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<document id="1C46A964FCF92D94D7CE6146818684A7" ID-CLB-Dataset="56065" ID-DOI="10.5852/ejt.2022.809.1719" ID-GBIF-Dataset="f76de735-01e3-454c-a7f1-bbafd26686f5" ID-ISSN="2118-9773" ID-Zenodo-Dep="6419425" ID-ZooBank="A754493E-5466-4479-B515-AABEDDE09D93" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649273451180" checkinUser="felipe" docAuthor="Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. &amp; Abakumov, Evgeny V." docDate="2022" docId="03FC8666556FFF8A5067BE00FDB3FDAB" docLanguage="en" docName="ejt-809_1-71.pdf" docOrigin="European Journal of Taxonomy 809" docSource="http://dx.doi.org/10.5852/ejt.2022.809.1719" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Apatopsis Semenov 1891" docType="treatment" docVersion="9" lastPageNumber="13" masterDocId="FFC5FE1E556AFF865230BB19FFBAFF8B" masterDocTitle="A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes" masterLastPageNumber="71" masterPageNumber="1" pageNumber="6" updateTime="1699319505913" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1D66FB76130AAB3AE9633B1975EE9AB9">A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes</mods:title>
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<mods:affiliation id="3DFB0F0FF9472455975C9F3489BC4A74">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str., 45, Makhachkala, Republic of Dagestan 367000 Russia. &amp; Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000 Russia. &amp; Severtsov Institute of ecology and evolution of the Russian Academy of sciences, Leninsky Prospect, 33, Moscow 119071 Russia.</mods:affiliation>
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Genus 
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, 
<figureCitation id="136E2BF5556FFF835097BE25FD4BFADD" box="[679,753,1340,1366]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[189,1398,395,1834]" captionTargetId="figure-18@14.[189,1398,395,1834]" captionTargetPageId="14" captionText="Fig. 8. ApatopsisSemenov, 1891andHabrobates Semenov, 1903,detailsofstructure.A. A.grombczewskii Semenov, 1891, ♂ (ZIN), pronotum. B. Ditto, scales and seta on pronotum. C. Ditto, elytral scales, setation and granulation. D. Ditto, granule, seta and scales on elytra. E. H. vernalis Semenov, 1903, ♂, paralectotype (ZIN), pronotum. F. Scales and spinulose setation on pronotum." figureDoi="http://doi.org/10.5281/zenodo.6419449" httpUri="https://zenodo.org/record/6419449/files/figure.png" pageId="5" pageNumber="6">8A–D</figureCitation>
, 
<figureCitation id="136E2BF5556FFF8350CDBE25FCA7FADD" box="[765,797,1340,1366]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[197,1398,757,1854]" captionTargetId="figure-148@35.[189,1398,757,1854]" captionTargetPageId="35" captionText="Fig. 14. Apatopsis grombczewskii Semenov, 1891 (ZIN), genitalia. A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419467" httpUri="https://zenodo.org/record/6419467/files/figure.png" pageId="5" pageNumber="6">14</figureCitation>
, 
<figureCitation id="136E2BF5556FFF835119BE25FCE4FADD" box="[809,862,1340,1366]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[208,1379,488,1777]" captionTargetId="figure-18@46.[200,1387,478,1783]" captionTargetPageId="46" captionText="Fig. 23. Female genital ducts of Pimeliini. A. Apatopsis grombczewskii Semenov, 1891 (ZIN). B. Habrobates vernalis Semenov, 1903 (ZIN). C. Dietomorpha pardalis Kühnelt, 1957 (ZIN). D. Przewalskia dilatata (Reitter, 1887) (ZIN). E. Mantichorula semenowi Reitter, 1889 (ZIN). F. Platyope leucogramma (Pallas, 1773) (ZIN). G. Platyope grumi (Semenov, 1893), paralectotype (ZIN). H. Earophanta serrata (Semenov, 1893) (ZIN). Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419493" httpUri="https://zenodo.org/record/6419493/files/figure.png" pageId="5" pageNumber="6">23A</figureCitation>
, 
<figureCitation id="136E2BF5556FFF83515ABE25FC23FADD" box="[874,921,1340,1366]" captionStart="Fig" captionStartId="52.[189,232,1822,1848]" captionTargetBox="[290,1286,773,1776]" captionTargetId="figure-71@52.[281,1307,763,1783]" captionTargetPageId="52" captionText="Fig. 26. Protibiae of Pimeliini, optic microscopy. A. Habrochiton vernus Semenov-Tjan-Shansky, 1907, ♂, lectotype (ZIN). B. Habrobates gabrieli (Schuster, 1935) comb. nov., paralectotype (NMB). C. Dietomorpha pardalis Kühnelt, 1957, ♂ (ZIN). D. Mantichorula semenowi Reitter, 1889, ♂ (ZIN). E. Przewalskia dilatata (Reitter, 1887), ♂ (ZIN). F. Apatopsis grombczewskii Semenov, 1891, ♀, paralectotype (ZIN). G. Earophanta serrata (Semenov, 1893), ♂ (ZIN). H. Platyope leucogramma (Pallas, 1773), ♂ (ZIN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6419501" httpUri="https://zenodo.org/record/6419501/files/figure.png" pageId="5" pageNumber="6">26F</figureCitation>
, 
<figureCitation id="136E2BF5556FFF835194BE25FC43FADD" box="[932,1017,1340,1366]" captionStart="Fig" captionStartId="53.[189,232,1857,1883]" captionTargetBox="[189,1398,449,1817]" captionTargetId="figure-26@53.[189,1398,449,1818]" captionTargetPageId="53" captionText="Fig. 27. Protibiae of Pimeliini, SEM. A. Apatopsis grombczewskii Semenov, 1891, ♀ (ZIN), dorsal view. B. Ditto, apical part. C. Habrobates vernalis Semenov, 1903, ♂, paralectotype (ZIN), ventral view. D. Ditto, dorsal view. E. Habrochiton vernus Semenov-Tjan-Shansky, 1907, dorsal view (ZIN). F. Mantichorula semenowi Reitter, 1889, ♂ (ZIN), ventral view. G. Ditto, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6419505" httpUri="https://zenodo.org/record/6419505/files/figure.png" pageId="5" pageNumber="6">27A–B</figureCitation>
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<bibRefCitation id="EFC44A81556FFF835309BE64FD96FA1C" author="Semenov A. P." box="[313,556,1405,1431]" pageId="5" pageNumber="6" pagination="262 - 382" refId="ref33160" refString="Semenov A. P. 1891. Diagnoses coleopterorum novorum ex Asia centrali et orientali. Horae Societatis Entomologicae Rossicae 25: 262 - 382." type="journal article" year="1891">Semenov, 1891: 368</bibRefCitation>
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, 370.
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– 
<treatmentCitation id="0AF41161556FFF83537EBEA6FD82FA52" author="Semenov A. P." box="[334,568,1471,1497]" page="12" pageId="5" pageNumber="6" year="1903">
<bibRefCitation id="EFC44A81556FFF83537EBEA6FD82FA52" author="Semenov A. P." box="[334,568,1471,1497]" pageId="5" pageNumber="6" pagination="8 - 13" refId="ref33217" refString="Semenov A. P. 1903 a. Coleoptera novae faunae turanicae. Revue russe dEntomologie 3: 8 - 13." type="journal article" year="1903">Semenov 1903a: 12</bibRefCitation>
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. — 
<treatmentCitation id="0AF41161556FFF835057BEA6FC45FA52" author="Semenov-Tjan-Shansky A. P." box="[615,1023,1471,1497]" page="177" pageId="5" pageNumber="6" year="1907">
<bibRefCitation id="EFC44A81556FFF835057BEA6FC45FA52" author="Semenov-Tjan-Shansky A. P." box="[615,1023,1471,1497]" pageId="5" pageNumber="6" pagination="1907 - 1908" refId="ref33261" refString="Semenov-Tjan-Shansky A. P. 1907. Synopsis generum tribus Platyopinorum (Coleoptera, Tenebrionidae Pimeliini). Horae Societatis Entomologicae Rossicae 38 [1907 - 1908]: 175 - 184." type="journal article" year="1907">Semenov-Tjan-Shansky 1907: 177</bibRefCitation>
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, 179. — 
<treatmentCitation id="0AF41161556FFF835659BEA6FA82FA52" author="Reitter E." box="[1129,1336,1471,1497]" page="204" pageId="5" pageNumber="6" year="1893">
<bibRefCitation id="EFC44A81556FFF835659BEA6FA82FA52" author="Reitter E." box="[1129,1336,1471,1497]" pageId="5" pageNumber="6" pagination="201 - 250" refId="ref32741" refString="Reitter E. 1893. Bestimmungs-Tabelle der unechten Pimeliden aus der palaearctischen Fauna. Verhandlungen des Naturforschenden Vereines in Brunn 31 (1892): 201 - 250." type="journal article" year="1893">Reitter 1893: 204</bibRefCitation>
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, 249. — 
<treatmentCitation id="0AF41161556FFF83533EBEFBFE7FFA77" author="Gebien H." box="[270,453,1506,1532]" page="3" pageId="5" pageNumber="6" year="1940">
<bibRefCitation id="EFC44A81556FFF83533EBEFBFE7FFA77" author="Gebien H." box="[270,453,1506,1532]" pageId="5" pageNumber="6" pagination="1 - 3" refId="ref30595" refString="Gebien H. 1940. Die von Dr. Trinkler in Tibet gesammelten Tenebrioniden (Col. Tenebr.). Mitteilungen der Munchener Entomologischen Gesellschaft 30: 1 - 3." type="journal article" year="1940">Gebien 1940: 3</bibRefCitation>
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. — 
<treatmentCitation id="0AF41161556FFF8353C6BEFBFD7DFA76" author="Kuhnelt W." box="[502,711,1506,1533]" page="85" pageId="5" pageNumber="6" year="1957">
<bibRefCitation id="EFC44A81556FFF8353C6BEFBFD7DFA76" author="Kuhnelt W." box="[502,711,1506,1533]" pageId="5" pageNumber="6" pagination="65 - 102" refId="ref31209" refString="Kuhnelt W. 1957. Ergebnisse der Osterreichischen Iran-Expedition 1949 / 50. Die Tenebrioniden Irans. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch- Naturwissenschaftliche Klasse 166 (2): 65 - 102." type="journal article" year="1957">Kühnelt 1957: 85</bibRefCitation>
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. — 
<treatmentCitation id="0AF41161556FFF8350C7BEFBFBB5FA77" author="Ren G. - D. &amp; Dong S." box="[759,1039,1506,1532]" page="269" pageId="5" pageNumber="6" year="2001">
