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<document id="874A46BB0FACF668460B6CFBBB23CA63" ID-DOI="10.5281/zenodo.13244881" ID-ISSN="2345-7600" ID-Zenodo-Dep="13244881" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722970250919" checkinUser="felipe" docAuthor="Hendrich, Lars &amp; Balke, Michael" docDate="2004" docId="03D687B7A73AFFB03804FAFEBB834A8F" docLanguage="en" docName="RafflesBZool.52.1.97-145.pdf" docOrigin="Raffles Bulletin of Zoology 52 (1)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Copelatus oblitus Sharp 1882" docType="treatment" docVersion="2" lastPageNumber="118" masterDocId="FFEFFFCFA72FFFA53903FF87BF1E4871" masterDocTitle="Aquatic Coleoptera Of Singapore: Species Richness, Ecology And Conservation #" masterLastPageNumber="145" masterPageNumber="97" pageNumber="118" updateTime="1722975545229" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B471C804B4F057CB6D73D665DF83560C">Aquatic Coleoptera Of Singapore: Species Richness, Ecology And Conservation #</mods:title>
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<mods:date id="A3C4399673BADFFF550AFB867E8E94EA">2004</mods:date>
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<mods:identifier id="D55FB6769AED7541011F76CAC8D773A5" type="ISSN">2345-7600</mods:identifier>
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<treatment id="03D687B7A73AFFB03804FAFEBB834A8F" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB03804FAFEBB834A8F" httpUri="http://treatment.plazi.org/id/03D687B7A73AFFB03804FAFEBB834A8F" lastPageNumber="118" pageId="21" pageNumber="118">
<subSubSection id="C365652AA73AFFB03804FAFEBD454DE0" box="[263,603,1401,1425]" pageId="21" pageNumber="118" type="nomenclature">
<paragraph id="8BC036A1A73AFFB03804FAFEBD454DE0" blockId="21.[263,603,1401,1425]" box="[263,603,1401,1425]" pageId="21" pageNumber="118">
<heading id="D08881CDA73AFFB03804FAFEBD454DE0" bold="true" box="[263,603,1401,1425]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="118" reason="2">
<taxonomicName id="4C7F4D22A73AFFB03804FAFEBD454DE0" ID-CoL="Y3DC" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[263,603,1401,1425]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="oblitus">
<emphasis id="B90BEAB3A73AFFB03804FAFEBD454DE0" bold="true" box="[263,603,1401,1425]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03804FAFEBED64DE0" bold="true" box="[263,456,1401,1425]" italics="true" pageId="21" pageNumber="118">Copelatus oblitus</emphasis>
<bibRefCitation id="EFEE4B50A73AFFB038D3FAFEBD454DE0" author="Sharp, D" box="[464,603,1401,1425]" pageId="21" pageNumber="118" pagination="6 - 18" refId="ref41075" refString="Sharp, D., 1882. On aquatic carnivorous Coleoptera or Dytiscidae. Transactions of the Royal Dublin Society, (2) 2: 179 - 1003, pl. 6 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C365652AA73AFFB03972FA31BDEE4D96" pageId="21" pageNumber="118" type="reference_group">
<paragraph id="8BC036A1A73AFFB03972FA31BDEE4DBB" blockId="21.[113,753,1461,1713]" box="[113,752,1461,1483]" pageId="21" pageNumber="118">
<treatmentCitationGroup id="AB6F118FA73AFFB03972FA31BDEE4DBB" box="[113,752,1461,1483]" pageId="21" pageNumber="118">
<treatmentCitation id="0ADE10B0A73AFFB03972FA31BEEF4DBB" author="Sharp, D" box="[113,497,1461,1483]" page="582" pageId="21" pageNumber="118" year="1882">
<taxonomicName id="4C7F4D22A73AFFB03972FA31BEEF4DBB" ID-CoL="Y3DC" authority="Sharp, 1882: 582" authorityName="Sharp" authorityPageNumber="582" authorityYear="1882" box="[113,497,1461,1483]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="oblitus">
<emphasis id="B90BEAB3A73AFFB03972FA31BE354DBA" box="[113,299,1462,1483]" italics="true" pageId="21" pageNumber="118">Copelatus oblitus</emphasis>
<bibRefCitation id="EFEE4B50A73AFFB03835FA32BEEF4DBB" author="Sharp, D" box="[310,497,1461,1482]" pageId="21" pageNumber="118" pagination="6 - 18" refId="ref41075" refString="Sharp, D., 1882. On aquatic carnivorous Coleoptera or Dytiscidae. Transactions of the Royal Dublin Society, (2) 2: 179 - 1003, pl. 6 - 18." type="journal article" year="1882">Sharp, 1882: 582</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73AFFB03B03FA32BDF24DBB" author="Regimbart, M" box="[512,748,1461,1482]" page="298" pageId="21" pageNumber="118" year="1899">
<bibRefCitation id="EFEE4B50A73AFFB03B03FA32BDF24DBB" author="Regimbart, M" box="[512,748,1461,1482]" pageId="21" pageNumber="118" pagination="186 - 367" refId="ref40540" refString="Regimbart, M., 1899. Revision des Dytiscidae de la region indosino-malaise. Annales de la Societe Entomologique de France, 68: 186 - 367." type="journal article" year="1899">Régimbart, 1899: 298</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC036A1A73AFFB03994FA55BDEE4D96" blockId="21.[113,753,1461,1713]" box="[151,752,1490,1511]" pageId="21" pageNumber="118">
<bibRefCitation id="EFEE4B50A73AFFB03994FA55BEB44D96" author="Zimmermann, A" box="[151,426,1490,1511]" pageId="21" pageNumber="118" pagination="1 - 326" refId="ref42405" refString="Zimmermann, A., 1920. Dytiscidae. In: Junk, W. &amp; S. Schenkling (eds.), Coleopterorum Catatalogus, Berlin, 4, pars 71, 1 - 326." type="journal article" year="1920">Zimmermann, 1920: 141</bibRefCitation>
;
<bibRefCitation id="EFEE4B50A73AFFB038B9FA55BDF24D96" author="Balfour-Browne, J" box="[442,748,1490,1511]" pageId="21" pageNumber="118" pagination="57 - 88" refId="ref35364" refString="Balfour-Browne, J., 1939 a. On Copelatus Erichson and Leiopterus Stephens (Col. Dytiscidae) with description of new species. Transactions of the Royal Entomological Society of London, 88 (2): 57 - 88." type="journal article" year="1939">Balfour-Browne, 1939a: 79</bibRefCitation>
;
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73AFFB03994FA68BD1B4EC0" pageId="21" pageNumber="118" type="description">
<paragraph id="8BC036A1A73AFFB03994FA68BDEF4E50" blockId="21.[113,753,1461,1713]" pageId="21" pageNumber="118">
<bibRefCitation id="EFEE4B50A73AFFB03994FA68BE714E75" author="Gueorguiev, V" box="[151,367,1519,1540]" pageId="21" pageNumber="118" pagination="5 - 45" refId="ref37343" refString="Gueorguiev, V., 1968. Essai de classification des Coleopteres Dytiscidae, Tribus Copelatini (Colymbetinae). Izvestija na Zoologitjeskija Institut s Musei Sofia, 28: 5 - 45." type="journal article" year="1968">Guéorguiev, 1968: 22</bibRefCitation>
;
<bibRefCitation id="EFEE4B50A73AFFB03879FA68BD2D4E75" author="Vazirani, T" box="[378,563,1519,1540]" pageId="21" pageNumber="118" refId="ref41335" refString="Vazirani, T. G., 1977. Catalogue of Oriental Dytiscidae. Records of the Zoological Survey of India, Occasional Paper, 6: ii + 111 pp." type="book" year="1977">Vazirani, 1977: 56</bibRefCitation>
;
<bibRefCitation id="EFEE4B50A73AFFB03B3EFA68BDF34E75" author="Nilsson, A" box="[573,749,1519,1540]" pageId="21" pageNumber="118" refId="ref39046" refString="Nilsson, A. N., 2001. Dytiscidae (Coleoptera). In: World Catalogue of Insects Volume 3. Apollo Books, Stenstrup, Denmark, 395 pp." type="book" year="2001">Nilsson, 2001: 70</bibRefCitation>
.
<taxonomicName id="4C7F4D22A73AFFB03972F98BBD2D4E50" authority="Regimbart, 1899: 302" authorityName="Regimbart" authorityPageNumber="302" authorityYear="1899" box="[113,563,1548,1569]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="andamanicus">
<emphasis id="B90BEAB3A73AFFB03972F98BBE474E50" box="[113,345,1548,1569]" italics="true" pageId="21" pageNumber="118">Copelatus andamanicus</emphasis>
<bibRefCitation id="EFEE4B50A73AFFB0385CF98BBD2D4E50" author="Regimbart, M" box="[351,563,1548,1569]" pageId="21" pageNumber="118" pagination="186 - 367" refId="ref40540" refString="Regimbart, M., 1899. Revision des Dytiscidae de la region indosino-malaise. Annales de la Societe Entomologique de France, 68: 186 - 367." type="journal article" year="1899">Régimbart, 1899: 302</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EFEE4B50A73AFFB03B3FF98BBDEF4E50" author="Gueorguiev, V" box="[572,753,1548,1569]" pageId="21" pageNumber="118" pagination="5 - 45" refId="ref37343" refString="Gueorguiev, V., 1968. Essai de classification des Coleopteres Dytiscidae, Tribus Copelatini (Colymbetinae). Izvestija na Zoologitjeskija Institut s Musei Sofia, 28: 5 - 45." type="journal article" year="1968">Guéorguiev, 1968:</bibRefCitation>
</paragraph>
<paragraph id="8BC036A1A73AFFB03994F9AEBDEE4E4F" blockId="21.[113,753,1461,1713]" box="[151,752,1577,1598]" pageId="21" pageNumber="118">
20;
<bibRefCitation id="EFEE4B50A73AFFB039BDF9AEBE984E4F" author="Vazirani, T" box="[190,390,1577,1598]" pageId="21" pageNumber="118" pagination="303 - 362" refId="ref41292" refString="Vazirani, T. G., 1971. Contributions to the study of aquatic beetles (Coleoptera). VII. A revision of Indian Colymbetinae (Dytiscidae). Oriental Insects, 4 (3): 303 - 362." type="journal article" year="1971">Vazirani, 1971: 310</bibRefCitation>
;
<bibRefCitation id="EFEE4B50A73AFFB03891F9AEBEF34E4F" author="Vazirani, T" box="[402,493,1577,1598]" pageId="21" pageNumber="118" refId="ref41335" refString="Vazirani, T. G., 1977. Catalogue of Oriental Dytiscidae. Records of the Zoological Survey of India, Occasional Paper, 6: ii + 111 pp." type="book" year="1977">1977: 53</bibRefCitation>
;
<bibRefCitation id="EFEE4B50A73AFFB038FBF9AEBD984E4F" author="Sato, M" box="[504,646,1577,1598]" pageId="21" pageNumber="118" pagination="35 - 41" refId="ref40808" refString="Sato, M., 1983. Distribution of the genus Copelatus in Japan (Dytiscidae). Special Issue Aquatic Coleoptera Workshop XVI. International Congress of Entomology, Kyoto, 1983: 35 - 41." type="journal article" year="1983">Satô 1983: 35</bibRefCitation>
;
<bibRefCitation id="EFEE4B50A73AFFB03B91F9AEBDF24E4F" author="Sato, M" box="[658,748,1577,1598]" pageId="21" pageNumber="118" pagination="57 - 67" refId="ref40846" refString="Sato, M., 1985. The Genus Copelatus of Japan (Coleoptera: Dytiscidae). Transactions of the Shikoku Entomological Society, 17 (1 / 2): 57 - 67." type="journal article" year="1985">1985: 58</bibRefCitation>
;
</paragraph>
<paragraph id="8BC036A1A73AFFB03994F9C2BD1B4EC0" blockId="21.[113,753,1461,1713]" pageId="21" pageNumber="118">
<bibRefCitation id="EFEE4B50A73AFFB03994F9C2BFEF4E2B" author="Sato, M" box="[151,241,1605,1626]" pageId="21" pageNumber="118" refId="ref40882" refString="Sato, M., 1990. Synonymic Notes on Copelatus Species (Dytiscidae). Elytra, 18 (1): 81." type="journal volume" year="1990">1990: 81</bibRefCitation>
; Balke et al., 1999: 325.
<emphasis id="B90BEAB3A73AFFB038E9F9C2BD504E2B" bold="true" box="[490,590,1605,1626]" pageId="21" pageNumber="118">Syn. Nov.</emphasis>
<taxonomicName id="4C7F4D22A73AFFB03972F9E5BD504E06" authority="Zimmermann, 1919 b: 76" authorityName="Zimmermann" authorityPageNumber="76" authorityYear="1919" box="[113,590,1634,1655]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="subfasciatus">
<emphasis id="B90BEAB3A73AFFB03972F9E5BE4D4E06" box="[113,339,1634,1655]" italics="true" pageId="21" pageNumber="118">Copelatus subfasciatus</emphasis>
<bibRefCitation id="EFEE4B50A73AFFB0385AF9E5BD504E06" author="Zimmermann, A" box="[345,590,1634,1655]" pageId="21" pageNumber="118" pagination="75 - 77" refId="ref42373" refString="Zimmermann, A., 1919 b. Sauters' Formosa-Ausbeute: Haliplidae et Dytiscidae (Coleoptera). Entomologische Mitteilungen, 8 (4 / 6): 75 - 77." type="journal article" year="1919">Zimmermann, 1919b: 76</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C7F4D22A73AFFB03972F9F8BD214EE5" authority="Sato, 1961: 8" authorityName="Sato" authorityPageNumber="8" authorityYear="1961" box="[113,575,1663,1684]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="118" phylum="Arthropoda" rank="subSpecies" species="socienus" subSpecies="ryukuyuensis">
<emphasis id="B90BEAB3A73AFFB03972F9F8BEAA4EE5" box="[113,436,1663,1684]" italics="true" pageId="21" pageNumber="118">Copelatus socienus ryukuyuensis</emphasis>
<bibRefCitation id="EFEE4B50A73AFFB038B8F9F8BD214EE5" author="Sato, M" box="[443,575,1663,1684]" pageId="21" pageNumber="118" pagination="7 - 10" refId="ref40744" refString="Sato, M., 1961. Aquatic Coleoptera from Amami-Oshima of the Ryukyu Islands (II). Akitu, Kyoto, 10: 7 - 10." type="journal article" year="1961">Satô, 1961: 8</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C7F4D22A73AFFB03972F91BBD1F4EC0" authority="Nakane, 1963: 25" authorityName="Nakane" authorityPageNumber="25" authorityYear="1963" box="[113,513,1692,1713]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="tokaraensis">
<emphasis id="B90BEAB3A73AFFB03972F91BBE554EC0" box="[113,331,1692,1713]" italics="true" pageId="21" pageNumber="118">Copelatus tokaraensis</emphasis>
<bibRefCitation id="EFEE4B50A73AFFB03851F91BBD1F4EC0" author="Nakane, T" box="[338,513,1692,1713]" pageId="21" pageNumber="118" pagination="23 - 26" refId="ref38865" refString="Nakane, T., 1963. New or little-known Coleoptera from Japan and ist adjacent regions, 18. Fragmenta Coleopterologica, 6: 23 - 26." type="journal article" year="1963">Nakane, 1963: 25</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73AFFB03972F952BC6B49F3" pageId="21" pageNumber="118" type="materials_examined">
<paragraph id="8BC036A1A73AFFB03972F952BFCC4F2F" blockId="21.[113,753,1749,1886]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03972F952BE594E9B" bold="true" box="[113,327,1749,1770]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03972F952BE2F4E9B" bold="true" box="[113,305,1749,1770]" italics="true" pageId="21" pageNumber="118">Material examined</emphasis>
. -
</emphasis>
<materialsCitation id="3B173CFCA73AFFB0384CF952BFD04F2F" collectionCode="BMNH, MNHN" country="Singapore" location="Type" pageId="21" pageNumber="118" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
Type material of
<taxonomicName id="4C7F4D22A73AFFB038FEF951BDBC4E9B" authority="Sharp" authorityName="Sharp" authorityYear="1882" box="[509,674,1749,1771]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="oblitus">
<emphasis id="B90BEAB3A73AFFB038FEF951BD404E9A" box="[509,606,1750,1771]" italics="true" pageId="21" pageNumber="118">C. oblitus</emphasis>
Sharp
</taxonomicName>
:
<specimenCount id="9D79FD28A73AFFB03BAEF952BDEE4E9B" box="[685,752,1749,1770]" count="1" pageId="21" pageNumber="118" type="male">1 male</specimenCount>
(
<typeStatus id="54C48803A73AFFB0397BF975BFCE4F76" box="[120,208,1778,1799]" pageId="21" pageNumber="118" type="holotype">holotype</typeStatus>
) - “
<collectingCountry id="F3687631A73AFFB039FBF975BE7F4F76" box="[248,353,1778,1799]" name="Singapore" pageId="21" pageNumber="118">Singapore</collectingCountry>
”, “
<location id="8EA0607AA73AFFB0387DF975BEAA4F76" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB03804FAFEBB834A8F:8EA0607AA73AFFB0387DF975BEAA4F76" box="[382,436,1778,1799]" country="Singapore" name="Type" pageId="21" pageNumber="118">Type</location>
” (
<collectionCode id="ED6EAE64A73AFFB038CFF975BD044F76" box="[460,538,1778,1799]" pageId="21" pageNumber="118">BMNH</collectionCode>
). Type material of
<taxonomicName id="4C7F4D22A73AFFB03BD9F975BE794F55" authority="Regimbart" authorityName="Regimbart" authorityYear="1899" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="andamanicus">
<emphasis id="B90BEAB3A73AFFB03BD9F975BFED4F55" italics="true" pageId="21" pageNumber="118">C. andamanicus</emphasis>
