treatments-xml/data/03/A4/92/03A4923D7D62AD3915FFFC63F07F4A35.xml

<document id="ECB4879B5A1B0BD064C13CBCCAEB4D2E" ID-DOI="10.5281/zenodo.5312012" ID-ISSN="0374-1036" ID-Zenodo-Dep="5312012" ID-ZooBank="D72D1DC9-87D1-4209-A01F-8185A4D21F88" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630118846529" checkinUser="marcus" docAuthor="Sehnal, Richard, Král, David &amp; Bezděk, Aleš" docDate="2014" docId="03A4923D7D62AD3915FFFC63F07F4A35" docLanguage="en" docName="ActaEntMusNatPra.2014.54.suppl.139-171.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Canuschiza skand Sehnal &amp; Král &amp; Bezděk 2014, sp. nov." docType="treatment" docVersion="4" lastPageNumber="168" masterDocId="FF9DEA457D7AAD241413FFF8F470496C" masterDocTitle="Canuschiza of Socotra Island (Coleoptera: Scarabaeidae: Melolonthinae) Part 1. Canuschiza insularis species group" masterLastPageNumber="171" masterPageNumber="139" pageNumber="163" updateTime="1699158890131" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4656652E232B680B17D99B7ABF5D18B1">Canuschiza of Socotra Island (Coleoptera: Scarabaeidae: Melolonthinae) Part 1. Canuschiza insularis species group</mods:title>
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<mods:namePart id="F20DAC37F92CFA3DC7B88373D4CDE328">Sehnal, Richard</mods:namePart>
<mods:affiliation id="A1D0469DAD789F4E96805F8257C368A6">Czech University of Life Sciences Prague, Faculty of Agrobiology, Food and Natural Resources, Department of Zoology and Fisheries, Kamýcká 129, CZ- 165 21 Praha 6, Czech Republic; e-mail: richard. sehnal @ seznam. cz</mods:affiliation>
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<mods:namePart id="C87E147D1C6891E5FC6A029659243C32">Král, David</mods:namePart>
<mods:affiliation id="82CEEB8BD65504CD52E237D25EE57FD1">Charles University in Prague, Faculty of Science, Department of Zoology, Viničná 7, CZ- 128 43 Praha 2, Czech Republic; e-mail: kral @ natur. cuni. cz</mods:affiliation>
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<mods:namePart id="88FA1B15A150554468DE3B12217331CE">Bezděk, Aleš</mods:namePart>
<mods:affiliation id="8EF02687E9BBADFA5A027325A34BAD40">Biology Centre ASCR, Institute of Entomology, Branišovská 31, CZ- 370 05 České Budějovice, Czech Republic; e-mail: bezdek @ entu. cas. cz</mods:affiliation>
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<subSubSection id="C31770A07D62AD3C15FFFC63F7614ADF" box="[492,785,923,947]" pageId="24" pageNumber="163" type="nomenclature">
<paragraph id="8BB2232B7D62AD3C15FFFC63F7614ADF" blockId="24.[492,785,923,947]" box="[492,785,923,947]" pageId="24" pageNumber="163">
<heading id="D0FA94477D62AD3C15FFFC63F7614ADF" bold="true" box="[492,785,923,947]" centered="true" fontSize="8" level="2" pageId="24" pageNumber="163" reason="3">
<emphasis id="B979FF397D62AD3C15FFFC63F7614ADF" bold="true" box="[492,785,923,947]" pageId="24" pageNumber="163">
<taxonomicName id="4C0D58A87D62AD3C15FFFC63F6C54ADF" authority="Sehnal &amp; Král &amp; Bezděk, 2014" authorityName="Sehnal &amp; Král &amp; Bezděk" authorityYear="2014" box="[492,693,923,947]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="163" phylum="Arthropoda" rank="species" species="skand" status="sp. nov.">
<emphasis id="B979FF397D62AD3C15FFFC63F6C54ADF" bold="true" box="[492,693,923,947]" italics="true" pageId="24" pageNumber="163">Canuschiza skand</emphasis>
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<taxonomicNameLabel id="A24A42427D62AD3C16AFFC64F7614ADF" box="[700,785,924,947]" pageId="24" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C31770A07D62AD3C1630FC42F6AA4AA1" box="[547,730,954,973]" pageId="24" pageNumber="163" type="description">
<paragraph id="8BB2232B7D62AD3C1630FC42F6AA4AA1" blockId="24.[547,730,954,973]" box="[547,730,954,973]" pageId="24" pageNumber="163">
(
<figureCitation id="13363FAE7D62AD3C163AFC42F61F4AA1" box="[553,623,954,973]" captionStart="Figs 1" captionStartId="2.[148,185,1540,1559]" captionTargetBox="[237,1038,641,1515]" captionTargetId="figure-123@2.[232,1044,633,1522]" captionTargetPageId="2" captionText="Figs 1A–D. Canuschiza Lacroix, 1999:A–B – head, dorsal view; C–D – metathoracic wing.A – Canuschiza insularis Lacroix, 1999, holotype; B – C. minuta Lacroix, 1999, holotype; C – C. hagher sp. nov., holotype; D – C. skand sp. nov., holotype. Scale bar: 2.0 mm. Figs A, B not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312014" httpUri="https://zenodo.org/record/5312014/files/figure.png" pageId="24" pageNumber="163">Figs 1D</figureCitation>
, 
<figureCitation id="13363FAE7D62AD3C166AFC42F6D74AA1" box="[633,679,954,973]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="24" pageNumber="163">9A–I</figureCitation>
, 