<bibRefCitation id="EFC44A81556FFF8350C7BEFBFBB5FA77" author="Ren G. - D. &amp; Dong S." box="[759,1039,1506,1532]" pageId="5" pageNumber="6" pagination="296 - 302" refId="ref32868" refString="Ren G. - D. &amp; Dong S. 2001. The beetles of the tribe Platyopini in China and description of two new species (Coleoptera: Tenebrionidae). Journal of Hebei University (Natural Science Edition) 21 (3): 296 - 302." type="journal article" year="2001">Ren &amp; Dong 2001: 269</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34F64FB556FFF83528DBD3AFB72F9E7" pageId="5" pageNumber="6" type="type_taxon">
<paragraph id="8BEA3770556FFF83528DBD3AFEE3F9B6" blockId="5.[189,345,1571,1597]" box="[189,345,1571,1597]" pageId="5" pageNumber="6">
<heading id="D0A2801C556FFF83528DBD3AFEE3F9B6" bold="true" box="[189,345,1571,1597]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B921EB62556FFF83528DBD3AFEE3F9B6" bold="true" box="[189,345,1571,1597]" pageId="5" pageNumber="6">
<typeStatus id="54EE89D2556FFF83528DBD3AFF41F9B6" box="[189,251,1571,1597]" pageId="5" pageNumber="6">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA3770556FFF83528DBD4BFB72F9E7" blockId="5.[189,1224,1618,1644]" box="[189,1224,1618,1644]" pageId="5" pageNumber="6">
<taxonomicName id="4C554CF3556FFF83528DBD4BFD1DF9E7" authority="Semenov, 1891" authorityName="Semenov" authorityYear="1891" box="[189,679,1618,1644]" class="Insecta" family="Tenebrionidae" genus="Apatopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="grombczewskii">
<emphasis id="B921EB62556FFF83528DBD4BFE52F9E7" box="[189,488,1618,1644]" italics="true" pageId="5" pageNumber="6">Apatopsis grombczewskii</emphasis>
<bibRefCitation id="EFC44A81556FFF8353C0BD4BFD1DF9E7" author="Semenov A. P." box="[496,679,1618,1644]" pageId="5" pageNumber="6" pagination="262 - 382" refId="ref33160" refString="Semenov A. P. 1891. Diagnoses coleopterorum novorum ex Asia centrali et orientali. Horae Societatis Entomologicae Rossicae 25: 262 - 382." type="journal article" year="1891">Semenov, 1891</bibRefCitation>
</taxonomicName>
, by subsequent designation (
<bibRefCitation id="EFC44A81556FFF8351C8BD4BFB01F9E7" author="Semenov A. P." box="[1016,1211,1618,1644]" pageId="5" pageNumber="6" pagination="8 - 13" refId="ref33217" refString="Semenov A. P. 1903 a. Coleoptera novae faunae turanicae. Revue russe dEntomologie 3: 8 - 13." type="journal article" year="1903">Semenov 1903a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F64FB556FFF83528DBD8AFB0BF957" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="8BEA3770556FFF83528DBD8AFE34F926" blockId="5.[189,398,1683,1709]" box="[189,398,1683,1709]" pageId="5" pageNumber="6">
<heading id="D0A2801C556FFF83528DBD8AFE34F926" bold="true" box="[189,398,1683,1709]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B921EB62556FFF83528DBD8AFE34F926" bold="true" box="[189,398,1683,1709]" pageId="5" pageNumber="6">Species included</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA3770556FFF83528DBDDBFB0BF957" blockId="5.[189,1201,1729,1756]" box="[189,1201,1729,1756]" pageId="5" pageNumber="6">
<taxonomicName id="4C554CF3556FFF83528DBDDBFD1FF957" authority="Semenov, 1891" authorityName="Semenov" authorityYear="1891" box="[189,677,1730,1756]" class="Insecta" family="Tenebrionidae" genus="Apatopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="grombczewskii">
<emphasis id="B921EB62556FFF83528DBDDBFE52F957" box="[189,488,1730,1756]" italics="true" pageId="5" pageNumber="6">Apatopsis grombczewskii</emphasis>
<bibRefCitation id="EFC44A81556FFF8353C0BDDBFD1FF957" author="Semenov A. P." box="[496,677,1730,1756]" pageId="5" pageNumber="6" pagination="262 - 382" refId="ref33160" refString="Semenov A. P. 1891. Diagnoses coleopterorum novorum ex Asia centrali et orientali. Horae Societatis Entomologicae Rossicae 25: 262 - 382." type="journal article" year="1891">Semenov, 1891</bibRefCitation>
</taxonomicName>
[= 
<taxonomicName id="4C554CF3556FFF8350FEBDDBFBA8F957" authority="Semenov, 1891" authorityName="Semenov" authorityYear="1891" box="[718,1042,1730,1756]" class="Insecta" family="Tenebrionidae" genus="Apatopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="conradti">
<emphasis id="B921EB62556FFF8350FEBDDBFCE9F957" box="[718,851,1730,1756]" italics="true" pageId="5" pageNumber="6">A. conradti</emphasis>
<bibRefCitation id="EFC44A81556FFF83516BBDDBFBA8F957" author="Semenov A. P." box="[859,1042,1730,1756]" pageId="5" pageNumber="6" pagination="262 - 382" refId="ref33160" refString="Semenov A. P. 1891. Diagnoses coleopterorum novorum ex Asia centrali et orientali. Horae Societatis Entomologicae Rossicae 25: 262 - 382." type="journal article" year="1891">Semenov, 1891</bibRefCitation>
</taxonomicName>
, syn. resurr.].
</paragraph>
</subSubSection>
<subSubSection id="C34F64FB556FFF80528DBC1AFCFFFE06" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="discussion">
<paragraph id="8BEA3770556FFF83528DBC1AFEBEF896" blockId="5.[189,260,1795,1821]" box="[189,260,1795,1821]" pageId="5" pageNumber="6">
<heading id="D0A2801C556FFF83528DBC1AFEBEF896" bold="true" box="[189,260,1795,1821]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B921EB62556FFF83528DBC1AFEBEF896" bold="true" box="[189,260,1795,1821]" pageId="5" pageNumber="6">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA3770556FFF80528DBC2BFCFFFE06" blockId="5.[188,1399,1842,2009]" lastBlockId="6.[189,1399,265,397]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Semenov described two species in this genus but did not designate a type species. Later, he placed 
<taxonomicName id="4C554CF3556FFF83528DBC4CFDE2F8E4" authority="Semenov, 1891" authorityName="Semenov" authorityYear="1891" box="[189,600,1877,1903]" class="Insecta" family="Tenebrionidae" genus="Apatopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="conradti">
<emphasis id="B921EB62556FFF83528DBC4CFE27F8E4" box="[189,413,1877,1903]" italics="true" pageId="5" pageNumber="6">Apatopsis conradti</emphasis>
<bibRefCitation id="EFC44A81556FFF835393BC4CFDE2F8E4" author="Semenov A. P." box="[419,600,1877,1903]" pageId="5" pageNumber="6" pagination="262 - 382" refId="ref33160" refString="Semenov A. P. 1891. Diagnoses coleopterorum novorum ex Asia centrali et orientali. Horae Societatis Entomologicae Rossicae 25: 262 - 382." type="journal article" year="1891">Semenov, 1891</bibRefCitation>
</taxonomicName>
as a junior synonym of 
<taxonomicName id="4C554CF3556FFF83515EBC4CFB43F8E4" authority="Semenov, 1891" authorityName="Semenov" authorityYear="1891" box="[878,1273,1877,1903]" class="Insecta" family="Tenebrionidae" genus="Apatopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="grombczewskii">
<emphasis id="B921EB62556FFF83515EBC4CFB84F8E4" box="[878,1086,1877,1903]" italics="true" pageId="5" pageNumber="6">A. grombczewskii</emphasis>
<bibRefCitation id="EFC44A81556FFF835675BC4CFB43F8E4" author="Semenov A. P." box="[1093,1273,1877,1903]" pageId="5" pageNumber="6" pagination="262 - 382" refId="ref33160" refString="Semenov A. P. 1891. Diagnoses coleopterorum novorum ex Asia centrali et orientali. Horae Societatis Entomologicae Rossicae 25: 262 - 382." type="journal article" year="1891">Semenov, 1891</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFC44A81556FFF835737BC4CFEB2F819" author="Semenov A. P." pageId="5" pageNumber="6" pagination="8 - 13" refId="ref33217" refString="Semenov A. P. 1903 a. Coleoptera novae faunae turanicae. Revue russe dEntomologie 3: 8 - 13." type="journal article" year="1903">Semenov 1903a</bibRefCitation>
; 
<bibRefCitation id="EFC44A81556FFF835324BC61FDC8F819" author="Semenov-Tjan-Shansky A. P." box="[276,626,1912,1938]" pageId="5" pageNumber="6" pagination="1907 - 1908" refId="ref33261" refString="Semenov-Tjan-Shansky A. P. 1907. Synopsis generum tribus Platyopinorum (Coleoptera, Tenebrionidae Pimeliini). Horae Societatis Entomologicae Rossicae 38 [1907 - 1908]: 175 - 184." type="journal article" year="1907">Semenov-Tjan-Shansky 1907</bibRefCitation>
) and clearly indicated that the genus 
<taxonomicName id="4C554CF3556FFF835600BC60FB1FF818" authorityName="Semenov" authorityYear="1891" box="[1072,1189,1913,1939]" class="Insecta" family="Tenebrionidae" genus="Apatopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB62556FFF835600BC60FB1FF818" box="[1072,1189,1913,1939]" italics="true" pageId="5" pageNumber="6">Apatopsis</emphasis>
</taxonomicName>
includes only the one latter species. This indication (
<bibRefCitation id="EFC44A81556FFF83506FBC85FC9AF83D" author="Semenov A. P." box="[607,800,1948,1974]" pageId="5" pageNumber="6" pagination="8 - 13" refId="ref33217" refString="Semenov A. P. 1903 a. Coleoptera novae faunae turanicae. Revue russe dEntomologie 3: 8 - 13." type="journal article" year="1903">Semenov 1903a</bibRefCitation>
) corresponds to the subsequent designation of the type species of the genus according to article 69.1.1 of 
<bibRefCitation id="EFC44A81556FFF835104BCA6FC6AF852" author="ICZN" box="[820,976,1983,2009]" pageId="5" pageNumber="6" refId="ref30699" refString="ICZN. 1999. International Code of Zoological Nomenclature. 4 th Edition. Adopted by the International Union of Biological Sciences. The International Trust for Zoological Nomenclature. London, UK. Available from http: // iczn. org [accessed 9 Mar. 2022]." type="url" year="1999">ICZN (1999)</bibRefCitation>
. 
