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<mods:title id="FF4A52CFA45875887B57E5A37D9E3B22">Description of a new genus and species of Hemisphaeriini from Brunei with an identification key to the Bornean species of the tribe (Hemiptera: Fulgoroidea: Issidae)</mods:title>
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<treatment id="03E12E350732657380AAFDC62E323AD6" ID-DOI="http://doi.org/10.5281/zenodo.4504054" ID-GBIF-Taxon="180328967" ID-Zenodo-Dep="4504054" LSID="urn:lsid:plazi:treatment:03E12E350732657380AAFDC62E323AD6" httpUri="http://treatment.plazi.org/id/03E12E350732657380AAFDC62E323AD6" lastPageNumber="16" pageId="7" pageNumber="16">
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<heading id="D0BF284F0732657380AAFDC629C63CBC" bold="true" box="[254,948,628,654]" centered="true" fontSize="11" level="1" pageId="7" pageNumber="16" reason="2">
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Key to species of
<taxonomicName id="4C48E4A00732657381B0FDC628D43CBC" authorityName="Melichar" authorityYear="1906" box="[484,678,628,654]" class="Insecta" family="Issidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="16" phylum="Arthropoda" rank="tribe" tribe="Hemisphaeriini">Hemisphaeriini</taxonomicName>
known from Borneo
</emphasis>
</heading>
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<subSubSection id="C352CCA8073265738025FD1F2E323AD6" pageId="7" pageNumber="16" type="key">
<keyStep id="30BC8786073265738025FD1F2E323D36" pageId="7" pageNumber="16">
<paragraph id="8BF79F23073265738025FD1F2E323CD6" blockId="7.[113,1089,685,1252]" pageId="7" pageNumber="16">
<keyLead id="30B93C16073265738025FD1F2E323CD6" pageId="7" pageNumber="16">
1 Metope narrow, dark brown red with pale lateral margins (
<figureCitation id="137383A6073265738352FD1F29273CF6" box="[774,853,685,708]" captionStart="Figs 1519. 1516" captionStartId="5.[113,150,1460,1479]" captionTargetBox="[113,1088,220,1404]" captionTargetId="figure-11@5.[111,1092,220,904]" captionTargetPageId="5" captionText="Figs 1519. 1516 Hemisphaerius rufovarius Walker, 1858, holotype (BMNH), body length 3.5 mm; 1718 H. maculatus Melichar, 1906, syntype (NHRS), body length 4.5 mm (15, 17 dorsal view; 16, 18 lateral view). 19 Gergithus niger (Walker, 1857), syntype (BMNH), frontal view." figureDoi="http://doi.org/10.5281/zenodo.4503871" httpUri="https://zenodo.org/record/4503871/files/figure.png" pageId="7" pageNumber="16">Fig. 19</figureCitation>
). ................................ ..............................................................................
