177 lines
24 KiB
XML
177 lines
24 KiB
XML
<document id="415D23052612C27FC75D3D5AFEFF1CF2" ID-DOI="10.26879/1323" ID-ISSN="1094-8074" ID-Zenodo-Dep="10962296" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712859958310" checkinUser="felipe" docAuthor="Villa, Andrea, Macaluso, Loredana & Mörs, Thomas" docDate="2024" docId="03F52665D05CFF81FED59CB4FDD47309" docLanguage="en" docName="PalaeontolElectron.27.1.a3.pdf" docOrigin="Palaeontologia Electronica (a 3) 27 (1)" docSource="http://dx.doi.org/10.26879/1323" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.3:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="3" docTitle="Mioproteus wezei Estes 1984" docType="treatment" docVersion="3" lastPageNumber="9" masterDocId="FFCC5E1DD054FF88FFA69832FFDE760C" masterDocTitle="Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe" masterLastPageNumber="56" masterPageNumber="1" pageNumber="9" updateTime="1713792755732" updateUser="felipe" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="7EC0E08AD695138CB0DF29B8A36CEE14">Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe</mods:title>
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<mods:name id="6CE916A373B72AC47054953A2217052D" type="personal">
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<mods:namePart id="AA6A670B011FFC147C498BF2E79CA51C">Villa, Andrea</mods:namePart>
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<mods:namePart id="F3CF0EE0CE5F465FFEFC0DBBCBF70076">Macaluso, Loredana</mods:namePart>
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<mods:namePart id="F44F9387D020710C246747664ED6D602">Mörs, Thomas</mods:namePart>
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<mods:typeOfResource id="8F8A70487DA9A7DE67F3CA98931A0968">text</mods:typeOfResource>
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<mods:title id="1F5BC6802504895950E006027FC560B9">Palaeontologia Electronica</mods:title>
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<mods:part id="1F8102C91F69BCA5BB8D5F264F054587">
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<mods:date id="1F919D778D6A6FCC6460CF378F8593AD">2024</mods:date>
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<mods:detail id="E5AE2F5C801E4F4C07DCAC4174FE7EDB" type="series">
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<mods:title id="D714E44542ECC9A9DE64B5F03F13FDD9">a 3</mods:title>
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<mods:detail id="05069ED648F4F546E709ABF6E50BF92A" type="volume">
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<mods:number id="8F507915B92AF58F2921832E612FE5A9">27</mods:number>
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<mods:start id="1FE4827C29F597E85613F3A70E462B54">1</mods:start>
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<mods:url id="5B66B6116453F89F6587ACD71564A0B7">http://dx.doi.org/10.26879/1323</mods:url>
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<mods:classification id="BF81B531311B040E4B379A87DBB4347D">journal article</mods:classification>
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<mods:identifier id="F68F7C497E8FD25B0815DF20081B2469" type="DOI">10.26879/1323</mods:identifier>
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<mods:identifier id="0953E4524205D569EDDA0A09E4D69A80" type="ISSN">1094-8074</mods:identifier>
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<treatment id="03F52665D05CFF81FED59CB4FDD47309" LSID="urn:lsid:plazi:treatment:03F52665D05CFF81FED59CB4FDD47309" httpUri="http://treatment.plazi.org/id/03F52665D05CFF81FED59CB4FDD47309" lastPageId="9" lastPageNumber="9" pageId="8" pageNumber="9">
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<subSubSection id="C346C4F8D05CFF80FED59CB4FDBB7293" box="[371,613,1158,1183]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8BE39773D05CFF80FED59CB4FDBB7293" blockId="8.[215,770,1062,1215]" box="[371,613,1158,1183]" pageId="8" pageNumber="9">
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<heading id="D0AB201FD05CFF80FED59CB4FDBB7293" box="[371,613,1158,1183]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="9" reason="7">
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<taxonomicName id="4C5CECF0D05CFF80FED59CB4FDBB7293" authority="Estes, 1984" authorityName="Estes" authorityYear="1984" box="[371,613,1158,1183]" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="wezei">
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<emphasis id="B9284B61D05CFF80FED59CB4FE297293" box="[371,503,1158,1183]" italics="true" pageId="8" pageNumber="9">Mioproteus</emphasis>
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cf.