Régimbart
</taxonomicName>
:
<specimenCount id="9D79FD28A73AFFB03872F888BEAB4F55" box="[369,437,1807,1828]" count="1" pageId="21" pageNumber="118" type="male">1 male</specimenCount>
(
<typeStatus id="54C48803A73AFFB038C7F888BD3F4F55" box="[452,545,1807,1828]" pageId="21" pageNumber="118" type="lectotype">lectotype</typeStatus>
) -: “Ills Andaman”, “Cotype”, “
<taxonomicName id="4C7F4D22A73AFFB039E4F8ABBD7F4F30" authority="Reg. Cotype" authorityName="Reg. Cotype" box="[231,609,1836,1857]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="andamanicus">Copelatus andamanicus Rég. Cotype</taxonomicName>
”, “
<typeStatus id="54C48803A73AFFB03B82F8ABBDF64F30" box="[641,744,1836,1857]" pageId="21" pageNumber="118" type="lectotype">Lectotype</typeStatus>
” (
<collectionCode id="ED6EAE64A73AFFB0397BF8CEBFD94F2F" box="[120,199,1865,1886]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="118">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC036A1A73AFFB03972F805BC6B49F3" blockId="21.[113,753,1922,2116]" lastBlockId="21.[813,1452,193,386]" pageId="21" pageNumber="118">
Additional material:
<materialsCitation id="3B173CFCA73AFFB03838F805BDD84FE6" box="[315,710,1922,1943]" location="Central" pageId="21" pageNumber="118" specimenCount="44">
<specimenCount id="9D79FD28A73AFFB03838F805BEA14FE6" box="[315,447,1922,1943]" count="44" pageId="21" pageNumber="118" type="generic">44 specimens</specimenCount>
-
<emphasis id="B90BEAB3A73AFFB038D2F805BDD84FE6" bold="true" box="[465,710,1922,1943]" pageId="21" pageNumber="118">
<location id="8EA0607AA73AFFB038D2F805BD3F4FE6" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB03804FAFEBB834A8F:8EA0607AA73AFFB038D2F805BD3F4FE6" box="[465,545,1922,1943]" name="Central" pageId="21" pageNumber="118">Central</location>
catchment area
</emphasis>
</materialsCitation>
-
<materialsCitation id="3B173CFCA73AFFB03BDBF805BE9C4FA0" collectingDate="1996-12-16" collectionCode="CBH, ZRC" collectorName="L. Hendrich" location="Nee Soon Swamp Forest" pageId="21" pageNumber="118" specimenCount="11">
<specimenCount id="9D79FD28A73AFFB03BDBF805BF8F4FC5" count="11" pageId="21" pageNumber="118" type="generic">11 ex.</specimenCount>
,
<location id="8EA0607AA73AFFB03998F818BE914FC5" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB03804FAFEBB834A8F:8EA0607AA73AFFB03998F818BE914FC5" box="[155,399,1951,1972]" name="Nee Soon Swamp Forest" pageId="21" pageNumber="118">Nee Soon Swamp Forest</location>
, puddle on track,
<date id="FFC11061A73AFFB03B3FF818BDA24FC5" box="[572,700,1951,1972]" pageId="21" pageNumber="118" value="1996-12-16">
<collectingDate id="EF85E989A73AFFB03B3FF818BDA24FC5" box="[572,700,1951,1972]" pageId="21" pageNumber="118" value="1996-12-16">16 Dec.1996</collectingDate>
</date>
, coll.
<collectorName id="268A5377A73AFFB03972F83BBFF14FA0" box="[113,239,1980,2001]" pageId="21" pageNumber="118">L. Hendrich</collectorName>
(
<collectionCode id="ED6EAE64A73AFFB03802F83BBE264FA0" box="[257,312,1980,2001]" pageId="21" pageNumber="118">CBH</collectionCode>
,
<collectionCode id="ED6EAE64A73AFFB03845F83BBE654FA0" box="[326,379,1980,2001]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="118">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73AFFB03892F83BBDF34F9F" collectingDate="1997-04-21" collectionCode="CBH, ZRC" collectorName="M. Balke &amp; L. Hendrich &amp; C. M. Yang" location="MacRitchie Reservoir" pageId="21" pageNumber="118" specimenCount="16">
<specimenCount id="9D79FD28A73AFFB03892F83BBECA4FA0" box="[401,468,1980,2001]" count="16" pageId="21" pageNumber="118" type="generic">16 ex.</specimenCount>
,
<location id="8EA0607AA73AFFB038E0F83BBDD74FA0" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB03804FAFEBB834A8F:8EA0607AA73AFFB038E0F83BBDD74FA0" box="[483,713,1980,2001]" name="MacRitchie Reservoir" pageId="21" pageNumber="118">MacRitchie Reservoir</location>
,
<date id="FFC11061A73AFFB03BD4F83BBFD34F9F" pageId="21" pageNumber="118" value="1997-04-21">
<collectingDate id="EF85E989A73AFFB03BD4F83BBFD34F9F" pageId="21" pageNumber="118" value="1997-04-21">21 Apr.1997</collectingDate>
</date>
, coll.