<bibRefCitation id="EFC44A81556FFF8351E9BCA6FB38F852" author="Gebien H." box="[985,1154,1983,2009]" pageId="5" pageNumber="6" pagination="505 - 883" refId="ref30566" refString="Gebien H. 1937. Katalog der Tenebrioniden (Col. Heteromera) Teil I. Pubblicazioni del Museo Entomologico ' Pietro Rossi' 2: 505 - 883." type="journal article" year="1937">Gebien (1937)</bibRefCitation>
also listed 
<taxonomicName id="4C554CF3556FFF835730BCA6FACFF852" authorityName="Semenov" authorityYear="1891" box="[1280,1397,1983,2009]" class="Insecta" family="Tenebrionidae" genus="Apatopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB62556FFF835730BCA6FACFF852" box="[1280,1397,1983,2009]" italics="true" pageId="5" pageNumber="6">Apatopsis</emphasis>
</taxonomicName>
as monotypical genus. We do not know why these two species of 
<taxonomicName id="4C554CF3556CFF80518DBA10FB88FEA8" authorityName="Semenov" authorityYear="1891" box="[957,1074,265,291]" class="Insecta" family="Tenebrionidae" genus="Apatopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB62556CFF80518DBA10FB88FEA8" box="[957,1074,265,291]" italics="true" pageId="6" pageNumber="7">Apatopsis</emphasis>
</taxonomicName>
were given as valid in both editions of the Palaearctic Catalogue (
<bibRefCitation id="EFC44A81556CFF8050BBBA35FCECFECD" author="Lobl I. &amp; Merkl O. &amp; Ando K. &amp; Bouchard P. &amp; Lillig M. &amp; Masumoto K. &amp; Schawaller W." box="[651,854,300,326]" pageId="6" pageNumber="7" pagination="105 - 352" refId="ref31535" refString="Lobl I., Merkl O., Ando K., Bouchard P., Lillig M., Masumoto K. &amp; Schawaller W. 2008. Family Tenebrionidae Latreille, 1802. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Vol. 5: Tenebrionoidea: 105 - 352. Apollo books, Stenstrup." type="book chapter" year="2008">
Löbl 
<emphasis id="B921EB62556CFF8050FEBA34FCB7FECD" box="[718,781,300,326]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
2008
</bibRefCitation>
; 
<bibRefCitation id="EFC44A81556CFF805155BA34FB8AFECD" author="Iwan D. &amp; Lobl I. &amp; Bouchard P. &amp; Bousquet Y. &amp; Kaminski M. J. &amp; Merkl O. &amp; Ando K. &amp; Schawaller W." box="[869,1072,300,327]" pageId="6" pageNumber="7" pagination="104 - 476" refId="ref30795" refString="Iwan D., Lobl I., Bouchard P., Bousquet Y., Kaminski M. J., Merkl O., Ando K. &amp; Schawaller W. 2020. Family Tenebrionidae Latreille, 1802. In: Iwan D. &amp; Lobl I. (eds) Catalogue of Palaearctic Coleoptera. Vol. 5: Tenebrionoidea: 104 - 476. Brill, Leiden. https: // doi. org / 10.1163 / 9789004434998" type="book chapter" year="2020">
Iwan 
<emphasis id="B921EB62556CFF80519ABA34FC53FECD" box="[938,1001,300,326]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
2020
</bibRefCitation>
) and we here resurrect the synonymy after the examination of the 
<typeStatus id="54EE89D2556CFF8050B5BA56FD46FEE2" box="[645,764,335,361]" pageId="6" pageNumber="7" type="lectotype">lectotypes</typeStatus>
of these conspecific taxa: 
<taxonomicName id="4C554CF3556CFF80561EBA49FF42FE06" authority="Semenov, 1891" authorityName="Semenov" authorityYear="1891" class="Insecta" family="Tenebrionidae" genus="Apatopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="grombczewskii">
<emphasis id="B921EB62556CFF80561EBA49FB47FEE2" box="[1070,1277,335,362]" italics="true" pageId="6" pageNumber="7">A. grombczewskii</emphasis>
<bibRefCitation id="EFC44A81556CFF805733BA49FF42FE06" author="Semenov A. P." pageId="6" pageNumber="7" pagination="262 - 382" refId="ref33160" refString="Semenov A. P. 1891. Diagnoses coleopterorum novorum ex Asia centrali et orientali. Horae Societatis Entomologicae Rossicae 25: 262 - 382." type="journal article" year="1891">Semenov, 1891</bibRefCitation>
</taxonomicName>
= 
<taxonomicName id="4C554CF3556CFF805327BA6AFD0DFE06" authority="Semenov, 1891" authorityName="Semenov" authorityYear="1891" box="[279,695,370,397]" class="Insecta" family="Tenebrionidae" genus="Apatopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="conradti">
<emphasis id="B921EB62556CFF805327BA6AFE42FE07" box="[279,504,370,397]" italics="true" pageId="6" pageNumber="7">Apatopsis conradti</emphasis>
<bibRefCitation id="EFC44A81556CFF805030BA6AFD0DFE06" author="Semenov A. P." box="[512,695,371,397]" pageId="6" pageNumber="7" pagination="262 - 382" refId="ref33160" refString="Semenov A. P. 1891. Diagnoses coleopterorum novorum ex Asia centrali et orientali. Horae Societatis Entomologicae Rossicae 25: 262 - 382." type="journal article" year="1891">Semenov, 1891</bibRefCitation>
</taxonomicName>
, syn. resurr.
</paragraph>
</subSubSection>
<subSubSection id="C34F64FB556CFF80528DBAADFC6AFDEC" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BEA3770556CFF80528DBAADFE8DFE45" blockId="6.[189,311,436,462]" box="[189,311,436,462]" pageId="6" pageNumber="7">
<heading id="D0A2801C556CFF80528DBAADFE8DFE45" bold="true" box="[189,311,436,462]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B921EB62556CFF80528DBAADFE8DFE45" bold="true" box="[189,311,436,462]" pageId="6" pageNumber="7">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA3770556CFF80528DBAFBFC6AFDEC" blockId="6.[188,1399,482,615]" pageId="6" pageNumber="7">
Body brown, elongate-oval (
<figureCitation id="136E2BF5556CFF805025BAFBFD23FE76" box="[533,665,482,509]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[292,1296,745,1810]" captionTargetId="figure-129@6.[285,1303,741,1820]" captionTargetPageId="6" captionText="Fig. 1. Species of the genera Apatopsis Semenov, 1891, Habrochiton Semenov Tjan-Shansky, 1907 and Habrobates Semenov, 1903, habitus. A. A. grombczewskii Semenov, 1891, ♀, paralectotype (ZIN), dorsal view. B. Ditto, ventral view. C. Habrochiton vernus Semenov-Tjan-Shansky, 1907, ♀, paralectotype (ZIN), dorsal view. D. Ditto, ventral view. E. Habrobates vernalis Semenov, 1903, ♂, paralectotype (ZIN), dorsal view. F. Ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6419427" httpUri="https://zenodo.org/record/6419427/files/figure.png" pageId="6" pageNumber="7">Fig. 1A–B</figureCitation>
), completely covered with goldish, lanceolate, longitudinally striated scales (
<figureCitation id="136E2BF5556CFF805347B91FFE41FDAB" box="[375,507,518,544]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[189,1398,395,1834]" captionTargetId="figure-18@14.[189,1398,395,1834]" captionTargetPageId="14" captionText="Fig. 8. ApatopsisSemenov, 1891andHabrobates Semenov, 1903,detailsofstructure.A. A.grombczewskii Semenov, 1891, ♂ (ZIN), pronotum. B. Ditto, scales and seta on pronotum. C. Ditto, elytral scales, setation and granulation. D. Ditto, granule, seta and scales on elytra. E. H. vernalis Semenov, 1903, ♂, paralectotype (ZIN), pronotum. F. Scales and spinulose setation on pronotum." figureDoi="http://doi.org/10.5281/zenodo.6419449" httpUri="https://zenodo.org/record/6419449/files/figure.png" pageId="6" pageNumber="7">Fig. 8A–D</figureCitation>
) and sparse, long, erected, yellowish setae (
<figureCitation id="136E2BF5556CFF805625B91FFB23FDAB" box="[1045,1177,518,544]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[189,1398,395,1834]" captionTargetId="figure-18@14.[189,1398,395,1834]" captionTargetPageId="14" captionText="Fig. 8. ApatopsisSemenov, 1891andHabrobates Semenov, 1903,detailsofstructure.A. A.grombczewskii Semenov, 1891, ♂ (ZIN), pronotum. B. Ditto, scales and seta on pronotum. C. Ditto, elytral scales, setation and granulation. D. Ditto, granule, seta and scales on elytra. E. H. vernalis Semenov, 1903, ♂, paralectotype (ZIN), pronotum. F. Scales and spinulose setation on pronotum." figureDoi="http://doi.org/10.5281/zenodo.6419449" httpUri="https://zenodo.org/record/6419449/files/figure.png" pageId="6" pageNumber="7">Fig. 8C–D</figureCitation>
), body length 
<quantity id="4CAD9A95556CFF80577AB91FFEABFDC8" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="6" pageNumber="7" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10– 12 mm</quantity>
. Eyes small, circular, located almost dorsally; surface behind eyes not covered by anterior margin of pronotum. Head covered with short, sparse projecting thin setae.
