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<document id="C91E14C464DC8FCD6E916FACE0C28E28" ID-CLB-Dataset="24929" ID-DOI="10.5281/zenodo.3532762" ID-GBIF-Dataset="db23a775-d25c-4253-abda-b5a007110f6d" ID-ISSN="0374-1036" ID-Zenodo-Dep="3532762" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573206962882" checkinUser="plazi" docAuthor="Irmler, Ulrich" docDate="2013" docId="03A3FB753F5B1E63FE76FE4644A60F52" docLanguage="en" docName="ActaEntMusNatPra.53.1.155-176.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Holotrochus tortilis Irmler 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="166" masterDocId="FF9A830D3F511E68FF93FFED46660A2F" masterDocTitle="New species of the genus Mimogonia and Holotrochus from South America (Coleoptera: Staphylindae: Osoriinae)" masterLastPageNumber="176" masterPageNumber="155" pageNumber="165" updateTime="1698764282444" updateUser="ExternalLinkService">
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<mods:title id="9105F880BCA94D7D1D5A0071830445A0">New species of the genus Mimogonia and Holotrochus from South America (Coleoptera: Staphylindae: Osoriinae)</mods:title>
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<mods:namePart id="2948ABA990AFA59E59BD43E9CFAA6C65">Irmler, Ulrich</mods:namePart>
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<mods:title id="3BAF44191AF139AE111781B7235370B7">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="B5529D365B38A8B2E559CB97AF90EC6C">2013</mods:date>
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<treatment id="03A3FB753F5B1E63FE76FE4644A60F52" ID-DOI="http://doi.org/10.5281/zenodo.4427163" ID-GBIF-Taxon="160218244" ID-Zenodo-Dep="4427163" LSID="urn:lsid:plazi:treatment:03A3FB753F5B1E63FE76FE4644A60F52" httpUri="http://treatment.plazi.org/id/03A3FB753F5B1E63FE76FE4644A60F52" lastPageId="11" lastPageNumber="166" pageId="10" pageNumber="165">
<subSubSection id="C31019E83F5B1E62FE76FE46457F0BEB" box="[485,793,427,452]" pageId="10" pageNumber="165" type="nomenclature">
<paragraph id="8BB54A633F5B1E62FE76FE46457F0BEB" blockId="10.[485,793,427,477]" box="[485,793,427,452]" pageId="10" pageNumber="165">
<heading id="D0FDFD0F3F5B1E62FE76FE46457F0BEB" bold="true" box="[485,793,427,452]" centered="true" fontSize="10" level="3" pageId="10" pageNumber="165" reason="4">
<emphasis id="B97E96713F5B1E62FE76FE46457F0BEB" bold="true" box="[485,793,427,452]" pageId="10" pageNumber="165">
<taxonomicName id="4C0A31E03F5B1E62FE76FE4644DB0BEC" ID-CoL="6LYPL" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[485,701,427,451]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="165" phylum="Arthropoda" rank="species" species="tortilis" status="sp. nov.">
<emphasis id="B97E96713F5B1E62FE76FE4644DB0BEC" bold="true" box="[485,701,427,451]" italics="true" pageId="10" pageNumber="165">Holotrochus tortilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5B1E62FD50FE41457F0BEB" box="[707,793,428,452]" pageId="10" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5B1E62FDD2FE2744DB0BF2" box="[577,701,458,477]" pageId="10" pageNumber="165" type="description">
<paragraph id="8BB54A633F5B1E62FDD2FE2744DB0BF2" blockId="10.[485,793,427,477]" box="[577,701,458,477]" pageId="10" pageNumber="165">
(