<taxonomicName id="4C48E4A00732657382FFFD7E2E323CD6" ID-CoL="3FWG4" authority="niger (Walker, 1857)" authorityName="niger (Walker" authorityYear="1857" baseAuthorityName="Walker" baseAuthorityYear="1857" box="[683,1088,716,740]" class="Insecta" family="Issidae" genus="Gergithus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="16" phylum="Arthropoda" rank="subSpecies" species="niger" subSpecies="niger">
<emphasis id="B93C43310732657382FFFD7E29E83CD6" bold="true" box="[683,922,716,740]" italics="true" pageId="7" pageNumber="16">Gergithus niger niger</emphasis>
(
<bibRefCitation id="EFD9E2D20732657383F3FD7F2E4B3CD6" author="WALKER F." box="[935,1081,717,740]" pageId="7" pageNumber="16" pagination="141 - 175" refId="ref3902" refString="WALKER F. 1857: Catalogue of the Homopterous insects collected at Sarawak, Borneo, by Mr. A. R. Wallace, with descriptions of new species. Journal of the Proceedings of the Linnean Society 1: 141 - 175 + plates 7 - 8." type="journal article" year="1857">Walker, 1857</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF79F23073265738025FD5F2E323D36" blockId="7.[113,1089,685,1252]" box="[113,1088,749,772]" pageId="7" pageNumber="16">
<keyLead id="30B93C16073265738025FD5F2E323D36" box="[113,1088,749,772]" pageId="7" pageNumber="16">
Metope wide (
<figureCitation id="137383A6073265738166FD5F2BF23D36" box="[306,384,749,772]" captionStart="Figs 1114" captionStartId="4.[151,188,1054,1073]" captionTargetBox="[167,1126,218,992]" captionTargetId="figure-10@4.[151,1129,218,1000]" captionTargetPageId="4" captionText="Figs 1114. Bruneastrum cardinale gen. &amp; sp. nov., holotype (BMNH), body length 4.1 mm. 11 dorsal view; 12 lateral view; 13 frontal view; 14 head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4503869" httpUri="https://zenodo.org/record/4503869/files/figure.png" pageId="7" pageNumber="16">Fig. 13</figureCitation>
), with different coloration. .......................................................... 2
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<keyStep id="30BC8786073265738025FCBF2E323DB6" pageId="7" pageNumber="16">
<paragraph id="8BF79F23073265738025FCBF2E323D76" blockId="7.[113,1089,685,1252]" pageId="7" pageNumber="16">
<keyLead id="30B93C16073265738025FCBF2E323D76" pageId="7" pageNumber="16">
2 Fore wings elongate, each with three separate light spots (
<figureCitation id="137383A6073265738355FCBF290E3D16" box="[769,892,781,804]" captionStart="Figs 1114" captionStartId="4.[151,188,1054,1073]" captionTargetBox="[167,1126,218,992]" captionTargetId="figure-10@4.[151,1129,218,1000]" captionTargetPageId="4" captionText="Figs 1114. Bruneastrum cardinale gen. &amp; sp. nov., holotype (BMNH), body length 4.1 mm. 11 dorsal view; 12 lateral view; 13 frontal view; 14 head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4503869" httpUri="https://zenodo.org/record/4503869/files/figure.png" pageId="7" pageNumber="16">Figs 11, 12</figureCitation>
). .......................... ..........................................................................
<emphasis id="B93C43310732657382C5FC9E2E323D76" bold="true" box="[657,1088,812,836]" pageId="7" pageNumber="16">
<taxonomicName id="4C48E4A00732657382C5FC9E29E03D76" ID-CoL="NFHT" authorityName="Gnezdilov" authorityYear="2015" box="[657,914,812,836]" class="Insecta" family="Issidae" genus="Bruneastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="16" phylum="Arthropoda" rank="species" species="cardinale">
<emphasis id="B93C43310732657382C5FC9E29E03D76" bold="true" box="[657,914,812,836]" italics="true" pageId="7" pageNumber="16">Bruneastrum cardinale</emphasis>
</taxonomicName>
gen. &amp; sp. nov.