</paragraph>
</subSubSection>
<caption id="DF2A67F8556CFF80528DBC58FCF4F863" ID-DOI="http://doi.org/10.5281/zenodo.6419427" ID-Zenodo-Dep="6419427" httpUri="https://zenodo.org/record/6419427/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1857,1883]" targetBox="[292,1296,745,1810]" targetPageId="6">
<paragraph id="8BEA3770556CFF80528DBC58FCF4F863" blockId="6.[189,1399,1857,2025]" pageId="6" pageNumber="7">
<emphasis id="B921EB62556CFF80528DBC58FEB5F8D0" bold="true" box="[189,271,1857,1883]" pageId="6" pageNumber="7">Fig. 1.</emphasis>
Species of the genera 
<taxonomicName id="4C554CF3556CFF805017BC5BFCDBF8D7" authority="Semenov, 1891" authorityName="Semenov" authorityYear="1891" box="[551,865,1858,1884]" class="Insecta" family="Tenebrionidae" genus="Apatopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB62556CFF805017BC5BFD26F8D7" box="[551,668,1858,1884]" italics="true" pageId="6" pageNumber="7">Apatopsis</emphasis>
<bibRefCitation id="EFC44A81556CFF805097BC5BFCDBF8D7" author="Semenov A. P." box="[679,865,1858,1884]" pageId="6" pageNumber="7" pagination="262 - 382" refId="ref33160" refString="Semenov A. P. 1891. Diagnoses coleopterorum novorum ex Asia centrali et orientali. Horae Societatis Entomologicae Rossicae 25: 262 - 382." type="journal article" year="1891">Semenov, 1891</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C554CF3556CFF805141BC58FACDF8D7" authority="Semenov Tjan-Shansky, 1907" authorityName="Semenov Tjan-Shansky" authorityYear="1907" box="[881,1399,1857,1884]" class="Insecta" family="Tenebrionidae" genus="Habrochiton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB62556CFF805141BC58FBBDF8D0" box="[881,1031,1857,1883]" italics="true" pageId="6" pageNumber="7">Habrochiton</emphasis>
Semenov Tjan-Shansky, 1907
</taxonomicName>
and 
<taxonomicName id="4C554CF3556CFF8052C6BC7CFDF7F8F4" authority="Semenov, 1903" authorityName="Semenov" authorityYear="1903" box="[246,589,1893,1919]" class="Insecta" family="Tenebrionidae" genus="Habrobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB62556CFF8052C6BC7CFE3BF8F4" box="[246,385,1893,1919]" italics="true" pageId="6" pageNumber="7">Habrobates</emphasis>
Semenov, 1903
</taxonomicName>
, habitus. 
<emphasis id="B921EB62556CFF8050FBBC7DFD5AF8F5" bold="true" box="[715,736,1892,1918]" pageId="6" pageNumber="7">A</emphasis>
. 
<taxonomicName id="4C554CF3556CFF8050C6BC7CFB2DF8F4" authority="Semenov, 1891" authorityName="Semenov" authorityYear="1891" box="[758,1175,1893,1919]" class="Insecta" family="Tenebrionidae" genus="Apatopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="grombczewskii">
<emphasis id="B921EB62556CFF8050C6BC7CFC77F8F4" box="[758,973,1893,1919]" italics="true" pageId="6" pageNumber="7">A. grombczewskii</emphasis>
<bibRefCitation id="EFC44A81556CFF8051EBBC7CFB2DF8F4" author="Semenov A. P." box="[987,1175,1893,1919]" pageId="6" pageNumber="7" pagination="262 - 382" refId="ref33160" refString="Semenov A. P. 1891. Diagnoses coleopterorum novorum ex Asia centrali et orientali. Horae Societatis Entomologicae Rossicae 25: 262 - 382." type="journal article" year="1891">Semenov, 1891</bibRefCitation>
</taxonomicName>
, ♀, paralectotype (ZIN), dorsal view. 
<emphasis id="B921EB62556CFF80539DBC91FE7BF829" bold="true" box="[429,449,1928,1954]" pageId="6" pageNumber="7">B</emphasis>
. Ditto, ventral view. 
<emphasis id="B921EB62556CFF8050F3BC91FD62F829" bold="true" box="[707,728,1928,1954]" pageId="6" pageNumber="7">C</emphasis>
. 
<taxonomicName id="4C554CF3556CFF8050DABC91FC60F829" authorityName="Semenov Tjan-Shansky" authorityYear="1907" box="[746,986,1928,1954]" class="Insecta" family="Tenebrionidae" genus="Habrochiton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vernus">
<emphasis id="B921EB62556CFF8050DABC91FC60F829" box="[746,986,1928,1954]" italics="true" pageId="6" pageNumber="7">Habrochiton vernus</emphasis>
</taxonomicName>
<bibRefCitation id="EFC44A81556CFF8051D4BC91FAF0F829" author="Semenov-Tjan-Shansky A. P." box="[996,1354,1928,1954]" pageId="6" pageNumber="7" pagination="1907 - 1908" refId="ref33261" refString="Semenov-Tjan-Shansky A. P. 1907. Synopsis generum tribus Platyopinorum (Coleoptera, Tenebrionidae Pimeliini). Horae Societatis Entomologicae Rossicae 38 [1907 - 1908]: 175 - 184." type="journal article" year="1907">Semenov-Tjan-Shansky, 1907</bibRefCitation>
, ♀, paralectotype (ZIN), dorsal view. 
<emphasis id="B921EB62556CFF805065BCB2FDD0F84E" bold="true" box="[597,618,1963,1989]" pageId="6" pageNumber="7">D</emphasis>
. Ditto, ventral view. 
<emphasis id="B921EB62556CFF80515DBCB2FC3BF84E" bold="true" box="[877,897,1963,1989]" pageId="6" pageNumber="7">E</emphasis>
. 
<taxonomicName id="4C554CF3556CFF8051A2BCB2FAF0F84E" authority="Semenov, 1903" authorityName="Semenov" authorityYear="1903" box="[914,1354,1963,1989]" class="Insecta" family="Tenebrionidae" genus="Habrobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vernalis">
<emphasis id="B921EB62556CFF8051A2BCB2FB3DF84E" box="[914,1159,1963,1989]" italics="true" pageId="6" pageNumber="7">Habrobates vernalis</emphasis>
Semenov, 1903
</taxonomicName>
, ♂, paralectotype (ZIN), dorsal view. 
<emphasis id="B921EB62556CFF80507ABCD7FDE6F863" bold="true" box="[586,604,1998,2024]" pageId="6" pageNumber="7">F</emphasis>
. Ditto, ventral view.
</paragraph>
</caption>
<subSubSection id="C34F64FB556DFF8A528DBA10FBDAFE24" lastPageId="12" lastPageNumber="13" pageId="7" pageNumber="8" type="description">
<paragraph id="8BEA3770556DFF81528DBA10FC79FE58" blockId="7.[189,1401,265,467]" pageId="7" pageNumber="8">
Pronotum transverse (width 1.8× length). Anterolateral angles of pronotum weakly projecting, obtuse, not extending to posterior margin of eyes. Disc of pronotum (
<figureCitation id="136E2BF5556DFF815196BA35FBB0FECD" box="[934,1034,300,326]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[292,1296,745,1810]" captionTargetId="figure-129@6.[285,1303,741,1820]" captionTargetPageId="6" captionText="Fig. 1. Species of the genera Apatopsis Semenov, 1891, Habrochiton Semenov Tjan-Shansky, 1907 and Habrobates Semenov, 1903, habitus. A. A. grombczewskii Semenov, 1891, ♀, paralectotype (ZIN), dorsal view. B. Ditto, ventral view. C. Habrochiton vernus Semenov-Tjan-Shansky, 1907, ♀, paralectotype (ZIN), dorsal view. D. Ditto, ventral view. E. Habrobates vernalis Semenov, 1903, ♂, paralectotype (ZIN), dorsal view. F. Ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6419427" httpUri="https://zenodo.org/record/6419427/files/figure.png" pageId="7" pageNumber="8">Figs 1A</figureCitation>
, 
<figureCitation id="136E2BF5556DFF815628BA35FB84FECD" box="[1048,1086,300,326]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[189,1398,395,1834]" captionTargetId="figure-18@14.[189,1398,395,1834]" captionTargetPageId="14" captionText="Fig. 8. ApatopsisSemenov, 1891andHabrobates Semenov, 1903,detailsofstructure.A. A.grombczewskii Semenov, 1891, ♂ (ZIN), pronotum. B. Ditto, scales and seta on pronotum. C. Ditto, elytral scales, setation and granulation. D. Ditto, granule, seta and scales on elytra. E. H. vernalis Semenov, 1903, ♂, paralectotype (ZIN), pronotum. F. Scales and spinulose setation on pronotum." figureDoi="http://doi.org/10.5281/zenodo.6419449" httpUri="https://zenodo.org/record/6419449/files/figure.png" pageId="7" pageNumber="8">8A</figureCitation>
) moderately convex, with suture-like longitudinal median line, surface of pronotum without depressions. Prothorax almost cylindrical. Prosternum without transverse triangular depression along anterior margin, short, 1.8× as long as transverse diameter of one procoxa. Prosternal process short (ratio of length of prosternal process to its maximum width: 3:2), not raised between procoxae.
