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<document id="C9A3044829C9DA84A310F2790A8D2BC3" ID-CLB-Dataset="7873" ID-DOI="10.5281/zenodo.4564890" ID-GBIF-Dataset="d9ee6109-a271-4440-8a93-dd53f043be8e" ID-ISSN="1942-1354" ID-Zenodo-Dep="4564890" ID-ZooBank="CF7B0E89-C497-408C-B359-B33939AB5C58" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614351828702" checkinUser="felipe" docAuthor="Atkinson, Thomas H." docDate="2020" docId="03A4926FFF958840FF6161B6BAA2A501" docLanguage="en" docName="InsectaMundi.2020.0792.1-25.pdf" docOrigin="Insecta Mundi 2020 (792)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Corthylus cristatus Atkinson 2020, new species" docType="treatment" docVersion="8" lastPageNumber="9" masterDocId="FF9DEA17FF92884AFFA1655DBE24A645" masterDocTitle="New species, new records and synonymy of Mexican Corthylus Erichson, 1834 (Coleoptera: Curculionidae: Scolytinae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="6" updateTime="1698931197508" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="076694567AEFCAEEF600243D7CD08817">New species, new records and synonymy of Mexican Corthylus Erichson, 1834 (Coleoptera: Curculionidae: Scolytinae)</mods:title>
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<mods:namePart id="3E497545D178C97445D270FF929DEFE2">Atkinson, Thomas H.</mods:namePart>
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<mods:title id="90FBCD0C70E634025324F5F316A2FFE1">Insecta Mundi</mods:title>
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<mods:date id="69691BBC92EB295ECDEE174867736533">2020</mods:date>
<mods:detail id="F61C6B0E667208A779724596BED6D75C" type="pubDate">
<mods:number id="56F7B8866E33DA9282ABE774ADD36556">2020-09-25</mods:number>
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<mods:number id="F6B1FA4BDC171328AF00ACCB5713C315">2020</mods:number>
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<mods:number id="999E29774C0189271F63C88AC80050B8">792</mods:number>
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<mods:classification id="58013F91AC6E9DA3D0FC120B554A56A8">journal article</mods:classification>
<mods:identifier id="B1BD7AFC382856ED0E6D4879BD4AF643" type="CLB-Dataset">7873</mods:identifier>
<mods:identifier id="45A159FAB9DF6D259DA4F55A04A71359" type="DOI">10.5281/zenodo.4564890</mods:identifier>
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<treatment id="03A4926FFF958840FF6161B6BAA2A501" ID-DOI="http://doi.org/10.5281/zenodo.4586059" ID-GBIF-Taxon="180298697" ID-Zenodo-Dep="4586059" LSID="urn:lsid:plazi:treatment:03A4926FFF958840FF6161B6BAA2A501" httpUri="http://treatment.plazi.org/id/03A4926FFF958840FF6161B6BAA2A501" lastPageId="10" lastPageNumber="9" pageId="7" pageNumber="6">
<subSubSection id="C31770F2FF95884DFF6161B6BCEBA34C" box="[192,719,1259,1289]" pageId="7" pageNumber="6" type="nomenclature">
<paragraph id="8BB22379FF95884DFF6161B6BCEBA34C" blockId="7.[192,719,1259,1323]" box="[192,719,1259,1289]" pageId="7" pageNumber="6">
<heading id="D0FA9415FF95884DFF6161B6BCEBA34C" box="[192,719,1259,1289]" fontSize="12" level="2" pageId="7" pageNumber="6" reason="2">
<emphasis id="B979FF6BFF95884DFF6161B6BCEBA34C" bold="true" box="[192,719,1259,1289]" pageId="7" pageNumber="6">
<taxonomicName id="4C0D58FAFF95884DFF6161B6BC0AA34C" authority="Atkinson" authorityName="Atkinson" authorityYear="2020" box="[192,558,1259,1289]" class="Insecta" family="Curculionidae" genus="Corthylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="cristatus" status="sp. nov.">
<emphasis id="B979FF6BFF95884DFF6161B6BF8AA34D" bold="true" box="[192,430,1259,1288]" italics="true" pageId="7" pageNumber="6">Corthylus cristatus</emphasis>
Atkinson
</taxonomicName>
,
<taxonomicNameLabel id="A24A4210FF95884DFD9B61B1BCEBA34C" box="[570,719,1260,1289]" pageId="7" pageNumber="6" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31770F2FF95884DFF61604CBF9EA36E" box="[192,442,1296,1323]" pageId="7" pageNumber="6" type="description">
<paragraph id="8BB22379FF95884DFF61604CBF9EA36E" blockId="7.[192,719,1259,1323]" box="[192,442,1296,1323]" pageId="7" pageNumber="6">
<figureCitation id="13363FFCFF95884DFF61604CBF12A36E" box="[192,310,1297,1323]" captionStart="Figure 4" captionStartId="8.[192,268,1672,1698]" captionTargetBox="[305,1326,213,1641]" captionTargetId="figure-10@8.[305,1326,213,1641]" captionTargetPageId="8" captionText="Figure 4. Species of Corthylus with acute spines on declivity or with interstriae 3 curved and serrate (2 of 2). A, B) C. cristatus new species, holotype. C, D) C. cristatulus new species, holotype. E, F) C. granulocristatus new species, holotype. G, H) C. poblanus new species, holotype. All photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.4564899" httpUri="https://zenodo.org/record/4564899/files/figure.png" pageId="7" pageNumber="6">Fig. 4 A, B</figureCitation>
;
