treatments-xml/data/03/F5/FA/03F5FA39BC7DFFD9B1D0FDCF30F9FC6E.xml

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<mods:title>A revision of Canarium L. (Burseraceae) in Madagascar</mods:title>
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<mods:namePart>Daly, Douglas C.</mods:namePart>
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<mods:namePart>Raharimampionona, Jeannie</mods:namePart>
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<mods:namePart>Federman, Sarah</mods:namePart>
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<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[135,680,597,624]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="37" pageNumber="314" phylum="Tracheophyta" rank="species" species="lamianum" status="sp. nov.">
<emphasis bold="true" box="[135,364,597,624]" italics="true" pageId="37" pageNumber="314">Canarium lamianum</emphasis>
Daly, Raharim. &amp; Federman
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(
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; 
<figureCitation box="[482,513,630,656]" captionStart="FIG" captionStartId="36.[132,143,1962,1980]" captionTargetBox="[208,1379,215,1865]" captionTargetId="figure-19@36.[202,1386,207,1870]" captionTargetPageId="36" captionText="FIG. 18. — Canarium lamianum Daly, Raharim. &amp; Federman, sp. nov.: A, flowering branchlet; B, staminate flower with bracteole; C, adaxial view of petal; D, sta- mens and ovariodisk; E, portion of infructescence; F, fruiting branchlet; G, leaf with detail of stipules (G’) and leaflet margin (G’’); A-D, Daly et al. 13065 (NY); E, Treuer et al. 44 (NY); F, McPherson &amp; van der Werff 16378 (MO); G, Antilahimena et al. 663 (NY). Scale bars: A, E-G, 2 cm; B, 3 mm; C, D, 2 mm; G’, G’’, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5208937" httpUri="https://zenodo.org/record/5208937/files/figure.png" pageId="37" pageNumber="314">18</figureCitation>
).
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Small to large trees, leaves (2)3-4-jugate, leaflet secondary veins in 6-10 pairs and on adaxial side very broadly prominulous to flat, the spacing (slightly) decreasing toward the apex, and fruits 
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long, (oblong-) ovoid to (oblong-)obovoid, with small raised lenticels; distinguished from
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C. obtusifolium 
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<emphasis italics="true" pageId="37" pageNumber="314">), and the secondary veins not wavy and concolorous (vs wavy and discolorous), and branching close to the flat margin (vs not branching close to the flat margin).</emphasis>
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TYPUS. — 
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Forêt 
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<geoCoordinate box="[657,771,1232,1256]" degrees="15" direction="south" minutes="54" orientation="latitude" pageId="37" pageNumber="314" precision="15" seconds="03" value="-15.900833">15°54’03”S</geoCoordinate>
, 
<geoCoordinate box="[131,247,1258,1282]" degrees="49" direction="east" minutes="27" orientation="longitude" pageId="37" pageNumber="314" precision="15" seconds="44" value="49.462223">49°27’44”E</geoCoordinate>
, 
<date box="[257,382,1258,1282]" pageId="37" pageNumber="314" value="2007-07-23">
<collectingDate box="[257,382,1258,1282]" pageId="37" pageNumber="314" value="2007-07-23">23.VII.2007</collectingDate>
</date>
, 
<emphasis box="[391,647,1258,1282]" italics="true" pageId="37" pageNumber="314">
P. 