</paragraph>
<paragraph id="8BEA3770556DFF81528DBAE3FACEFDD0" blockId="7.[187,1399,506,603]" pageId="7" pageNumber="8">
Elytra translucent (see in transmitted light), with wide, oval depressions on lateral sides, surface of depressions densely covered with very long yellowish setae (
<figureCitation id="136E2BF5556DFF81514AB907FC6FFDB3" box="[890,981,542,568]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[292,1296,745,1810]" captionTargetId="figure-129@6.[285,1303,741,1820]" captionTargetPageId="6" captionText="Fig. 1. Species of the genera Apatopsis Semenov, 1891, Habrochiton Semenov Tjan-Shansky, 1907 and Habrobates Semenov, 1903, habitus. A. A. grombczewskii Semenov, 1891, ♀, paralectotype (ZIN), dorsal view. B. Ditto, ventral view. C. Habrochiton vernus Semenov-Tjan-Shansky, 1907, ♀, paralectotype (ZIN), dorsal view. D. Ditto, ventral view. E. Habrobates vernalis Semenov, 1903, ♂, paralectotype (ZIN), dorsal view. F. Ditto, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6419427" httpUri="https://zenodo.org/record/6419427/files/figure.png" pageId="7" pageNumber="8">Fig. 1A</figureCitation>
). Surface of elytra with longitudinal rows of very small sparse granules; each granule with long, erect, yellowish setae (
<figureCitation id="136E2BF5556DFF8156A7B958FB4CFDD0" box="[1175,1270,577,603]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[189,1398,395,1834]" captionTargetId="figure-18@14.[189,1398,395,1834]" captionTargetPageId="14" captionText="Fig. 8. ApatopsisSemenov, 1891andHabrobates Semenov, 1903,detailsofstructure.A. A.grombczewskii Semenov, 1891, ♂ (ZIN), pronotum. B. Ditto, scales and seta on pronotum. C. Ditto, elytral scales, setation and granulation. D. Ditto, granule, seta and scales on elytra. E. H. vernalis Semenov, 1903, ♂, paralectotype (ZIN), pronotum. F. Scales and spinulose setation on pronotum." figureDoi="http://doi.org/10.5281/zenodo.6419449" httpUri="https://zenodo.org/record/6419449/files/figure.png" pageId="7" pageNumber="8">Fig. 8D</figureCitation>
). Scutellar
</paragraph>
<caption id="DF2A67F8556DFF81528EBC7DFCE7F862" ID-DOI="http://doi.org/10.5281/zenodo.6419431" ID-Zenodo-Dep="6419431" httpUri="https://zenodo.org/record/6419431/files/figure.png" pageId="7" pageNumber="8" startId="7.[190,233,1892,1918]" targetBox="[189,1398,684,1851]" targetPageId="7">
<paragraph id="8BEA3770556DFF81528EBC7DFCE7F862" blockId="7.[189,1399,1892,2025]" pageId="7" pageNumber="8">
<emphasis id="B921EB62556DFF81528EBC7DFEA9F8F4" bold="true" box="[190,275,1892,1919]" pageId="7" pageNumber="8">Fig. 2.</emphasis>
<taxonomicName id="4C554CF3556DFF81532FBC7CFD53F80B" authority="(Schuster, 1935)" authorityName="Chigray &amp; Nabozhenko &amp; Chigray &amp; Abakumov" authorityYear="2022" baseAuthorityName="Schuster" baseAuthorityYear="1935" box="[287,745,1893,1920]" class="Insecta" family="Tenebrionidae" genus="Habrobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gabrieli" status="comb. nov.">
<emphasis id="B921EB62556DFF81532FBC7CFDAFF8F4" box="[287,533,1893,1919]" italics="true" pageId="7" pageNumber="8">Habrobates gabrieli</emphasis>
(
<bibRefCitation id="EFC44A81556DFF81501BBC7CFD5BF80B" author="Schuster A." box="[555,737,1893,1920]" pageId="7" pageNumber="8" pagination="20 - 29" refId="ref33019" refString="Schuster A. 1935. Die Tenebrioniden-Ausbeute Dr. Gabriels in Persien. Koleopterologische Rundschau 21: 20 - 29." type="journal article" year="1935">Schuster, 1935</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2125619556DFF8150C7BC7CFC38F8F4" box="[759,898,1893,1919]" pageId="7" pageNumber="8" rank="species">comb. nov.</taxonomicNameLabel>
, type specimens, habitus, labels (NHB). 
<emphasis id="B921EB62556DFF81528DBC91FF68F829" bold="true" box="[189,210,1928,1954]" pageId="7" pageNumber="8">A</emphasis>
. Lectotype, dorsal view. 
<emphasis id="B921EB62556DFF8153CEBC91FDABF829" bold="true" box="[510,529,1928,1954]" pageId="7" pageNumber="8">B</emphasis>
. Ditto, ventral view. 
<emphasis id="B921EB62556DFF81513CBC91FC93F829" bold="true" box="[780,809,1928,1954]" pageId="7" pageNumber="8">C.</emphasis>
Ditto, lateral view. 
<emphasis id="B921EB62556DFF815625BC91FB90F829" bold="true" box="[1045,1066,1928,1954]" pageId="7" pageNumber="8">D</emphasis>
. Paralectotype, dorsal view. 
<emphasis id="B921EB62556DFF81528EBCB2FF6BF84E" bold="true" box="[190,209,1963,1989]" pageId="7" pageNumber="8">E</emphasis>
. Ditto, lateral view. 
<emphasis id="B921EB62556DFF8153FCBCB2FE64F84E" bold="true" box="[460,478,1963,1989]" pageId="7" pageNumber="8">F</emphasis>
. Labels of the lectotype. 
<emphasis id="B921EB62556DFF815125BCB2FC96F84E" bold="true" box="[789,812,1963,1989]" pageId="7" pageNumber="8">G</emphasis>
. Labels of the paralectotype. Photos courtesy of © Naturhistorisches Museum Basel (Basel, Switzerland).
</paragraph>
</caption>
<paragraph id="8BEA37705562FF8E528DBA10FD8FFECD" blockId="8.[188,1398,265,327]" pageId="8" pageNumber="9">shield almost concealed by base of pronotum. Transverse length of metacoxae 1.2× as long as intercoxal process of abdominal ventrite 1.</paragraph>
<paragraph id="8BEA37705562FF8F528DBA77FE63FE58" blockId="8.[189,1399,366,568]" lastBlockId="9.[189,1398,265,467]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
Trochanters with long sparse setae. Meso- and metafemora weakly curved outward, protibiae straight, meso- and metatibiae weakly curved outward, covered with short, yellowish setae. Protibiae (
<figureCitation id="136E2BF55562FF8E5739BA88FAC9FE20" box="[1289,1395,401,427]" captionStart="Fig" captionStartId="52.[189,232,1822,1848]" captionTargetBox="[290,1286,773,1776]" captionTargetId="figure-71@52.[281,1307,763,1783]" captionTargetPageId="52" captionText="Fig. 26. Protibiae of Pimeliini, optic microscopy. A. Habrochiton vernus Semenov-Tjan-Shansky, 1907, ♂, lectotype (ZIN). B. Habrobates gabrieli (Schuster, 1935) comb. nov., paralectotype (NMB). C. Dietomorpha pardalis Kühnelt, 1957, ♂ (ZIN). D. Mantichorula semenowi Reitter, 1889, ♂ (ZIN). E. Przewalskia dilatata (Reitter, 1887), ♂ (ZIN). F. Apatopsis grombczewskii Semenov, 1891, ♀, paralectotype (ZIN). G. Earophanta serrata (Semenov, 1893), ♂ (ZIN). H. Platyope leucogramma (Pallas, 1773), ♂ (ZIN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6419501" httpUri="https://zenodo.org/record/6419501/files/figure.png" pageId="8" pageNumber="9">Figs 26F</figureCitation>
, 
<figureCitation id="136E2BF55562FF8E528DBAADFEA9FE45" box="[189,275,436,462]" captionStart="Fig" captionStartId="53.[189,232,1857,1883]" captionTargetBox="[189,1398,449,1817]" captionTargetId="figure-26@53.[189,1398,449,1818]" captionTargetPageId="53" captionText="Fig. 27. Protibiae of Pimeliini, SEM. A. Apatopsis grombczewskii Semenov, 1891, ♀ (ZIN), dorsal view. B. Ditto, apical part. C. Habrobates vernalis Semenov, 1903, ♂, paralectotype (ZIN), ventral view. D. Ditto, dorsal view. E. Habrochiton vernus Semenov-Tjan-Shansky, 1907, dorsal view (ZIN). F. Mantichorula semenowi Reitter, 1889, ♂ (ZIN), ventral view. G. Ditto, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6419505" httpUri="https://zenodo.org/record/6419505/files/figure.png" pageId="8" pageNumber="9">27A–B</figureCitation>
) widened at apex, outer anterior margin forms projecting process with outer apical tooth. Outer margin of protibiae covered with conical spinose tubercles, ventral surface of protibiae with long dense setae. Protibial terminal spurs elongated, extending to base of protarsomere 5, mesotibial terminal spursextending to apex of mesotarsomere 2, metatibial terminal spurs extending to midlength of metatarsomere 1. Outer and inner protibial terminal spurs subequal in length, inner meso- and metatibial terminal spurs longer than outer ones, spurs flattened, acutely angulate at apex. Tarsi elongated, flattened from sides, protarsi with short spinose setae ventrally and more elongate fine dense setae dorsally, mesotarsi with very long erected setae dorsally and shorter setae ventrally, metatarsi with very long and dense suberected setae dorsally and very short recumbent setae ventrally. Tarsal claws elongated, thin, weakly curved outward.
</paragraph>
<caption id="DF2A67F85562FF8E528DBC7DFB90F863" ID-DOI="http://doi.org/10.5281/zenodo.6419433" ID-Zenodo-Dep="6419433" httpUri="https://zenodo.org/record/6419433/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1892,1918]" targetBox="[249,1340,656,1847]" targetPageId="8">
<paragraph id="8BEA37705562FF8E528DBC7DFB90F863" blockId="8.[189,1399,1892,2025]" pageId="8" pageNumber="9">
<emphasis id="B921EB625562FF8E528DBC7DFEB4F8F4" bold="true" box="[189,270,1892,1919]" pageId="8" pageNumber="9">Fig. 3.</emphasis>
Species of the genus 
<taxonomicName id="4C554CF35562FF8E502DBC7CFCC9F80B" authority="Kuhnelt, 1957" authorityName="Kuhnelt" authorityYear="1957" box="[541,883,1893,1920]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB625562FF8E502DBC7CFD01F8F4" box="[541,699,1893,1919]" italics="true" pageId="8" pageNumber="9">Dietomorpha</emphasis>
<bibRefCitation id="EFC44A815562FF8E50F6BC7CFCC9F80B" author="Kuhnelt W." box="[710,883,1893,1920]" pageId="8" pageNumber="9" pagination="65 - 102" refId="ref31209" refString="Kuhnelt W. 1957. Ergebnisse der Osterreichischen Iran-Expedition 1949 / 50. Die Tenebrioniden Irans. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch- Naturwissenschaftliche Klasse 166 (2): 65 - 102." type="journal article" year="1957">Kühnelt, 1957</bibRefCitation>
</taxonomicName>
, habitus. 
<emphasis id="B921EB625562FF8E51D9BC7DFC44F8F5" bold="true" box="[1001,1022,1892,1918]" pageId="8" pageNumber="9">A</emphasis>
. 
<taxonomicName id="4C554CF35562FF8E5620BC7FFAEBF80B" authority="Kuhnelt, 1957" authorityName="Kuhnelt" authorityYear="1957" box="[1040,1361,1893,1920]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pardalis">
<emphasis id="B921EB625562FF8E5620BC7FFB23F8F4" box="[1040,1177,1893,1919]" italics="true" pageId="8" pageNumber="9">D. pardalis</emphasis>
<bibRefCitation id="EFC44A815562FF8E5694BC7CFAEBF80B" author="Kuhnelt W." box="[1188,1361,1893,1920]" pageId="8" pageNumber="9" pagination="65 - 102" refId="ref31209" refString="Kuhnelt W. 1957. Ergebnisse der Osterreichischen Iran-Expedition 1949 / 50. Die Tenebrioniden Irans. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch- Naturwissenschaftliche Klasse 166 (2): 65 - 102." type="journal article" year="1957">Kühnelt, 1957</bibRefCitation>
</taxonomicName>
, ♂ (ZIN), dorsal view. 
