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<document id="77482B620516C6EAACEE8A3A3E7AB1EF" ID-CLB-Dataset="29010" ID-DOI="10.11646/zootaxa.4459.3.7" ID-GBIF-Dataset="81db55ed-b19e-42e9-930e-f9697834cc1b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458879" ID-ZooBank="8CA301B7-A24A-4CB1-8231-FF0BAFF175DA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539290198072" checkinUser="plazi" docAuthor="Semenyuk, Irina, Golovatch, Sergei I. &amp; Wesener, Thomas" docDate="2018" docId="03D7EC02FFD02428CBBDE69FDDA8FF66" docLanguage="en" docName="zootaxa.4459.3.7.pdf" docOrigin="Zootaxa 4459 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphaerobelum Verhoeff 1924" docType="treatment" docVersion="8" lastPageNumber="543" masterDocId="FFEE947AFFD82421CB2AE274D834FFAC" masterDocTitle="Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae)" masterLastPageNumber="550" masterPageNumber="535" pageNumber="543" updateTime="1698578940799" updateUser="ExternalLinkService">
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<mods:title id="47AB2B0CC736FBE11B05D984712B38CC">Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:namePart id="35B4AFB9F50A007A32418F0C3318B765">Semenyuk, Irina</mods:namePart>
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<mods:namePart id="E40A560AF7D94B2615541861FB72A5EE">Golovatch, Sergei I.</mods:namePart>
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<treatment id="03D7EC02FFD02428CBBDE69FDDA8FF66" ID-DOI="http://doi.org/10.5281/zenodo.6495766" ID-GBIF-Taxon="148692869" ID-Zenodo-Dep="6495766" LSID="urn:lsid:plazi:treatment:03D7EC02FFD02428CBBDE69FDDA8FF66" httpUri="http://treatment.plazi.org/id/03D7EC02FFD02428CBBDE69FDDA8FF66" lastPageId="9" lastPageNumber="543" pageId="8" pageNumber="543">
<subSubSection id="C3640E9FFFD02429CBBDE69FDBE8FAA9" box="[151,988,1258,1285]" pageId="8" pageNumber="543" type="nomenclature">
<paragraph id="8BC15D14FFD02429CBBDE69FDBE8FAA9" blockId="8.[151,988,1258,1285]" box="[151,988,1258,1285]" pageId="8" pageNumber="543">
<heading id="D089EA78FFD02429CBBDE69FDBE8FAA9" bold="true" box="[151,988,1258,1285]" fontSize="11" level="1" pageId="8" pageNumber="543" reason="1">
<emphasis id="B90A8106FFD02429CBBDE69FDBE8FAA9" bold="true" box="[151,988,1258,1285]" pageId="8" pageNumber="543">
Key to species of
<taxonomicName id="4C7E2697FFD02429CA47E69EDA29FAA8" ID-CoL="7Q2C9" ID-ENA="1173041" authorityName="Verhoeff" authorityYear="1924" box="[365,541,1258,1284]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="B90A8106FFD02429CA47E69EDA29FAA8" bold="true" box="[365,541,1258,1284]" italics="true" pageId="8" pageNumber="543">Sphaerobelum</emphasis>
</taxonomicName>
(based mainly on male characters):
</emphasis>
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</paragraph>
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<keyStep id="308A45B1FFD02429CBBDE745DDAFFACF" pageId="8" pageNumber="543">
<subSubSection id="C3640E9FFFD02429CBBDE745DDAFFAEA" box="[151,1435,1329,1350]" pageId="8" pageNumber="543" type="key">
<paragraph id="8BC15D14FFD02429CBBDE745DDAFFAEA" blockId="8.[151,1436,1329,2025]" box="[151,1435,1329,1350]" pageId="8" pageNumber="543">
<keyLead id="308FFE21FFD02429CBBDE745DDAFFAEA" box="[151,1435,1329,1350]" pageId="8" pageNumber="543">1. Anterior telopod with 3 telopoditomeres................................................................... 2</keyLead>
</paragraph>
</subSubSection>
<paragraph id="8BC15D14FFD02429CBBDE73ADDAFFACF" blockId="8.[151,1436,1329,2025]" box="[151,1435,1358,1379]" pageId="8" pageNumber="543">
<keyLead id="308FFE21FFD02429CBBDE73ADDAFFACF" box="[151,1435,1358,1379]" pageId="8" pageNumber="543">- Anterior telopod with 4 telopoditomeres.................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="308A45B1FFD02429CBBDE718DDAFFA75" pageId="8" pageNumber="543">
<paragraph id="8BC15D14FFD02429CBBDE718DDAFFA32" blockId="8.[151,1436,1329,2025]" pageId="8" pageNumber="543">
<keyLead id="308FFE21FFD02429CBBDE718DDAFFA32" pageId="8" pageNumber="543">
2. Tergites shiny, glabrous. Process of podomere 2 of posterior telopod swollen only at inner margin. Endotergum with 34 dense rows of short marginal bristles....................................................