<figureCitation id="133156E63F5B1E62FDD4FE2744EF0BF2" box="[583,649,458,477]" captionStart="Figs 1215. 12" captionStartId="11.[113,149,1034,1053]" captionTargetBox="[126,1068,225,1013]" captionTargetId="figure@11.[113,1088,216,1020]" captionTargetPageId="11" captionText="Figs 1215. 12 Holotrochus tortilis sp. nov.; 13 H. lobatus sp. nov.; 14 H. agostii sp.nov.; 15 H.pseudoleticiae sp. nov. Body parts:A aedeagus; B antenna; C last sternite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532770" httpUri="https://zenodo.org/record/3532770/files/figure.png" pageId="10" pageNumber="165">Figs 12</figureCitation>
,
<figureCitation id="133156E63F5B1E62FD01FE2744D00BF2" box="[658,694,458,477]" captionStart="Fig" captionStartId="17.[113,144,1541,1560]" captionTargetBox="[121,1067,217,1507]" captionTargetId="figure@17.[113,1086,211,1515]" captionTargetPageId="17" captionText="Fig. 18. Head, pronotum, and elytra of Holotrochus Erichson,1839 showing shape, punctation, and microsculpture.A H. paraguensis sp. nov.; B H.tortilis sp. nov.; C H.hamatus sp. nov.; D H.yanuniensis sp. nov.; E H. mrazi sp. nov.; F H. lobatus sp. nov. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3532776" httpUri="https://zenodo.org/record/3532776/files/figure.png" pageId="10" pageNumber="165">18B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5B1E62FF04FE1946B4085E" pageId="10" pageNumber="165" type="materials_examined">
<paragraph id="8BB54A633F5B1E62FF04FE1944020829" blockId="10.[151,1127,499,625]" box="[151,612,499,519]" pageId="10" pageNumber="165">
<materialsCitation id="3B62403E3F5B1E62FF04FE1944060829" ID-GBIF-Occurrence="2449305645" box="[151,608,499,519]" country="Bolivia" location="Guayaramerin" municipality="Province" pageId="10" pageNumber="165" specimenCount="1" stateProvince="Beni" typeStatus="holotype">
<emphasis id="B97E96713F5B1E62FF04FE1947760829" bold="true" box="[151,272,499,519]" pageId="10" pageNumber="165">
<typeStatus id="54B1F4C13F5B1E62FF04FE1946A20828" box="[151,196,500,519]" pageId="10" pageNumber="165">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F31D0AF33F5B1E62FE86FE1E47300829" box="[277,342,499,518]" name="Bolivia" pageId="10" pageNumber="165">Bolivia</collectingCountry>
,
<collectingMunicipality id="6BD1D0193F5B1E62FECCFE1E47CD0829" box="[351,427,499,518]" pageId="10" pageNumber="165">Province</collectingMunicipality>
<collectingRegion id="49CE84813F5B1E62FE23FE1E47BC0829" box="[432,474,499,518]" country="Bolivia" name="Beni" pageId="10" pageNumber="165">Beni</collectingRegion>
,
<location id="8ED51CB83F5B1E62FE70FE1E44060829" LSID="urn:lsid:plazi:treatment:03A3FB753F5B1E63FE76FE4644A60F52:8ED51CB83F5B1E62FE70FE1E44060829" box="[483,608,499,518]" country="Bolivia" municipality="Province" name="Guayaramerin" pageId="10" pageNumber="165" stateProvince="Beni">Guayaramerin</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB54A633F5B1E62FF04FDE346B4085E" blockId="10.[151,1127,499,625]" pageId="10" pageNumber="165">
<emphasis id="B97E96713F5B1E62FF04FDE34779080E" bold="true" box="[151,287,526,545]" pageId="10" pageNumber="165">Type material.</emphasis>
<materialsCitation id="3B62403E3F5B1E62FEB7FDE245230813" ID-GBIF-Occurrence="2449305663" collectingDate="1966-11-26" collectionCode="HNHM" collectorName="J. Balogh'" country="Bolivia" county="Guayaramerin" latitude="-10.820833" location="Berlese" longLatPrecision="13" longitude="-65.367836" municipality="Rio Marmore" pageId="10" pageNumber="165" specimenCount="1" stateProvince="Beni" typeStatus="holotype">
<typeStatus id="54B1F4C13F5B1E62FEB7FDE2471C0830" box="[292,378,527,546]" pageId="10" pageNumber="165" type="holotype">HOLOTYPE</typeStatus>
:,
<collectingCountry id="F31D0AF33F5B1E62FE37FDE3478C080E" box="[420,490,526,545]" name="Bolivia" pageId="10" pageNumber="165">Bolivia</collectingCountry>
, Prov.
<collectingRegion id="49CE84813F5B1E62FDB5FDE34437080E" box="[550,593,526,545]" country="Bolivia" name="Beni" pageId="10" pageNumber="165">Beni</collectingRegion>
,
<collectingCounty id="62D432EF3F5B1E62FDC8FDE344BE080E" box="[603,728,526,545]" pageId="10" pageNumber="165">Guayaramerin</collectingCounty>
(
<geoCoordinate id="EE3E2CA43F5B1E62FD70FDE34532080D" box="[739,852,526,546]" degrees="65" direction="west" minutes="22.07" orientation="longitude" pageId="10" pageNumber="165" precision="9" value="-65.367836">65°22.07 W</geoCoordinate>
,
<geoCoordinate id="EE3E2CA43F5B1E62FCCEFDE345A3080E" box="[861,965,526,545]" degrees="10" direction="south" minutes="49.25" orientation="latitude" pageId="10" pageNumber="165" precision="9" value="-10.820833">10°49.25 S</geoCoordinate>
), banks of the
<collectingMunicipality id="6BD1D0193F5B1E62FBD4FDE3468C0813" pageId="10" pageNumber="165">Rio Marmoré</collectingMunicipality>
, sampled by
<location id="8ED51CB83F5B1E62FEF0FDC447C20813" LSID="urn:lsid:plazi:treatment:03A3FB753F5B1E63FE76FE4644A60F52:8ED51CB83F5B1E62FEF0FDC447C20813" box="[355,420,553,572]" country="Bolivia" county="Guayaramerin" latitude="-10.820833" longLatPrecision="13" longitude="-65.367836" municipality="Rio Marmore" name="Berlese" pageId="10" pageNumber="165" stateProvince="Beni">Berlese</location>
method,
<date id="FFB46CA33F5B1E62FE69FDC4443B0813" box="[506,605,553,572]" pageId="10" pageNumber="165" value="1966-11-26">
<collectingDate id="EFF0954B3F5B1E62FE69FDC4443B0813" box="[506,605,553,572]" pageId="10" pageNumber="165" value="1966-11-26">26.11.1966</collectingDate>
</date>
, leg.
<collectorName id="26FF2FB53F5B1E62FD03FDC4448D0813" box="[656,747,553,572]" pageId="10" pageNumber="165">J. Balogh</collectorName>
(
<collectionCode id="ED1BD2A63F5B1E62FD6BFDC445590813" box="[760,831,553,572]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="165">HNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B62403E3F5B1E62FCC3FDC446A8085E" ID-GBIF-Occurrence="2449305664" collectingDate="1981-01-06" collectionCode="HNHM, UIC" collectorName="D. Balazs'" country="Suriname" county="Guayaramerin" latitude="5.5006666" location="Albina" longLatPrecision="13" longitude="-54.054165" municipality="Rio Marmore" pageId="10" pageNumber="165" specimenCount="1" specimenCount-female="1" stateProvince="Beni" typeStatus="paratype">
<typeStatus id="54B1F4C13F5B1E62FCC3FDC445CF0815" box="[848,937,553,572]" pageId="10" pageNumber="165" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D0C81EA3F5B1E62FC25FDC445BF0813" box="[950,985,549,572]" pageId="10" pageNumber="165" type="female">1 ♀</specimenCount>
, from the same location as the holotype (
<collectionCode id="ED1BD2A63F5B1E62FEFCFDA947D00878" box="[367,438,580,599]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="165">HNHM</collectionCode>
); 1,
<collectingCountry id="F31D0AF33F5B1E62FE63FDAE442C0879" box="[496,586,579,598]" name="Suriname" pageId="10" pageNumber="165">Suriname</collectingCountry>
,
<location id="8ED51CB83F5B1E62FDC3FDAE44ED0879" LSID="urn:lsid:plazi:treatment:03A3FB753F5B1E63FE76FE4644A60F52:8ED51CB83F5B1E62FDC3FDAE44ED0879" box="[592,651,579,598]" country="Suriname" latitude="5.5006666" longLatPrecision="13" longitude="-54.054165" name="Albina" pageId="10" pageNumber="165">Albina</location>
(
<geoCoordinate id="EE3E2CA43F5B1E62FD05FDAE45600878" box="[662,774,579,599]" degrees="54" direction="west" minutes="03.25" orientation="longitude" pageId="10" pageNumber="165" precision="9" value="-54.054165">54°03.25 W</geoCoordinate>
,
<geoCoordinate id="EE3E2CA43F5B1E62FC9EFDAE45080878" box="[781,878,579,599]" degrees="5" direction="north" minutes="30.04" orientation="latitude" pageId="10" pageNumber="165" precision="9" value="5.5006666">5°30.04 N</geoCoordinate>
),
<date id="FFB46CA33F5B1E62FCEEFDAE45870879" box="[893,993,579,598]" pageId="10" pageNumber="165" value="1981-01-06">
<collectingDate id="EFF0954B3F5B1E62FCEEFDAE45870879" box="[893,993,579,598]" pageId="10" pageNumber="165" value="1981-01-06">01.06.1981</collectingDate>
</date>
, leg.