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BF79F23073265738025FCFF2E323DB6" blockId="7.[113,1089,685,1252]" pageId="7" pageNumber="16">
<keyLead id="30B93C16073265738025FCFF2E323DB6" pageId="7" pageNumber="16">
Fore wings relatively wider (
<figureCitation id="137383A6073265738193FCFF28323D56" box="[455,576,845,868]" captionStart="Figs 1519. 1516" captionStartId="5.[113,150,1460,1479]" captionTargetBox="[113,1088,220,1404]" captionTargetId="figure-11@5.[111,1092,220,904]" captionTargetPageId="5" captionText="Figs 1519. 1516 Hemisphaerius rufovarius Walker, 1858, holotype (BMNH), body length 3.5 mm; 1718 H. maculatus Melichar, 1906, syntype (NHRS), body length 4.5 mm (15, 17 dorsal view; 16, 18 lateral view). 19 Gergithus niger (Walker, 1857), syntype (BMNH), frontal view." figureDoi="http://doi.org/10.5281/zenodo.4503871" httpUri="https://zenodo.org/record/4503871/files/figure.png" pageId="7" pageNumber="16">Figs 16, 18</figureCitation>
); if spots are present, then two of them are partly fused posteriorly (
<figureCitation id="137383A607326573810EFCDF2BDB3DB6" box="[346,425,877,900]" captionStart="Figs 1519. 1516" captionStartId="5.[113,150,1460,1479]" captionTargetBox="[113,1088,220,1404]" captionTargetId="figure-11@5.[111,1092,220,904]" captionTargetPageId="5" captionText="Figs 1519. 1516 Hemisphaerius rufovarius Walker, 1858, holotype (BMNH), body length 3.5 mm; 1718 H. maculatus Melichar, 1906, syntype (NHRS), body length 4.5 mm (15, 17 dorsal view; 16, 18 lateral view). 19 Gergithus niger (Walker, 1857), syntype (BMNH), frontal view." figureDoi="http://doi.org/10.5281/zenodo.4503871" httpUri="https://zenodo.org/record/4503871/files/figure.png" pageId="7" pageNumber="16">Fig. 18</figureCitation>
). ............................................................................................. 3
</keyLead>
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<keyStep id="30BC8786073265738025FC3F2E323DF6" pageId="7" pageNumber="16">
<paragraph id="8BF79F23073265738025FC3F2E323D96" blockId="7.[113,1089,685,1252]" box="[113,1088,909,932]" pageId="7" pageNumber="16">
<keyLead id="30B93C16073265738025FC3F2E323D96" box="[113,1088,909,932]" pageId="7" pageNumber="16">3 Fore wings unicoloured, without spots or bands. ........................................................... 4</keyLead>
</paragraph>
<paragraph id="8BF79F23073265738025FC1F2E323DF6" blockId="7.[113,1089,685,1252]" box="[113,1088,941,964]" pageId="7" pageNumber="16">
<keyLead id="30B93C16073265738025FC1F2E323DF6" box="[113,1088,941,964]" pageId="7" pageNumber="16"> Fore wings with spots or bands. ..................................................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BC8786073265738025FC7F2E323A36" pageId="7" pageNumber="16">
<paragraph id="8BF79F23073265738025FC7F2E323DD6" blockId="7.[113,1089,685,1252]" box="[113,1088,972,996]" pageId="7" pageNumber="16">
<keyLead id="30B93C16073265738025FC7F2E323DD6" box="[113,1088,972,996]" pageId="7" pageNumber="16">
4 General coloration dark brown to black. .........
<taxonomicName id="4C48E4A00732657382DBFC7E2E323DD6" ID-CoL="3KSRS" authority="Butler, 1875" authorityName="Butler" authorityYear="1875" box="[655,1088,972,996]" class="Insecta" family="Issidae" genus="Hemisphaerius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="16" phylum="Arthropoda" rank="species" species="bacculinus">
<emphasis id="B93C43310732657382DBFC7E29C73DD6" bold="true" box="[655,949,972,996]" italics="true" pageId="7" pageNumber="16">Hemisphaerius bacculinus</emphasis>
<bibRefCitation id="EFD9E2D20732657383E8FC7F2E323DD6" author="BUTLER A. G." box="[956,1088,973,996]" pageId="7" pageNumber="16" pagination="92 - 100" refId="ref3121" refString="BUTLER A. G. 1875: List of the species of homopterous genus Hemisphaerius, with descriptions of new forms in the collection of the British Museum. Annales and Magazine of Natural History, Series 4 16: 92 - 100." type="journal article" year="1875">Butler, 1875</bibRefCitation>
</taxonomicName>
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<keyLead id="30B93C16073265738025FC5F2E323A36" box="[113,1088,1005,1028]" pageId="7" pageNumber="16"> General coloration pale. ................................................................................................. 5</keyLead>
</paragraph>
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<keyStep id="30BC8786073265738025FBBF2E323A56" pageId="7" pageNumber="16">
<paragraph id="8BF79F23073265738025FBBF2E323A76" blockId="7.[113,1089,685,1252]" pageId="7" pageNumber="16">
<keyLead id="30B93C16073265738025FBBF2E323A76" pageId="7" pageNumber="16">
5 Fulvous, mesonotum with two wide red stripes (
<figureCitation id="137383A60732657382CDFBBF29673A16" box="[665,789,1037,1060]" captionStart="Figs 1519. 1516" captionStartId="5.[113,150,1460,1479]" captionTargetBox="[113,1088,220,1404]" captionTargetId="figure-11@5.[111,1092,220,904]" captionTargetPageId="5" captionText="Figs 1519. 1516 Hemisphaerius rufovarius Walker, 1858, holotype (BMNH), body length 3.5 mm; 1718 H. maculatus Melichar, 1906, syntype (NHRS), body length 4.5 mm (15, 17 dorsal view; 16, 18 lateral view). 19 Gergithus niger (Walker, 1857), syntype (BMNH), frontal view." figureDoi="http://doi.org/10.5281/zenodo.4503871" httpUri="https://zenodo.org/record/4503871/files/figure.png" pageId="7" pageNumber="16">Figs 15, 16</figureCitation>
). ......................................... .........................................................................
<taxonomicName id="4C48E4A00732657382DEFB9E2E323A76" ID-CoL="3KSTZ" authority="Walker, 1858" authorityName="Walker" authorityYear="1858" box="[650,1088,1068,1092]" class="Insecta" family="Issidae" genus="Hemisphaerius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="16" phylum="Arthropoda" rank="species" species="rufovarius">
<emphasis id="B93C43310732657382DEFB9E29DE3A76" bold="true" box="[650,940,1068,1092]" italics="true" pageId="7" pageNumber="16">Hemisphaerius rufovarius</emphasis>
<bibRefCitation id="EFD9E2D20732657383E6FB9F2E323A76" author="WALKER F." box="[946,1088,1069,1092]" pageId="7" pageNumber="16" refId="ref3952" refString="WALKER F. 1858: List of the specimens of Homopterous insects in the collection of the British Museum. Supplement. Printed by order of the Trustees, London, 369 pp." type="book" year="1858">Walker, 1858</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="8BF79F23073265738025FBFF2E323A56" blockId="7.[113,1089,685,1252]" box="[113,1088,1100,1124]" pageId="7" pageNumber="16">
<keyLead id="30B93C16073265738025FBFF2E323A56" box="[113,1088,1100,1124]" pageId="7" pageNumber="16">
Brown yellowish, mesonotum without stripes. ....
<taxonomicName id="4C48E4A00732657382F7FBFE2E323A56" ID-CoL="6M32D" authority="Walker, 1857" authorityName="Walker" authorityYear="1857" box="[675,1088,1100,1124]" class="Insecta" family="Issidae" genus="Hemisphaerius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="16" phylum="Arthropoda" rank="species" species="torpidus">
<emphasis id="B93C43310732657382F7FBFE29DE3A56" bold="true" box="[675,940,1100,1124]" italics="true" pageId="7" pageNumber="16">Hemisphaerius torpidus</emphasis>
<bibRefCitation id="EFD9E2D20732657383E6FBFF2E323A56" author="WALKER F." box="[946,1088,1101,1124]" pageId="7" pageNumber="16" pagination="141 - 175" refId="ref3902" refString="WALKER F. 1857: Catalogue of the Homopterous insects collected at Sarawak, Borneo, by Mr. A. R. Wallace, with descriptions of new species. Journal of the Proceedings of the Linnean Society 1: 141 - 175 + plates 7 - 8." type="journal article" year="1857">Walker, 1857</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
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<keyStep id="30BC8786073265738025FBDF2E323AD6" pageId="7" pageNumber="16">
<paragraph id="8BF79F23073265738025FBDF2E323A96" blockId="7.[113,1089,685,1252]" pageId="7" pageNumber="16">
<keyLead id="30B93C16073265738025FBDF2E323A96" pageId="7" pageNumber="16">
6 Fore wings testaceous with two wide, irregular, parallel, transverse dark brown to black lines (see
<bibRefCitation id="EFD9E2D2073265738156FB3F2BFD3A96" author="BUTLER A. G." box="[258,399,1165,1188]" pageId="7" pageNumber="16" pagination="92 - 100" refId="ref3121" refString="BUTLER A. G. 1875: List of the species of homopterous genus Hemisphaerius, with descriptions of new forms in the collection of the British Museum. Annales and Magazine of Natural History, Series 4 16: 92 - 100." type="journal article" year="1875">BUTLER 1875</bibRefCitation>
: pl. 4, fig. 14). .................
<taxonomicName id="4C48E4A00732657382E4FB3E2E323A96" ID-CoL="6LR26" authority="Walker, 1857" authorityName="Walker" authorityYear="1857" box="[688,1088,1164,1188]" class="Insecta" family="Issidae" genus="Hemisphaerius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="16" phylum="Arthropoda" rank="species" species="typicus">
<emphasis id="B93C43310732657382E4FB3E29D93A96" bold="true" box="[688,939,1164,1188]" italics="true" pageId="7" pageNumber="16">Hemisphaerius typicus</emphasis>
<bibRefCitation id="EFD9E2D20732657383E6FB3F2E323A96" author="WALKER F." box="[946,1088,1165,1188]" pageId="7" pageNumber="16" pagination="141 - 175" refId="ref3902" refString="WALKER F. 1857: Catalogue of the Homopterous insects collected at Sarawak, Borneo, by Mr. A. R. Wallace, with descriptions of new species. Journal of the Proceedings of the Linnean Society 1: 141 - 175 + plates 7 - 8." type="journal article" year="1857">Walker, 1857</bibRefCitation>
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<paragraph id="8BF79F23073265738025FB1F2E323AD6" blockId="7.[113,1089,685,1252]" pageId="7" pageNumber="16">
<keyLead id="30B93C16073265738025FB1F2E323AD6" pageId="7" pageNumber="16">
Fore wings dark brown, each with three large light spots, two of which are partly fused posteriorly (
<figureCitation id="137383A607326573814DFB7F2BE63AD6" box="[281,404,1229,1252]" captionStart="Figs 1519. 1516" captionStartId="5.[113,150,1460,1479]" captionTargetBox="[113,1088,220,1404]" captionTargetId="figure-11@5.[111,1092,220,904]" captionTargetPageId="5" captionText="Figs 1519. 1516 Hemisphaerius rufovarius Walker, 1858, holotype (BMNH), body length 3.5 mm; 1718 H. maculatus Melichar, 1906, syntype (NHRS), body length 4.5 mm (15, 17 dorsal view; 16, 18 lateral view). 19 Gergithus niger (Walker, 1857), syntype (BMNH), frontal view." figureDoi="http://doi.org/10.5281/zenodo.4503871" httpUri="https://zenodo.org/record/4503871/files/figure.png" pageId="7" pageNumber="16">Figs 17, 18</figureCitation>
). .............................
<taxonomicName id="4C48E4A0073265738223FB7E2E323AD6" ID-CoL="3KSTD" authority="Melichar, 1906" authorityName="Melichar" authorityYear="1906" box="[631,1088,1228,1252]" class="Insecta" family="Issidae" genus="Hemisphaerius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="16" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B93C4331073265738223FB7E29E53AD6" bold="true" box="[631,919,1228,1252]" italics="true" pageId="7" pageNumber="16">Hemisphaerius maculatus</emphasis>
<bibRefCitation id="EFD9E2D20732657383CAFB7F2E323AD6" author="MELICHAR L." box="[926,1088,1229,1252]" pageId="7" pageNumber="16" pagination="1 - 327" refId="ref3790" refString="MELICHAR L. 1906: Monographie der Issiden (Homoptera). Abhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 3 (4): 1 - 327." type="journal article" year="1906">Melichar, 1906</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>