<taxonomicName id="4C7E2697FFD02429CFA8E7FDDDAFFA32" authority="Verhoeff, 1924" authorityName="Verhoeff" authorityYear="1924" box="[1154,1435,1417,1438]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="clavigerum">
<emphasis id="B90A8106FFD02429CFA8E7FDDD32FA32" box="[1154,1286,1417,1438]" italics="true" pageId="8" pageNumber="543">S. clavigerum</emphasis>
<bibRefCitation id="EFEF20E5FFD02429CE26E7FDDDAFFA32" box="[1292,1435,1417,1438]" pageId="8" pageNumber="543" refId="ref9896" refString="Verhoeff, K. W. (1924) Results of Dr. E. Mjoberg's Swedish scientific expedition to Australia 1910 - 1913. 34. Myriapoda: Diplopoda. Arkif for zoologi, 16 (5), 1 - 142." type="journal article">Verhoeff, 1924</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC15D14FFD02429CBBDE7D2DDAFFA75" blockId="8.[151,1436,1329,2025]" pageId="8" pageNumber="543">
<keyLead id="308FFE21FFD02429CBBDE7D2DDAFFA75" pageId="8" pageNumber="543">- Tergites hairy, process of podomere 2 of posterior telopod strongly swollen apically, spherical, broader than base of podomere.................................................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="308A45B1FFD02429CBBDE795DDAFF9C7" pageId="8" pageNumber="543">
<paragraph id="8BC15D14FFD02429CBBDE795DDA8F99D" blockId="8.[151,1436,1329,2025]" pageId="8" pageNumber="543">
<keyLead id="308FFE21FFD02429CBBDE795DDA8F99D" pageId="8" pageNumber="543">
3. Inner margin of femur extended, with teeth. Lateral margin of prefemur with a sharp projection juxtaposed to coxal process. Body coloration black. Head smooth. Process of podomere 2 of posterior telopods truncated. Operculum of vulva two-horned, both horns blunt....................................................................
<taxonomicName id="4C7E2697FFD02429CF99E468DDA8F99D" authority="Attems, 1938" authorityName="Attems" authorityYear="1938" box="[1203,1436,1564,1585]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="bicorne">
<emphasis id="B90A8106FFD02429CF99E468DD20F99D" box="[1203,1300,1564,1585]" italics="true" pageId="8" pageNumber="543">S. bicorne</emphasis>
<bibRefCitation id="EFEF20E5FFD02429CE31E468DDA8F99D" box="[1307,1436,1564,1585]" pageId="8" pageNumber="543" refId="ref9195" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in franzosisch Indochina gesammelten Myriopoden. Memoires du Museum national d'Histoire naturelle, New Series, 6 (2), 187 - 353." type="journal article">Attems, 1938</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC15D14FFD02429CBBDE44DDDAFF9C7" blockId="8.[151,1436,1329,2025]" pageId="8" pageNumber="543">
<keyLead id="308FFE21FFD02429CBBDE44DDDAFF9C7" pageId="8" pageNumber="543">
- Inner margin of femur smooth, prefemur without a process. Body coloration dirty olive-brown; head punctured and densely hirsute.........................................................................
<taxonomicName id="4C7E2697FFD02429CFB3E422DDAFF9C7" authority="Verhoeff, 1924" authorityName="Verhoeff" authorityYear="1924" box="[1177,1435,1622,1643]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="hirsutum">
<emphasis id="B90A8106FFD02429CFB3E422DD31F9C7" box="[1177,1285,1622,1643]" italics="true" pageId="8" pageNumber="543">S. hirsutum</emphasis>
<bibRefCitation id="EFEF20E5FFD02429CE26E422DDAFF9C7" box="[1292,1435,1622,1643]" pageId="8" pageNumber="543" refId="ref9896" refString="Verhoeff, K. W. (1924) Results of Dr. E. Mjoberg's Swedish scientific expedition to Australia 1910 - 1913. 34. Myriapoda: Diplopoda. Arkif for zoologi, 16 (5), 1 - 142." type="journal article">Verhoeff, 1924</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308A45B1FFD02429CBBDE400DDAFF90A" pageId="8" pageNumber="543">
<paragraph id="8BC15D14FFD02429CBBDE400DDA8F925" blockId="8.[151,1436,1329,2025]" box="[151,1436,1652,1673]" pageId="8" pageNumber="543">
<keyLead id="308FFE21FFD02429CBBDE400DDA8F925" box="[151,1436,1652,1673]" pageId="8" pageNumber="543">
4. Inner margin of femur extended, with teeth (
<figureCitation id="13454191FFD02429C9A0E400DAEEF925" box="[650,730,1652,1673]" captionStart="FIGURE 2" captionStartId="4.[151,249,1919,1941]" captionTargetBox="[191,1429,686,1892]" captionTargetId="figure@4.[176,1429,657,1899]" captionTargetPageId="4" captionText="FIGURE 2. Sphaerobelum cattiense sp. nov. AG = male paratypes, I = female paratype (all ZMUM). A. left anterior telopod, anterior view; B. left anterior telopod, lateral view; C. left anterior telopod, posterior view; D. leg 9, right, anterior view; E. right posterior telopod, posterior view; F. right posterior telopod, anterior view; G. telopoditomere 4, lateral view; H. right male antenna; I. left coxa and prefemur with vulva. Scale bar = 1 mm." httpUri="https://zenodo.org/record/1458883/files/figure.png" pageId="8" pageNumber="543">Figs 2D</figureCitation>
,
<figureCitation id="13454191FFD02429C9CEE400DB30F925" box="[740,772,1652,1673]" captionStart="FIGURE 5" captionStartId="7.[151,249,1727,1749]" captionTargetBox="[174,1411,508,1704]" captionTargetId="figure@7.[160,1430,477,1735]" captionTargetPageId="7" captionText="FIGURE 5. Sphaerobelum konkakinhense sp. nov. AF, H = male holotype (ZMUM), G = female paratype (ZFMK MYR6612). A: right anterior telopod, posterior view; B and C: right anterior telopod, anterior and lateral views, respectively; right anterior telopod; D. left leg 9, posterior view; E. right male antenna; F. posterior telopod, posterior view; G. posterior telopod, anterior view; H: vulva. Not to scale." httpUri="https://zenodo.org/record/1458889/files/figure.png" pageId="8" pageNumber="543">5D</figureCitation>
). Locking carina on anal shield short......................... 6
</keyLead>
</paragraph>
<paragraph id="8BC15D14FFD02429CBBDE4E5DDAFF90A" blockId="8.[151,1436,1329,2025]" box="[151,1435,1681,1702]" pageId="8" pageNumber="543">
<keyLead id="308FFE21FFD02429CBBDE4E5DDAFF90A" box="[151,1435,1681,1702]" pageId="8" pageNumber="543">- Inner margin of femur smooth. Locking carina on anal shield very long........................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="308A45B1FFD02429CBBDE4DADDA8F8D8" pageId="8" pageNumber="543">
<paragraph id="8BC15D14FFD02429CBBDE4DADDA8F953" blockId="8.[151,1436,1329,2025]" pageId="8" pageNumber="543">
<keyLead id="308FFE21FFD02429CBBDE4DADDA8F953" pageId="8" pageNumber="543">
5. Body coloration dark olive, posterior margin uniformly light brown, antennae and legs conspicuously orange (
<figureCitation id="13454191FFD02429CE29E4DADD7BF96F" box="[1283,1359,1710,1731]" captionStart="FIGURE 1" captionStartId="3.[151,249,1736,1758]" captionTargetBox="[152,1435,447,1716]" captionTargetId="figure@3.[152,1435,447,1716]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map with pictures of live specimens. A. Sphaerobelum cattiense sp. nov.; B. Sphaerobelum konkakinhense sp. nov.; C. Zephronia konkakinhensis sp. nov.; D. Zephronia montis sp. nov." httpUri="https://zenodo.org/record/1458881/files/figure.png" pageId="8" pageNumber="543">Fig. 1A</figureCitation>
). Prefemur without lateral projection (
<figureCitation id="13454191FFD02429C926E4B8DA6CF94D" box="[524,600,1740,1761]" captionStart="FIGURE 2" captionStartId="4.[151,249,1919,1941]" captionTargetBox="[191,1429,686,1892]" captionTargetId="figure@4.[176,1429,657,1899]" captionTargetPageId="4" captionText="FIGURE 2. Sphaerobelum cattiense sp. nov. AG = male paratypes, I = female paratype (all ZMUM). A. left anterior telopod, anterior view; B. left anterior telopod, lateral view; C. left anterior telopod, posterior view; D. leg 9, right, anterior view; E. right posterior telopod, posterior view; F. right posterior telopod, anterior view; G. telopoditomere 4, lateral view; H. right male antenna; I. left coxa and prefemur with vulva. Scale bar = 1 mm." httpUri="https://zenodo.org/record/1458883/files/figure.png" pageId="8" pageNumber="543">Fig. 2D</figureCitation>
). Apical margins of vulval operculum equally converging towards a mesal tip (
<figureCitation id="13454191FFD02429CE67E4B8DDA4F94D" box="[1357,1424,1740,1761]" captionStart="FIGURE 2" captionStartId="4.[151,249,1919,1941]" captionTargetBox="[191,1429,686,1892]" captionTargetId="figure@4.[176,1429,657,1899]" captionTargetPageId="4" captionText="FIGURE 2. Sphaerobelum cattiense sp. nov. AG = male paratypes, I = female paratype (all ZMUM). A. left anterior telopod, anterior view; B. left anterior telopod, lateral view; C. left anterior telopod, posterior view; D. leg 9, right, anterior view; E. right posterior telopod, posterior view; F. right posterior telopod, anterior view; G. telopoditomere 4, lateral view; H. right male antenna; I. left coxa and prefemur with vulva. Scale bar = 1 mm." httpUri="https://zenodo.org/record/1458883/files/figure.png" pageId="8" pageNumber="543">
Fig.
<date id="FFC07BD4FFD02429CE53E4B8DDA4F94D" box="[1401,1424,1740,1761]" pageId="8" pageNumber="543">2I</date>
</figureCitation>
). Tip of laterotergite 1 slightly extended, well-rounded. Male antennomere 6 thickened (
<figureCitation id="13454191FFD02429CF34E49DDC5CF952" box="[1054,1128,1769,1790]" captionStart="FIGURE 3" captionStartId="5.[151,249,662,684]" captionTargetBox="[166,1425,189,646]" captionTargetId="figure@5.[166,1426,187,647]" captionTargetPageId="5" captionText="FIGURE 3. Sphaerobelum cattiense sp. nov. Male paratype (ZFMK MYR6516), right antenna. A. lateral view; B. apical view of disc; C. details of apical cone. Abbreviations: sc = sclerotized scales; sb = sensilla basiconica. Scale bars: A = 1000 µm; B = 200 µm; C = 10 µm." httpUri="https://zenodo.org/record/1458885/files/figure.png" pageId="8" pageNumber="543">Fig. 3A</figureCitation>
).........
<taxonomicName id="4C7E2697FFD02429CFF6E49DDD7DF952" authority="Semenyuk &amp; Golovatch &amp; Wesener, 2018" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2018" box="[1244,1353,1769,1790]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="cattiense" status="sp. nov.">
<emphasis id="B90A8106FFD02429CFF6E49DDD7DF952" box="[1244,1353,1769,1790]" italics="true" pageId="8" pageNumber="543">S. cattiense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2393C7DFFD02429CE65E49DDDA8F953" box="[1359,1436,1769,1791]" pageId="8" pageNumber="543" rank="species">
<emphasis id="B90A8106FFD02429CE65E49DDDA8F953" bold="true" box="[1359,1436,1769,1791]" pageId="8" pageNumber="543">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC15D14FFD02429CBBDE572DDA8F8D8" blockId="8.[151,1436,1329,2025]" pageId="8" pageNumber="543">
<keyLead id="308FFE21FFD02429CBBDE572DDA8F8D8" pageId="8" pageNumber="543">
- Body coloration dark chestnut brown, posterior tergal margins black, legs dark brown, tips light brown (
<figureCitation id="13454191FFD02429CFE2E572DD26F8B7" box="[1224,1298,1798,1819]" captionStart="FIGURE 1" captionStartId="3.[151,249,1736,1758]" captionTargetBox="[152,1435,447,1716]" captionTargetId="figure@3.[152,1435,447,1716]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map with pictures of live specimens. A. Sphaerobelum cattiense sp. nov.; B. Sphaerobelum konkakinhense sp. nov.; C. Zephronia konkakinhensis sp. nov.; D. Zephronia montis sp. nov." httpUri="https://zenodo.org/record/1458881/files/figure.png" pageId="8" pageNumber="543">Fig. 1B</figureCitation>
). Lateral margin of prefemur with a sharp projection juxtaposed to coxal process (
<figureCitation id="13454191FFD02429C8A2E550DBE3F895" box="[904,983,1828,1849]" captionStart="FIGURE 5" captionStartId="7.[151,249,1727,1749]" captionTargetBox="[174,1411,508,1704]" captionTargetId="figure@7.[160,1430,477,1735]" captionTargetPageId="7" captionText="FIGURE 5. Sphaerobelum konkakinhense sp. nov. AF, H = male holotype (ZMUM), G = female paratype (ZFMK MYR6612). A: right anterior telopod, posterior view; B and C: right anterior telopod, anterior and lateral views, respectively; right anterior telopod; D. left leg 9, posterior view; E. right male antenna; F. posterior telopod, posterior view; G. posterior telopod, anterior view; H: vulva. Not to scale." httpUri="https://zenodo.org/record/1458889/files/figure.png" pageId="8" pageNumber="543">Fig. 5D</figureCitation>
). Apical margin of vulval operculum recessed centrally (
<figureCitation id="13454191FFD02429CA68E535D9BBF8FA" box="[322,399,1857,1878]" captionStart="FIGURE 5" captionStartId="7.[151,249,1727,1749]" captionTargetBox="[174,1411,508,1704]" captionTargetId="figure@7.[160,1430,477,1735]" captionTargetPageId="7" captionText="FIGURE 5. Sphaerobelum konkakinhense sp. nov. AF, H = male holotype (ZMUM), G = female paratype (ZFMK MYR6612). A: right anterior telopod, posterior view; B and C: right anterior telopod, anterior and lateral views, respectively; right anterior telopod; D. left leg 9, posterior view; E. right male antenna; F. posterior telopod, posterior view; G. posterior telopod, anterior view; H: vulva. Not to scale." httpUri="https://zenodo.org/record/1458889/files/figure.png" pageId="8" pageNumber="543">Fig. 5H</figureCitation>
). Tip of laterotergite 1 slightly extended, sharp. Male antennomere 6 axe-shaped (
<figureCitation id="13454191FFD02429CFB5E535DCDFF8FA" box="[1183,1259,1857,1878]" captionStart="FIGURE 6" captionStartId="8.[151,249,1125,1147]" captionTargetBox="[211,1382,375,1107]" captionTargetId="figure@8.[211,1383,375,1107]" captionTargetPageId="8" captionText="FIGURE 6. Sphaerobelum konkakinhense sp. nov., male holotype (ZMUM), right antenna. A. lateral view; B. apical view of disc. Abbreviations: sc = sclerotized scales; sb = sensilla basiconica. Scale bars: A = 1000 µm; B = 100 µm." httpUri="https://zenodo.org/record/1458891/files/figure.png" pageId="8" pageNumber="543">Fig. 6A</figureCitation>
)..............................................................................................
<taxonomicName id="4C7E2697FFD02429CF8EE52ADD7CF8DF" authority="Semenyuk &amp; Golovatch &amp; Wesener, 2018" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2018" box="[1188,1352,1886,1907]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="konkakinhense" status="sp. nov.">
<emphasis id="B90A8106FFD02429CF8EE52ADD7CF8DF" box="[1188,1352,1886,1907]" italics="true" pageId="8" pageNumber="543">S. konkakinhense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2393C7DFFD02429CE65E52ADDA8F8D8" box="[1359,1436,1886,1908]" pageId="8" pageNumber="543" rank="species">
<emphasis id="B90A8106FFD02429CE65E52ADDA8F8D8" bold="true" box="[1359,1436,1886,1908]" pageId="8" pageNumber="543">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BC15D14FFD02429CBBDE508DDA8F867" blockId="8.[151,1436,1329,2025]" pageId="8" pageNumber="543">
<keyStep id="308A45B1FFD02429CBBDE508DDA8F867" pageId="8" pageNumber="543">
<keyLead id="308FFE21FFD02429CBBDE508DDA8F867" pageId="8" pageNumber="543">
6. Podomere 3 of anterior telopod distally with a lateral extension; podomere 4 of posterior telopod distally with an incurved process overlapping with process of podomere 2. Marginal setae of terga not reaching posterior margin....................................................................................................
<taxonomicName id="4C7E2697FFD02429CFBDE5C2DDA8F867" authority="Attems, 1953" authorityName="Attems" authorityYear="1953" box="[1175,1436,1974,1995]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="separatum">
<emphasis id="B90A8106FFD02429CFBDE5C2DD20F867" box="[1175,1300,1974,1995]" italics="true" pageId="8" pageNumber="543">S. separatum</emphasis>
<bibRefCitation id="EFEF20E5FFD02429CE31E5C2DDA8F867" box="[1307,1436,1974,1995]" pageId="8" pageNumber="543" refId="ref9234" refString="Attems, C. (1953) Myriopoden von Indochina, Expedition von Dr. C. Dawydoff (1938 - 1939) Memoires du Museum national d'Histoire naturelle, New Series, A, 5, 133 - 230." type="journal article">Attems, 1953</bibRefCitation>
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BC15D14FFD02428CBBDE5A0DDA8FF66" blockId="8.[151,1436,1329,2025]" lastBlockId="9.[225,1436,151,202]" lastPageId="9" lastPageNumber="544" pageId="8" pageNumber="543">
<keyLead id="308FFE21FFD02429CBBDE5A0DDA8F845" box="[151,1436,2004,2025]" pageId="8" pageNumber="543">- Podomere 3 of anterior telopod cylindrical, without lateral extension; podomere 4 of posterior telopod without incurved pro-</keyLead>
<subSection id="E2F144FFFFD12428CBCBE2E3DDA8FF66" pageId="9" pageNumber="544" type="multiple">
cess, but with 3 separate spines. Marginal setae of terga reaching the posterior margin...................................................................................................
<taxonomicName id="4C7E2697FFD12428C8D1E2C1DDA8FF66" authority="Wongthamwanich et al., 2012" authorityName="Wongthamwanich et al." authorityYear="2012" box="[1019,1436,181,202]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="544" phylum="Arthropoda" rank="species" species="truncatum">
<emphasis id="B90A8106FFD12428C8D1E2C1DC42FF66" box="[1019,1142,181,202]" italics="true" pageId="9" pageNumber="544">S. truncatum</emphasis>
<bibRefCitation id="EFEF20E5FFD12428CF56E2C1DDA8FF66" box="[1148,1436,181,202]" pageId="9" pageNumber="544" refId="ref10201" refString="Wongthamwanich, N., Panha, S., Sierwald, P., Wesener, T. &amp; Thirakupt, K. (2012) A new species of the giant pill-millipede genus Sphaerobelum Verhoeff, 1924 from northern Thailand, with an extensive description and molecular characters (Diplopoda: Sphaerotheriida: Zephroniidae). Zootaxa, 3220, 29 - 43." type="journal article">
Wongthamwanich
<emphasis id="B90A8106FFD12428CE1AE2C1DD6EFF66" box="[1328,1370,181,202]" italics="true" pageId="9" pageNumber="544">et al</emphasis>
., 2012
</bibRefCitation>
</taxonomicName>
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</paragraph>
</treatment>
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