<collectorName box="[412,541,1258,1282]" pageId="37" pageNumber="314">Antilahimena</collectorName>
et al. 5631
</emphasis>
(
<collectionCode box="[660,706,1258,1282]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="37" pageNumber="314" type="Herbarium">MO</collectionCode>
, 
<collectionCode box="[715,750,1259,1282]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="37" pageNumber="314" type="Herbarium">NY</collectionCode>
, P, 
<collectionCode box="[131,184,1285,1309]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="37" pageNumber="314" type="Herbarium">TAN</collectionCode>
, 
<collectionCode box="[193,242,1285,1309]" country="Madagascar" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="37" pageNumber="314" type="Herbarium">TEF</collectionCode>
); 
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC7DFFD9B1D0FDCF30F9FC6E:8E831DF4BC7DFFDBB053FA9C30B0FABC" box="[260,442,1285,1309]" country="United States of America" county="Reserve Naturelle Integrale Betampona" latitude="-17.904999" longLatPrecision="21" longitude="49.206112" municipality="Atsinanana" name="Masoala Peninsula" pageId="37" pageNumber="314" stateProvince="Fianarantsoa">Masoala Peninsula</location>
, 
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC7DFFD9B1D0FDCF30F9FC6E:8E831DF4BC7DFFDBB092FA9C3335FABC" box="[453,575,1285,1309]" country="United States of America" county="Reserve Naturelle Integrale Betampona" latitude="-17.904999" longLatPrecision="21" longitude="49.206112" municipality="Atsinanana" name="Mahalevona" pageId="37" pageNumber="314" stateProvince="Fianarantsoa">Mahalevona</location>
, 
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC7DFFD9B1D0FDCF30F9FC6E:8E831DF4BC7DFFDBB31EFA9C33C7FABD" box="[585,717,1285,1308]" country="United States of America" county="Reserve Naturelle Integrale Betampona" latitude="-17.904999" longLatPrecision="21" longitude="49.206112" municipality="Atsinanana" name="Maroantsetra" pageId="37" pageNumber="314" stateProvince="Fianarantsoa">Maroantsetra</location>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.93" metricValueMax="10.11" metricValueMin="3.75" pageId="37" pageNumber="314" unit="m" value="693.0" valueMax="1011.0" valueMin="375.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.93" metricValueMax="10.11" metricValueMin="3.75" pageId="37" pageNumber="314" unit="m" value="693.0" valueMax="1011.0" valueMin="375.0">375- 1011 m</elevation>
</quantity>
, 
<geoCoordinate box="[216,328,1311,1335]" degrees="15" direction="south" minutes="17" orientation="latitude" pageId="37" pageNumber="314" precision="15" seconds="22" value="-15.289445">15°17’22”S</geoCoordinate>
, 
<geoCoordinate box="[336,450,1311,1335]" degrees="50" direction="east" minutes="00" orientation="longitude" pageId="37" pageNumber="314" precision="15" seconds="58" value="50.01611">50°00’58”E</geoCoordinate>
, 
<date box="[459,563,1311,1335]" pageId="37" pageNumber="314" value="1997-10-08">
<collectingDate box="[459,563,1311,1335]" pageId="37" pageNumber="314" value="1997-10-08">08.X.1997</collectingDate>
</date>
, 
<emphasis italics="true" pageId="37" pageNumber="314">
<collectorName box="[570,702,1311,1335]" pageId="37" pageNumber="314">C. Birkinshaw</collectorName>
, 
<collectorName pageId="37" pageNumber="314">H. Ralimanana</collectorName>
&amp; 
<collectorName box="[254,411,1338,1361]" pageId="37" pageNumber="314">R. Ranaivojaona</collectorName>
493
</emphasis>
(
<collectionCode box="[468,514,1338,1362]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="37" pageNumber="314" type="Herbarium">MO</collectionCode>
, 
<collectionCode box="[523,558,1338,1361]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="37" pageNumber="314" type="Herbarium">NY</collectionCode>
, P, 
<collectionCode box="[588,642,1338,1362]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="37" pageNumber="314" type="Herbarium">TAN</collectionCode>