<collectorName id="268A5377A73AFFB03801F85EBE474F9F" box="[258,345,2009,2030]" pageId="21" pageNumber="118">M. Balke</collectorName>
&amp;
<collectorName id="268A5377A73AFFB03877F85EBEFB4F9F" box="[372,485,2009,2030]" pageId="21" pageNumber="118">L. Hendrich</collectorName>
&amp;
<collectorName id="268A5377A73AFFB03B03F85EBD724F9F" box="[512,620,2009,2030]" pageId="21" pageNumber="118">C. M. Yang</collectorName>
(
<collectionCode id="ED6EAE64A73AFFB03B7BF85EBDB24F9F" box="[632,684,2009,2030]" pageId="21" pageNumber="118">CBH</collectionCode>
,
<collectionCode id="ED6EAE64A73AFFB03BB7F85EBDF84F9F" box="[692,742,2009,2030]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="118">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73AFFB03972F872BEFD4056" collectingDate="1997-04-22" collectionCode="ZRC" collectorName="H. K. Lua" location="Lower Peirce Reservoir" pageId="21" pageNumber="118" specimenCount="1">
<specimenCount id="9D79FD28A73AFFB03972F872BFBD407B" box="[113,163,2037,2058]" count="1" pageId="21" pageNumber="118" type="generic">1 ex.</specimenCount>
,
<location id="8EA0607AA73AFFB039AEF872BE86407B" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB03804FAFEBB834A8F:8EA0607AA73AFFB039AEF872BE86407B" box="[173,408,2037,2058]" name="Lower Peirce Reservoir" pageId="21" pageNumber="118">Lower Peirce Reservoir</location>
, after the concrete water runoff,
<date id="FFC11061A73AFFB03BDBF872BFD14056" pageId="21" pageNumber="118" value="1997-04-22">
<collectingDate id="EF85E989A73AFFB03BDBF872BFD14056" pageId="21" pageNumber="118" value="1997-04-22">22 Apr.1997</collectingDate>
</date>
, 230a, coll.
<collectorName id="268A5377A73AFFB0383CF795BE834056" box="[319,413,2066,2087]" pageId="21" pageNumber="118">H. K. Lua</collectorName>
(
<collectionCode id="ED6EAE64A73AFFB038AAF795BEC34056" box="[425,477,2066,2087]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="118">ZRC</collectionCode>
)
</materialsCitation>
.
<emphasis id="B90BEAB3A73AFFB038E8F795BD734056" bold="true" box="[491,621,2066,2087]" pageId="21" pageNumber="118">Bukit Timah</emphasis>
–
<materialsCitation id="3B173CFCA73AFFB03B80F795BD294035" collectionCode="ZRC" collectorName="C. F. Lim" location="Bukit Timah Nature Reserve" pageId="21" pageNumber="118" specimenCount="6">
<specimenCount id="9D79FD28A73AFFB03B80F795BDAC4056" box="[643,690,2066,2087]" count="6" pageId="21" pageNumber="118" type="generic">6 ex.</specimenCount>
,
<location id="8EA0607AA73AFFB03BB8F795BE4A4035" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB03804FAFEBB834A8F:8EA0607AA73AFFB03BB8F795BE4A4035" name="Bukit Timah Nature Reserve" pageId="21" pageNumber="118">Bukit Timah Nature Reserve</location>
, coll.
<collectorName id="268A5377A73AFFB03891F7A8BEF24035" box="[402,492,2095,2116]" pageId="21" pageNumber="118">C.F. Lim</collectorName>
(
<collectionCode id="ED6EAE64A73AFFB038F8F7A8BD314035" box="[507,559,2095,2116]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="118">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73AFFB03B40F7A8BC6C4883" collectingDate="1997-04-09" collectionCode="CBH" collectorName="L. Hendrich" pageId="21" pageNumber="118" specimenCount="1">
<specimenCount id="9D79FD28A73AFFB03B40F7A8BD684035" box="[579,630,2095,2116]" count="1" pageId="21" pageNumber="118" type="generic">1 ex.</specimenCount>
, at pipeline, MTB track, swampy area near pond,
<date id="FFC11061A73AFFB03D8CFF46BBE148A7" box="[1167,1279,193,214]" pageId="21" pageNumber="118" value="1997-04-09">
<collectingDate id="EF85E989A73AFFB03D8CFF46BBE148A7" box="[1167,1279,193,214]" pageId="21" pageNumber="118" value="1997-04-09">9 Apr.1997</collectingDate>
</date>
, coll.
<collectorName id="268A5377A73AFFB03C34FF46BAB548A7" box="[1335,1451,193,214]" pageId="21" pageNumber="118">L. Hendrich</collectorName>
(
<collectionCode id="ED6EAE64A73AFFB03A37FF5ABC754883" box="[820,875,221,242]" pageId="21" pageNumber="118">CBH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73AFFB03A7FFF5ABB18497E" collectingDate="2001-12-22" collectionCode="CBH" collectorName="M. Balke &amp; C. M. Yang" pageId="21" pageNumber="118" specimenCount="4">
<specimenCount id="9D79FD28A73AFFB03A7FFF5ABCB34883" box="[892,941,221,242]" count="4" pageId="21" pageNumber="118" type="generic">4 ex.</specimenCount>
, MTB track at pipeline,
<date id="FFC11061A73AFFB03D9AFF5ABA064883" box="[1177,1304,221,242]" pageId="21" pageNumber="118" value="2001-12-22">
<collectingDate id="EF85E989A73AFFB03D9AFF5ABA064883" box="[1177,1304,221,242]" pageId="21" pageNumber="118" value="2001-12-22">22 Dec.2001</collectingDate>
</date>
, coll.