<emphasis id="B921EB625562FF8E5381BC91FE7FF829" bold="true" box="[433,453,1928,1954]" pageId="8" pageNumber="9">B</emphasis>
. Ditto, ventral view. 
<emphasis id="B921EB625562FF8E50FDBC91FD58F829" bold="true" box="[717,738,1928,1954]" pageId="8" pageNumber="9">C</emphasis>
. 
<taxonomicName id="4C554CF35562FF8E50C5BC90FB66F829" authority="S. Chigray &amp; Nabozhenko" authorityName="S. Chigray &amp; Nabozhenko" authorityYear="2022" box="[757,1244,1928,1954]" class="Insecta" family="Tenebrionidae" genus="Dietomorpha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="gonzalesi" status="sp. nov.">
<emphasis id="B921EB625562FF8E50C5BC90FC34F829" box="[757,910,1928,1954]" italics="true" pageId="8" pageNumber="9">D. gonzalesi</emphasis>
S. Chigray &amp; Nabozhenko
</taxonomicName>
<taxonomicNameLabel id="A21256195562FF8E56D8BC91FAFDF829" box="[1256,1351,1928,1954]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, ♀, holotype (MNHP), dorsal view. 
<emphasis id="B921EB625562FF8E5008BCB2FDF7F84E" bold="true" box="[568,589,1963,1989]" pageId="8" pageNumber="9">D</emphasis>
. Ditto, ventral view. 
<emphasis id="B921EB625562FF8E5176BCB2FCE0F84E" bold="true" box="[838,858,1963,1989]" pageId="8" pageNumber="9">E</emphasis>
. Ditto, anterior view. 
<emphasis id="B921EB625562FF8E566CBCB2FBD4F84E" bold="true" box="[1116,1134,1963,1989]" pageId="8" pageNumber="9">F</emphasis>
. Ditto, posterior view. 
<emphasis id="B921EB625562FF8E528DBCD7FF6EF863" bold="true" box="[189,212,1998,2024]" pageId="8" pageNumber="9">G</emphasis>
. Ditto, dorso-antero-lateral view. C–G: photographs by David Gonzales.
</paragraph>
</caption>
<caption id="DF2A67F85563FF8F528DBC7DFB0FF863" ID-DOI="http://doi.org/10.5281/zenodo.6419435" ID-Zenodo-Dep="6419435" httpUri="https://zenodo.org/record/6419435/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1892,1918]" targetBox="[233,1353,649,1847]" targetPageId="9">
<paragraph id="8BEA37705563FF8F528DBC7DFB0FF863" blockId="9.[189,1399,1892,2025]" pageId="9" pageNumber="10">
<emphasis id="B921EB625563FF8F528DBC7DFEB4F8F4" bold="true" box="[189,270,1892,1919]" pageId="9" pageNumber="10">Fig. 4.</emphasis>
Species of the genera 
<taxonomicName id="4C554CF35563FF8F5014BC7CFCC2F8F4" authority="Semenov, 1893" authorityName="Semenov" authorityYear="1893" box="[548,888,1893,1919]" class="Magnoliopsida" family="Solanaceae" genus="Przewalskia" kingdom="Plantae" order="Solanales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">
<emphasis id="B921EB625563FF8F5014BC7CFD0FF8F4" box="[548,693,1893,1919]" italics="true" pageId="9" pageNumber="10">Przewalskia</emphasis>
<bibRefCitation id="EFC44A815563FF8F508FBC7CFCC2F8F4" author="Semenov A. P." box="[703,888,1893,1919]" pageId="9" pageNumber="10" pagination="1892 - 1893" refId="ref33186" refString="Semenov A. P. 1893. Symbolae ad cognitionem pimeliidarum. I - III. Horae Societatis Entomologicae Rossicae 27 [1892 - 1893]: 249 - 264." type="journal article" year="1893">Semenov, 1893</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C554CF35563FF8F51B7BC7CFB75F8F4" authority="Reitter, 1889" authorityName="Semenovi Rtt. Coll. Reitter" authorityYear="1889" box="[903,1231,1893,1919]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB625563FF8F51B7BC7CFB90F8F4" box="[903,1066,1893,1919]" italics="true" pageId="9" pageNumber="10">Mantichorula</emphasis>
<bibRefCitation id="EFC44A815563FF8F5604BC7CFB75F8F4" author="Reitter E." box="[1076,1231,1893,1919]" pageId="9" pageNumber="10" pagination="678 - 710" refId="ref32704" refString="Reitter E. 1889. Insecta, a cl. G. N. Potanin in China et in Mongolia novissime lecta. XIII. Tenebrionidae. Horae Societatis Entomologicae Rossicae 23: 678 - 710." type="journal article" year="1889">Reitter, 1889</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C554CF35563FF8F573DBC7CFDA2F829" authority="Fischer von Waldheim, 1820" authorityName="Fischer von Waldheim" authorityYear="1820" class="Insecta" family="Tenebrionidae" genus="Platyope" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB625563FF8F573DBC7CFACCF8F4" box="[1293,1398,1893,1919]" italics="true" pageId="9" pageNumber="10">Platyope</emphasis>
<bibRefCitation id="EFC44A815563FF8F528DBC91FDA2F829" author="Fischer von Waldheim G." box="[189,536,1928,1954]" pageId="9" pageNumber="10" refId="ref30454" refString="Fischer von Waldheim G. 1820. Entomographie de la Russie (Entomographia Imperii Rossici). Auctoritate Societatis Caesareae Mosquensis naturae scrutatorum collecta et in lucem edita. Vol. 1. A. Semen, Moscow." type="book" year="1820">Fischer von Waldheim, 1820</bibRefCitation>
</taxonomicName>
, habitus. 
<emphasis id="B921EB625563FF8F50BEBC91FD1BF829" bold="true" box="[654,673,1928,1954]" pageId="9" pageNumber="10">A</emphasis>
. 
<taxonomicName id="4C554CF35563FF8F5082BC91FBDEF829" authority="(Reitter, 1887)" authorityName="ZIN" baseAuthorityName="Reitter" baseAuthorityYear="1887" box="[690,1124,1928,1954]" class="Magnoliopsida" family="Solanaceae" genus="Przewalskia" kingdom="Plantae" order="Solanales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="dilatata">
<emphasis id="B921EB625563FF8F5082BC91FC13F829" box="[690,937,1928,1954]" italics="true" pageId="9" pageNumber="10">Przewalskia dilatata</emphasis>
(
<bibRefCitation id="EFC44A815563FF8F518BBC91FBE0F829" author="Reitter E." box="[955,1114,1928,1954]" pageId="9" pageNumber="10" pagination="355 - 389" refId="ref32671" refString="Reitter E. 1887. Insecta in itinere Cl. N. Przewalskii in Asia Centrali novissime lecta. IX. Tenebrionidae. Horae Societatis Entomologicae Rossicae 21: 355 - 389." type="journal article" year="1887">Reitter, 1887</bibRefCitation>
)
</taxonomicName>
, ♀ (ZIN), dorsal view. 
<emphasis id="B921EB625563FF8F528DBCB2FF6BF84E" bold="true" box="[189,209,1963,1989]" pageId="9" pageNumber="10">B</emphasis>
. Ditto, ventral view. 
<emphasis id="B921EB625563FF8F53E6BCB2FE51F84E" bold="true" box="[470,491,1963,1989]" pageId="9" pageNumber="10">C</emphasis>
. 
<taxonomicName id="4C554CF35563FF8F53CDBCB5FCFFF84E" authority="Reitter, 1889" authorityName="Reitter" authorityYear="1889" box="[509,837,1963,1989]" class="Insecta" family="Tenebrionidae" genus="Mantichorula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="semenowi">
<emphasis id="B921EB625563FF8F53CDBCB5FD27F84E" box="[509,669,1964,1989]" italics="true" pageId="9" pageNumber="10">M. semenowi</emphasis>
<bibRefCitation id="EFC44A815563FF8F5097BCB2FCFFF84E" author="Reitter E." box="[679,837,1963,1989]" pageId="9" pageNumber="10" pagination="678 - 710" refId="ref32704" refString="Reitter E. 1889. Insecta, a cl. G. N. Potanin in China et in Mongolia novissime lecta. XIII. Tenebrionidae. Horae Societatis Entomologicae Rossicae 23: 678 - 710." type="journal article" year="1889">Reitter, 1889</bibRefCitation>
</taxonomicName>
, ♂ (ZIN), dorsal view. 
<emphasis id="B921EB625563FF8F5657BCB2FBC6F84E" bold="true" box="[1127,1148,1963,1989]" pageId="9" pageNumber="10">D</emphasis>
. Ditto, ventral view. 
<emphasis id="B921EB625563FF8F528DBCD7FF6BF863" bold="true" box="[189,209,1998,2024]" pageId="9" pageNumber="10">E</emphasis>
. 
<taxonomicName id="4C554CF35563FF8F52EFBCD7FD1BF863" authority="(Pallas, 1773)" authorityName="ZIN" baseAuthorityName="Pallas" baseAuthorityYear="1773" box="[223,673,1998,2024]" class="Insecta" family="Tenebrionidae" genus="Platyope" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="leucogramma">
<emphasis id="B921EB625563FF8F52EFBCD7FE49F863" box="[223,499,1998,2024]" italics="true" pageId="9" pageNumber="10">Platyope leucogramma</emphasis>
(Pallas, 1773)
</taxonomicName>
, ♂ (ZIN), dorsal view. 
<emphasis id="B921EB625563FF8F5181BCD7FC79F863" bold="true" box="[945,963,1998,2024]" pageId="9" pageNumber="10">F</emphasis>
. Ditto, ventral view.