); 
<collectingMunicipality box="[659,771,1338,1362]" pageId="37" pageNumber="314">Atsinanana</collectingMunicipality>
, 
<collectingCounty box="[132,520,1364,1388]" pageId="37" pageNumber="314">Réserve Naturelle Intégrale Betampona</collectingCounty>
, 
<quantity box="[530,644,1364,1388]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="37" pageNumber="314" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">
<elevation box="[530,644,1364,1388]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="37" pageNumber="314" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">400-600 m</elevation>
</quantity>
, 
<geoCoordinate box="[656,770,1364,1388]" degrees="17" direction="south" minutes="54" orientation="latitude" pageId="37" pageNumber="314" precision="15" seconds="18" value="-17.904999">17°54’18”S</geoCoordinate>
, 
<geoCoordinate box="[133,249,1391,1415]" degrees="49" direction="east" minutes="12" orientation="longitude" pageId="37" pageNumber="314" precision="15" seconds="22" value="49.206112">49°12’22”E</geoCoordinate>
, 
<date box="[258,373,1391,1415]" pageId="37" pageNumber="314" value="2008-03-07">
<collectingDate box="[258,373,1391,1415]" pageId="37" pageNumber="314" value="2008-03-07">07.III.2008</collectingDate>
</date>
, 
<emphasis box="[379,712,1390,1415]" italics="true" pageId="37" pageNumber="314">
<collectorName box="[379,496,1390,1414]" pageId="37" pageNumber="314">S. Federman</collectorName>
, 
<collectorName box="[504,586,1391,1414]" pageId="37" pageNumber="314">T.Treuer</collectorName>
&amp; 
<collectorName box="[614,682,1390,1414]" pageId="37" pageNumber="314">Sylvain</collectorName>
40
</emphasis>
(
<collectionCode box="[725,764,1391,1415]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="37" pageNumber="314" type="Herbarium">NY</collectionCode>
)
</materialsCitation>
; 
<materialsCitation collectingDate="2008-04-07" collectionCode="NY" collectorName="Treuer &amp; S. Federman &amp; Sylvain" country="United States of America" elevation="500" latitude="-17.903055" location="Atsinanana" longLatPrecision="21" longitude="49.218334" municipality="Reserve Naturelle Integrale Betampona" pageId="37" pageNumber="314" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC7DFFD9B1D0FDCF30F9FC6E:8E831DF4BC7DFFDBB1D4FA1031F9FA00" box="[131,243,1417,1441]" country="United States of America" latitude="-17.903055" longLatPrecision="21" longitude="49.218334" municipality="Reserve Naturelle Integrale Betampona" name="Atsinanana" pageId="37" pageNumber="314" stateProvince="Fianarantsoa">Atsinanana</location>
, 
<collectingMunicipality box="[254,643,1417,1441]" pageId="37" pageNumber="314">Réserve Naturelle Intégrale Betampona</collectingMunicipality>
, 
<quantity box="[655,771,1417,1441]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="37" pageNumber="314" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">
<elevation box="[655,771,1417,1441]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="37" pageNumber="314" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">400-600 m</elevation>
</quantity>
, 
<geoCoordinate box="[134,251,1443,1467]" degrees="17" direction="south" minutes="54" orientation="latitude" pageId="37" pageNumber="314" precision="15" seconds="11" value="-17.903055">17°54’11”S</geoCoordinate>
, 
<geoCoordinate box="[263,383,1443,1467]" degrees="49" direction="east" minutes="13" orientation="longitude" pageId="37" pageNumber="314" precision="15" seconds="06" value="49.218334">49°13’06”E</geoCoordinate>
, 
<date box="[395,512,1444,1468]" pageId="37" pageNumber="314" value="2008-04-07">
<collectingDate box="[395,512,1444,1468]" pageId="37" pageNumber="314" value="2008-04-07">07.IV.2008</collectingDate>
</date>
, 
<emphasis italics="true" pageId="37" pageNumber="314">
T. 