<collectorName id="268A5377A73AFFB03C51FF5ABAB24883" box="[1362,1452,221,242]" pageId="21" pageNumber="118">M. Balke</collectorName>
&amp;
<collectorName id="268A5377A73AFFB03A45FF7DBCA5497E" box="[838,955,250,271]" pageId="21" pageNumber="118">C. M. Yang</collectorName>
(
<collectionCode id="ED6EAE64A73AFFB03ACBFF7DBCE1497E" box="[968,1023,250,271]" pageId="21" pageNumber="118">CBH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B173CFCA73AFFB03D13FF7DBB92497E" box="[1040,1164,250,271]" location="Other" pageId="21" pageNumber="118" specimenCount="1">
<emphasis id="B90BEAB3A73AFFB03D13FF7DBB92497E" bold="true" box="[1040,1164,250,271]" pageId="21" pageNumber="118">
<location id="8EA0607AA73AFFB03D13FF7DBB50497E" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB03804FAFEBB834A8F:8EA0607AA73AFFB03D13FF7DBB50497E" box="[1040,1102,250,271]" name="Other" pageId="21" pageNumber="118">Other</location>
areas
</emphasis>
</materialsCitation>
-
<materialsCitation id="3B173CFCA73AFFB03DA1FF7DBAB6495D" collectingDate="1992-01-10" collectionCode="ZRC" collectorName="C. M. Yang &amp; K. L. Yeo" location="Sembawang Senoko" pageId="21" pageNumber="118" specimenCount="1">
<specimenCount id="9D79FD28A73AFFB03DA1FF7DBBCA497E" box="[1186,1236,250,271]" count="1" pageId="21" pageNumber="118" type="generic">1 ex.</specimenCount>
,
<location id="8EA0607AA73AFFB03DDCFF7DBAB6497E" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB03804FAFEBB834A8F:8EA0607AA73AFFB03DDCFF7DBAB6497E" box="[1247,1448,250,271]" name="Sembawang Senoko" pageId="21" pageNumber="118">Sembawang Senoko</location>
, mud track,
<date id="FFC11061A73AFFB03A96FE90BB17495D" box="[917,1033,279,300]" pageId="21" pageNumber="118" value="1992-01-10">
<collectingDate id="EF85E989A73AFFB03A96FE90BB17495D" box="[917,1033,279,300]" pageId="21" pageNumber="118" value="1992-01-10">10 Jan.1992</collectingDate>
</date>
, 177 b, coll.
<collectorName id="268A5377A73AFFB03D7EFE90BBF5495D" box="[1149,1259,279,300]" pageId="21" pageNumber="118">C. M. Yang</collectorName>
&amp;
<collectorName id="268A5377A73AFFB03C05FE90BA7D495D" box="[1286,1379,279,300]" pageId="21" pageNumber="118">K. L. Yeo</collectorName>
(
<collectionCode id="ED6EAE64A73AFFB03C6DFE90BABF495D" box="[1390,1441,279,300]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="118">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B173CFCA73AFFB03A2EFEB3BC6F49F3" collectingDate="2001-12-22" collectionCode="CBH" collectorName="M. Balke &amp; C. M. Yang" location="Singapor" municipality="Bukit Timah" pageId="21" pageNumber="118" specimenCount="4">
<specimenCount id="9D79FD28A73AFFB03A2EFEB3BC404938" box="[813,862,308,329]" count="4" pageId="21" pageNumber="118" type="generic">4 ex.</specimenCount>
, ditch on grassy patch near road,
<location id="8EA0607AA73AFFB03DA0FEB3BBE54938" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB03804FAFEBB834A8F:8EA0607AA73AFFB03DA0FEB3BBE54938" box="[1187,1275,308,329]" municipality="Bukit Timah" name="Singapor" pageId="21" pageNumber="118">Singapor</location>
gun club, close to
<collectingMunicipality id="6BA4ACDBA73AFFB03A2EFED6BCBA4917" box="[813,932,337,358]" pageId="21" pageNumber="118">Bukit Timah</collectingMunicipality>
MTB track,
<date id="FFC11061A73AFFB03D18FED6BB8A4917" box="[1051,1172,337,358]" pageId="21" pageNumber="118" value="2001-12-22">
<collectingDate id="EF85E989A73AFFB03D18FED6BB8A4917" box="[1051,1172,337,358]" pageId="21" pageNumber="118" value="2001-12-22">22 Dec.2001</collectingDate>
</date>
, coll.