</paragraph>
</caption>
<caption id="DF2A67F85560FF8C528DBC58FB68F863" ID-DOI="http://doi.org/10.5281/zenodo.6419437" ID-Zenodo-Dep="6419437" httpUri="https://zenodo.org/record/6419437/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1857,1883]" targetBox="[189,1398,349,1805]" targetPageId="10">
<paragraph id="8BEA37705560FF8C528DBC58FB68F863" blockId="10.[188,1399,1857,2025]" pageId="10" pageNumber="11">
<emphasis id="B921EB625560FF8C528DBC58FEB4F8D7" bold="true" box="[189,270,1857,1884]" pageId="10" pageNumber="11">Fig. 5.</emphasis>
Species of the genus 
<taxonomicName id="4C554CF35560FF8C502CBC58FC56F8D7" authority="Fischer von Waldheim, 1820" authorityName="Fischer von Waldheim" authorityYear="1820" box="[540,1004,1857,1884]" class="Insecta" family="Tenebrionidae" genus="Platyope" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB625560FF8C502CBC58FD3FF8D0" box="[540,645,1857,1883]" italics="true" pageId="10" pageNumber="11">Platyope</emphasis>
<bibRefCitation id="EFC44A815560FF8C50A0BC58FC56F8D7" author="Fischer von Waldheim G." box="[656,1004,1857,1884]" pageId="10" pageNumber="11" refId="ref30454" refString="Fischer von Waldheim G. 1820. Entomographie de la Russie (Entomographia Imperii Rossici). Auctoritate Societatis Caesareae Mosquensis naturae scrutatorum collecta et in lucem edita. Vol. 1. A. Semen, Moscow." type="book" year="1820">Fischer von Waldheim, 1820</bibRefCitation>
</taxonomicName>
, details of structure and habitus. 
<emphasis id="B921EB625560FF8C528DBC7DFF68F8F5" bold="true" box="[189,210,1892,1918]" pageId="10" pageNumber="11">A</emphasis>
. 
<taxonomicName id="4C554CF35560FF8C52D4BC7FFDE3F8F4" authority="(Pallas, 1773)" authorityName="ZIN" baseAuthorityName="Pallas" baseAuthorityYear="1773" box="[228,601,1893,1919]" class="Insecta" family="Tenebrionidae" genus="Platyope" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="leucogramma">
<emphasis id="B921EB625560FF8C52D4BC7FFE1DF8F4" box="[228,423,1893,1919]" italics="true" pageId="10" pageNumber="11">P. leucogramma</emphasis>
(Pallas, 1773)
</taxonomicName>
, ♂ (ZIN), prosternal process, ventral view. 
<emphasis id="B921EB625560FF8C565ABC7CFBC4F8F4" bold="true" box="[1130,1150,1893,1919]" pageId="10" pageNumber="11">B</emphasis>
. Ditto, ventral view. 
<emphasis id="B921EB625560FF8C528DBC91FF68F829" bold="true" box="[189,210,1928,1954]" pageId="10" pageNumber="11">C</emphasis>
. Ditto, dorso-lateral view. 
<emphasis id="B921EB625560FF8C5021BC91FD9CF829" bold="true" box="[529,550,1928,1954]" pageId="10" pageNumber="11">D</emphasis>
. 
<taxonomicName id="4C554CF35560FF8C5005BC90FCD5F829" authority="(Semenov, 1893)" baseAuthorityName="Semenov" baseAuthorityYear="1893" box="[565,879,1928,1954]" class="Insecta" family="Tenebrionidae" genus="Homopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="grumi">
<emphasis id="B921EB625560FF8C5005BC90FD20F829" box="[565,666,1929,1954]" italics="true" pageId="10" pageNumber="11">P. grumi</emphasis>
(
<bibRefCitation id="EFC44A815560FF8C509BBC91FCDFF829" author="Semenov A. P." box="[683,869,1928,1954]" pageId="10" pageNumber="11" pagination="1892 - 1893" refId="ref33186" refString="Semenov A. P. 1893. Symbolae ad cognitionem pimeliidarum. I - III. Horae Societatis Entomologicae Rossicae 27 [1892 - 1893]: 249 - 264." type="journal article" year="1893">Semenov, 1893</bibRefCitation>
)
</taxonomicName>
, ♂, paralectotype (ZIN), prosternal process, ventral view. 
<emphasis id="B921EB625560FF8C536ABCB2FED4F84E" bold="true" box="[346,366,1963,1989]" pageId="10" pageNumber="11">E</emphasis>
. Ditto, lateral view. 
<emphasis id="B921EB625560FF8C506CBCB2FDD4F84E" bold="true" box="[604,622,1963,1989]" pageId="10" pageNumber="11">F</emphasis>
. Ditto, dorso-lateral view. 
<emphasis id="B921EB625560FF8C5197BCB2FC7FF84E" bold="true" box="[935,965,1963,1989]" pageId="10" pageNumber="11">G.</emphasis>
Ditto, habitus, dorsal view. 
<emphasis id="B921EB625560FF8C573DBCB2FA9EF84E" bold="true" box="[1293,1316,1963,1989]" pageId="10" pageNumber="11">H</emphasis>
. Ditto, ventral view. 
<emphasis id="B921EB625560FF8C536CBCD7FEDDF863" bold="true" box="[348,359,1998,2024]" pageId="10" pageNumber="11">I</emphasis>
. 
<taxonomicName id="4C554CF35560FF8C5346BCD6FCDEF863" authority="Fischer von Waldheim, 1820" authorityName="Fischer von Waldheim" authorityYear="1820" box="[374,868,1998,2025]" class="Insecta" family="Tenebrionidae" genus="Platyope" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="granulata">
<emphasis id="B921EB625560FF8C5346BCD6FDB2F863" box="[374,520,1998,2024]" italics="true" pageId="10" pageNumber="11">P. granulata</emphasis>
<bibRefCitation id="EFC44A815560FF8C5020BCD7FCDEF863" author="Fischer von Waldheim G." box="[528,868,1998,2025]" pageId="10" pageNumber="11" refId="ref30454" refString="Fischer von Waldheim G. 1820. Entomographie de la Russie (Entomographia Imperii Rossici). Auctoritate Societatis Caesareae Mosquensis naturae scrutatorum collecta et in lucem edita. Vol. 1. A. Semen, Moscow." type="book" year="1820">Fischer von Waldheim, 1820</bibRefCitation>
</taxonomicName>
, ♂ (ZIN), habitus, dorsal view.
</paragraph>
</caption>
<paragraph id="8BEA37705561FF8D52DABA10FE25FEA9" blockId="11.[188,1399,264,397]" box="[234,415,264,291]" pageId="11" pageNumber="12">
<emphasis id="B921EB625561FF8D52DABA10FE25FEA9" bold="true" box="[234,415,264,291]" pageId="11" pageNumber="12">Male genitalia</emphasis>
</paragraph>
<paragraph id="8BEA37705561FF8D528DBA35FD20FE06" blockId="11.[188,1399,264,397]" pageId="11" pageNumber="12">
Inner sternite VIII (
<figureCitation id="136E2BF55561FF8D53AFBA35FDB3FECD" box="[415,521,300,326]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[197,1398,757,1854]" captionTargetId="figure-148@35.[189,1398,757,1854]" captionTargetPageId="35" captionText="Fig. 14. Apatopsis grombczewskii Semenov, 1891 (ZIN), genitalia. A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419467" httpUri="https://zenodo.org/record/6419467/files/figure.png" pageId="11" pageNumber="12">Fig. 14H</figureCitation>
) weakly sclerotized on lateral margins and membranous in middle, densely covered with short setae; setae longer along posterior margin; anterior margin arcuately emarginated; gland absent (at least was not observed).
</paragraph>
<paragraph id="8BEA37705561FF8D528DBAAAFDF2FE7A" blockId="11.[189,1399,435,497]" pageId="11" pageNumber="12">
Rods of spiculum gastrale (
<figureCitation id="136E2BF55561FF8D53CCBAAAFDDCFE45" box="[508,614,435,462]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[197,1398,757,1854]" captionTargetId="figure-148@35.[189,1398,757,1854]" captionTargetPageId="35" captionText="Fig. 14. Apatopsis grombczewskii Semenov, 1891 (ZIN), genitalia. A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419467" httpUri="https://zenodo.org/record/6419467/files/figure.png" pageId="11" pageNumber="12">Fig. 14G</figureCitation>
) arcuately connected at apex; derivatives of inner sternite IX ladleshaped covered with sparse setae.
</paragraph>
<paragraph id="8BEA37705561FF8D528DB901FE6FFD69" blockId="11.[189,1399,536,739]" pageId="11" pageNumber="12">
Basal piece of tegmen (
<figureCitation id="136E2BF55561FF8D53E4B901FDD8FDB9" box="[468,610,536,562]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[197,1398,757,1854]" captionTargetId="figure-148@35.[189,1398,757,1854]" captionTargetPageId="35" captionText="Fig. 14. Apatopsis grombczewskii Semenov, 1891 (ZIN), genitalia. A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419467" httpUri="https://zenodo.org/record/6419467/files/figure.png" pageId="11" pageNumber="12">Fig. 14A–C</figureCitation>
) 1.2–1.4× as long as apical piece, rounded at base, widest in basal third. Apical piece bare, rounded at apex, strongly curved (
<figureCitation id="136E2BF55561FF8D5141B922FC61FDDE" box="[881,987,571,597]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[197,1398,757,1854]" captionTargetId="figure-148@35.[189,1398,757,1854]" captionTargetPageId="35" captionText="Fig. 14. Apatopsis grombczewskii Semenov, 1891 (ZIN), genitalia. A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419467" httpUri="https://zenodo.org/record/6419467/files/figure.png" pageId="11" pageNumber="12">Fig. 14C</figureCitation>
), ventral apophyses of apical piece very short (
<figureCitation id="136E2BF55561FF8D5379B947FE03FDF2" box="[329,441,606,633]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[197,1398,757,1854]" captionTargetId="figure-148@35.[189,1398,757,1854]" captionTargetPageId="35" captionText="Fig. 14. Apatopsis grombczewskii Semenov, 1891 (ZIN), genitalia. A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419467" httpUri="https://zenodo.org/record/6419467/files/figure.png" pageId="11" pageNumber="12">Fig. 14A</figureCitation>
), projecting and elevated, dorsal apophyses long (
<figureCitation id="136E2BF55561FF8D5629B947FB3DFDF2" box="[1049,1159,606,633]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[197,1398,757,1854]" captionTargetId="figure-148@35.[189,1398,757,1854]" captionTargetPageId="35" captionText="Fig. 14. Apatopsis grombczewskii Semenov, 1891 (ZIN), genitalia. A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419467" httpUri="https://zenodo.org/record/6419467/files/figure.png" pageId="11" pageNumber="12">Fig. 14B</figureCitation>
), triangular, acutely angulate at apex, not merged; median lobe (
<figureCitation id="136E2BF55561FF8D508EB99BFCF2FD17" box="[702,840,642,668]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[197,1398,757,1854]" captionTargetId="figure-148@35.[189,1398,757,1854]" captionTargetPageId="35" captionText="Fig. 14. Apatopsis grombczewskii Semenov, 1891 (ZIN), genitalia. A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419467" httpUri="https://zenodo.org/record/6419467/files/figure.png" pageId="11" pageNumber="12">Fig. 14D–F</figureCitation>
) strongly curved, bifurcated at base, membrane between baculi weakly sclerotized in basal third, baсuli at apex dorsally rolled into tube, not merged, apex narrowly rounded.
</paragraph>
<paragraph id="8BEA37705561FF8D52DAB813FE01FCA8" blockId="11.[189,1398,777,875]" box="[234,443,777,804]" pageId="11" pageNumber="12">
<emphasis id="B921EB625561FF8D52DAB813FE01FCA8" bold="true" box="[234,443,777,804]" pageId="11" pageNumber="12">Female genitalia</emphasis>
</paragraph>
<paragraph id="8BEA37705561FF8D528DB834FB27FCE0" blockId="11.[189,1398,777,875]" pageId="11" pageNumber="12">
Spiculum ventrale (
<figureCitation id="136E2BF55561FF8D539AB834FDB6FCCC" box="[426,524,813,839]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[197,1398,757,1854]" captionTargetId="figure-148@35.[189,1398,757,1854]" captionTargetPageId="35" captionText="Fig. 14. Apatopsis grombczewskii Semenov, 1891 (ZIN), genitalia. A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419467" httpUri="https://zenodo.org/record/6419467/files/figure.png" pageId="11" pageNumber="12">Fig. 14I</figureCitation>
) very short and thickened, with short common stem, rods connected by membranous plate at apex; apophyses of sternite VIII long, acutely angulate at apex.
</paragraph>
<paragraph id="8BEA37705561FF8D528DB888FAFEFBB3" blockId="11.[189,1400,913,1080]" pageId="11" pageNumber="12">
Ovipositor (
<figureCitation id="136E2BF55561FF8D537AB888FE76FC27" box="[330,460,913,940]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[197,1398,757,1854]" captionTargetId="figure-148@35.[189,1398,757,1854]" captionTargetPageId="35" captionText="Fig. 14. Apatopsis grombczewskii Semenov, 1891 (ZIN), genitalia. A. Tegmen of aedeagus, ventrally. B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII. I. ♀, inner sternite VIII and spiculum ventrale. J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419467" httpUri="https://zenodo.org/record/6419467/files/figure.png" pageId="11" pageNumber="12">Fig. 14J–L</figureCitation>
) weakly sclerotized, very short, transverse. Paraproct short, with narrow baculi; coxite lobes not merged; lobe IV transformed into weakly sclerotized and very densely pubescent pair plate, rounded at apex; lobe III forms additional elongate apical process, similar to lobe IV; lobes I–II with shorter setae; lobe I with oblique, narrow baculi, widened at apices. Proctiger wide, with straight apical margin and densely pubescent middle, baculi of proctiger strongly widened along all length.
</paragraph>
<caption id="DF2A67F85561FF8D528DBC91FEB3F863" ID-DOI="http://doi.org/10.5281/zenodo.6419441" ID-Zenodo-Dep="6419441" httpUri="https://zenodo.org/record/6419441/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1928,1954]" targetBox="[273,1325,1150,1881]" targetPageId="11">
<paragraph id="8BEA37705561FF8D528DBC91FEB3F863" blockId="11.[186,1400,1928,2024]" pageId="11" pageNumber="12">
<emphasis id="B921EB625561FF8D528DBC91FEB2F829" bold="true" box="[189,264,1928,1954]" pageId="11" pageNumber="12">Fig. 6.</emphasis>
Species of the genus 
<taxonomicName id="4C554CF35561FF8D53C8BC91FC0EF829" authority="Fischer von Waldheim, 1820" authorityName="Fischer von Waldheim" authorityYear="1820" box="[504,948,1928,1954]" class="Insecta" family="Tenebrionidae" genus="Platyope" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B921EB625561FF8D53C8BC91FDDBF829" box="[504,609,1928,1954]" italics="true" pageId="11" pageNumber="12">Platyope</emphasis>
<bibRefCitation id="EFC44A815561FF8D5058BC91FC0EF829" author="Fischer von Waldheim G." box="[616,948,1928,1954]" pageId="11" pageNumber="12" refId="ref30454" refString="Fischer von Waldheim G. 1820. Entomographie de la Russie (Entomographia Imperii Rossici). Auctoritate Societatis Caesareae Mosquensis naturae scrutatorum collecta et in lucem edita. Vol. 1. A. Semen, Moscow." type="book" year="1820">Fischer von Waldheim, 1820</bibRefCitation>
</taxonomicName>
, habitus, dorsal view. 
<emphasis id="B921EB625561FF8D569BBC91FB7AF829" bold="true" box="[1195,1216,1928,1954]" pageId="11" pageNumber="12">A</emphasis>
. 
<taxonomicName id="4C554CF35561FF8D56FFBC90FD16F84E" authority="proctoleuca Fischer von Waldheim, 1820" authorityName="proctoleuca Fischer von Waldheim" authorityYear="1820" class="Insecta" family="Tenebrionidae" genus="Platyope" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="proctoleuca" subSpecies="proctoleuca">
<emphasis id="B921EB625561FF8D56FFBC90FEFDF84E" italics="true" pageId="11" pageNumber="12">P. proctoleuca proctoleuca</emphasis>
<bibRefCitation id="EFC44A815561FF8D5363BCB2FD16F84E" author="Fischer von Waldheim G." box="[339,684,1963,1990]" pageId="11" pageNumber="12" refId="ref30454" refString="Fischer von Waldheim G. 1820. Entomographie de la Russie (Entomographia Imperii Rossici). Auctoritate Societatis Caesareae Mosquensis naturae scrutatorum collecta et in lucem edita. Vol. 1. A. Semen, Moscow." type="book" year="1820">Fischer von Waldheim, 1820</bibRefCitation>
</taxonomicName>
, ♂ (ZIN). 
<emphasis id="B921EB625561FF8D5100BCB2FCF8F84E" bold="true" box="[816,834,1963,1989]" pageId="11" pageNumber="12">B</emphasis>
. 
<taxonomicName id="4C554CF35561FF8D5162BCB5FC59F84E" box="[850,995,1963,1989]" class="Insecta" family="Tenebrionidae" genus="Platyope" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ordossica">
<emphasis id="B921EB625561FF8D5162BCB5FC59F84E" box="[850,995,1963,1989]" italics="true" pageId="11" pageNumber="12">P. ordossica</emphasis>
</taxonomicName>
<bibRefCitation id="EFC44A815561FF8D51DFBCB2FAEEF84E" author="Semenov-Tjan-Shansky A. P." box="[1007,1364,1963,1989]" pageId="11" pageNumber="12" pagination="1907 - 1908" refId="ref33261" refString="Semenov-Tjan-Shansky A. P. 1907. Synopsis generum tribus Platyopinorum (Coleoptera, Tenebrionidae Pimeliini). Horae Societatis Entomologicae Rossicae 38 [1907 - 1908]: 175 - 184." type="journal article" year="1907">Semenov-Tjan-Shansky, 1907</bibRefCitation>
, ♂ (ZIN).
</paragraph>
</caption>
<paragraph id="8BEA37705566FF8A52DABA10FDC8FEA8" blockId="12.[186,1399,264,432]" box="[234,626,264,291]" pageId="12" pageNumber="13">
<emphasis id="B921EB625566FF8A52DABA10FE54FEA8" bold="true" box="[234,494,264,291]" pageId="12" pageNumber="13">Female genital ducts</emphasis>
(
<figureCitation id="136E2BF55566FF8A53CFBA11FDD0FEA8" box="[511,618,264,291]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[208,1379,488,1777]" captionTargetId="figure-18@46.[200,1387,478,1783]" captionTargetPageId="46" captionText="Fig. 23. Female genital ducts of Pimeliini. A. Apatopsis grombczewskii Semenov, 1891 (ZIN). B. Habrobates vernalis Semenov, 1903 (ZIN). C. Dietomorpha pardalis Kühnelt, 1957 (ZIN). D. Przewalskia dilatata (Reitter, 1887) (ZIN). E. Mantichorula semenowi Reitter, 1889 (ZIN). F. Platyope leucogramma (Pallas, 1773) (ZIN). G. Platyope grumi (Semenov, 1893), paralectotype (ZIN). H. Earophanta serrata (Semenov, 1893) (ZIN). Not to scale. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.6419493" httpUri="https://zenodo.org/record/6419493/files/figure.png" pageId="12" pageNumber="13">Fig. 23A</figureCitation>
)
</paragraph>
<paragraph id="8BEA37705566FF8A528DBA35FBDAFE24" blockId="12.[186,1399,264,432]" pageId="12" pageNumber="13">Vagina sacciform, narrower before oviduct and after spermatheca, apical part of vagina elastically curved. Spermatheca consists of six tubes: two tubes flow into vagina independently, other four tubes combined into bundle with common base; two tubes (one independent, one in bundle) bifurcated. Accessory gland with four constrictions, one (as narrow tube) at base and three others in middle.</paragraph>
</subSubSection>
<subSubSection id="C34F64FB5566FF8A528DBACFFDB3FDAB" box="[189,521,470,544]" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8BEA37705566FF8A528DBACFFEE2FE7B" blockId="12.[189,344,470,496]" box="[189,344,470,496]" pageId="12" pageNumber="13">
<heading id="D0A2801C5566FF8A528DBACFFEE2FE7B" bold="true" box="[189,344,470,496]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B921EB625566FF8A528DBACFFEE2FE7B" bold="true" box="[189,344,470,496]" pageId="12" pageNumber="13">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA37705566FF8A528DB91CFDB3FDAB" blockId="12.[189,521,517,544]" box="[189,521,517,544]" pageId="12" pageNumber="13">
<collectingCountry id="F34277E05566FF8A528DB91CFEB9FD94" box="[189,259,517,543]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
(Taklamakan Desert).
</paragraph>
</subSubSection>
</treatment>
</document>