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<emphasis id="B9284B61D05CFF80FD879CB4FDBB7293" box="[545,613,1158,1183]" italics="true" pageId="8" pageNumber="9">wezei</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C346C4F8D05CFF80FE1D9C94FDC372B3" box="[443,541,1190,1215]" pageId="8" pageNumber="9" type="description">
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<paragraph id="8BE39773D05CFF80FE1D9C94FDC372B3" blockId="8.[215,770,1062,1215]" box="[443,541,1190,1215]" pageId="8" pageNumber="9">
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<figureCitation id="13678BF6D05CFF80FE1D9C94FDC372B3" box="[443,541,1190,1215]" captionStart="FIGURE 6" captionStartId="9.[202,293,718,741]" captionTargetBox="[204,1428,183,698]" captionTargetId="figure-452@9.[200,1437,173,702]" captionTargetPageId="9" captionText="FIGURE 6. Mioproteus cf. wezei from Hambach: trunk vertebra (IPB-HaR 2000) in dorsal (A), ventral (B) and right lateral (C) views; trunk vertebra (IPB-HaR 2006) in dorsal (D), ventral (E) and right lateral (F) views; trunk vertebra (IPB-HaR 2007) in dorsal (G), ventral (H) and left lateral (I) views; trunk vertebra (IPB-HaR 2403) in dorsal (J) and ventral (K) views. Scale bars equal 1 mm. Abbreviations: b, basapophysis; k, keel; n, neurapophysis; ps, posterior spine; tp, transverse process; vl, ventral lamina." figureDoi="http://doi.org/10.5281/zenodo.10962309" httpUri="https://zenodo.org/record/10962309/files/figure.png" pageId="8" pageNumber="9">Figure 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C346C4F8D05CFF80FF669CE3FE917346" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph id="8BE39773D05CFF80FF669CE3FE917346" blockId="8.[192,793,1233,1898]" pageId="8" pageNumber="9">
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<emphasis id="B9284B61D05CFF80FF669CE3FEF272E6" bold="true" box="[192,300,1233,1258]" pageId="8" pageNumber="9">Material.</emphasis>
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Hambach 11: nine trunk vertebrae (IPB-HaR 2000/2007, IPB-HaR 2013). Hambach 11C: six trunk vertebrae (IPB-HaR 2400/2403, IPB-HaR 2427/2428).
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</paragraph>
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</subSubSection>
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<subSubSection id="C346C4F8D05CFF80FF669D63FB377293" pageId="8" pageNumber="9" type="description">
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<paragraph id="8BE39773D05CFF80FF669D63FB377293" blockId="8.[192,793,1233,1898]" lastBlockId="8.[840,1441,166,1887]" pageId="8" pageNumber="9">
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<emphasis id="B9284B61D05CFF80FF669D63FE847366" bold="true" box="[192,346,1361,1386]" pageId="8" pageNumber="9">Description.</emphasis>
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These vertebrae (
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<figureCitation id="13678BF6D05CFF80FD989D63FD767366" box="[574,680,1361,1386]" captionStart="FIGURE 6" captionStartId="9.[202,293,718,741]" captionTargetBox="[204,1428,183,698]" captionTargetId="figure-452@9.[200,1437,173,702]" captionTargetPageId="9" captionText="FIGURE 6. Mioproteus cf. wezei from Hambach: trunk vertebra (IPB-HaR 2000) in dorsal (A), ventral (B) and right lateral (C) views; trunk vertebra (IPB-HaR 2006) in dorsal (D), ventral (E) and right lateral (F) views; trunk vertebra (IPB-HaR 2007) in dorsal (G), ventral (H) and left lateral (I) views; trunk vertebra (IPB-HaR 2403) in dorsal (J) and ventral (K) views. Scale bars equal 1 mm. Abbreviations: b, basapophysis; k, keel; n, neurapophysis; ps, posterior spine; tp, transverse process; vl, ventral lamina." figureDoi="http://doi.org/10.5281/zenodo.10962309" httpUri="https://zenodo.org/record/10962309/files/figure.png" pageId="8" pageNumber="9">Figure 6</figureCitation>
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) reach a large size and are robustly ossified. The largest one has a centrum length of about
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<quantity id="4CA43A96D05CFF80FDD69DA3FD1173A6" box="[624,719,1425,1450]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="9" unit="mm" value="6.5">6.5 mm</quantity>
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. They display a range of different degree of preservational statuses, from very fragmentary to well preserved. The centrum is amphicoelous and pierced by a wide notochordal canal. Ventrally, the centrum bears a robust and high keel, which is more or less narrow (from sharp to slightly thicker). By the anterior and posterior ends, the keel expands into triangular and flat ventral surfaces. The keel is flanked by small subcentral foramina, sometimes more than one per side. Posteriorly, poorly-developed basapophyses are present. The neural arch is low, without a significant rising posteriorly, and dorsally flat. A neurapophysis is present, being either rather low or slightly higher. It runs almost entirely along the dorsal surface of the arch. The anterior margin of the arch appears straight in dorsal view, even though it is never completely preserved. It is located roughly at midlength of the prezygapophyses. The posterior margin is rather straight (but very slightly wavy, even though with no median notch) and ends well before the end of the postzygapophyses. By each side of the neurapophysis, the posterior end of the dorsal surface of the neural arch carries a well-developed spine, which projects well beyond the posterior margin, but not beyond the postzygapophyses. The spines are well spaced in dorsal view, not strongly extended anteriorly (they do not reach the level of the transverse processes) and not connected to the neurapophysis medially. The zygapophyses are suboval and almost horizontal. The ventral lamina has a subtrapezoidal shape, originated by very well-developed anterior ventral crests and less-developed posterior ones. The zygapophyseal crests are also well developed, but not as much as the anterior ventral crests. The posterior zygapophyseal crests are distinctly convex dorsally in lateral view. Due to the strong development of both the ventral lamina and the zygapophyseal crests, the vertebrae assume a wide appearance in both ventral and dorsal views. The transverse processes are reduced to a small single structure, which is slender and posterolaterally directed. Individualized
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<typeStatus id="54E729D1D05CFF80FCEE9C74FC587253" box="[840,902,1094,1119]" pageId="8" pageNumber="9" type="paratype">para-</typeStatus>
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and diapophyses are not clearly discernible. A more-or-less large foramen is visible anterior to the base of the transverse process.