<collectorName box="[551,614,1444,1467]" pageId="37" pageNumber="314">Treuer</collectorName>
, 
<collectorName box="[626,748,1443,1467]" pageId="37" pageNumber="314">S. Federman</collectorName>
&amp; 
<collectorName box="[134,203,1470,1494]" pageId="37" pageNumber="314">Sylvain</collectorName>
84
</emphasis>
(
<collectionCode box="[246,286,1470,1494]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="37" pageNumber="314" type="Herbarium">NY</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="314" type="distribution">
<paragraph blockId="37.[130,780,687,1874]" pageId="37" pageNumber="314">
DISTRIBUTION AND ECOLOGY. — 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[477,683,1517,1541]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="37" pageNumber="314" phylum="Tracheophyta" rank="species" species="lamianum" status="sp. nov.">
<emphasis box="[477,683,1517,1541]" italics="true" pageId="37" pageNumber="314">Canarium lamianum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[698,776,1518,1541]" pageId="37" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
is one of the most broadly distributed, variable, and ecologically versatile species of the genus in 
<collectingCountry box="[449,566,1570,1594]" name="Madagascar" pageId="37" pageNumber="314">Madagascar</collectingCountry>
. It has been recorded in many of the moister parts of the country except the southeast. It tends to occur in dense forest on slopes and dense lowland forest, but it ranges between 
<quantity box="[357,458,1650,1674]" metricMagnitude="2" metricUnit="m" metricValue="7.525" metricValueMax="15.0" metricValueMin="0.05" pageId="37" pageNumber="314" unit="m" value="752.5" valueMax="1500.0" valueMin="5.0">5-1500 m</quantity>
elevation. Flowering Jan.-Mar., fruiting all year.
</paragraph>
<paragraph blockId="37.[130,780,687,1874]" pageId="37" pageNumber="314">
COMMON NAME AND USES. — Ramy. Used for caulking canoes, illumination, and in religious (
<emphasis box="[432,499,1750,1774]" italics="true" pageId="37" pageNumber="314">tromba</emphasis>
) ceremonies.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="314" type="etymology">
<paragraph blockId="37.[130,780,687,1874]" pageId="37" pageNumber="314">
ETYMOLOGY. — We name this species in honor of H. J. Lam, who contributed so much to our understanding of paleotropical 
<taxonomicName box="[134,251,1851,1874]" class="Magnoliopsida" family="Burseraceae" kingdom="Plantae" order="Sapindales" pageId="37" pageNumber="314" phylum="Tracheophyta" rank="family">Burseraceae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="39" lastPageNumber="316" pageId="37" pageNumber="314" type="description">
<paragraph blockId="37.[132,775,1906,2028]" box="[132,277,1906,1932]" pageId="37" pageNumber="314">DESCRIPTION</paragraph>
<paragraph blockId="37.[132,775,1906,2028]" lastBlockId="39.[130,777,215,819]" lastPageId="39" lastPageNumber="316" pageId="37" pageNumber="314">
Trees, reproductive size 
<quantity box="[372,447,1937,1964]" metricMagnitude="1" metricUnit="m" metricValue="1.8" metricValueMax="3.0" metricValueMin="0.6" pageId="37" pageNumber="314" unit="m" value="18.0" valueMax="30.0" valueMin="6.0">6-30 m</quantity>
tall 
<emphasis box="[489,505,1938,1963]" italics="true" pageId="37" pageNumber="314">×</emphasis>
<quantity box="[507,606,1937,1964]" metricMagnitude="-1" metricUnit="m" metricValue="3.6" metricValueMax="5.7" metricValueMin="1.5" pageId="37" pageNumber="314" unit="cm" value="36.0" valueMax="57.0" valueMin="15.0">15-57 cm</quantity>
diam, with simple or branched plank buttresses 
<quantity box="[469,583,1969,1996]" metricMagnitude="0" metricUnit="m" metricValue="1.16" metricValueMax="2.0" metricValueMin="0.32" pageId="37" pageNumber="314" unit="cm" value="116.0" valueMax="200.0" valueMin="32.0">32-200 cm</quantity>
tall and extending 
<quantity box="[132,197,2001,2028]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="37" pageNumber="314" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">2-4 m</quantity>
from trunk. Outer bark (brownish-)gray, thin or thick, scalloped,sometimes with hoop marks, sometime finely fissured, with dense to sparse raised lenticels, shed as irregular plates; inner bark pale red, light orange, or reddish orange. Resin clear when fresh. Leaves 
<quantity box="[1006,1125,311,338]" metricMagnitude="-1" metricUnit="m" metricValue="2.375" metricValueMax="3.3" metricValueMin="1.45" pageId="37" pageNumber="314" unit="cm" value="23.75" valueMax="33.0" valueMin="14.5">14.5-33 cm</quantity>
long, (2) 3-4-jugate; petiole 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="6.5" metricValueMin="2.5" pageId="37" pageNumber="314" unit="cm" value="4.5" valueMax="6.5" valueMin="2.5">2.5- 6.5 cm</quantity>
, petiole and rachis usually with scattered thick, appressed golden hairs to 
<quantity box="[965,1060,374,401]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="314" unit="mm" value="0.05">0.05 mm</quantity>
long, also with scattered (rarely sparse) capitate glands; stipules 4-13 (19) mm from petiole insertion, 
<quantity box="[813,899,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="37" pageNumber="314" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3-7 mm</quantity>
long, orbicular to ovate, base constricted, fleshy, usually persistent, with dense ascending to appressed thick golden hairs to 
<quantity box="[899,997,502,529]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="314" unit="mm" value="0.15">0.15 mm</quantity>
long (sometimes sparse, or on older leaflets subglabrous); the scar 
<quantity box="[1041,1165,534,561]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="37" pageNumber="314" unit="mm" value="1.45" valueMax="1.6" valueMin="1.3">1.3-1.6 mm</quantity>
long, raised; basal and other lateral petiolules 
<quantity box="[984,1083,566,593]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="12.0" metricValueMin="4.0" pageId="37" pageNumber="314" unit="mm" value="8.0" valueMax="12.0" valueMin="4.0">4-12 mm</quantity>
long, terminal one 
<quantity box="[1285,1396,566,593]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="3.0" metricValueMin="1.3" pageId="37" pageNumber="314" unit="mm" value="21.5" valueMax="30.0" valueMin="13.0">13-30 mm</quantity>
long, petiolules slightly canaliculate, lateral pulvinuli conspicuous; basal leaflets 2.7-11.1 
<emphasis box="[1034,1050,631,656]" italics="true" pageId="37" pageNumber="314">×</emphasis>
<quantity box="[1051,1166,630,656]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="4.1" metricValueMin="1.7" pageId="37" pageNumber="314" unit="cm" value="2.9" valueMax="4.1" valueMin="1.7">1.7-4.1 cm</quantity>
, (ob)ovate to broadly elliptic or rarely lanceolate, other laterals 4 
<emphasis box="[1183,1199,662,687]" italics="true" pageId="37" pageNumber="314">×</emphasis>
13.5 
<emphasis box="[1260,1276,662,687]" italics="true" pageId="37" pageNumber="314">×</emphasis>
1.8-4.0 cm, (oblong) elliptic to slightly oblong-obovate or oblong-oblanceolate, terminal one 4.5-11.5 
<emphasis box="[1053,1069,726,751]" italics="true" pageId="37" pageNumber="314">×</emphasis>
2-35, obovate or sometimes elliptic; leaflet apex abruptly and broadly short-acuminate, the acumen 2-7 (11) mm long (very rarely some leaflets with rounded apex or the apex tip sometimes retuse); lateral leaflet base usually subsymmetrical, obtuse to truncate to acute, sometimes short-attenuate; margin flat; leaflets coriaceous, drying dark brown, dull; secondary vein framework brochidodromous but secondaries branching near margin, secondaries in 6-10 pairs, slightly arcuate, concolorous, spacing slightly decreasing toward apex and base, often irregular, angle acute to subperpendicular, increasing toward apex and decreasing toward base or uniform, perpendicular epimedial tertiaries sometimes present; intercostal tertiaries random-reticulate with some admedial branching; on abaxial side the midvein broadly prominent, rest nearly flat, the surface subglabrous but often with a few scattered capitate glands and sometimes a few appressed fine hairs to 
<quantity box="[1274,1358,1204,1230]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="314" unit="mm" value="0.1">0.1 mm</quantity>