<collectorName id="26FF2FB53F5B1E62FB9EFDA942010879" box="[1037,1127,579,599]" pageId="10" pageNumber="165">D. Balázs</collectorName>
(
<collectionCode id="ED1BD2A63F5B1E62FF0DFDB346AE085E" box="[158,200,606,625]" pageId="10" pageNumber="165">UIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5B1E62FF04FD644782096E" pageId="10" pageNumber="165" type="diagnosis">
<paragraph id="8BB54A633F5B1E62FF04FD644782096E" blockId="10.[151,1127,649,1570]" pageId="10" pageNumber="165">
<emphasis id="B97E96713F5B1E62FF04FD64476A088E" bold="true" box="[151,268,649,673]" pageId="10" pageNumber="165">Diagnosis.</emphasis>
This species certainly belongs to the
<taxonomicName id="4C0A31E03F5B1E62FD30FD674509088E" authorityName="Fauvel" authorityYear="1863" box="[675,879,650,673]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="165" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B97E96713F5B1E62FD30FD674509088E" box="[675,879,650,673]" italics="true" pageId="10" pageNumber="165">Holotrochus minor</emphasis>
</taxonomicName>
group due to the same combination of character as mentioned above under
<taxonomicName id="4C0A31E03F5B1E62FD2CFD47450708EE" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[703,865,682,705]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="165" phylum="Arthropoda" rank="species" species="paraguensis" status="sp. nov.">
<emphasis id="B97E96713F5B1E62FD2CFD47450708EE" box="[703,865,682,705]" italics="true" pageId="10" pageNumber="165">H. paraguensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5B1E62FCF4FD4745D208EE" box="[871,948,682,705]" pageId="10" pageNumber="165" rank="species">sp. nov</taxonomicNameLabel>
.. Concerning the structure of the fore-body, the two species
<taxonomicName id="4C0A31E03F5B1E62FDC4FD2744D808CE" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[599,702,714,737]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="165" phylum="Arthropoda" rank="species" species="tortilis" status="sp. nov.">
<emphasis id="B97E96713F5B1E62FDC4FD2744D808CE" box="[599,702,714,737]" italics="true" pageId="10" pageNumber="165">H. tortilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5B1E62FD57FD27457008CE" box="[708,790,714,737]" pageId="10" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C0A31E03F5B1E62FCD4FD27458F08CE" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[839,1001,714,737]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="165" phylum="Arthropoda" rank="species" species="paraguensis" status="sp. nov.">
<emphasis id="B97E96713F5B1E62FCD4FD27458F08CE" box="[839,1001,714,737]" italics="true" pageId="10" pageNumber="165">H. paraguensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5B1E62FC7CFD27422708CE" box="[1007,1089,714,737]" pageId="10" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
are mainly distinguished by the shape of the pronotum and the shape of the shoulders of elytra. Furthermore, the elytra in
<taxonomicName id="4C0A31E03F5B1E62FE23FCE74471090E" authorityName="Irmler" authorityYear="2013" box="[432,535,778,801]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="165" phylum="Arthropoda" rank="species" species="tortilis">
<emphasis id="B97E96713F5B1E62FE23FCE74471090E" box="[432,535,778,801]" italics="true" pageId="10" pageNumber="165">H. tortilis</emphasis>
</taxonomicName>
are slightly wider and the punctation is slightly coarser than in
<taxonomicName id="4C0A31E03F5B1E62FF74FCC747EC096E" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[231,394,810,833]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="165" phylum="Arthropoda" rank="species" species="paraguensis" status="sp. nov.">
<emphasis id="B97E96713F5B1E62FF74FCC747EC096E" box="[231,394,810,833]" italics="true" pageId="10" pageNumber="165">H. paraguensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5B1E62FE02FCC74782096E" box="[401,484,810,833]" pageId="10" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5B1E63FF04FCA445680F32" lastPageId="11" lastPageNumber="166" pageId="10" pageNumber="165" type="description">
<paragraph id="8BB54A633F5B1E62FF04FCA4443009AD" blockId="10.[151,1127,649,1570]" pageId="10" pageNumber="165">
<emphasis id="B97E96713F5B1E62FF04FCA44744094E" bold="true" box="[151,290,841,865]" pageId="10" pageNumber="165">Description.</emphasis>
Length:
<quantity id="4CF2E7863F5B1E62FE12FCA747B0094E" box="[385,470,842,866]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="10" pageNumber="165" unit="mm" value="3.2">3.2 mm</quantity>
. Colouration dark brown; small posterior margin of pronotum lighter reddish; legs red; antennae yellow.
</paragraph>
<paragraph id="8BB54A633F5B1E62FF26FC6745E00E4D" blockId="10.[151,1127,649,1570]" pageId="10" pageNumber="165">
Head
<quantity id="4CF2E7863F5B1E62FF61FC674728098E" box="[242,334,906,930]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="10" pageNumber="165" unit="mm" value="0.38">0.38 mm</quantity>
long,
<quantity id="4CF2E7863F5B1E62FE1EFC67478E098E" box="[397,488,906,930]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="165" unit="mm" value="0.55">0.55 mm</quantity>
wide; eyes slightly prominent, as long as temples; with supraocular margin continuing to base of antennae; within margin, row of 4 setiferous punctures; on a line between posterior edge of eyes, four setiferous punctures; interstices between inner setiferous punctures nearly twice as wide as distance between inner and outer punctures; punctation moderately deep and dense; at clypeus denser than on vertex; on average, interstices between punctures as wide as diameter of punctures; with sparse micro-punctation between normal punctures; surface without microsculpture, polished and shiny.
</paragraph>
<paragraph id="8BB54A633F5B1E62FF26FB8744A70F2D" blockId="10.[151,1127,649,1570]" pageId="10" pageNumber="165">Antennae slightly longer than head; second antennomere globular; distinctly smaller than first and distinctly wider than conical third antennomere; third antennomere 1.5 times longer than second; antennomere 4 to 6 more or less quadrate only slightly increasing in width; seventh antennomere distinctly wider than sixth; following antennomeres increasing in width; tenth antennomere more than twice as wide as long.</paragraph>
<paragraph id="8BB54A633F5B1E62FF26FAE744010FED" blockId="10.[151,1127,649,1570]" pageId="10" pageNumber="165">
Pronotum
<quantity id="4CF2E7863F5B1E62FEB4FAE747E10F0E" box="[295,391,1290,1314]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="165" unit="mm" value="0.55">0.55 mm</quantity>
long,
<quantity id="4CF2E7863F5B1E62FE5DFAE744480F0E" box="[462,558,1290,1314]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="10" pageNumber="165" unit="mm" value="0.71">0.71 mm</quantity>
wide; widest in middle; more narrowed to posterior angles than to anterior angles; sides distinctly margined from posterior angles to anterior angles; lateral margin visible throughout its total length and continuing to front edge; punctation deep and dense; interstices between punctures on most parts as wide as diameter of punctures; near front angles with sparser punctation; narrow midline in posterior half impunctate; without microsculpture; surface polished and shiny.
</paragraph>
<paragraph id="8BB54A633F5B1E62FF26FA2747980C0E" blockId="10.[151,1127,649,1570]" pageId="10" pageNumber="165">
Elytra
<quantity id="4CF2E7863F5B1E62FF6FFA27473F0FCE" box="[252,345,1482,1506]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="165" unit="mm" value="0.65">0.65 mm</quantity>
long,
<quantity id="4CF2E7863F5B1E62FE0AFA2747900FCE" box="[409,502,1482,1506]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="10" pageNumber="165" unit="mm" value="0.73">0.73 mm</quantity>
wide; shoulders more or less rectangular, with short prominent tooth; punctation deep, dense, and partly coriaceous; with coriaceous ground sculpture; surface less shiny than pronotum.
</paragraph>
<caption id="DF751AEB3F5A1E63FFE2FBE745260E17" ID-DOI="http://doi.org/10.5281/zenodo.3532770" ID-Zenodo-Dep="3532770" httpUri="https://zenodo.org/record/3532770/files/figure.png" pageId="11" pageNumber="166" startId="11.[113,149,1034,1053]" targetBox="[126,1068,225,1013]" targetPageId="11">
<paragraph id="8BB54A633F5A1E63FFE2FBE745260E17" blockId="11.[113,1088,1034,1080]" pageId="11" pageNumber="166">
Figs 1215. 12
<taxonomicName id="4C0A31E03F5A1E63FE93FBE647C30E32" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[256,421,1035,1053]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="166" phylum="Arthropoda" rank="species" species="tortilis" status="sp. nov.">
<emphasis id="B97E96713F5A1E63FE93FBE647C30E32" box="[256,421,1035,1053]" italics="true" pageId="11" pageNumber="166">Holotrochus tortilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5A1E63FE3AFBE6478B0E31" box="[425,493,1035,1054]" pageId="11" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
; 13
<taxonomicName id="4C0A31E03F5A1E63FD89FBE644170E32" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[538,625,1035,1053]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="166" phylum="Arthropoda" rank="species" species="lobatus" status="sp. nov.">
<emphasis id="B97E96713F5A1E63FD89FBE644170E32" box="[538,625,1035,1053]" italics="true" pageId="11" pageNumber="166">H. lobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5A1E63FDE6FBE644DF0E31" box="[629,697,1035,1054]" pageId="11" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
; 14
<taxonomicName id="4C0A31E03F5A1E63FD75FBE6455F0E32" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[742,825,1035,1053]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="166" phylum="Arthropoda" rank="species" species="agostii" status="sp. nov.">
<emphasis id="B97E96713F5A1E63FD75FBE6455F0E32" box="[742,825,1035,1053]" italics="true" pageId="11" pageNumber="166">H. agostii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5A1E63FCAFFBE645E60E31" box="[828,896,1035,1054]" pageId="11" pageNumber="166" rank="species">sp.nov.</taxonomicNameLabel>
; 15
<taxonomicName id="4C0A31E03F5A1E63FC3DFBE642260E32" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[942,1088,1035,1053]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="166" phylum="Arthropoda" rank="species" species="pseudoleticiae" status="sp. nov.">
<emphasis id="B97E96713F5A1E63FC3DFBE642260E32" box="[942,1088,1035,1053]" italics="true" pageId="11" pageNumber="166">H.pseudoleticiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5A1E63FFE2FBC846D20E17" box="[113,180,1061,1080]" pageId="11" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
Body parts:A aedeagus; B antenna; C last sternite. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BB54A633F5A1E63FF03FB4842400EF2" blockId="11.[113,1089,1189,1405]" pageId="11" pageNumber="166">Abdomen finely and densely punctate; with deep and dense net-like microsculpture, in particular on tergite VII; surface matte; last sternite triangular with few lateral punctures.</paragraph>
<paragraph id="8BB54A633F5A1E63FF03FB0845680F32" blockId="11.[113,1089,1189,1405]" pageId="11" pageNumber="166">Aedeagus with acute apex pointing reverse; endophallus with two large torsions; parameres more or less straight; slightly outreaching apex of central lobe.</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5A1E63FFE2FAC844470F72" pageId="11" pageNumber="166" type="etymology">
<paragraph id="8BB54A633F5A1E63FFE2FAC844470F72" blockId="11.[113,1089,1189,1405]" pageId="11" pageNumber="166">
<emphasis id="B97E96713F5A1E63FFE2FAC846940F12" bold="true" box="[113,242,1317,1341]" pageId="11" pageNumber="166">Etymology.</emphasis>
The specific name is the Latin adjective
<taxonomicName id="4C0A31E03F5A1E63FD31FAC8448F0F13" authorityName="Irmler" authorityYear="2013" box="[674,745,1317,1340]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="166" phylum="Arthropoda" rank="species" species="tortilis">
<emphasis id="B97E96713F5A1E63FD31FAC8448F0F13" box="[674,745,1317,1340]" italics="true" pageId="11" pageNumber="166">tortilis</emphasis>
</taxonomicName>
meaning “twisted” and refers to the twisted endophallus of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5A1E63FFE2FA8844A60F52" box="[113,704,1381,1405]" pageId="11" pageNumber="166" type="distribution">
<paragraph id="8BB54A633F5A1E63FFE2FA8844A60F52" blockId="11.[113,1089,1189,1405]" box="[113,704,1381,1405]" pageId="11" pageNumber="166">
<emphasis id="B97E96713F5A1E63FFE2FA8847620F52" bold="true" box="[113,260,1381,1405]" pageId="11" pageNumber="166">Distribution.</emphasis>
Known only from
<collectingCountry id="F31D0AF33F5A1E63FE43FA8844790F52" box="[464,543,1381,1405]" name="Bolivia" pageId="11" pageNumber="166">Bolivia</collectingCountry>
and
<collectingCountry id="F31D0AF33F5A1E63FDC1FA8844DA0F52" box="[594,700,1381,1405]" name="Suriname" pageId="11" pageNumber="166">Suriname</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>