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</paragraph>
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</subSubSection>
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<subSubSection id="C346C4F8D05CFF81FCEE9C94FDD47309" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="discussion">
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<paragraph id="8BE39773D05CFF81FCEE9C94FDD47309" blockId="8.[840,1441,166,1887]" lastBlockId="9.[192,793,908,1285]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
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<emphasis id="B9284B61D05CFF80FCEE9C94FC1E72B3" bold="true" box="[840,960,1190,1215]" pageId="8" pageNumber="9">Remarks.</emphasis>
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These vertebrae are referred to
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<taxonomicName id="4C5CECF0D05CFF80FAE09C94FCA472D3" authorityName="Estes and Darevsky" authorityYear="1977" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
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<emphasis id="B9284B61D05CFF80FAE09C94FCA472D3" italics="true" pageId="8" pageNumber="9">Mioproteus</emphasis>
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</taxonomicName>
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because of the following combination of features (
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<bibRefCitation id="EFCDEA82D05CFF80FC349CD4FB0F72F3" author="Estes and Darevsky" box="[914,1233,1254,1279]" firstAuthor="Estes" pageId="8" pageNumber="9" pagination="164 - 169" refId="ref31723" refString="Estes, R. and Darevsky, I. 1977. Fossil amphibians from the Miocene of the North Caucasus, U. S. S. R. Journal of the Palaeontological Society of India, 20: 164 - 169." type="journal article" year="1977">Estes and Darevsky, 1977</bibRefCitation>
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): robust and wellossified aspect; well-spaced posterior spines; wide appearance in dorsal and ventral views (i.e., wide neural arch, wide ventral lamina); presence of basapophyses. Three species of
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<taxonomicName id="4C5CECF0D05CFF80FB4E9D54FAB27373" authorityName="Estes and Darevsky" authorityYear="1977" box="[1256,1388,1382,1407]" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
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<emphasis id="B9284B61D05CFF80FB4E9D54FAB27373" box="[1256,1388,1382,1407]" italics="true" pageId="8" pageNumber="9">Mioproteus</emphasis>
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</taxonomicName>
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are currently known:
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<taxonomicName id="4C5CECF0D05CFF80FBB49DB4FBD673B3" authority="Estes and Darevsky, 1977" authorityName="Estes and Darevsky" authorityYear="1977" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="caucasicus">
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<emphasis id="B9284B61D05CFF80FBB49DB4FAFC7393" box="[1042,1314,1414,1439]" italics="true" pageId="8" pageNumber="9">Mioproteus caucasicus</emphasis>
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<bibRefCitation id="EFCDEA82D05CFF80FA8F9DB4FBD673B3" author="Estes and Darevsky" firstAuthor="Estes" pageId="8" pageNumber="9" pagination="164 - 169" refId="ref31723" refString="Estes, R. and Darevsky, I. 1977. Fossil amphibians from the Miocene of the North Caucasus, U. S. S. R. Journal of the Palaeontological Society of India, 20: 164 - 169." type="journal article" year="1977">Estes and Darevsky, 1977</bibRefCitation>
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</taxonomicName>
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,
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<emphasis id="B9284B61D05CFF80FBBE9D94FB7573B3" box="[1048,1195,1446,1471]" italics="true" pageId="8" pageNumber="9">
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<taxonomicName id="4C5CECF0D05CFF80FBBE9D94FB7973B3" authorityName="Venczel and Codrea" authorityYear="2018" box="[1048,1191,1446,1471]" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="gardneri">M. gardneri</taxonomicName>
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,
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</emphasis>
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and
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<taxonomicName id="4C5CECF0D05CFF80FB569D94FA8073B3" box="[1264,1374,1446,1471]" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="wezei">
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<emphasis id="B9284B61D05CFF80FB569D94FA8073B3" box="[1264,1374,1446,1471]" italics="true" pageId="8" pageNumber="9">M. wezei</emphasis>
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</taxonomicName>
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. The vertebrae from Hambach are concordant in size with both
