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<mods:title id="5E8B2F4C3E98C1AC95BFD41B8E867C9A">Taxonomic notes on the tribe Dryptini Bonelli, 1810 with description of a new genus and species from China (Coleoptera: Carabidae: Dryptini)</mods:title>
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<treatment id="03D77E1287106B1CFF0EFAB661E4F81C" ID-DOI="http://doi.org/10.5281/zenodo.3665217" ID-GBIF-Taxon="162074195" ID-Zenodo-Dep="3665217" LSID="urn:lsid:plazi:treatment:03D77E1287106B1CFF0EFAB661E4F81C" httpUri="http://treatment.plazi.org/id/03D77E1287106B1CFF0EFAB661E4F81C" lastPageNumber="524" pageId="2" pageNumber="524">
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<paragraph id="8BC1CF0487106B1CFF0EFAB660E0FAFE" blockId="2.[151,488,1314,1341]" box="[151,488,1314,1341]" pageId="2" pageNumber="524">
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<taxonomicName id="4C7EB48787106B1CFF0EFAB660E0FAFE" ID-CoL="8KZP6" authority="Jeannel, 1949" authorityName="Jeannel" authorityYear="1949" box="[151,488,1314,1341]" class="Insecta" family="Carabidae" genus="Prionodrypta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="B90A131687106B1CFF0EFAB660E0FAFE" bold="true" box="[151,488,1314,1341]" pageId="2" pageNumber="524">
<emphasis id="B90A131687106B1CFF0EFAB6603FFAFF" bold="true" box="[151,311,1314,1341]" italics="true" pageId="2" pageNumber="524">Prionodrypta</emphasis>
<bibRefCitation id="EFEFB2F587106B1CFEA7FAB760E0FAFE" author="Jeannel, R." box="[318,488,1314,1341]" pageId="2" pageNumber="524" pagination="767 - 1146" refId="ref5904" refString="Jeannel, R. (1949) Faune de l'empire francais. XI. Coleopteres carabiques de la region Malgache. Troisieme Partie. Libriarie Larose Paris, 380 pp. [pp. 767 - 1146]" type="book chapter" year="1949">Jeannel, 1949</bibRefCitation>
</emphasis>
</taxonomicName>
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<paragraph id="8BC1CF0487106B1CFF0EFAF163D4FABD" blockId="2.[151,732,1381,1407]" box="[151,732,1381,1407]" pageId="2" pageNumber="524">
<typeStatus id="54C571A687106B1CFF0EFAF161C7FABD" box="[151,207,1381,1407]" pageId="2" pageNumber="524">Type</typeStatus>
species:
<taxonomicName id="4C7EB48787106B1CFEACFAF263D4FABD" authority="Castelnau, 1897" authorityName="Castelnau" authorityYear="1897" box="[309,732,1381,1407]" class="Insecta" family="Carabidae" genus="Drypta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="524" phylum="Arthropoda" rank="species" species="mandibularis">
<emphasis id="B90A131687106B1CFEACFAF26329FABD" box="[309,545,1381,1407]" italics="true" pageId="2" pageNumber="524">Drypta mandibularis</emphasis>
Castelnau, 1897
</taxonomicName>
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<paragraph id="8BC1CF0487106B1CFF0EFA33602EF9EC" blockId="2.[151,1436,1447,2014]" pageId="2" pageNumber="524">
This genus, very synthetically described by
<bibRefCitation id="EFEFB2F587106B1CFD1AFA336222FA00" author="Jeannel, R." box="[643,810,1447,1474]" pageId="2" pageNumber="524" pagination="767 - 1146" refId="ref5904" refString="Jeannel, R. (1949) Faune de l'empire francais. XI. Coleopteres carabiques de la region Malgache. Troisieme Partie. Libriarie Larose Paris, 380 pp. [pp. 767 - 1146]" type="book chapter" year="1949">Jeannel (1949)</bibRefCitation>
, is scarcely known, so that none of the few authors who have mentioned it has noticed its peculiar characters and its complete independence from the “true”
<taxonomicName id="4C7EB48787106B1CFB64FA586444FA27" authorityName="Latreille" authorityYear="1796" box="[1277,1356,1484,1509]" class="Insecta" family="Carabidae" genus="Drypta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="B90A131687106B1CFB64FA586444FA27" box="[1277,1356,1484,1509]" italics="true" pageId="2" pageNumber="524">Drypta</emphasis>
</taxonomicName>
, rather treating it as a simple subgenus of
<taxonomicName id="4C7EB48787106B1CFD80FA646211F9C8" authority="(Lorenz, 2005)" baseAuthorityName="Lorenz" baseAuthorityYear="2005" box="[537,793,1519,1546]" class="Insecta" family="Carabidae" genus="Drypta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="B90A131687106B1CFD80FA646360F9CB" box="[537,616,1520,1545]" italics="true" pageId="2" pageNumber="524">Drypta</emphasis>
(
<bibRefCitation id="EFEFB2F587106B1CFDEEFA646219F9C8" author="Lorenz, W." box="[631,785,1519,1546]" pageId="2" pageNumber="524" refId="ref6110" refString="Lorenz, W. (2005) A systematic list of extant ground beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), 2 nd edition. W. Lorenz, Tutzing, 530 pp." type="book" year="2005">Lorenz, 2005</bibRefCitation>
)
</taxonomicName>
, or implicitly considering it a synonym of
<taxonomicName id="4C7EB48787106B1CFB6DFA64644BF9CB" baseAuthorityName="Lorenz" baseAuthorityYear="2005" box="[1268,1347,1520,1545]" class="Insecta" family="Carabidae" genus="Drypta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="B90A131687106B1CFB6DFA64644BF9CB" box="[1268,1347,1520,1545]" italics="true" pageId="2" pageNumber="524">Drypta</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFEFB2F587106B1CFACBFA7B61DAF9EC" author="Baehr, M." pageId="2" pageNumber="524" pagination="1 - 819" refId="ref5599" refString="Baehr, M. (2003) Archostemata-Myxophaga-Adephaga. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Apollo Books, Stenstrup, pp. 1 - 819." type="book chapter" year="2003">Baehr, 2003</bibRefCitation>
,
<bibRefCitation id="EFEFB2F587106B1CFF44F9806012F9EC" author="Baehr, M." box="[221,282,1556,1582]" pageId="2" pageNumber="524" pagination="1 - 1446" refId="ref5644" refString="Baehr, M. (2017) Archostemata-Myxophaga-Adephaga. Vol. 1. Revised and updated edition. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. I. Archostemata-Myxophaga-Adephaga. Brill, Leiden, pp. 1 - 1446." type="book chapter" year="2017">2017</bibRefCitation>
).
</paragraph>
<paragraph id="8BC1CF0487106B1CFF5EF9AC629CF97F" blockId="2.[151,1436,1447,2014]" pageId="2" pageNumber="524">
In our opinion it is a well distinguished genus, probably related to
<taxonomicName id="4C7EB48787106B1CFC5BF9AC6543F990" authorityName="Jeannel" authorityYear="1949" box="[962,1099,1592,1618]" class="Insecta" family="Carabidae" genus="Nesiodrypta" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="B90A131687106B1CFC5BF9AC6543F990" box="[962,1099,1592,1618]" italics="true" pageId="2" pageNumber="524">Nesiodrypta</emphasis>
</taxonomicName>
and forming with it and with
<taxonomicName id="4C7EB48787106B1CFF0EF9C86028F9B4" authority="Sciaky &amp; Anichtchenko, 2020" authorityName="Sciaky &amp; Anichtchenko" authorityYear="2020" box="[151,288,1628,1654]" class="Insecta" family="Carabidae" genus="Megadrypta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90A131687106B1CFF0EF9C86028F9B4" box="[151,288,1628,1654]" italics="true" pageId="2" pageNumber="524">Megadrypta</emphasis>
</taxonomicName>
, here described, a group of Gondwanian origin, The completely beaded sides of pronotum seem to us a more primitive character compared to the unbeaded sides of
<taxonomicName id="4C7EB48787106B1CFCCCF91462ACF95B" baseAuthorityName="Lorenz" baseAuthorityYear="2005" box="[853,932,1664,1689]" class="Insecta" family="Carabidae" genus="Drypta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="B90A131687106B1CFCCCF91462ACF95B" box="[853,932,1664,1689]" italics="true" pageId="2" pageNumber="524">Drypta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7EB48787106B1CFC42F9146579F958" authorityName="Schmidt-Goebel" authorityYear="1846" box="[987,1137,1664,1690]" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="B90A131687106B1CFC42F9146579F958" box="[987,1137,1664,1690]" italics="true" pageId="2" pageNumber="524">Dendrocellus</emphasis>
</taxonomicName>
, indicating a more ancient group from which probably the other two genera have later evolved.
</paragraph>
<paragraph id="8BC1CF0487106B1CFF5EF95C60CEF8C7" blockId="2.[151,1436,1447,2014]" pageId="2" pageNumber="524">
It contains 8 species from South-East Asia (
<collectingCountry id="F3698F9487106B1CFD32F95363E1F923" box="[683,745,1735,1761]" name="India" pageId="2" pageNumber="524">India</collectingCountry>
,
<collectingCountry id="F3698F9487106B1CFD6AF95C626AF920" box="[755,866,1736,1762]" name="Myanmar" pageId="2" pageNumber="524">Myanmar</collectingCountry>
,
<collectingCountry id="F3698F9487106B1CFCF5F95362D4F923" box="[876,988,1735,1761]" name="Indonesia" pageId="2" pageNumber="524">Indonesia</collectingCountry>
,
<collectingCountry id="F3698F9487106B1CFC7FF9536524F923" box="[998,1068,1735,1761]" name="China" pageId="2" pageNumber="524">China</collectingCountry>
,
<collectingCountry id="F3698F9487106B1CFBAFF95C6578F920" box="[1078,1136,1736,1762]" name="Laos" pageId="2" pageNumber="524">Laos</collectingCountry>
), still inadequately known; a revision would be useful.
</paragraph>
<paragraph id="8BC1CF0487106B1CFF5EF89B61E4F81C" blockId="2.[151,1436,1447,2014]" pageId="2" pageNumber="524">
The unique combination of characters distinguishing this genus is: pronotal bead well marked, with margin serrate (fig. 11); punctuation on head sparse, often leaving the frons smooth, on pronotum dense and irregular, with the punctures often confluent; pronotum evidently constricted towards base; elytral microsculpture almost absent, intervals smooth; elytral pubescence only aligned along the striae; single scutellar pore; intervals rather convex; tarsal claws thick, strongly curved, smooth on inner side; no setae, or a rudimental one, on outer side of stylomere (
<figureCitation id="1345D38187106B1CFF06F85761D6F81C" box="[159,222,1987,2014]" captionStart="FIGS" captionStartId="5.[151,208,1725,1750]" captionTargetBox="[350,1235,1327,1678]" captionTargetId="figure@5.[340,1247,1314,1700]" captionText="FIGS. 48. Stylomere structure of: 4—Megadrypta mirabilis gen. n. sp. n.; 5—Nesiodrypta waterhousei (Oberthür, 1881); 6—Prionodrypta lugens (Schmidt-Goebel, 1846); 7—Drypta lineola W. S. MacLeay, 1825; 8—Dendrocellus australis (Peringuey, 1896)." figureDoi="http://doi.org/10.5281/zenodo.3690687" httpUri="https://zenodo.org/record/3690687/files/figure.png" pageId="2" pageNumber="524">fig. 6</figureCitation>
).
</paragraph>
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