<collectorName id="268A5377A73AFFB03DC8FED6BA3C4917" box="[1227,1314,337,358]" pageId="21" pageNumber="118">M. Balke</collectorName>
&amp;
<collectorName id="268A5377A73AFFB03C3DFED6BAB24917" box="[1342,1452,337,358]" pageId="21" pageNumber="118">C. M. Yang</collectorName>
(
<collectionCode id="ED6EAE64A73AFFB03A37FEEABC7449F3" box="[820,874,365,386]" pageId="21" pageNumber="118">CBH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73AFFB03A2EFE20BCBC4A4F" pageId="21" pageNumber="118" type="distribution">
<paragraph id="8BC036A1A73AFFB03A2EFE20BCBC4A4F" blockId="21.[813,1452,423,574]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03A2EFE20BCD449CE" bold="true" box="[813,970,423,447]" italics="true" pageId="21" pageNumber="118">Distribution. –</emphasis>
<collectingCountry id="F3687631A73AFFB03ACCFE20BB1B49CF" box="[975,1029,423,446]" name="India" pageId="21" pageNumber="118">India</collectingCountry>
[Andaman Islands],
<collectingCountry id="F3687631A73AFFB03DE3FE20BA0249CF" box="[1248,1308,423,446]" name="Japan" pageId="21" pageNumber="118">Japan</collectingCountry>
[Yaku-shima and Ryukus Islands],
<collectingCountry id="F3687631A73AFFB03D21FE40BB9B49AF" box="[1058,1157,455,478]" name="Malaysia" pageId="21" pageNumber="118">Malaysia</collectingCountry>
[
<collectingCountry id="F3687631A73AFFB03D95FE40BBE549AF" box="[1174,1275,455,478]" name="Malaysia" pageId="21" pageNumber="118">Sarawak</collectingCountry>
],
<collectingCountry id="F3687631A73AFFB03C13FE40BA7F49AF" box="[1296,1377,455,478]" name="Taiwan" pageId="21" pageNumber="118">Taiwan</collectingCountry>
(
<bibRefCitation id="EFEE4B50A73AFFB03C70FE40BC7B498F" author="Sato, M" pageId="21" pageNumber="118" pagination="57 - 67" refId="ref40846" refString="Sato, M., 1985. The Genus Copelatus of Japan (Coleoptera: Dytiscidae). Transactions of the Shikoku Entomological Society, 17 (1 / 2): 57 - 67." type="journal article" year="1985">Satô, 1985</bibRefCitation>
,
<bibRefCitation id="EFEE4B50A73AFFB03A6CFE60BCB8498F" author="Sato, M" box="[879,934,487,510]" pageId="21" pageNumber="118" refId="ref40882" refString="Sato, M., 1990. Synonymic Notes on Copelatus Species (Dytiscidae). Elytra, 18 (1): 81." type="journal volume" year="1990">1990</bibRefCitation>
) and
<collectingCountry id="F3687631A73AFFB03AE2FE60BB50498F" box="[993,1102,487,510]" name="Singapore" pageId="21" pageNumber="118">Singapore</collectingCountry>
(
<bibRefCitation id="EFEE4B50A73AFFB03D58FE60BA1E498F" author="Vazirani, T" box="[1115,1280,487,510]" pageId="21" pageNumber="118" refId="ref41335" refString="Vazirani, T. G., 1977. Catalogue of Oriental Dytiscidae. Records of the Zoological Survey of India, Occasional Paper, 6: ii + 111 pp." type="book" year="1977">Vazirani, 1977</bibRefCitation>
). The record for
<collectingCountry id="F3687631A73AFFB03A2EFD80BC844A6F" box="[813,922,519,542]" name="Indonesia" pageId="21" pageNumber="118">Indonesia</collectingCountry>
:
<collectingRegion id="49BBF843A73AFFB03AA4FD80BB164A6F" box="[935,1032,519,542]" country="Indonesia" name="Sulawesi Utara" pageId="21" pageNumber="118">Sulawesi</collectingRegion>
(
<bibRefCitation id="EFEE4B50A73AFFB03D19FD80BA044A6F" author="Balfour-Browne, J" box="[1050,1306,519,542]" pageId="21" pageNumber="118" pagination="57 - 88" refId="ref35364" refString="Balfour-Browne, J., 1939 a. On Copelatus Erichson and Leiopterus Stephens (Col. Dytiscidae) with description of new species. Transactions of the Royal Entomological Society of London, 88 (2): 57 - 88." type="journal article" year="1939">Balfour-Browne 1939a</bibRefCitation>
) needs to be confirmed.
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73AFFB03A2EFDE0BB834A8F" pageId="21" pageNumber="118" type="biology_ecology">
<paragraph id="8BC036A1A73AFFB03A2EFDE0BB834A8F" blockId="21.[813,1452,615,766]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03A2EFDE0BCBF4A0E" bold="true" box="[813,929,615,639]" italics="true" pageId="21" pageNumber="118">Ecology. –</emphasis>
The species is found among leaf litter, roots and submerged plants of very small, temporary, shallow puddles in or near forested areas. The species seems to be restricted to the second growth forests of the Central Catchment Areas and Bukit Timah Nature Reserve.
</paragraph>
</subSubSection>
</treatment>
</document>
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