<figureCitation id="13363FFCFF95884DFEE0604CBF5BA36E" box="[321,383,1296,1323]" captionStart="Figure 5" captionStartId="9.[193,269,1560,1586]" captionTargetBox="[337,1262,213,1529]" captionTargetId="figure-122@9.[337,1262,213,1529]" captionTargetPageId="9" captionText="Figure 5. Corthylus cristatus, new species, female holotype. A) Dorsal habitus. B) Lateral habitus. C) Frons and antenna. D) Declivity. Note: anterior portion of pronotum is abnormal, whether from damage to the specimen or abnormal development. All photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.4564901" httpUri="https://zenodo.org/record/4564901/files/figure.png" pageId="7" pageNumber="6">Fig. 5</figureCitation>
A–D
</paragraph>
</subSubSection>
<subSubSection id="C31770F2FF958840FF616066BF4DA441" lastPageId="10" lastPageNumber="9" pageId="7" pageNumber="6" type="diagnosis">
<paragraph id="8BB22379FF95884DFF616066BA56A0F1" blockId="7.[191,1441,1339,1927]" pageId="7" pageNumber="6">
<emphasis id="B979FF6BFF95884DFF616066BF18A310" bold="true" box="[192,316,1339,1365]" pageId="7" pageNumber="6">Diagnosis.</emphasis>
This species,
<taxonomicName id="4C0D58FAFF95884DFE7D6061BC55A313" authority="Atkinson, 2020" authorityName="Atkinson" authorityYear="2020" box="[476,625,1340,1366]" class="Insecta" family="Curculionidae" genus="Corthylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="cristatulus" status="sp. nov.">
<emphasis id="B979FF6BFF95884DFE7D6061BFCAA313" box="[476,494,1340,1366]" italics="true" pageId="7" pageNumber="6">C</emphasis>
.
<emphasis id="B979FF6BFF95884DFE5C6061BC55A313" box="[509,625,1340,1366]" italics="true" pageId="7" pageNumber="6">cristatulus</emphasis>
</taxonomicName>
<emphasis id="B979FF6BFF95884DFDD06061BC51A313" box="[625,629,1340,1366]" italics="true" pageId="7" pageNumber="6">,</emphasis>
<emphasis id="B979FF6BFF95884DFDDF6061BD2DA310" bold="true" box="[638,777,1339,1366]" pageId="7" pageNumber="6">
<taxonomicNameLabel id="A24A4210FF95884DFDDF6061BD2DA310" box="[638,777,1339,1366]" pageId="7" pageNumber="6" rank="species">new species</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C0D58FAFF95884DFCEA6061BA3BA313" authority="Atkinson, 2020" authorityName="Atkinson" authorityYear="2020" box="[843,1055,1340,1366]" class="Insecta" family="Curculionidae" genus="Corthylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="granulocristatus" status="sp. nov.">
<emphasis id="B979FF6BFF95884DFCEA6061BD79A313" box="[843,861,1340,1366]" italics="true" pageId="7" pageNumber="6">C</emphasis>
.
<emphasis id="B979FF6BFF95884DFCCD6061BA3BA313" box="[876,1055,1340,1366]" italics="true" pageId="7" pageNumber="6">granulocristatus</emphasis>
</taxonomicName>
<emphasis id="B979FF6BFF95884DFBBE6061BA07A313" box="[1055,1059,1340,1366]" italics="true" pageId="7" pageNumber="6">,</emphasis>
<emphasis id="B979FF6BFF95884DFB8D6061BA92A310" bold="true" box="[1068,1206,1339,1366]" pageId="7" pageNumber="6">
<taxonomicNameLabel id="A24A4210FF95884DFB8D6061BA92A310" box="[1068,1206,1339,1366]" pageId="7" pageNumber="6" rank="species">new species</taxonomicNameLabel>
</emphasis>
, are quite similar in general appearance (
<figureCitation id="13363FFCFF95884DFE026002BFC7A33C" box="[419,483,1375,1401]" captionStart="Figure 4" captionStartId="8.[192,268,1672,1698]" captionTargetBox="[305,1326,213,1641]" captionTargetId="figure-10@8.[305,1326,213,1641]" captionTargetPageId="8" captionText="Figure 4. Species of Corthylus with acute spines on declivity or with interstriae 3 curved and serrate (2 of 2). A, B) C. cristatus new species, holotype. C, D) C. cristatulus new species, holotype. E, F) C. granulocristatus new species, holotype. G, H) C. poblanus new species, holotype. All photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.4564899" httpUri="https://zenodo.org/record/4564899/files/figure.png" pageId="7" pageNumber="6">Fig. 4</figureCitation>
). They can be distinguished from each other by body size, size and density of punctures on the declivity, and by variations in the size of the elevated, serrate crest on the declivity. This crest is the sharply elevated third interstriae. This crest is strongly curved towards the median line in the middle of the declivity, strongly reducing the width of interstriae 1 and 2, but in “normal” position at the base and apex of the declivity. The posterolateral area of the declivity is strongly elevated (couplet 1,
<bibRefCitation id="EF9C5E88FF95884DFC5F60B6BAA0A040" author="Wood SL" box="[1022,1156,1515,1541]" pageId="7" pageNumber="6" pagination="1 - 1356" refId="ref9695" refString="Wood SL. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Naturalist Memoirs 6: 1 - 1356." type="journal article" year="1982">Wood 1982</bibRefCitation>
). The combination of the curved, serrate declivital crest, elevated posterolateral line on the declivity which is otherwise convex, and the deep confused punctures on the elytra are unique to this group among the known species of the genus.
<taxonomicName id="4C0D58FAFF95884DFA94636CBF38A02B" authorityName="Atkinson" authorityYear="2020" class="Insecta" family="Curculionidae" genus="Corthylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="B979FF6BFF95884DFA94636CBF38A02B" italics="true" pageId="7" pageNumber="6">Corthylus cristatus</emphasis>
</taxonomicName>
is distinguished from the other two by the closely spaced, large confused punctures on the elytral disc, separated by their own diameter. The declivital crest on interstriae 3 is the most pronounced in height and curvature to the point that interstriae 2 are basically obsolete in the middle of the declivity.
</paragraph>
<paragraph id="8BB22379FF95884DFEA163E0BC07A158" blockId="7.[191,1441,1339,1927]" pageId="7" pageNumber="6">
<emphasis id="B979FF6BFF95884DFEA163E0BF7FA092" bold="true" box="[256,347,1725,1751]" pageId="7" pageNumber="6">Female.</emphasis>
Length:
<quantity id="4CF58E9CFF95884DFE6063E0BC01A092" box="[449,549,1725,1751]" metricMagnitude="-3" metricUnit="m" metricValue="3.12" pageId="7" pageNumber="6" unit="mm" value="3.12">3.12 mm</quantity>
(2.97–3.33), maximum width:
<quantity id="4CF58E9CFF95884DFC2763E0BDCFA092" box="[902,1003,1725,1751]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="7" pageNumber="6" unit="mm" value="1.23">1.23 mm</quantity>
(1.13–1.27); length of elytra:
<quantity id="4CF58E9CFF95884DFA9A63E0BB84A092" box="[1339,1440,1725,1751]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="7" pageNumber="6" unit="mm" value="1.23">1.23 mm</quantity>
(1.67–1.90); length to width: 2.5; elytral length/total length: 0.58; elytral length/width: 1.47. (n = 4,
<specimenCount id="9D0BE8F0FF95884DFA8E63BDBBACA0BF" box="[1327,1416,1760,1786]" pageId="7" pageNumber="6" type="male">3 males</specimenCount>
,
<specimenCount id="9D0BE8F0FF95884DFA3563BDBF35A15B" pageId="7" pageNumber="6" type="female">1 female</specimenCount>
). Color uniformly black.
</paragraph>
<paragraph id="8BB22379FF958843FEA1627ABC82A09B" blockId="7.[191,1441,1339,1927]" lastBlockId="9.[191,1440,1698,1927]" lastPageId="9" lastPageNumber="8" pageId="7" pageNumber="6">
Frons evenly concave from epistoma to vertex. Lower surface with widely spaced, low granules, each associated with a short seta. Setae becoming denser on sides and upper portion of frons, marginal setae only slightly longer than those in center, not forming incurved margins. Area just above epistoma impunctate. Antenna slightly asymmetrical, 1
<superScript id="7C788E31FF9B8843FED063FFBF58A0F5" attach="left" box="[369,380,1698,1712]" fontSize="6" pageId="9" pageNumber="8">st</superScript>
and 2
<superScript id="7C788E31FF9B8843FE6063FFBFF6A0F5" attach="left" box="[449,466,1698,1712]" fontSize="6" pageId="9" pageNumber="8">nd</superScript>
sutures indicated by external grooves, 1
<superScript id="7C788E31FF9B8843FC3063FFBDB8A0F5" attach="left" box="[913,924,1698,1712]" fontSize="6" pageId="9" pageNumber="8">st</superScript>
partly septate. Cirrus very short, longest setae less than 1/2 the width of the antennal club.
</paragraph>
<caption id="DF7273F1FF9A8842FF6163D5BA8AA0A0" ID-DOI="http://doi.org/10.5281/zenodo.4564899" ID-Zenodo-Dep="4564899" httpUri="https://zenodo.org/record/4564899/files/figure.png" pageId="8" pageNumber="7" startId="8.[192,268,1672,1698]" targetBox="[305,1326,213,1641]" targetPageId="8">
<paragraph id="8BB22379FF9A8842FF6163D5BA8AA0A0" blockId="8.[192,1440,1672,1766]" pageId="8" pageNumber="7">
<emphasis id="B979FF6BFF9A8842FF6163D5BF0EA0E7" bold="true" box="[192,298,1672,1698]" pageId="8" pageNumber="7">Figure 4.</emphasis>
Species of
<taxonomicName id="4C0D58FAFF9A8842FE0D63D5BC33A0E7" authorityName="Erichson" authorityYear="1834" box="[428,535,1672,1698]" class="Insecta" family="Curculionidae" genus="Corthylus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B979FF6BFF9A8842FE0D63D5BC33A0E7" box="[428,535,1672,1698]" italics="true" pageId="8" pageNumber="7">Corthylus</emphasis>
</taxonomicName>
with acute spines on declivity or with interstriae 3 curved and serrate (2 of 2).
<emphasis id="B979FF6BFF9A8842FF6163F4BED8A086" bold="true" box="[192,252,1705,1731]" pageId="8" pageNumber="7">A, B)</emphasis>
<taxonomicName id="4C0D58FAFF9A8842FEA263F7BF5AA081" authority="Atkinson, 2020" authorityName="Atkinson" authorityYear="2020" box="[259,382,1706,1732]" class="Insecta" family="Curculionidae" genus="Corthylus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="cristatus" status="sp. nov.">
<emphasis id="B979FF6BFF9A8842FEA263F7BF31A081" box="[259,277,1706,1732]" italics="true" pageId="8" pageNumber="7">C</emphasis>
.
<emphasis id="B979FF6BFF9A8842FE8363F6BF5AA081" box="[290,382,1707,1732]" italics="true" pageId="8" pageNumber="7">cristatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A4210FF9A8842FE2563F7BC2EA081" box="[388,522,1706,1732]" pageId="8" pageNumber="7" rank="species">new species</taxonomicNameLabel>
, holotype.
<emphasis id="B979FF6BFF9A8842FD2563F4BC9DA081" bold="true" box="[644,697,1705,1732]" pageId="8" pageNumber="7">C, D</emphasis>
)
<taxonomicName id="4C0D58FAFF9A8842FD6863F7BD7EA081" authority="Atkinson, 2020" authorityName="Atkinson" authorityYear="2020" box="[713,858,1706,1732]" class="Insecta" family="Curculionidae" genus="Corthylus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="cristatulus" status="sp. nov.">
<emphasis id="B979FF6BFF9A8842FD6863F7BCFFA081" box="[713,731,1706,1732]" italics="true" pageId="8" pageNumber="7">C</emphasis>
.
<emphasis id="B979FF6BFF9A8842FD4963F7BD7EA081" box="[744,858,1706,1732]" italics="true" pageId="8" pageNumber="7">cristatulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A4210FF9A8842FCC163F7BDC2A081" box="[864,998,1706,1732]" pageId="8" pageNumber="7" rank="species">new species</taxonomicNameLabel>
, holotype.
<emphasis id="B979FF6BFF9A8842FBFE63F7BAB2A086" bold="true" box="[1119,1174,1705,1732]" pageId="8" pageNumber="7">E, F)</emphasis>
<taxonomicName id="4C0D58FAFF9A8842FB3D63F7BB4FA081" authority="Atkinson, 2020" authorityName="Atkinson" authorityYear="2020" box="[1180,1387,1706,1732]" class="Insecta" family="Curculionidae" genus="Corthylus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="granulocristatus" status="sp. nov.">
<emphasis id="B979FF6BFF9A8842FB3D63F7BA8AA081" box="[1180,1198,1706,1732]" italics="true" pageId="8" pageNumber="7">C</emphasis>
.
<emphasis id="B979FF6BFF9A8842FB1A63F7BB4FA081" box="[1211,1387,1706,1732]" italics="true" pageId="8" pageNumber="7">granulocristatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A4210FF9A8842FAD063F7BF35A0A0" pageId="8" pageNumber="7" rank="species">new species</taxonomicNameLabel>
, holotype.
<emphasis id="B979FF6BFF9A8842FE2B6396BFEFA0A0" bold="true" box="[394,459,1739,1765]" pageId="8" pageNumber="7">G, H)</emphasis>
<taxonomicName id="4C0D58FAFF9A8842FE736391BC77A0A0" authority="Atkinson, 2020" authorityName="Atkinson" authorityYear="2020" box="[466,595,1739,1766]" class="Insecta" family="Curculionidae" genus="Corthylus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="poblanus" status="sp. nov.">
<emphasis id="B979FF6BFF9A8842FE736391BFC0A0A3" box="[466,484,1740,1766]" italics="true" pageId="8" pageNumber="7">C</emphasis>
.
<emphasis id="B979FF6BFF9A8842FE516396BC77A0A0" box="[496,595,1739,1765]" italics="true" pageId="8" pageNumber="7">poblanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A4210FF9A8842FDF86391BCFBA0A0" box="[601,735,1739,1766]" pageId="8" pageNumber="7" rank="species">new species</taxonomicNameLabel>
, holotype. All photographs by the author.
</paragraph>
</caption>
<caption id="DF7273F1FF9B8843FF606345BD01A030" ID-DOI="http://doi.org/10.5281/zenodo.4564901" ID-Zenodo-Dep="4564901" httpUri="https://zenodo.org/record/4564901/files/figure.png" pageId="9" pageNumber="8" startId="9.[193,269,1560,1586]" targetBox="[337,1262,213,1529]" targetPageId="9">
<paragraph id="8BB22379FF9B8843FF606345BD01A030" blockId="9.[191,1440,1560,1653]" pageId="9" pageNumber="8">
<emphasis id="B979FF6BFF9B8843FF606345BF0DA077" bold="true" box="[193,297,1560,1586]" pageId="9" pageNumber="8">Figure 5.</emphasis>
<taxonomicName id="4C0D58FAFF9B8843FE916345BFDAA077" authority="Atkinson, 2020" authorityName="Atkinson" authorityYear="2020" box="[304,510,1560,1586]" class="Insecta" family="Curculionidae" genus="Corthylus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="cristatus" status="sp. nov.">
<emphasis id="B979FF6BFF9B8843FE916345BFDAA077" box="[304,510,1560,1586]" italics="true" pageId="9" pageNumber="8">Corthylus cristatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24A4210FF9B8843FDAD6344BCB7A077" box="[524,659,1560,1587]" pageId="9" pageNumber="8" rank="species">new species</taxonomicNameLabel>
, female holotype.
<emphasis id="B979FF6BFF9B8843FCC16345BD5AA077" bold="true" box="[864,894,1560,1586]" pageId="9" pageNumber="8">A)</emphasis>
Dorsal habitus.
<emphasis id="B979FF6BFF9B8843FB976345BA75A077" bold="true" box="[1078,1105,1560,1586]" pageId="9" pageNumber="8">B)</emphasis>
Lateral habitus.
<emphasis id="B979FF6BFF9B8843FAAD6345BB0DA077" bold="true" box="[1292,1321,1560,1586]" pageId="9" pageNumber="8">C)</emphasis>
Frons and antenna.
<emphasis id="B979FF6BFF9B8843FE876364BF61A016" bold="true" box="[294,325,1593,1619]" pageId="9" pageNumber="8">D)</emphasis>
Declivity. Note: anterior portion of pronotum is abnormal, whether from damage to the specimen or abnormal development. All photographs by the author.
</paragraph>
</caption>
<paragraph id="8BB22379FF9B8843FEA163BBBB84A164" blockId="9.[191,1440,1698,1927]" pageId="9" pageNumber="8">Anterior margin of pronotum broadly rounded, unarmed. Asperities on anterior slope not elevated, weakly developed, obsolete at summit. Pronotal disc shining, with deep punctures, separated by about 3× their diameters.</paragraph>
<paragraph id="8BB22379FF9B8840FF5F6274BC3DA7DC" blockId="9.[191,1440,1698,1927]" lastBlockId="10.[191,1441,214,409]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">Striae and interstriae indistinct on elytral disc, all punctures similar in size and completely confused, those at base of elytral about twice as large as punctures on pronotal disc and separated by about twice their diameters. Declivity abruptly curved in profile. Elevated postero-lateral crest at elytral apex weakly developed, partly obscured by deep confused punctures. Striae and interstriae indistinct on declivity, but short, yellow setae present on odd-numbered interstriae. Interstriae 1 slightly costate, without or with very weakly developed granules. Interstriae 2 impressed, extremely narrow in declivital face. Interstriae 3 strongly elevated, with contiguous, pointed granules, giving a serrate impression, strongly curving towards declivital suture in middle, curving outwards again towards apex, serrate elevation not reaching bottom 1/3 of declivity. Much smaller granules irregularly spaced exterior to interstriae 3.</paragraph>
<paragraph id="8BB22379FF988840FF6164FBBF4DA441" blockId="10.[192,1440,422,516]" pageId="10" pageNumber="9">
<emphasis id="B979FF6BFF988840FF6164FBBF25A785" bold="true" box="[192,257,422,448]" pageId="10" pageNumber="9">Male.</emphasis>
Similar to female except for frons and antenna. Frons evenly convex, shining, with widely separated, deep punctures, vestiture absent. A weak transverse impression above epistoma. Antenna suboval, without longer, marginal setae.
</paragraph>
</subSubSection>
<subSubSection id="C31770F2FF988840FF61674CBB72A491" pageId="10" pageNumber="9" type="materials_examined">
<paragraph id="8BB22379FF988840FF61674CBB72A491" blockId="10.[192,1441,529,797]" pageId="10" pageNumber="9">
<emphasis id="B979FF6BFF988840FF61674CBF49A46E" bold="true" box="[192,365,529,555]" pageId="10" pageNumber="9">Type material.</emphasis>
<materialsCitation id="3B652924FF988840FED6674CBCFCA408" ID-GBIF-Occurrence="3046457318" collectingDate="2016-03-28" collectionCode="CNIN" country="Mexico" location="El Santuario" pageId="10" pageNumber="9" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<typeStatus id="54B69DDBFF988840FED6674CBFFBA46E" box="[375,479,529,555]" pageId="10" pageNumber="9" type="holotype">Holotype</typeStatus>
female.
<collectingCountry id="F31A63E9FF988840FDE7674CBCB8A46E" box="[582,668,529,555]" name="Mexico" pageId="10" pageNumber="9">Mexico</collectingCountry>
:
<collectingRegion id="49C9ED9BFF988840FD0B674CBD35A46E" box="[682,785,529,555]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="10" pageNumber="9">Veracruz</collectingRegion>
,
<collectorName id="26F846AFFF988840FC81674CBD8BA46E" box="[800,943,529,555]" pageId="10" pageNumber="9">
<location id="8ED275A2FF988840FC81674CBD8BA46E" LSID="urn:lsid:plazi:treatment:03A4926FFF958840FF6161B6BAA2A501:8ED275A2FF988840FC81674CBD8BA46E" box="[800,943,529,555]" country="Mexico" name="El Santuario" pageId="10" pageNumber="9" stateProvince="Veracruz de Ignacio de la Llave">El Santuario</location>
</collectorName>
, campus INECOL,
<date id="FFB305B9FF988840FB34674CBB3DA46E" box="[1173,1305,529,555]" pageId="10" pageNumber="9" value="2016-03-28">
<collectingDate id="EFF7FC51FF988840FB34674CBB3DA46E" box="[1173,1305,529,555]" pageId="10" pageNumber="9" value="2016-03-28">28-III-2016</collectingDate>
</date>
,
<collectingMethod id="524C5B6EFF988840FA86674CBB85A46E" box="[1319,1441,529,555]" pageId="10" pageNumber="9">bottle trap</collectingMethod>
with ethanol,
<collectorName id="26F846AFFF988840FEFD676EBFFCA408" box="[348,472,563,589]" pageId="10" pageNumber="9">L.A. Ibarra</collectorName>
and
<collectorName id="26F846AFFF988840FDB2676EBC73A408" box="[531,599,563,589]" pageId="10" pageNumber="9">A. Gil</collectorName>
V. (
<collectionCode id="ED1CBBBCFF988840FD24676EBCF5A408" box="[645,721,563,589]" country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="10" pageNumber="9">CNIN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B652924FF988840FD45676EBCF3A42A" ID-GBIF-Occurrence="3046457309" collectingDate="2016-05-09" collectionCode="CNIN" collectorName="El Santuario &amp; A. Gil" country="Mexico" location="El Santuario" pageId="10" pageNumber="9" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="allotype">
<typeStatus id="54B69DDBFF988840FD45676EBD60A408" box="[740,836,563,589]" pageId="10" pageNumber="9" type="allotype">Allotype</typeStatus>
male.
<collectingRegion id="49C9ED9BFF988840FC31676EBDD3A408" box="[912,1015,563,589]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="10" pageNumber="9">Veracruz</collectingRegion>
,
<collectorName id="26F846AFFF988840FBA2676EBAABA408" box="[1027,1167,563,589]" pageId="10" pageNumber="9">
<location id="8ED275A2FF988840FBA2676EBAABA408" LSID="urn:lsid:plazi:treatment:03A4926FFF958840FF6161B6BAA2A501:8ED275A2FF988840FBA2676EBAABA408" box="[1027,1167,563,589]" country="Mexico" name="El Santuario" pageId="10" pageNumber="9" stateProvince="Veracruz de Ignacio de la Llave">El Santuario</location>
</collectorName>
, campus INECOL,
<date id="FFB305B9FF988840FACC676EBEDCA42B" pageId="10" pageNumber="9" value="2016-05-09">
<collectingDate id="EFF7FC51FF988840FACC676EBEDCA42B" pageId="10" pageNumber="9" value="2016-05-09">9-V- 2016</collectingDate>
</date>
,
<collectingMethod id="524C5B6EFF988840FEA26709BF53A42A" box="[259,375,596,623]" pageId="10" pageNumber="9">bottle trap</collectingMethod>
with ethanol,
<collectorName id="26F846AFFF988840FDB76708BC7EA42B" box="[534,602,596,623]" pageId="10" pageNumber="9">A. Gil</collectorName>
V. (
<collectionCode id="ED1CBBBCFF988840FD256708BCEBA42A" box="[644,719,597,623]" country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="10" pageNumber="9">CNIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B652924FF988840FD436708BA3AA4D5" ID-GBIF-Occurrence="3046457303" collectingDate="2016-02-03" collectionCode="IEXA, UTIC" collectorName="L. A. Ibarra &amp; A. Gil" country="Mexico" elevation="1" location="El Santuario" pageId="10" pageNumber="9" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<typeStatus id="54B69DDBFF988840FD436708BD75A42A" box="[738,849,597,623]" pageId="10" pageNumber="9" type="paratype">Paratypes</typeStatus>
.
<collectingRegion id="49C9ED9BFF988840FCFA6708BDE6A42A" box="[859,962,597,623]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="10" pageNumber="9">Veracruz</collectingRegion>
,
<location id="8ED275A2FF988840FC6D6709BA73A42B" LSID="urn:lsid:plazi:treatment:03A4926FFF958840FF6161B6BAA2A501:8ED275A2FF988840FC6D6709BA73A42B" box="[972,1111,596,622]" country="Mexico" name="El Santuario" pageId="10" pageNumber="9" stateProvince="Veracruz de Ignacio de la Llave">El Santuario</location>
, campus INECOL,
<date id="FFB305B9FF988840FA8E6709BBB8A42B" box="[1327,1436,596,622]" pageId="10" pageNumber="9" value="2016-02-03">
<collectingDate id="EFF7FC51FF988840FA8E6709BBB8A42B" box="[1327,1436,596,622]" pageId="10" pageNumber="9" value="2016-02-03">3-II-2016</collectingDate>
</date>
,
<collectingMethod id="524C5B6EFF988840FF61672BBF13A4D4" box="[192,311,630,657]" pageId="10" pageNumber="9">bottle trap</collectingMethod>
with ethanol,
<collectorName id="26F846AFFF988840FE43672ABC44A4D5" box="[482,608,630,657]" pageId="10" pageNumber="9">L.A. Ibarra</collectorName>
and
<collectorName id="26F846AFFF988840FD3F672ABCC1A4D5" box="[670,741,630,657]" pageId="10" pageNumber="9">A. Gil</collectorName>
V. (
<collectionCode id="ED1CBBBCFF988840FCB7672BBD7EA4D5" box="[790,858,630,656]" pageId="10" pageNumber="9">IEXA</collectionCode>
,
<quantity id="4CF58E9CFF988840FCC9672BBDAAA4D5" box="[872,910,630,656]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="9" unit="m" value="1.0">
<elevation id="0020C44AFF988840FCC9672BBDAAA4D5" box="[872,910,630,656]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="9" unit="m" value="1.0">1m</elevation>
</quantity>
,
<collectionCode id="ED1CBBBCFF988840FC3C672BBDC5A4D5" box="[925,993,630,656]" pageId="10" pageNumber="9">UTIC</collectionCode>
,
<quantity id="4CF58E9CFF988840FC4E672BBA32A4D5" box="[1007,1046,630,656]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="9" unit="m" value="1.0">
<elevation id="0020C44AFF988840FC4E672BBA32A4D5" box="[1007,1046,630,656]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="9" unit="m" value="1.0">1m</elevation>
</quantity>
)
</materialsCitation>
;
<materialsCitation id="3B652924FF988840FB8F672BBBB8A4D4" ID-GBIF-Occurrence="3046457304" box="[1070,1436,630,657]" collectorName="Union Juarez" country="Mexico" location="Chiapas" pageId="10" pageNumber="9" specimenCount="1" stateProvince="Chiapas" typeStatus="paratype">
<collectingRegion id="49C9ED9BFF988840FB8F672BBAAFA4D5" box="[1070,1163,630,656]" country="Mexico" name="Chiapas" pageId="10" pageNumber="9">Chiapas</collectingRegion>
:
<collectingRegion id="49C9ED9BFF988840FB39672BBAD1A4D5" box="[1176,1269,630,656]" country="Mexico" name="Chiapas" pageId="10" pageNumber="9">Chiapas</collectingRegion>
,
<collectorName id="26F846AFFF988840FAA2672BBBB8A4D4" box="[1283,1436,630,657]" pageId="10" pageNumber="9">Union Juarez</collectorName>
</materialsCitation>
;
<geoCoordinate id="EE3945BEFF988840FF6167C5BF12A4F7" box="[192,310,664,691]" degrees="15.1597" direction="north" orientation="latitude" pageId="10" pageNumber="9" precision="5" value="15.1597">15.1597°N</geoCoordinate>
,
<geoCoordinate id="EE3945BEFF988840FE9867C5BFB7A4F7" box="[313,403,664,690]" degrees="92.17" direction="east" orientation="longitude" pageId="10" pageNumber="9" precision="555" value="92.17">92.17°E</geoCoordinate>
;
<date id="FFB305B9FF988840FE0067C5BFDBA4F7" box="[417,511,664,690]" pageId="10" pageNumber="9" value="2013-04">IV-2013</date>
, cloud forest with pines and coffee;
<materialsCitation id="3B652924FF988840FC3C67C5BF04A490" ID-GBIF-Occurrence="3046457315" collectionCode="UFFE" country="Mexico" location="Francisco Infante" pageId="10" pageNumber="9" specimenCount="1" stateProvince="Chiapas" typeStatus="paratype">
<location id="8ED275A2FF988840FC3C67C5BA40A4F7" LSID="urn:lsid:plazi:treatment:03A4926FFF958840FF6161B6BAA2A501:8ED275A2FF988840FC3C67C5BA40A4F7" box="[925,1124,664,690]" country="Mexico" name="Francisco Infante" pageId="10" pageNumber="9" stateProvince="Chiapas">Francisco Infante</location>
, UFFE:11049 (
<collectionCode id="ED1CBBBCFF988840FAAF67C5BB71A4F7" box="[1294,1365,664,690]" pageId="10" pageNumber="9">UFFE</collectionCode>
, 2 f in ethanol)
</materialsCitation>
;
<materialsCitation id="3B652924FF988840FE8B67E7BB76A491" ID-GBIF-Occurrence="3046457314" box="[298,1362,698,725]" collectingDate="1990-04-21" collectionCode="TAMU" collectorName="N. San Augustin &amp; J. E. Wappes" country="Guatemala" elevation="1" location="El Progreso" pageId="10" pageNumber="9" specimenCount="1" stateProvince="El Progreso" typeStatus="paratype">
<collectingCountry id="F31A63E9FF988840FE8B67E7BF82A491" box="[298,422,698,724]" name="Guatemala" pageId="10" pageNumber="9">Guatemala</collectingCountry>
:
<collectingRegion id="49C9ED9BFF988840FE1167E7BC17A491" box="[432,563,698,724]" country="Guatemala" name="El Progreso" pageId="10" pageNumber="9">El Progreso</collectingRegion>
,
<quantity id="4CF58E9CFF988840FD9F67E7BCA0A491" box="[574,644,698,724]" metricMagnitude="4" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="9" unit="km" value="29.0">29 km</quantity>
<collectorName id="26F846AFFF988840FD2B67E7BD60A491" box="[650,836,698,724]" pageId="10" pageNumber="9">N. San Augustín</collectorName>
,
<date id="FFB305B9FF988840FCEF67E7BDF4A491" box="[846,976,698,724]" pageId="10" pageNumber="9" value="1990-04-21">
<collectingDate id="EFF7FC51FF988840FCEF67E7BDF4A491" box="[846,976,698,724]" pageId="10" pageNumber="9" value="1990-04-21">
21-IV-
<quantity id="4CF58E9CFF988840FC3667E7BDF4A491" box="[919,976,698,724]" metricMagnitude="6" metricUnit="m" metricValue="1.99065" pageId="10" pageNumber="9" unit="m" value="1990650.0">1990</quantity>
</collectingDate>
</date>
, 650 m,
<collectorName id="26F846AFFF988840FB8C67E7BA94A491" box="[1069,1200,698,724]" pageId="10" pageNumber="9">J.E. Wappes</collectorName>
(
<collectionCode id="ED1CBBBCFF988840FB1C67E7BB34A491" box="[1213,1296,698,724]" pageId="10" pageNumber="9">TAMU</collectionCode>
,
<quantity id="4CF58E9CFF988840FABA67E7BB6CA491" box="[1307,1352,698,724]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="9" unit="m" value="1.0">
<elevation id="0020C44AFF988840FABA67E7BB6CA491" box="[1307,1352,698,724]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="9" unit="m" value="1.0">1 m</elevation>
</quantity>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31770F2FF988840FF6167BCBCE8A558" pageId="10" pageNumber="9" type="discussion">
<paragraph id="8BB22379FF988840FF6167BCBCE8A558" blockId="10.[192,1441,529,797]" pageId="10" pageNumber="9">
<emphasis id="B979FF6BFF988840FF6167BCBF6CA4BE" bold="true" box="[192,328,737,763]" pageId="10" pageNumber="9">Comments.</emphasis>
The anterior portion of pronotum in the female
<typeStatus id="54B69DDBFF988840FCC967BCBDEEA4BE" box="[872,970,737,763]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
in
<figureCitation id="13363FFCFF988840FC4F67BCBA0FA4B9" box="[1006,1067,737,764]" captionStart="Figure 5" captionStartId="9.[193,269,1560,1586]" captionTargetBox="[337,1262,213,1529]" captionTargetId="figure-122@9.[337,1262,213,1529]" captionTargetPageId="9" captionText="Figure 5. Corthylus cristatus, new species, female holotype. A) Dorsal habitus. B) Lateral habitus. C) Frons and antenna. D) Declivity. Note: anterior portion of pronotum is abnormal, whether from damage to the specimen or abnormal development. All photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.4564901" httpUri="https://zenodo.org/record/4564901/files/figure.png" pageId="10" pageNumber="9">Fig. 5</figureCitation>
is abnormal, whether from damage to the specimen or abnormal development.
</paragraph>
</subSubSection>
<subSubSection id="C31770F2FF988840FF616677BAA2A501" box="[192,1158,810,837]" pageId="10" pageNumber="9" type="etymology">
<paragraph id="8BB22379FF988840FF616677BAA2A501" blockId="10.[192,1158,810,837]" box="[192,1158,810,837]" pageId="10" pageNumber="9">
<emphasis id="B979FF6BFF988840FF616677BF60A501" bold="true" box="[192,324,810,836]" pageId="10" pageNumber="9">Etymology.</emphasis>
The name is chosen for the prominent curved crest on the elytral declivity.
</paragraph>
</subSubSection>
</treatment>
</document>
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