwork by JvT
</paragraph>
</caption>
<paragraph blockId="66.[159,1036,254,317]" pageId="66" pageNumber="365">proximately the diameter of post-ocular spot; somewhat smaller paired spot behind lateral ocelli; remainder of head velvet black.</paragraph>
<paragraph blockId="66.[159,1036,365,616]" pageId="66" pageNumber="365">Thorax — Pronotum with anterior lobe with paired oval pale spot; lateral lobe with large pale spots, whitish with bluish tones.</paragraph>
<paragraph blockId="66.[159,1036,365,616]" pageId="66" pageNumber="365">
Synthorax. Brownish black with yellowish or creamish markings.Long ante- humeral stripe reaching from anterior side or mesepisternum beyond level of short stripe on mesepimeron; last stripe
<emphasis box="[657,682,516,541]" italics="true" pageId="66" pageNumber="365">ca</emphasis>
⅓ the length of metepimeron; pale stripe over synthorax from mesokatepisternum to posterior corner of metepimeron, somewhat irregular, but straight.
</paragraph>
<paragraph blockId="66.[159,441,664,691]" box="[159,441,664,691]" pageId="66" pageNumber="365">Legs — Brownish black.</paragraph>
<paragraph blockId="66.[159,1036,739,840]" pageId="66" pageNumber="365">Wings (colours not fully developed) — Opaque brown transverse band from Px 15 to distal side of pterostigma; tip opaque white, base of wing transparent with brown fume.</paragraph>
<paragraph blockId="66.[159,1037,888,1101]" pageId="66" pageNumber="365">Abdomen — S1 with sub-triangular spot,pointed anteriad;S2 with three spots, one sub-oval against anterior border of segment, one sub-triangular against hind margin, and a smaller oblong spot against ventral margin of tergite; S3 with L-shaped marking extending along anterior and dorsal margin of side of tergite; shorter rectangular spot ventro-posteriorly, sub-quadrangular spot against posterior border; S4 too crushed to evaluate, rest of abdomen missing.</paragraph>
<paragraph blockId="66.[159,1035,1149,1251]" box="[159,431,1149,1176]" pageId="66" pageNumber="365">
<heading bold="true" box="[159,431,1149,1176]" fontSize="11" level="3" pageId="66" pageNumber="365" reason="6">
<emphasis bold="true" box="[159,431,1149,1176]" pageId="66" pageNumber="365">Variation in paratypes</emphasis>
</heading>
</paragraph>
<paragraph blockId="66.[159,1035,1149,1251]" pageId="66" pageNumber="365">Mesurements [mm]. Males (n = 5). Hind wing length: x= 21.5 (20.523.0); hind wing width: x= 5.0 (5.05.0), abdomen (n =4): x= 18.0 (17.018.5).</paragraph>
</subSubSection>
<subSubSection lastPageId="67" lastPageNumber="366" pageId="66" pageNumber="365" type="distribution">
<paragraph blockId="66.[159,1036,1299,1549]" box="[159,459,1299,1325]" pageId="66" pageNumber="365">
<heading bold="true" box="[159,459,1299,1325]" fontSize="11" level="3" pageId="66" pageNumber="365" reason="6">
<emphasis bold="true" box="[159,459,1299,1325]" pageId="66" pageNumber="365">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph blockId="66.[159,1036,1299,1549]" pageId="66" pageNumber="365">This species is only known from a small series collected by F.G. Rozendaal on Manganitu (Sangihe Islands) in 1985. The habitat indicated on the labels includes »riverine gardens, coconut plantations and secondary forest«. A recent promotion film on the internet on Manganitu suggests that the hills above the town are still covered with remnants of forest, interspersed with gardens and plantations. Some parts of the island are set apart to protect</paragraph>
<paragraph blockId="67.[159,1036,254,429]" pageId="67" pageNumber="366">
endemic bird species. We presume that the streams will still be inhabited by this and other endemic odonate species, including
<emphasis box="[763,1035,291,317]" italics="true" pageId="67" pageNumber="366">
<taxonomicName authorityName="Lieftinck" authorityYear="1936" box="[763,922,291,317]" class="Insecta" family="Cordulegasteridae" genus="Celebophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="366" phylum="Arthropoda" rank="genus">Celebophlebia</taxonomicName>
carolinae
</emphasis>
van Tol, 1987,
<emphasis box="[335,629,328,354]" italics="true" pageId="67" pageNumber="366">
<taxonomicName box="[335,458,329,354]" class="Insecta" family="Platystictidae" genus="Protosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="366" phylum="Arthropoda" rank="genus">Protosticta</taxonomicName>
rozendalorum
</emphasis>
van Tol, 2000 and
<emphasis italics="true" pageId="67" pageNumber="366">
<taxonomicName authorityName="Selys" authorityYear="1840" box="[862,964,328,354]" class="Insecta" family="Chlorocyphidae" genus="Libellago" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="366" phylum="Arthropoda" rank="genus">Libellago</taxonomicName>
manganitu
</emphasis>
van Tol, 2007 (VAN TOL 1987b, 2000, 2007). These were all taken by the same collector during the same field trip.
</paragraph>
</subSubSection>
</treatment>
</document>