<figureCitation id="13363FAE7D62AD3C16BCFC42F6A44AA1" box="[687,724,954,973]" captionStart="Figs 11" captionStartId="28.[150,187,1487,1506]" captionTargetBox="[159,1117,227,1393]" captionTargetId="figure-11@28.[151,1126,218,1396]" captionTargetPageId="28" captionText="Figs 11A–D. Aedeagus in dorsal (left) and lateral (right) view: A – Canuschiza hagher sp. nov., holotype; B – C. insularis Lacroix, 1999, holotype; C – C. jatropha sp. nov., holotype; D – C. skand sp. nov., holotype. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5312044" httpUri="https://zenodo.org/record/5312044/files/figure.png" pageId="24" pageNumber="163">11D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31770A07D62AD3C1484FC1CF5C94D8B" pageId="24" pageNumber="163" type="materials_examined">
<paragraph id="8BB2232B7D62AD3C1484FC1CF0604A9B" blockId="24.[151,1128,996,1590]" box="[151,1040,996,1015]" pageId="24" pageNumber="163">
<emphasis id="B979FF397D62AD3C1484FC1CF5604A9B" bold="true" box="[151,272,996,1015]" pageId="24" pageNumber="163">
<typeStatus id="54B69D897D62AD3C1484FC1CF4B44A9B" box="[151,196,996,1015]" pageId="24" pageNumber="163">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F31A63BB7D62AD3C1506FC1CF5234A9B" box="[277,339,996,1015]" name="Yemen" pageId="24" pageNumber="163">Yemen</collectingCountry>
, 
<collectingCountry id="F31A63BB7D62AD3C1548FC1CF5A84A9B" box="[347,472,996,1015]" name="Socotra Island" pageId="24" pageNumber="163">Socotra Island</collectingCountry>
, 
<taxonomicName id="4C0D58A87D62AD3C15F3FC1CF6DF4A9B" authority="Mts., Scand Mt." authorityName="Mts., Scand Mt." box="[480,687,996,1015]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="163" phylum="Arthropoda" rank="species" species="hagher">Hagher Mts., Scand Mt.</taxonomicName>
env., 
<quantity id="4CF58ECE7D62AD3C16F3FC1CF7514A9B" box="[736,801,996,1015]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="24" pageNumber="163" unit="m" value="1450.0">1450 m</quantity>
a.s.l., 
<geoCoordinate id="EE3945EC7D62AD3C174AFC1CF7DF4A9B" box="[857,943,996,1015]" degrees="12" direction="north" minutes="34.6" orientation="latitude" pageId="24" pageNumber="163" precision="92" value="12.576667">12°34.6′N</geoCoordinate>
<geoCoordinate id="EE3945EC7D62AD3C17A6FC1CF07C4A9B" box="[949,1036,996,1015]" degrees="54" direction="east" minutes="01.5" orientation="longitude" pageId="24" pageNumber="163" precision="92" value="54.025">54°01.5′E</geoCoordinate>
.
</paragraph>
<paragraph id="8BB2232B7D62AD3C1484FC06F6744DFB" blockId="24.[151,1128,996,1590]" pageId="24" pageNumber="163">
<emphasis id="B979FF397D62AD3C1484FC06F5664D7D" bold="true" box="[151,278,1022,1041]" pageId="24" pageNumber="163">Type material</emphasis>
(
<specimenCount id="9D0BE8A27D62AD3C1531FC06F5EB4D7D" box="[290,411,1022,1041]" pageId="24" pageNumber="163" type="generic">37 specimens</specimenCount>
). 
<materialsCitation id="3B6529767D62AD3C15BAFC06F7D14D2B" collectingDate="2012-06-16" collectingDateMax="2012-06-18" collectingDateMin="2012-06-16" collectorName="Bezdek &amp; J. Hajek &amp; Hula &amp; P. Kment &amp; I. Malenovsky &amp; J. Niedobova &amp; L. Purchart" country="Yemen" elevation="1450" latitude="12.576667" location="YEMEN" longLatPrecision="129" longitude="54.025" pageId="24" pageNumber="163" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B69D897D62AD3C15BAFC06F58C4D7C" box="[425,508,1022,1041]" pageId="24" pageNumber="163" type="holotype">HOLOTYPE</typeStatus>
: J, labelled: ‘ 
<collectingCountry id="F31A63BB7D62AD3C1667FC06F6B64D7D" box="[628,710,1022,1041]" name="Yemen" pageId="24" pageNumber="163">YEMEN</collectingCountry>
, SOCOTRA Island / 
<taxonomicName id="4C0D58A87D62AD3C1770FC06F0444D7D" authority="Mts., SCAND Mt." authorityName="Mts., SCAND Mt." box="[867,1076,1022,1041]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="163" phylum="Arthropoda" rank="species" species="hagher">Hagher Mts., SCAND Mt.</taxonomicName>
env. / montane evergreen woodland / 
<date id="FFB305EB7D62AD3C15B4FBE1F6544D40" box="[423,548,1049,1068]" pageId="24" pageNumber="163" value="2012-06-16" valueMax="2012-06-18" valueMin="2012-06-16">
<collectingDate id="EFF7FC037D62AD3C15B4FBE1F6544D40" box="[423,548,1049,1068]" pageId="24" pageNumber="163" value="2012-06-16" valueMax="2012-06-18" valueMin="2012-06-16">16.-18.vi.2012</collectingDate>
</date>
/ 
<geoCoordinate id="EE3945EC7D62AD3C1625FBE1F6FF4D40" box="[566,655,1049,1068]" degrees="12" direction="north" minutes="34.6" orientation="latitude" pageId="24" pageNumber="163" precision="92" value="12.576667">12°34.6′N</geoCoordinate>
, 
<geoCoordinate id="EE3945EC7D62AD3C168AFBE1F6804D40" box="[665,752,1049,1068]" degrees="54" direction="east" minutes="01.5" orientation="longitude" pageId="24" pageNumber="163" precision="92" value="54.025">54°01.5′E</geoCoordinate>
, 
<quantity id="4CF58ECE7D62AD3C16EAFBE1F74B4D40" box="[761,827,1049,1068]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="24" pageNumber="163" unit="m" value="1450.0">
<elevation id="0020C4187D62AD3C16EAFBE1F74B4D40" box="[761,827,1049,1068]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="24" pageNumber="163" unit="m" value="1450.0">1450 m</elevation>
</quantity>
// SOCOTRA expedition 2012 / J. 
<collectorName id="26F846FD7D62AD3C1484FBCCF4AA4D2B" box="[151,218,1076,1095]" pageId="24" pageNumber="163">Bezděk</collectorName>
, 
<collectorName id="26F846FD7D62AD3C14F0FBCCF55B4D2B" box="[227,299,1076,1095]" pageId="24" pageNumber="163">J. Hájek</collectorName>
, V. 
<collectorName id="26F846FD7D62AD3C155FFBCCF5054D2B" box="[332,373,1076,1095]" pageId="24" pageNumber="163">Hula</collectorName>
/ 
<collectorName id="26F846FD7D62AD3C159BFBCCF5A94D2B" box="[392,473,1076,1095]" pageId="24" pageNumber="163">P. Kment</collectorName>
, 
<collectorName id="26F846FD7D62AD3C15F1FBCCF6114D2B" box="[482,609,1076,1095]" pageId="24" pageNumber="163">I. Malenovský</collectorName>
, 
<collectorName id="26F846FD7D62AD3C167AFBCCF6AB4D2B" box="[617,731,1076,1095]" pageId="24" pageNumber="163">J. Niedobová</collectorName>
&amp; 
<collectorName id="26F846FD7D62AD3C16E4FBCCF7284D2B" box="[759,856,1076,1095]" pageId="24" pageNumber="163">L. Purchart</collectorName>
leg. [p]
</materialsCitation>
’. 
<materialsCitation id="3B6529767D62AD3C17BCFBCCF6644D0D" collectingDate="2012-06-16" collectingDateMax="2012-06-18" collectingDateMin="2012-06-16" collectorName="Bezdek &amp; J. Hajek &amp; Hula &amp; P. Kment &amp; I. Malenovsky &amp; J. Niedobova &amp; L. Purchart" country="Yemen" elevation="1450" latitude="12.576667" location="Nos." longLatPrecision="129" longitude="54.025" pageId="24" pageNumber="163" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
<typeStatus id="54B69D897D62AD3C17BCFBCCF0754D2B" box="[943,1029,1076,1095]" pageId="24" pageNumber="163" type="paratype">Paratypes</typeStatus>
: 
<location id="8ED275F07D62AD3C101DFBCCF0474D2B" LSID="urn:lsid:plazi:treatment:03A4923D7D62AD3915FFFC63F07F4A35:8ED275F07D62AD3C101DFBCCF0474D2B" box="[1038,1079,1076,1095]" name="Nos." pageId="24" pageNumber="163">Nos.</location>
1–34 (30 JJ 
<specimenCount id="9D0BE8A27D62AD3C14F3FBB6F5604D0D" box="[224,272,1096,1121]" pageId="24" pageNumber="163" type="female">4 ♀♀</specimenCount>
), same label data as holotype
</materialsCitation>
; 
<materialsCitation id="3B6529767D62AD3C160DFBB6F5414D10" collectingDate="2009-06-24" collectorName="Z. Hruby" country="Republic of Yemen / Socotra" location="No." municipality="Isl." pageId="24" pageNumber="163" specimenCount="1" typeStatus="paratype">
<location id="8ED275F07D62AD3C160DFBB6F64E4D0D" LSID="urn:lsid:plazi:treatment:03A4923D7D62AD3915FFFC63F07F4A35:8ED275F07D62AD3C160DFBB6F64E4D0D" box="[542,574,1102,1121]" country="Republic of Yemen / Socotra" municipality="Isl." name="No." pageId="24" pageNumber="163">No.</location>
35 (J): ‘ 
<collectingCountry id="F31A63BB7D62AD3C169FFBB6F7FA4D0D" box="[652,906,1102,1121]" isEnumeration="true" pageId="24" pageNumber="163">Republic of Yemen / Socotra</collectingCountry>
<collectingMunicipality id="6BD6B9517D62AD3C1782FBB6F7DC4D0D" box="[913,940,1102,1121]" pageId="24" pageNumber="163">Isl.</collectingMunicipality>
/ SKANT / 
<collectorName id="26F846FD7D62AD3C1004FBB6F0174D0D" box="[1047,1127,1102,1121]" pageId="24" pageNumber="163">Z. Hrubý</collectorName>
lgt. 
<date id="FFB305EB7D62AD3C14AAFB91F57E4D10" box="[185,270,1129,1148]" pageId="24" pageNumber="163" value="2009-06-24">
<collectingDate id="EFF7FC037D62AD3C14AAFB91F57E4D10" box="[185,270,1129,1148]" pageId="24" pageNumber="163" value="2009-06-24">24.6.2009</collectingDate>
</date>
[p]
</materialsCitation>
’; 
<materialsCitation id="3B6529767D62AD3C152CFB91F58C4DFB" collectingDate="2010-06-07" collectingDateMax="2010-06-08" collectingDateMin="2010-06-07" collectorName="V. Hula &amp; J. Niedobova" country="Yemen" location="No." municipality="Isl." pageId="24" pageNumber="163" specimenCount="1" typeStatus="paratype">
<location id="8ED275F07D62AD3C152CFB91F5104D10" LSID="urn:lsid:plazi:treatment:03A4923D7D62AD3915FFFC63F07F4A35:8ED275F07D62AD3C152CFB91F5104D10" box="[319,352,1129,1148]" country="Yemen" municipality="Isl." name="No." pageId="24" pageNumber="163">No.</location>
36 (J): ‘ 
<collectingCountry id="F31A63BB7D62AD3C15BFFB91F58E4D10" box="[428,510,1129,1148]" name="Yemen" pageId="24" pageNumber="163">YEMEN</collectingCountry>
, Socotra 
<collectingMunicipality id="6BD6B9517D62AD3C165EFB91F6194D10" box="[589,617,1129,1148]" pageId="24" pageNumber="163">Isl.</collectingMunicipality>
/ 
<taxonomicName id="4C0D58A87D62AD3C166AFB91F7524D10" authority="Mts., Skant" authorityName="Mts., Skant" box="[633,802,1129,1148]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="163" phylum="Arthropoda" rank="species" species="hagher">Hagher Mts., Skant</taxonomicName>
, / N 12°34,557′, E 054°01,514′ / 
<date id="FFB305EB7D62AD3C102CFB91F4A74DFB" pageId="24" pageNumber="163" value="2010-06-07" valueMax="2010-06-08" valueMin="2010-06-07">
<collectingDate id="EFF7FC037D62AD3C102CFB91F4A74DFB" pageId="24" pageNumber="163" value="2010-06-07" valueMax="2010-06-08" valueMin="2010-06-07">7.-8. vi.2010</collectingDate>
</date>
/ 
<collectorName id="26F846FD7D62AD3C14FBFB7CF5584DFB" box="[232,296,1156,1175]" pageId="24" pageNumber="163">V. Hula</collectorName>
&amp; 
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leg. [p]
</materialsCitation>
’.
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depositories.
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and 
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Nos. 1–6, 34, 
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, 
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Nos. 7–14, 31 in 
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, 
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Nos. 15–21, 32, 
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, 
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Nos. 22, 23 in 
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, 
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Nos. 24, 25 in 
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, 
<collectionCode id="ED1CBBEE7D62AD3C168EFB41F6C64DA0" box="[669,694,1209,1228]" pageId="24" pageNumber="163">PT</collectionCode>
Nos. 26, 27 in 
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, 
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No. 
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, 
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No. 
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<collectionCode id="ED1CBBEE7D62AD3C14DBFB2CF5724D8B" box="[200,258,1236,1255]" pageId="24" pageNumber="163">MLCP</collectionCode>
, 
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No. 
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<collectionCode id="ED1CBBEE7D62AD3C156EFB2CF5C54D8B" box="[381,437,1236,1255]" pageId="24" pageNumber="163">GSCT</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C31770A07D62AD391484FB07F4A94875" lastPageId="29" lastPageNumber="168" pageId="24" pageNumber="163" type="description">
<paragraph id="8BB2232B7D62AD3C1484FB07F56E4CBA" blockId="24.[151,1128,996,1590]" pageId="24" pageNumber="163">
<emphasis id="B979FF397D62AD3C1484FB07F5D54C7A" bold="true" box="[151,421,1279,1302]" pageId="24" pageNumber="163">
Description of 
<typeStatus id="54B69D897D62AD3C1550FB07F5D54C7A" box="[323,421,1279,1302]" pageId="24" pageNumber="163" type="holotype">holotype</typeStatus>
</emphasis>
(J). Body elongate, almost parallel, weakly convex. Dorsal and ventral surface moderately shiny, elytra somewhat alutaceous, chestnut brown, macrosetation pale (
<figureCitation id="13363FAE7D62AD3C14C7FAC7F55B4C3A" box="[212,299,1343,1366]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="24" pageNumber="163">Fig. 9A</figureCitation>
). Dorsal surface of head, pronotum, scutellar shield and elytra covered with white short recumbent scale-like macrosetae, ventral surface of thorax and abdomen with white recumbent scale-like macrosetae. Head appendages and legs covered with yellowish moderately long macrosetae, mixed with a few white recumbent scale-like macrosetae on metafemora.
</paragraph>
<paragraph id="8BB2232B7D62AD3D14A6FA27F5DA489F" blockId="24.[151,1128,996,1590]" lastBlockId="25.[113,1090,220,1587]" lastPageId="25" lastPageNumber="164" pageId="24" pageNumber="163">
Head. Labrum transverse, deeply bilobed, lobes rounded, irregularly and coarsely punctate, covered with long erect macrosetae. Clypeus transverse, anterior margin weakly upturned and shallowly sinuate medially, anterior angles rounded. Fronto-clypeal suture present, forming continuous shallowly bisinuate line. Clypeus and frons densely, irregularly, coarsely punctate, each puncture with short, minute erect macroseta. Occiput sparsely regularly, moderately punctate. Eye-canthus narrow, short, bare. Eyes medium sized (width of both eyes combined approximately the same as maximum width of frons between eyes), distinctly extending beyond eye-canthus. Antenna with ten antennomeres; club with three antennomeres, nearly straight (
<figureCitation id="13363FAE7D63AD3D14C0FE84F55C48FF" box="[211,300,380,403]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="25" pageNumber="164">Fig. 9D</figureCitation>
), distinctly shorter than antennal shaft (antennomeres I–VII combined). Antennomeres I–VII with sparse long macrosetae, club sparsely, shortly macrosetaceous. Terminal maxillary palpomere elongate, apically truncate, approximately as long as palpomeres II and III combined.
</paragraph>
<paragraph id="8BB2232B7D63AD3D1483FE04F79B4BBF" blockId="25.[113,1090,220,1587]" pageId="25" pageNumber="164">
Pronotum transverse, moderately convex, widest approximately at midlength. Basal and lateral borders complete, anterior border missing. Lateral outline regularly rounded, margins in anterior half minutely crenate and macrosetaceous (
<figureCitation id="13363FAE7D63AD3D16ACFDC4F7634B3F" box="[703,787,572,595]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="25" pageNumber="164">Fig. 9E</figureCitation>
). Anterior margin regularly, broadly sinuate. Anterior angles moderately produced, obtuse-angular; posterior angles weakly produced posteriad, approximately rectangular (
<figureCitation id="13363FAE7D63AD3D16DCFD84F7564BFF" box="[719,806,636,659]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="25" pageNumber="164">Fig. 9C</figureCitation>
). Punctation consisting of coarse, umbilicate, almost regularly spaced punctures becoming somewhat denser laterad; each puncture bearing short, narrow, white scale-like, almost recumbent macroseta.
</paragraph>
<paragraph id="8BB2232B7D63AD3D1483FD24F0784B9F" blockId="25.[113,1090,220,1587]" box="[144,1032,732,755]" pageId="25" pageNumber="164">Scutellar shield large, almost equilaterally triangular, sides and apex rounded; bare.</paragraph>
<paragraph id="8BB2232B7D63AD3D1483FD04F6154ADF" blockId="25.[113,1090,220,1587]" pageId="25" pageNumber="164">
Elytra weakly convex, parallel-sided, rounded apically, apical angle approximately rectangular. Striae missing, excepting feebly visible sutural stria. Humeral umbones present, vaguely swollen. Surface finely microsculptured (shagreened), alutaceous; punctation coarse, almost regularly spaced, punctures separated by 1–2 their diameter. Each puncture bearing short, narrow, white scale-like, almost recumbent macroseta. Epipleuron distinct, complete, narrow, bare laterally. Macropterous (
<figureCitation id="13363FAE7D63AD3D1610FC64F6274ADF" box="[515,599,924,947]" captionStart="Figs 1" captionStartId="2.[148,185,1540,1559]" captionTargetBox="[237,1038,641,1515]" captionTargetId="figure-123@2.[232,1044,633,1522]" captionTargetPageId="2" captionText="Figs 1A–D. Canuschiza Lacroix, 1999:A–B – head, dorsal view; C–D – metathoracic wing.A – Canuschiza insularis Lacroix, 1999, holotype; B – C. minuta Lacroix, 1999, holotype; C – C. hagher sp. nov., holotype; D – C. skand sp. nov., holotype. Scale bar: 2.0 mm. Figs A, B not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312014" httpUri="https://zenodo.org/record/5312014/files/figure.png" pageId="25" pageNumber="164">Fig. 1D</figureCitation>
).
</paragraph>
<paragraph id="8BB2232B7D63AD3D1483FC44F07C4D3F" blockId="25.[113,1090,220,1587]" pageId="25" pageNumber="164">
Legs. Femora narrow, shiny, irregularly punctate, macrosetaceous. Protibia narrow, distinctly tridentate, terminal calcar inserted against medial dent. Meso- and metatibia slightly expanded apicad, with two macrosetiferous longitudinal carinae. Upper terminal calcar of metatibia flattened, slightly curved, acute apically, almost two times as long as lower, apically trunctate chisel-shaped metatibial calcar. Claws bifid, with ventrobasal teeth (
<figureCitation id="13363FAE7D63AD3D17BFFBC4F0704D3F" box="[940,1024,1084,1107]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="25" pageNumber="164">Fig. 9B</figureCitation>
).
</paragraph>
<paragraph id="8BB2232B7D63AD3D1483FBA4F4DD4DBF" blockId="25.[113,1090,220,1587]" pageId="25" pageNumber="164">Ventral surface covered with mixture of short, recumbent, white scale-like macrosetae and long, semierect yellowish macrosetae. Pygidium slightly transverse, convex, all around bordered, apically broadly rounded, irregularly covered with coarse macrosetiferous punctures.</paragraph>
<paragraph id="8BB2232B7D63AD3D1483FB24F4984C5F" blockId="25.[113,1090,220,1587]" pageId="25" pageNumber="164">
Male genitalia. Parameres symmetrical, shorter than phallobasis, regularly curved in lateral aspect, rounded apically in dorsal aspect; apex with tuft of long yellowish macrosetae (
<figureCitation id="13363FAE7D63AD3D146AFAE4F4AB4C5F" box="[121,219,1308,1331]" captionStart="Figs 11" captionStartId="28.[150,187,1487,1506]" captionTargetBox="[159,1117,227,1393]" captionTargetId="figure-11@28.[151,1126,218,1396]" captionTargetPageId="28" captionText="Figs 11A–D. Aedeagus in dorsal (left) and lateral (right) view: A – Canuschiza hagher sp. nov., holotype; B – C. insularis Lacroix, 1999, holotype; C – C. jatropha sp. nov., holotype; D – C. skand sp. nov., holotype. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5312044" httpUri="https://zenodo.org/record/5312044/files/figure.png" pageId="25" pageNumber="164">Fig. 11D</figureCitation>
).
</paragraph>
<paragraph id="8BB2232B7D63AD3D1462FAC4F74F4CBF" blockId="25.[113,1090,220,1587]" pageId="25" pageNumber="164">
<emphasis id="B979FF397D63AD3D1462FAC4F5284C3F" bold="true" box="[113,344,1340,1363]" pageId="25" pageNumber="164">Sexual dimorphism.</emphasis>
Female differs from male in the following characters: body slightly broadened posteriad (
<figureCitation id="13363FAE7D63AD3D1540FAA4F5D24C1F" box="[339,418,1372,1395]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="25" pageNumber="164">Fig. 9F</figureCitation>
); antennal club straight, distinctly shorter (
<figureCitation id="13363FAE7D63AD3D1773FAA4F7B14C1F" box="[864,961,1372,1395]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="25" pageNumber="164">Figs 9F,I</figureCitation>
); eyes small (width of both eyes combined shorter than maximum width of frons between eyes) (
<figureCitation id="13363FAE7D63AD3D17F9FA84F40C4CDF" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="25" pageNumber="164">Figs 9F, I</figureCitation>
); scale-like macrosetae of pronotum and elytra short, narrow (
<figureCitation id="13363FAE7D63AD3D170FFA64F7FB4CDF" box="[796,907,1436,1459]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="25" pageNumber="164">Figs 9F,H</figureCitation>
); metatibia more strongly expanded apically; tarsomeres of all legs shorter (
<figureCitation id="13363FAE7D63AD3D16F2FA44F7434CBF" box="[737,819,1468,1491]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="25" pageNumber="164">Fig. 9F</figureCitation>
).
</paragraph>
<paragraph id="8BB2232B7D63AD3D1462FA24F4C54F5F" blockId="25.[113,1090,220,1587]" pageId="25" pageNumber="164">
<emphasis id="B979FF397D63AD3D1462FA24F4804C9F" bold="true" box="[113,240,1500,1523]" pageId="25" pageNumber="164">Variability.</emphasis>
<typeStatus id="54B69D897D63AD3D14EBFA24F5104C9F" box="[248,352,1500,1523]" pageId="25" pageNumber="164" type="paratype">Paratypes</typeStatus>
only slightly vary in length (see measurements), colour and punctation of dorsal surface, length and distribution of scale-like macrosetation of pronotum and elytra.
</paragraph>
<caption id="DF7273A37D60AD3E1484FA11F7D74F5E" ID-DOI="http://doi.org/10.5281/zenodo.5312038" ID-Zenodo-Dep="5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="26" pageNumber="165" startId="26.[151,188,1513,1532]" targetBox="[164,1117,226,1448]" targetPageId="26">
<paragraph id="8BB2232B7D60AD3E1484FA11F7D74F5E" blockId="26.[151,1127,1513,1586]" pageId="26" pageNumber="165">
Figs 9A–I. 
<taxonomicName id="4C0D58A87D60AD3E14E8FA11F5E74C90" authority="Sehnal &amp; Král &amp; Bezděk, 2014" authorityName="Sehnal &amp; Král &amp; Bezděk" authorityYear="2014" box="[251,407,1513,1532]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="165" phylum="Arthropoda" rank="species" species="skand" status="sp. nov.">
<emphasis id="B979FF397D60AD3E14E8FA11F5E74C90" box="[251,407,1513,1532]" italics="true" pageId="26" pageNumber="165">Canuschiza skand</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A42427D60AD3E158DFA11F5944C90" box="[414,484,1513,1532]" pageId="26" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
, dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale.
</paragraph>
</caption>
<caption id="DF7273A37D61AD3F1462FA11F0624F7A" ID-DOI="http://doi.org/10.5281/zenodo.5312042" ID-Zenodo-Dep="5312042" httpUri="https://zenodo.org/record/5312042/files/figure.png" pageId="27" pageNumber="166" startId="27.[113,150,1513,1532]" targetBox="[121,1075,225,1464]" targetPageId="27">
<paragraph id="8BB2232B7D61AD3F1462FA11F0624F7A" blockId="27.[113,1089,1512,1558]" pageId="27" pageNumber="166">
Figs 10A–D.Aedeagus in dorsal (left) and lateral (right) view:A – 
<taxonomicName id="4C0D58A87D61AD3F168EFA10F7404C97" authority="Sehnal &amp; Král &amp; Bezděk, 2014" authorityName="Sehnal &amp; Král &amp; Bezděk" authorityYear="2014" box="[669,816,1512,1531]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="166" phylum="Arthropoda" rank="species" species="adah" status="sp. nov.">
<emphasis id="B979FF397D61AD3F168EFA10F7404C97" box="[669,816,1512,1531]" italics="true" pageId="27" pageNumber="166">Canuschiza adah</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A42427D61AD3F1720FA11F7074C90" box="[819,887,1513,1532]" pageId="27" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
, holotype; B – 
<taxonomicName id="4C0D58A87D61AD3F17E1FA10F0314C97" authority="Sehnal &amp; Král &amp; Bezděk, 2014" authorityName="Sehnal &amp; Král &amp; Bezděk" authorityYear="2014" box="[1010,1089,1512,1531]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="166" phylum="Arthropoda" rank="species" species="croton" status="sp. nov.">
<emphasis id="B979FF397D61AD3F17E1FA10F0314C97" box="[1010,1089,1512,1531]" italics="true" pageId="27" pageNumber="166">C. croton</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A42427D61AD3F1462F9FBF4C64F7A" box="[113,182,1539,1558]" pageId="27" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
, holotype; C – 
<taxonomicName id="4C0D58A87D61AD3F1524F9FBF5D14F7A" authority="Sehnal &amp; Král &amp; Bezděk, 2014" authorityName="Sehnal &amp; Král &amp; Bezděk" authorityYear="2014" box="[311,417,1539,1558]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="166" phylum="Arthropoda" rank="species" species="dracaena" status="sp. nov.">
<emphasis id="B979FF397D61AD3F1524F9FBF5D14F7A" box="[311,417,1539,1558]" italics="true" pageId="27" pageNumber="166">C. dracaena</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A42427D61AD3F15B5F9FBF59B4F7A" box="[422,491,1539,1558]" pageId="27" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
, holotype; D – 
<taxonomicName id="4C0D58A87D61AD3F167EF9FBF6B94F7A" authority="Sehnal &amp; Král &amp; Bezděk, 2014" authorityName="Sehnal &amp; Král &amp; Bezděk" authorityYear="2014" box="[621,713,1539,1558]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="166" phylum="Arthropoda" rank="species" species="firmihin" status="sp. nov.">
<emphasis id="B979FF397D61AD3F167EF9FBF6B94F7A" box="[621,713,1539,1558]" italics="true" pageId="27" pageNumber="166">C. firmihin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A42427D61AD3F16DCF9FBF7634F7A" box="[719,787,1539,1558]" pageId="27" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
, holotype. Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption id="DF7273A37D66AD381485FA37F4AE4F7B" ID-DOI="http://doi.org/10.5281/zenodo.5312044" ID-Zenodo-Dep="5312044" httpUri="https://zenodo.org/record/5312044/files/figure.png" pageId="28" pageNumber="167" startId="28.[150,187,1487,1506]" targetBox="[159,1117,227,1393]" targetPageId="28">
<paragraph id="8BB2232B7D66AD381485FA37F4AE4F7B" blockId="28.[150,1127,1486,1559]" pageId="28" pageNumber="167">
Figs 11A–D. Aedeagus in dorsal (left) and lateral (right) view: A – 
<taxonomicName id="4C0D58A87D66AD3816CFFA36F7F24C8D" authority="Sehnal &amp; Král &amp; Bezděk, 2014" authorityName="Sehnal &amp; Král &amp; Bezděk" authorityYear="2014" box="[732,898,1486,1505]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="167" phylum="Arthropoda" rank="species" species="hagher" status="sp. nov.">
<emphasis id="B979FF397D66AD3816CFFA36F7F24C8D" box="[732,898,1486,1505]" italics="true" pageId="28" pageNumber="167">Canuschiza hagher</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A42427D66AD38179BFA37F7BE4C8E" box="[904,974,1487,1506]" pageId="28" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
, holotype; B – 
<taxonomicName id="4C0D58A87D66AD381041FA37F5124C90" authority="Lacroix, 1999" authorityName="Lacroix" authorityYear="1999" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="167" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="B979FF397D66AD381041FA37F4914C90" italics="true" pageId="28" pageNumber="167">C. insularis</emphasis>
<bibRefCitation id="EF9C5EDA7D66AD3814F4FA11F5124C90" author="LACROIX M." box="[231,354,1513,1532]" pageId="28" pageNumber="167" pagination="87 - 96" refId="ref15497" refString="LACROIX M. 1999: Melolonthidae de Socotra (Coleoptera, Scarabaeoidea). Revue Francaise d'Entomologie (Nouvelle Serie) 21: 87 - 96." type="journal article" year="1999">Lacroix, 1999</bibRefCitation>
</taxonomicName>
, holotype; C – 
<taxonomicName id="4C0D58A87D66AD3815F4FA11F63A4C90" authority="Sehnal &amp; Král &amp; Bezděk, 2014" authorityName="Sehnal &amp; Král &amp; Bezděk" authorityYear="2014" box="[487,586,1513,1532]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="167" phylum="Arthropoda" rank="species" species="jatropha" status="sp. nov.">
<emphasis id="B979FF397D66AD3815F4FA11F63A4C90" box="[487,586,1513,1532]" italics="true" pageId="28" pageNumber="167">C. jatropha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A42427D66AD38165CFA11F6E54C90" box="[591,661,1513,1532]" pageId="28" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
, holotype; D – 
<taxonomicName id="4C0D58A87D66AD381709FA11F7164C90" authority="Sehnal &amp; Král &amp; Bezděk, 2014" authorityName="Sehnal &amp; Král &amp; Bezděk" authorityYear="2014" box="[794,870,1513,1532]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="167" phylum="Arthropoda" rank="species" species="skand" status="sp. nov.">
<emphasis id="B979FF397D66AD381709FA11F7164C90" box="[794,870,1513,1532]" italics="true" pageId="28" pageNumber="167">C. skand</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A42427D66AD38177FFA11F7C24C90" box="[876,946,1513,1532]" pageId="28" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
, holotype. Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BB2232B7D67AD391462FF1AF4A94875" blockId="29.[113,1089,219,857]" pageId="29" pageNumber="168">
<emphasis id="B979FF397D67AD391462FF1AF5534995" bold="true" box="[113,291,226,249]" pageId="29" pageNumber="168">Measurements.</emphasis>
Total body length: JJ 
<quantity id="4CF58ECE7D67AD391625FF1AF6A84995" box="[566,728,226,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.345" metricValueMax="1.47" metricValueMin="1.22" pageId="29" pageNumber="168" unit="mm" value="13.45" valueMax="14.7" valueMin="12.2">12.2–14.7 mm</quantity>
(
<typeStatus id="54B69D897D67AD3916FAFF1AF73A4995" box="[745,842,226,249]" pageId="29" pageNumber="168" type="holotype">holotype</typeStatus>
<quantity id="4CF58ECE7D67AD391746FF1AF7CE4995" box="[853,958,226,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="29" pageNumber="168" unit="mm" value="12.5">12.5 mm</quantity>
); 
<specimenCount id="9D0BE8A27D67AD3917C7FF23F78A4996" box="[980,1018,219,250]" pageId="29" pageNumber="168" type="female">♀♀</specimenCount>
<quantity id="4CF58ECE7D67AD391017FF1AF4A54875" metricMagnitude="-2" metricUnit="m" metricValue="1.295" metricValueMax="1.42" metricValueMin="1.17" pageId="29" pageNumber="168" unit="mm" value="12.95" valueMax="14.2" valueMin="11.7">11.7– 14.2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C31770A07D67AD391462FEDAF5C24BF5" pageId="29" pageNumber="168" type="diagnosis">
<paragraph id="8BB2232B7D67AD391462FEDAF5C24BF5" blockId="29.[113,1089,219,857]" pageId="29" pageNumber="168">
<emphasis id="B979FF397D67AD391462FEDAF5004855" bold="true" box="[113,368,290,313]" pageId="29" pageNumber="168">Differential diagnosis.</emphasis>
<taxonomicName id="4C0D58A87D67AD391566FEDAF6494856" authority="Sehnal &amp; Král &amp; Bezděk, 2014" authorityName="Sehnal &amp; Král &amp; Bezděk" authorityYear="2014" box="[373,569,290,314]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="168" phylum="Arthropoda" rank="species" species="skand" status="sp. nov.">
<emphasis id="B979FF397D67AD391566FEDAF6494856" box="[373,569,290,314]" italics="true" pageId="29" pageNumber="168">Canuschiza skand</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A42427D67AD39162CFEDAF6E24855" box="[575,658,290,313]" pageId="29" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
can be differentiated from all species of the 
<taxonomicName id="4C0D58A87D67AD39148BFEBAF5654836" authorityName="Lacroix" authorityYear="1999" box="[152,277,322,346]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="168" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="B979FF397D67AD39148BFEBAF5654836" box="[152,277,322,346]" italics="true" pageId="29" pageNumber="168">C. insularis</emphasis>
</taxonomicName>
species group mainly by combination of the following diagnostic characters in males: antennae with ten antennomeres (