long; on adaxial side midvein narrowly prominulous but sunk in a groove, secondaries narrowly prominulous, sometimes immersed, rest nearly flat, the surface glabrous. Inflorescences 
<quantity box="[1305,1395,1300,1326]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="1.9" metricValueMin="0.7" pageId="37" pageNumber="314" unit="cm" value="13.0" valueMax="19.0" valueMin="7.0">7-19 cm</quantity>
long, 1.8-3.2 (4) mm diam near base (
<quantity box="[1151,1257,1331,1358]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="37" pageNumber="314" unit="mm" value="3.75" valueMax="5.0" valueMin="2.5">2.5-5 mm</quantity>
diam in fruit), secondary axes to 
<quantity box="[971,1055,1363,1390]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="37" pageNumber="314" unit="mm" value="6.5">6.5 mm</quantity>
long, axes with dense flexuous golden hairs to 
<quantity box="[894,990,1395,1422]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="37" pageNumber="314" unit="mm" value="0.25">0.25 mm</quantity>
long and dense capitate glands, or with sparse flexuous hairs to 
<quantity box="[986,1068,1427,1454]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="314" unit="mm" value="0.1">0.1 mm</quantity>
long and sparse capitate glands (both scattered in fruit); bracts on primary axes of staminate inflorescences 
<quantity box="[885,969,1491,1517]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="37" pageNumber="314" unit="mm" value="3.4">3.4 mm</quantity>
long, very broadly ovate, semi-clasping, fleshy; other bracts 
<quantity box="[947,1076,1522,1549]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="37" pageNumber="314" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5-4.5 mm</quantity>
long, (ob)ovate to lanceolate, often with constricted base, longer than buds; bracteoles 
<quantity box="[1332,1457,1555,1581]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="37" pageNumber="314" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.2-2.7 mm</quantity>
long, lanceolate; bracts mostly deciduous, unknown for pistillate inflorescences; in staminate inflorescences the pedicel 0.5-2 
<emphasis box="[1440,1456,1619,1644]" italics="true" pageId="37" pageNumber="314">×</emphasis>
<quantity box="[814,946,1650,1677]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="37" pageNumber="314" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9-1.1 mm</quantity>
, cylindrical. Staminate flowers (5.4) 6-8.6 (9.5) mm long at anthesis; calyx 3.6-4.5 
<emphasis box="[1173,1189,1683,1708]" italics="true" pageId="37" pageNumber="314">×</emphasis>
4-5.3 (6) mm overall, the lobes 1.2-2 (3) mm long, ovate to deltate, somewhat spreading, abaxial surface with dense flexuous hairs to 
<quantity box="[1269,1355,1746,1772]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="314" unit="mm" value="0.1">0.1 mm</quantity>
long and capitate glands; petals 5.6-8.1 (9) 
<emphasis box="[1163,1179,1779,1804]" italics="true" pageId="37" pageNumber="314">×</emphasis>
<quantity box="[1186,1314,1778,1804]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="3.3" metricValueMin="1.8" pageId="37" pageNumber="314" unit="mm" value="2.55" valueMax="3.3" valueMin="1.8">1.8-3.3 mm</quantity>
, exposed part longer than calyx, (oblong-) ovate to oblong-obovate, sometimes slightly acuminate (usually acute),with an inflexed apiculum (not always evident) 
<quantity box="[976,1116,1873,1900]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="37" pageNumber="314" unit="mm" value="0.375" valueMax="0.4" valueMin="0.35">0.35-0.4 mm</quantity>
long, abaxial surface with dense thick appressed hairs to 
<quantity box="[1059,1141,1905,1932]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="314" unit="mm" value="0.1">0.1 mm</quantity>
long (
<quantity box="[1207,1304,1905,1932]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="314" unit="mm" value="0.15">0.15 mm</quantity>
at apex), these retrorse near base, adaxial surface with dense capitate glands; two series of stamens inserted at two levels near base of ovariodisk, (3.2) 
<quantity box="[869,974,2001,2028]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="5.0" metricValueMin="3.8" pageId="37" pageNumber="314" unit="mm" value="4.4" valueMax="5.0" valueMin="3.8">3.8-5 mm</quantity>
long (
<quantity box="[1039,1123,2001,2028]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="37" pageNumber="314" unit="mm" value="6.6">6.6 mm</quantity>
in 
<emphasis box="[1155,1371,2001,2027]" italics="true" pageId="37" pageNumber="314">Service Forestier 5892</emphasis>
), some- times the two series somewhat unequal, the anthers 1.1-1.7 (2.3) mm long, lanceolate in dorsiventral view, (oblong-)lanceolate in lateral view; ovariodisk 0.9-2 
<emphasis box="[458,474,280,305]" italics="true" pageId="39" pageNumber="316">×</emphasis>
(0.7) 
<quantity box="[534,662,279,306]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="2.2" metricValueMin="0.9" pageId="39" pageNumber="316" unit="mm" value="1.55" valueMax="2.2" valueMin="0.9">0.9-2.2 mm</quantity>
, essentially columnar to slightly (depressed-)obovoid with craggy apex.Pedicel for pistillate flowers 3-3.7 
<emphasis box="[442,458,344,369]" italics="true" pageId="39" pageNumber="316">×</emphasis>
<quantity box="[466,595,343,369]" metricMagnitude="-3" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.4" metricValueMin="2.3" pageId="39" pageNumber="316" unit="mm" value="2.3499999999999996" valueMax="2.4" valueMin="2.3">2.3-2.4 mm</quantity>
, clavate; pistillate flowers (passed and incomplete) 
<emphasis box="[471,488,375,400]" italics="true" pageId="39" pageNumber="316">c.</emphasis>
<quantity box="[494,571,375,401]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="39" pageNumber="316" unit="mm" value="11.0">11 mm</quantity>
long; calyx 6.8-8 
<emphasis box="[759,775,375,400]" italics="true" pageId="39" pageNumber="316">×</emphasis>
<quantity box="[133,261,407,433]" metricMagnitude="-3" metricUnit="m" metricValue="7.55" metricValueMax="7.7" metricValueMin="7.4" pageId="39" pageNumber="316" unit="mm" value="7.55" valueMax="7.7" valueMin="7.4">7.4-7.7 mm</quantity>
overall, cupular, lobes 
<quantity box="[507,616,406,433]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="39" pageNumber="316" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2-2.5 mm</quantity>
long, rounded depressed-deltate; pistil with dense appressed stiff pale hairs to 
<quantity box="[133,217,470,497]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="39" pageNumber="316" unit="mm" value="0.3">0.3 mm</quantity>
long. Fruiting pedicel 2-4.5 (7) 
<emphasis box="[556,572,471,496]" italics="true" pageId="39" pageNumber="316">×</emphasis>
<quantity box="[580,689,470,497]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" metricValueMax="5.0" metricValueMin="3.4" pageId="39" pageNumber="316" unit="mm" value="4.2" valueMax="5.0" valueMin="3.4">3.4-5 mm</quantity>
, clavate, fruiting calyx 
<quantity box="[278,364,502,529]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="39" pageNumber="316" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4-5 mm</quantity>
long, usually slightly cupular, the lobes distinct, 
<quantity box="[222,305,534,561]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="39" pageNumber="316" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1-2 mm</quantity>
long, spreading. Fruits 3.3-4.6 
<emphasis box="[623,639,535,560]" italics="true" pageId="39" pageNumber="316">×</emphasis>
(1.5) 1.9-2.7 (3) cm (to 
<quantity box="[243,315,566,593]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="39" pageNumber="316" unit="cm" value="3.4">3.4 cm</quantity>
diam when fresh), (grayish-) green to (light) brown, usually slightly obovoid, sometimes oblong-elliptic or rarely slightly ovoid, sometimes trigonous, apex rounded, base obtuse to truncate, lenticels on surface dense, small,slightly raised, ferrugineous, surface also with dense ferrugineous capitate glands and dense erect to flexuous golden to whitish hairs to 
<quantity box="[676,773,725,752]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="39" pageNumber="316" unit="mm" value="0.25">0.25 mm</quantity>
long, rarely glabrescent on older fruits. Pyrene (oblong-)ovoid, operculum ¾ its length.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5208939" ID-Zenodo-Dep="5208939" httpUri="https://zenodo.org/record/5208939/files/figure.png" pageId="38" pageNumber="315" startId="38.[132,143,1939,1957]" targetBox="[205,1384,214,1869]" targetPageId="38">
<paragraph blockId="38.[132,1456,1939,2025]" pageId="38" pageNumber="315">
FIG. 19. — 
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[223,623,1939,1957]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="315" phylum="Tracheophyta" rank="species" species="laxiflorum" status="sp. nov.">
<emphasis box="[223,388,1939,1957]" italics="true" pageId="38" pageNumber="315">Canarium laxiflorum</emphasis>
Daly, Raharim. &amp; Federman
</taxonomicName>
, 
<taxonomicNameLabel box="[631,698,1939,1957]" pageId="38" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
: 
<emphasis bold="true" box="[707,720,1939,1957]" pageId="38" pageNumber="315">A</emphasis>
, branchlet with staminate buds, with detail of leaflet margin (
<emphasis bold="true" box="[1227,1244,1939,1957]" pageId="38" pageNumber="315">A’</emphasis>
) and detail of petiole and stipular scars (
<emphasis bold="true" box="[254,276,1962,1980]" pageId="38" pageNumber="315">A’’</emphasis>
); 
<emphasis bold="true" box="[290,303,1962,1980]" pageId="38" pageNumber="315">B</emphasis>
, detail of inflorescence; 
<emphasis bold="true" box="[507,521,1962,1980]" pageId="38" pageNumber="315">C</emphasis>
, lateral view of staminate bud with bracteole (left); 
<emphasis bold="true" box="[947,961,1962,1980]" pageId="38" pageNumber="315">D</emphasis>
, adaxial view of petal; 
<emphasis bold="true" box="[1152,1164,1962,1980]" pageId="38" pageNumber="315">E</emphasis>
, lateral view of staminate bud with one petal and part of calyx removed; 
<emphasis bold="true" box="[445,456,1985,2003]" pageId="38" pageNumber="315">F</emphasis>
, androecium and ovariodisk (left), ovariodisk (right); 
<emphasis bold="true" box="[892,906,1984,2002]" pageId="38" pageNumber="315">G</emphasis>
, ventral and dorsal views of stamen; 
<emphasis bold="true" box="[1218,1252,1984,2003]" pageId="38" pageNumber="315">A -G</emphasis>
, 
<emphasis box="[1263,1456,1985,2003]" italics="true" pageId="38" pageNumber="315">Rabevohitra et al. 3985</emphasis>
[P00501630]. Scale bars: A, 2 cm; A’’, 5 mm; B, 1 cm; C-F, 2 mm; G, 1 mm; A’, not to scale.
</paragraph>
</caption>
<paragraph blockId="39.[131,775,854,975]" box="[132,206,854,880]" pageId="39" pageNumber="316">NOTES</paragraph>
<paragraph blockId="39.[131,775,854,975]" pageId="39" pageNumber="316">
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[132,352,885,911]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="39" pageNumber="316" phylum="Tracheophyta" rank="species" species="lamianum" status="sp. nov.">
<emphasis box="[132,352,885,911]" italics="true" pageId="39" pageNumber="316">Canarium lamianum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[364,444,885,911]" pageId="39" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
is contrasted with 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[637,766,885,911]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="39" pageNumber="316" phylum="Tracheophyta" rank="species" species="arcuatum" status="sp. nov.">
<emphasis box="[637,766,885,911]" italics="true" pageId="39" pageNumber="316">C. arcuatum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[132,216,917,943]" pageId="39" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[223,362,917,943]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="39" pageNumber="316" phylum="Tracheophyta" rank="species" species="laxiflorum" status="sp. nov.">
<emphasis box="[223,362,917,943]" italics="true" pageId="39" pageNumber="316">C. laxiflorum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[376,456,917,943]" pageId="39" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName box="[507,666,917,943]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="39" pageNumber="316" phylum="Tracheophyta" rank="species" species="obtusifolium">
<emphasis box="[507,666,917,943]" italics="true" pageId="39" pageNumber="316">C. obtusifolium</emphasis>
</taxonomicName>
under the discussions for these latter species.
</paragraph>
</subSubSection>
</treatment>
</document>