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<taxonomicName id="4C5CECF0D05CFF80FC639DD4FBAD73F3" authorityName="Estes and Darevsky" authorityYear="1977" box="[965,1139,1510,1535]" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="caucasicus">
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<emphasis id="B9284B61D05CFF80FC639DD4FBAD73F3" box="[965,1139,1510,1535]" italics="true" pageId="8" pageNumber="9">M. caucasicus</emphasis>
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</taxonomicName>
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and
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<emphasis id="B9284B61D05CFF80FB1F9DD4FAF173F3" box="[1209,1327,1510,1535]" italics="true" pageId="8" pageNumber="9">
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<taxonomicName id="4C5CECF0D05CFF80FB1F9DD4FAF473F3" box="[1209,1322,1510,1535]" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="wezei">M. wezei</taxonomicName>
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,
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</emphasis>
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whereas they are larger than
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<emphasis id="B9284B61D05CFF80FBEE9E34FB047013" box="[1096,1242,1542,1567]" italics="true" pageId="8" pageNumber="9">
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<taxonomicName id="4C5CECF0D05CFF80FBEE9E34FB097013" authorityName="Venczel and Codrea" authorityYear="2018" box="[1096,1239,1542,1567]" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="gardneri">M. gardneri</taxonomicName>
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.
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</emphasis>
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Regarding their morphology, they differ from
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<taxonomicName id="4C5CECF0D05CFF80FB1F9E14FA967033" authorityName="Venczel and Codrea" authorityYear="2018" box="[1209,1352,1574,1599]" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="gardneri">
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<emphasis id="B9284B61D05CFF80FB1F9E14FA967033" box="[1209,1352,1574,1599]" italics="true" pageId="8" pageNumber="9">M. gardneri</emphasis>
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</taxonomicName>
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in the more developed zygapophyseal crests, the taller neurapophysis, and the more laterally-trending prezygapophyses (
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<bibRefCitation id="EFCDEA82D05CFF80FB949EB4FA497093" author="Venczel and Codrea" box="[1074,1431,1670,1696]" firstAuthor="Venczel" pageId="8" pageNumber="9" pagination="1508027" refId="ref39349" refString="Venczel, M. and Codrea, V. A. 2018. A new proteid salamander from the early Oligocene of Romania with notes on the paleobiogeography of Eurasian proteids. Journal of Vertebrate Paleontology, 38: e 1508027. https: // doi. org / 10.1080 / 02724634.2018.1508027" type="journal article" year="2018">Venczel and Codrea, 2018</bibRefCitation>
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). Most characters reported to be diagnostic for
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<taxonomicName id="4C5CECF0D05CFF80FA249E94FC5270D3" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="wezei">
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<emphasis id="B9284B61D05CFF80FA249E94FC5270D3" italics="true" pageId="8" pageNumber="9">M. wezei</emphasis>
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</taxonomicName>
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are variable, but at least the consistent absence of a connection between the posterior spines and the neurapophysis seems to be sufficiently reliable (
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<bibRefCitation id="EFCDEA82D05CFF80FBA59F14FA847133" author="Syromyatnikova" box="[1027,1370,1830,1856]" etAl="et al." firstAuthor="Syromyatnikova" pageId="8" pageNumber="9" pagination="125 - 128" refId="ref38008" refString="Syromyatnikova, E., Tesakov, A., Frolov, P., and Titov, V. 2021. A Pliocene Mioproteus (Urodela: Proteidae) from the Taman Peninsula (Russia). Russian Journal of Herpetology, 28: 125 - 128. https: // doi. org / 10.30906 / 1026 - 2296 - 2021 - 28 - 2 - 125 - 128" type="journal article" year="2021">Syromyatnikova et al., 2021</bibRefCitation>
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). This condition is shown by the Hambach material, whereas
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<taxonomicName id="4C5CECF0D05DFF81FE899BBEFE0575A9" authorityName="Estes and Darevsky" authorityYear="1977" box="[303,475,908,933]" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="caucasicus">
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<emphasis id="B9284B61D05DFF81FE899BBEFE0575A9" box="[303,475,908,933]" italics="true" pageId="9" pageNumber="10">M. caucasicus</emphasis>