<figureCitation id="13363FAE7D67AD391657FE9AF6EB4815" box="[580,667,354,377]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="29" pageNumber="168">Fig. 9D</figureCitation>
); antennal club almost straight, as long as antennal shaft (antennomeres I–VII combined) (
<figureCitation id="13363FAE7D67AD39168AFE7AF77F48F5" box="[665,783,386,409]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="29" pageNumber="168">Figs 9A,D</figureCitation>
); anterior margin of clypeus weakly upturned, almost straight medially (
<figureCitation id="13363FAE7D67AD39165AFE5AF6CF48D5" box="[585,703,418,441]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="29" pageNumber="168">Figs 9A,D</figureCitation>
); eyes medium sized (width of both eyes combined approximately the same as maximum width of frons between eyes) (
<figureCitation id="13363FAE7D67AD391003FE3AF4C14895" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="29" pageNumber="168">Figs 9A,D</figureCitation>
); lateral margins of pronotum in anterior half minutely crenate and macrosetaceous (
<figureCitation id="13363FAE7D67AD39146AFDFAF4BE4B75" box="[121,206,514,537]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="29" pageNumber="168">Fig. 9E</figureCitation>
); posterior angles of pronotum rectangular, moderately produced posteriad (
<figureCitation id="13363FAE7D67AD391003FDFAF4DF4B55" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="29" pageNumber="168">Figs 9A,C</figureCitation>
); elytra densely irregularly punctate, each puncture bearing scale-like macroseta, longer than puncture diameter; scale-like macrosetae of pronotum and elytra considerably short, broad (
<figureCitation id="13363FAE7D67AD391515FD9AF50D4B15" box="[262,381,610,633]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="29" pageNumber="168">Figs 9A,C</figureCitation>
); macropterous. For tentative differentiation of females refer to the identification keys below.
</paragraph>
</subSubSection>
<subSubSection id="C31770A07D67AD391462FD5AF07F4A35" pageId="29" pageNumber="168" type="etymology">
<paragraph id="8BB2232B7D67AD391462FD5AF6A74BB5" blockId="29.[113,1089,219,857]" pageId="29" pageNumber="168">
<emphasis id="B979FF397D67AD391462FD5AF4824BD5" bold="true" box="[113,242,674,697]" pageId="29" pageNumber="168">Etymology.</emphasis>
Derived from the area of origin of the new species, vicinity of the 
<taxonomicName id="4C0D58A87D67AD3917D9FD5AF04D4BD5" authority="Mt." authorityName="Mt." box="[970,1085,674,697]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="168" phylum="Arthropoda" rank="species" species="skand">Skand Mt.</taxonomicName>
, 
<collectingCountry id="F31A63BB7D67AD391462FD3AF4B24BB5" box="[113,194,706,729]" name="Yemen" pageId="29" pageNumber="168">Socotra</collectingCountry>
(
<collectingCountry id="F31A63BB7D67AD3914C2FD3AF5524BB5" box="[209,290,706,729]" name="Yemen" pageId="29" pageNumber="168">Yemen</collectingCountry>
); noun in nominative case, in apposition.
</paragraph>
<paragraph id="8BB2232B7D67AD391462FD1AF7184A75" blockId="29.[113,1089,219,857]" pageId="29" pageNumber="168">
<emphasis id="B979FF397D67AD391462FD1AF5E54B95" bold="true" box="[113,405,738,761]" pageId="29" pageNumber="168">Collecting circumstances.</emphasis>
All 
<typeStatus id="54B69D897D67AD3915D6FD1AF5834B95" box="[453,499,738,761]" pageId="29" pageNumber="168">type</typeStatus>
material was captured attracted to light trap placed on clearing in evergreen montane woodland (J. Hájek, pers. comm. 2014).
</paragraph>
<paragraph id="8BB2232B7D67AD391462FCDAF07F4A35" blockId="29.[113,1089,219,857]" pageId="29" pageNumber="168">
<emphasis id="B979FF397D67AD391462FCDAF5EF4A55" bold="true" box="[113,415,802,825]" pageId="29" pageNumber="168">Geographical distribution.</emphasis>
<typeStatus id="54B69D897D67AD3915B7FCDAF5A94A55" box="[420,473,802,825]" pageId="29" pageNumber="168">Type</typeStatus>
material originates from the vicinity of the 
<taxonomicName id="4C0D58A87D67AD3917B6FCDAF0664A55" authority="Mt." authorityName="Mt." box="[933,1046,802,825]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="168" phylum="Arthropoda" rank="species" species="skand">Skand Mt.</taxonomicName>
, the highest area of the 
<taxonomicName id="4C0D58A87D67AD39152FFCBAF5CA4A35" authority="Mts" authorityName="Mts" box="[316,442,834,857]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="168" phylum="Arthropoda" rank="species" species="hagher">Hagher Mts</taxonomicName>
and whole 
<collectingCountry id="F31A63BB7D67AD391624FCBAF6FB4A35" box="[567,651,834,857]" name="Yemen" pageId="29" pageNumber="168">Socotra</collectingCountry>
, for details see 
<bibRefCitation id="EF9C5EDA7D67AD391723FCBAF07B4A35" author="BEZDEK J. &amp; PURCHART L. &amp; KRAL K." box="[816,1035,834,857]" pageId="29" pageNumber="168" pagination="1 - 557" refId="ref15271" refString="BEZDEK J., PURCHART L., KRAL K. &amp; HULA. V. 2012: List of local Socotran geographical names used in entomological literature. Pp. 27 - 67. In: HAJEK J. &amp; BEZDEK J. (eds.): Insect biodiversity of Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52 (Supplementum 2): i - vi + 1 - 557." type="journal article" year="2012">
<subScript id="1789216E7D67AD391723FCBAF70F4A34" attach="right" box="[816,895,834,857]" fontSize="7" pageId="29" pageNumber="168">BEZDĚK</subScript>
et al. (2012)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>