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</taxonomicName>
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either has posterior spine that connect or not to the neurapophysis medially. Strongly dorsally-convex posterior zygapophyseal crests may also be typical for
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<taxonomicName id="4C5CECF0D05DFF81FD8C9BDEFD4B7209" box="[554,661,1004,1029]" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="wezei">
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<emphasis id="B9284B61D05DFF81FD8C9BDEFD4B7209" box="[554,661,1004,1029]" italics="true" pageId="9" pageNumber="10">M. wezei</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="EFCDEA82D05DFF81FD039BDEFE927229" author="Mlynarski" etAl="et al." firstAuthor="Mlynarski" pageId="9" pageNumber="10" pagination="209 - 229" refId="ref34852" refString="Mlynarski, M., Szyndlar, Z., Estes, R., and Sanchiz B. 1984. Amphibians and reptiles from the Pliocene locality of Weze II near Dzialoszyn (Poland). Acta Palaeontologica Polonica, 29: 209 - 229." type="journal article" year="1984">Młynarski et al., 1984</bibRefCitation>
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;
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<bibRefCitation id="EFCDEA82D05DFF81FEFF9C3EFE2D7229" author="Bailon" box="[345,499,1036,1061]" firstAuthor="Bailon" pageId="9" pageNumber="10" pagination="1 - 8" refId="ref29841" refString="Bailon, S. 1995. Presence de Mioproteus wezei (Amphibia, Proteidae) dans le Pliocene superieur (MN 16) d'Europe de l'Ouest. Neues Jahrbuch fur Geologie und Palaontologie, Monatshefte, 1995: 1 - 8. https: // doi. org / 10.1127 / njgpm / 1995 / 1995 / 1" type="journal article" year="1995">Bailon, 1995</bibRefCitation>
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; even though
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<bibRefCitation id="EFCDEA82D05DFF81FD399C3EFE757249" author="Syromyatnikova" etAl="et al." firstAuthor="Syromyatnikova" pageId="9" pageNumber="10" pagination="125 - 128" refId="ref38008" refString="Syromyatnikova, E., Tesakov, A., Frolov, P., and Titov, V. 2021. A Pliocene Mioproteus (Urodela: Proteidae) from the Taman Peninsula (Russia). Russian Journal of Herpetology, 28: 125 - 128. https: // doi. org / 10.30906 / 1026 - 2296 - 2021 - 28 - 2 - 125 - 128" type="journal article" year="2021">Syromyatnikova et al., 2021</bibRefCitation>
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, mentioned some variation of this feature in
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<taxonomicName id="4C5CECF0D05DFF81FED69C7EFE2A7269" authorityName="Estes and Darevsky" authorityYear="1977" box="[368,500,1100,1125]" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
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<emphasis id="B9284B61D05DFF81FED69C7EFE2A7269" box="[368,500,1100,1125]" italics="true" pageId="9" pageNumber="10">Mioproteus</emphasis>
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</taxonomicName>
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vertebrae). Considering this, we here tentatively attribute the Hambach
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<taxonomicName id="4C5CECF0D05DFF81FF669CBEFE9A72A9" authorityName="Estes and Darevsky" authorityYear="1977" box="[192,324,1164,1189]" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
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<emphasis id="B9284B61D05DFF81FF669CBEFE9A72A9" box="[192,324,1164,1189]" italics="true" pageId="9" pageNumber="10">Mioproteus</emphasis>
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</taxonomicName>
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vertebrae to
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<emphasis id="B9284B61D05DFF81FE519CBEFDB172A9" box="[503,623,1164,1189]" italics="true" pageId="9" pageNumber="10">
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<taxonomicName id="4C5CECF0D05DFF81FE519CBEFDB572A9" box="[503,619,1164,1189]" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="wezei">M. wezei</taxonomicName>
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,
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</emphasis>
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even though pointing out uncertainty over this identification pending a clarification of the diagnostic features within Neogene
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<taxonomicName id="4C5CECF0D05DFF81FED99CDEFDDD7309" authorityName="Estes and Darevsky" authorityYear="1977" box="[383,515,1260,1285]" class="Amphibia" family="Proteidae" genus="Mioproteus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
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<emphasis id="B9284B61D05DFF81FED99CDEFDDD7309" box="[383,515,1260,1285]" italics="true" pageId="9" pageNumber="10">Mioproteus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |