348 lines
52 KiB
XML
348 lines
52 KiB
XML
<document id="FCD4E79322AC9EEE02044BC716C90F0C" ID-CLB-Dataset="20032" ID-DOI="10.11646/zootaxa.5124.2.2" ID-GBIF-Dataset="35f6031f-a9a3-4d6c-ba9e-187e617c65c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="6404962" ID-ZooBank="78F24E28-48AB-4D69-B4D7-D1AA92731606" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648797562183" checkinUser="plazi" docAuthor="Khosravi, Erfan, Sari, Alireza, Mirziee-Ataabadi, Majid, Gholamalian, Hossein & Hyžný, Matúš" docDate="2022" docId="03A387C5FFD4FFDDFF1BFAD8FE0FFB9C" docLanguage="en" docName="zootaxa.5124.2.2.pdf" docOrigin="Zootaxa 5124 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Galene dashtbani Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný 2022, n. sp." docType="treatment" docVersion="7" lastPageNumber="148" masterDocId="FF9AFFBDFFD2FFD4FF8CFFEFFFACFF8D" masterDocTitle="A new species of Galene de Haan, 1833 (Galenidae: Brachyura) from the middle Miocene of Zagros Mountains, Iran" masterLastPageNumber="154" masterPageNumber="139" pageNumber="145" updateTime="1699319176041" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FA79AA4E8B8F86E2C5BD0FBDFCF65CAF">A new species of Galene de Haan, 1833 (Galenidae: Brachyura) from the middle Miocene of Zagros Mountains, Iran</mods:title>
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<mods:name id="B8BFA43409121C7CCAB3121577784357" type="personal">
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<mods:roleTerm id="0E83B274AC6F5F9F1312CBB28CC0BA1B">Author</mods:roleTerm>
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<mods:namePart id="CDC9F33B95EE55089DE10E6BB7A6D2F2">Khosravi, Erfan</mods:namePart>
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<mods:affiliation id="CF28BAAF60703B1AB064461F6C5BC7B4">School of Biology and Centre of Excellence in Phylogeny, College of Science, University of Tehran, Tehran, Iran.</mods:affiliation>
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<mods:name id="08B66716A2C05752779A69344C1C04AF" type="personal">
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<mods:roleTerm id="2DFF4BF0D9CD2B6F97E364018F867250">Author</mods:roleTerm>
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<mods:namePart id="080A687ABD48FAD526FC12726437337F">Sari, Alireza</mods:namePart>
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<mods:affiliation id="E6D1A1B4EE14787C3E932E84A23E9788">School of Biology and Centre of Excellence in Phylogeny, College of Science, University of Tehran, Tehran, Iran.</mods:affiliation>
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<mods:name id="7394CE17B920A6C7506CC427A53481B4" type="personal">
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<mods:roleTerm id="F680FE64A91360143771569C0DA1E3B1">Author</mods:roleTerm>
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<mods:namePart id="3EC354D6551983902E9F81BBBBA7179F">Mirziee-Ataabadi, Majid</mods:namePart>
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<mods:affiliation id="DEAFD148EC2933F68A77D21AC6225D18">Departments of Geology, Faculty of Science, University of Zanjan, Zanjan, Iran.</mods:affiliation>
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</mods:name>
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<mods:name id="0745AF8A377DC6E51E67E10786E1494E" type="personal">
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<mods:namePart id="9DCD1791A6D206444907905A42888FC3">Gholamalian, Hossein</mods:namePart>
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<mods:affiliation id="598CC8F5B4C0EC346E38ECED657EE273">Departments of Geology, Faculty of Sciences, University of Hormozgan, Bandar-Abbas, Iran.</mods:affiliation>
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<mods:namePart id="16AB2E683388AE9C464B771A4EF1AD53">Hyžný, Matúš</mods:namePart>
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<mods:affiliation id="18B8C2CAF00AF94399D6643144B32125">Department of Geology and Paleontology, Faculty of Natural Sciences, Comenius University, Ilkovičova 6, SVK- 842 15, Bratislava 4, Slovakia. & Department of Geology and Paleontology, Natural History Museum, Vienna, Burgring 7, A- 1010 Vienna, Austria. & School of Biology and Centre of Excellence in Phylogeny, College of Science, University of Tehran, Tehran, Iran.</mods:affiliation>
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<mods:typeOfResource id="F15716B9A607BB291C0603728FF32758">text</mods:typeOfResource>
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<mods:title id="B38DB4D19ECD336E4FBF6E77E52DB30B">Zootaxa</mods:title>
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<mods:part id="C472CE5BDE83B3963EE967FF164B36A2">
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<mods:date id="86095A3E710BBBA450E9E28E50790F22">2022</mods:date>
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<mods:number id="C0F53BB8D9817C40656636918E07626F">2022-04-01</mods:number>
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<mods:number id="BF61B664D7C51E638D6C940CA73E9EB1">5124</mods:number>
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<mods:classification id="26F9C757EFCFFB3250E18565A6436229">journal article</mods:classification>
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<mods:identifier id="540DD67910665D92E177CEC2879FA2A5" type="CLB-Dataset">20032</mods:identifier>
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<mods:identifier id="AC847AB4D8D66D037C3B32AE9CC6BBD6" type="DOI">10.11646/zootaxa.5124.2.2</mods:identifier>
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<treatment id="03A387C5FFD4FFDDFF1BFAD8FE0FFB9C" ID-DOI="http://doi.org/10.5281/zenodo.6404976" ID-GBIF-Taxon="194361018" ID-Zenodo-Dep="6404976" LSID="urn:lsid:plazi:treatment:03A387C5FFD4FFDDFF1BFAD8FE0FFB9C" httpUri="http://treatment.plazi.org/id/03A387C5FFD4FFDDFF1BFAD8FE0FFB9C" lastPageId="9" lastPageNumber="148" pageId="6" pageNumber="145">
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<subSubSection id="C3106558FFD4FFD2FF1BFAD8FE1AFADF" box="[151,438,1335,1362]" pageId="6" pageNumber="145" type="nomenclature">
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<paragraph id="8BB536D3FFD4FFD2FF1BFAD8FE1AFADF" blockId="6.[151,438,1335,1399]" box="[151,438,1335,1362]" pageId="6" pageNumber="145">
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<heading id="D0FD81BFFFD4FFD2FF1BFAD8FE1AFADF" bold="true" box="[151,438,1335,1362]" fontSize="11" level="1" pageId="6" pageNumber="145" reason="1">
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<emphasis id="B97EEAC1FFD4FFD2FF1BFAD8FE1AFADF" bold="true" box="[151,438,1335,1362]" pageId="6" pageNumber="145">
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<taxonomicName id="4C0A4D50FFD4FFD2FF1BFAD8FEC0FADF" authority="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný, 2022" authorityName="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný" authorityYear="2022" box="[151,364,1335,1362]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="6" pageNumber="145" phylum="Rhodophyta" rank="species" species="dashtbani" status="sp. nov.">
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<emphasis id="B97EEAC1FFD4FFD2FF1BFAD8FEC0FADF" bold="true" box="[151,364,1335,1362]" italics="true" pageId="6" pageNumber="145">Galene dashtbani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24D57BAFFD4FFD2FEFFFAD7FE1AFADF" box="[371,438,1336,1362]" pageId="6" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3106558FFD4FFD2FF1BFAB3FEBEFAFA" box="[151,274,1372,1399]" pageId="6" pageNumber="145" type="description">
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<paragraph id="8BB536D3FFD4FFD2FF1BFAB3FEBEFAFA" blockId="6.[151,438,1335,1399]" box="[151,274,1372,1399]" pageId="6" pageNumber="145">
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(
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<figureCitation id="13312A56FFD4FFD2FF13FAB3FF43FAFA" box="[159,239,1372,1399]" captionStart="FIGURE 5" captionStartId="7.[151,250,1898,1922]" captionTargetBox="[214,1374,181,1874]" captionTargetId="figure-19@7.[214,1374,181,1874]" captionTargetPageId="7" captionText="FIGURE 5. Galene dashtbani sp. nov.: A–B, dorsal (A) and front (B) stereo-pair views of holotype ZUTC6881, carapace; C–H, dorsal stereo-pair (C), front stereo-pair (D), left lateral (E), right lateral (F), ventral (G) and posterior (H) views of non-type HUIM195, carapace; I–J, dorsal (I) and front (J) views of non-type HUIM196, carapace; K, dorsal view of paratype HUIM197, carapace; L, dorsal view of paratype HUIM198, carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404978" httpUri="https://zenodo.org/record/6404978/files/figure.png" pageId="6" pageNumber="145">Figs. 5</figureCitation>
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,
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<figureCitation id="13312A56FFD4FFD2FF77FAB3FEA6FAFA" box="[251,266,1372,1399]" captionStart="FIGURE 6" captionStartId="8.[151,250,1864,1888]" captionTargetBox="[198,1389,181,1840]" captionTargetId="figure-23@8.[198,1389,181,1840]" captionTargetPageId="8" captionText="FIGURE 6. Galene dashtbani sp. nov.: A–C, dorsal (A), right lateral (B) and ventral (C) views of paratype HUIM199, eroded carapace and partial cheliped; D, dorsal view of paratype HUIM200, eroded carapace; E, dorsal view of paratype HUIM201, eroded carapace; F–G, dorsal (F) and ventral (G) views of paratype HUIM202, eroded carapace and fragmentary cheliped; H–I, ventral (H) and left lateral (I) views of paratype HUIM203, cheliped; J–K, ventral (J) and dorsal (K) views of paratype HUIM204, cheliped and partial carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404980" httpUri="https://zenodo.org/record/6404980/files/figure.png" pageId="6" pageNumber="145">6</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3106558FFD4FFD2FF1BFA72FA86F902" pageId="6" pageNumber="145" type="materials_examined">
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<paragraph id="8BB536D3FFD4FFD2FF1BFA72FD46F9AE" blockId="6.[151,1437,1437,2040]" pageId="6" pageNumber="145">
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||
<emphasis id="B97EEAC1FFD4FFD2FF1BFA72FE90FA3A" bold="true" box="[151,316,1437,1463]" pageId="6" pageNumber="145">Type material</emphasis>
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.
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<materialsCitation id="3B623C8EFFD4FFD2FEC4FA72FDE2FA56" ID-GBIF-Occurrence="3744360302" collectionCode="ZUTC" pageId="6" pageNumber="145" specimenCode="ZUTC6881" specimenCount="1" typeStatus="holotype">
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<emphasis id="B97EEAC1FFD4FFD2FEC4FA72FE03FA3A" box="[328,431,1437,1463]" italics="true" pageId="6" pageNumber="145">
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<typeStatus id="54B18871FFD4FFD2FEC4FA72FE03FA3A" box="[328,431,1437,1463]" pageId="6" pageNumber="145" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
:
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<specimenCode id="DBAC9EA8FFD4FFD2FE37FA72FD93FA3A" box="[443,575,1437,1463]" collectionCode="ZUTC" pageId="6" pageNumber="145">ZUTC6881</specimenCode>
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, a carapace from the Honguyeh section, from the Middle Miocene (Langhian), Guri member of the Mishan Formation
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</materialsCitation>
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.
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<materialsCitation id="3B623C8EFFD4FFD2FDD5FA2DFCF8FA51" ID-GBIF-Occurrence="3744360301" box="[601,852,1473,1500]" collectionCode="HUIM" pageId="6" pageNumber="145" specimenCode="HUIM195" specimenCount="1" typeStatus="paratype">
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<emphasis id="B97EEAC1FFD4FFD2FDD5FA2DFD67FA56" box="[601,715,1474,1499]" italics="true" pageId="6" pageNumber="145">
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<typeStatus id="54B18871FFD4FFD2FDD5FA2DFD67FA56" box="[601,715,1474,1499]" pageId="6" pageNumber="145" type="paratype">Paratypes</typeStatus>
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</emphasis>
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||
:
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<specimenCode id="DBAC9EA8FFD4FFD2FD55FA2EFCFCFA51" box="[729,848,1473,1500]" collectionCode="HUIM" pageId="6" pageNumber="145">HUIM195</specimenCode>
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,
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</materialsCitation>
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<materialsCitation id="3B623C8EFFD4FFD2FCD7FA2EFBAAFA56" ID-GBIF-Occurrence="3744360303" box="[859,1030,1473,1499]" collectionCode="HUIM" pageId="6" pageNumber="145" specimenCode="HUIM198-202" specimenCount="1" typeStatus="paratype">
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<specimenCode id="DBAC9EA8FFD4FFD2FCD7FA2EFBAAFA56" box="[859,1030,1473,1499]" collectionCode="HUIM" pageId="6" pageNumber="145">HUIM198-202</specimenCode>
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</materialsCitation>
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, carapaces from the Gohreh section, from the Middle Miocene (Langhian), Guri member of the Mishan Formation. Measurements of the carapace length and width are provided for each specimen in
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<tableCitation id="C6880368FFD4FFD2FD1CF9E6FD48F9AE" box="[656,740,1545,1571]" captionStart="TABLE 1" captionStartId="9.[151,239,1088,1112]" captionTargetBox="[163,1174,1129,1523]" captionTargetPageId="9" captionText="TABLE 1. Carapace Measurements Carapace Width (mm)" httpUri="http://table.plazi.org/id/DF75665BFFDBFFDDFF1BFBAFFDBBFBD5" pageId="6" pageNumber="145" tableUuid="DF75665BFFDBFFDDFF1BFBAFFDBBFBD5">Table 1</tableCitation>
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.
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</paragraph>
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<paragraph id="8BB536D3FFD4FFD2FF4BF9C2FA86F902" blockId="6.[151,1437,1437,2040]" pageId="6" pageNumber="145">
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||
<emphasis id="B97EEAC1FFD4FFD2FF4BF9C2FE1BF9CA" bold="true" box="[199,439,1581,1607]" pageId="6" pageNumber="145">Additional material</emphasis>
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||
. Non-types:
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<collectionCode id="ED1BAE16FFD4FFD2FDDCF9C2FD0CF9CA" box="[592,672,1581,1607]" pageId="6" pageNumber="145">HUIM</collectionCode>
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196-197, carapaces from the Gohreh section, the Middle Miocene (Langhian) of the Guri Member of the Mishan Formation.
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<collectionCode id="ED1BAE16FFD4FFD2FC83F9BEFCF2F9E1" box="[783,862,1617,1644]" pageId="6" pageNumber="145">HUIM</collectionCode>
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203-205, chelipeds (one with fragmentary carapace), from the Gohreh section, the Middle Miocene (Langhian) of the Guri Member of the Mishan Formation.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3106558FFD4FFD2FF4BF976FE7CF95A" pageId="6" pageNumber="145" type="diagnosis">
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||
<paragraph id="8BB536D3FFD4FFD2FF4BF976FE7CF95A" blockId="6.[151,1437,1437,2040]" pageId="6" pageNumber="145">
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<emphasis id="B97EEAC1FFD4FFD2FF4BF976FE96F93E" bold="true" box="[199,314,1689,1715]" pageId="6" pageNumber="145">Diagnosis</emphasis>
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. Carapace with smooth dorsal surface, no posterolateral spines, frontal and orbital lobes with swollen tuberosities, with no spines.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3106558FFD4FFDDFF4BF90EFB34FF58" lastPageId="9" lastPageNumber="148" pageId="6" pageNumber="145" type="description">
|
||
<paragraph id="8BB536D3FFD4FFD2FF4BF90EFA87F875" blockId="6.[151,1437,1437,2040]" pageId="6" pageNumber="145">
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||
<emphasis id="B97EEAC1FFD4FFD2FF4BF90EFEFDF976" bold="true" box="[199,337,1761,1787]" pageId="6" pageNumber="145">Description</emphasis>
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. Carapace subhexagonal, anterior margins more ovate; posterior margins with well-defined 45° angle between posterolateral and posterior margins; two anterolateral blunt teeth on epibranchial margins of carapace; not very sharp, but clearly distinguished (
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<figureCitation id="13312A56FFD4FFD2FD5BF8C6FCF9F8C9" box="[727,853,1833,1860]" captionStart="FIGURE 5" captionStartId="7.[151,250,1898,1922]" captionTargetBox="[214,1374,181,1874]" captionTargetId="figure-19@7.[214,1374,181,1874]" captionTargetPageId="7" captionText="FIGURE 5. Galene dashtbani sp. nov.: A–B, dorsal (A) and front (B) stereo-pair views of holotype ZUTC6881, carapace; C–H, dorsal stereo-pair (C), front stereo-pair (D), left lateral (E), right lateral (F), ventral (G) and posterior (H) views of non-type HUIM195, carapace; I–J, dorsal (I) and front (J) views of non-type HUIM196, carapace; K, dorsal view of paratype HUIM197, carapace; L, dorsal view of paratype HUIM198, carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404978" httpUri="https://zenodo.org/record/6404978/files/figure.png" pageId="6" pageNumber="145">Figs. 5E, F</figureCitation>
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), no posterolateral tuberosities (
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<figureCitation id="13312A56FFD4FFD2FB37F8C6FA34F8C9" box="[1211,1432,1833,1860]" captionStart="FIGURE 5" captionStartId="7.[151,250,1898,1922]" captionTargetBox="[214,1374,181,1874]" captionTargetId="figure-19@7.[214,1374,181,1874]" captionTargetPageId="7" captionText="FIGURE 5. Galene dashtbani sp. nov.: A–B, dorsal (A) and front (B) stereo-pair views of holotype ZUTC6881, carapace; C–H, dorsal stereo-pair (C), front stereo-pair (D), left lateral (E), right lateral (F), ventral (G) and posterior (H) views of non-type HUIM195, carapace; I–J, dorsal (I) and front (J) views of non-type HUIM196, carapace; K, dorsal view of paratype HUIM197, carapace; L, dorsal view of paratype HUIM198, carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404978" httpUri="https://zenodo.org/record/6404978/files/figure.png" pageId="6" pageNumber="145">Figs. 5A, C, E–I, K</figureCitation>
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,
|
||
<figureCitation id="13312A56FFD4FFD2FF1BF8A2FF53F8E5" box="[151,255,1869,1896]" captionStart="FIGURE 6" captionStartId="8.[151,250,1864,1888]" captionTargetBox="[198,1389,181,1840]" captionTargetId="figure-23@8.[198,1389,181,1840]" captionTargetPageId="8" captionText="FIGURE 6. Galene dashtbani sp. nov.: A–C, dorsal (A), right lateral (B) and ventral (C) views of paratype HUIM199, eroded carapace and partial cheliped; D, dorsal view of paratype HUIM200, eroded carapace; E, dorsal view of paratype HUIM201, eroded carapace; F–G, dorsal (F) and ventral (G) views of paratype HUIM202, eroded carapace and fragmentary cheliped; H–I, ventral (H) and left lateral (I) views of paratype HUIM203, cheliped; J–K, ventral (J) and dorsal (K) views of paratype HUIM204, cheliped and partial carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404980" httpUri="https://zenodo.org/record/6404980/files/figure.png" pageId="6" pageNumber="145">6A, D–F</figureCitation>
|
||
); frontal margin bilobed. Dorsal surface of carapace smooth, longitudinally slightly convex; cardiac and mesogasteric regions defined with prominent fissures; lateral borders of anterior mesogastric region sub-parallel, extending close to the frontal lobe with narrow furrow (
|
||
<figureCitation id="13312A56FFD4FFD2FC80F87AFC63F83D" box="[780,975,1941,1968]" captionStart="FIGURE 5" captionStartId="7.[151,250,1898,1922]" captionTargetBox="[214,1374,181,1874]" captionTargetId="figure-19@7.[214,1374,181,1874]" captionTargetPageId="7" captionText="FIGURE 5. Galene dashtbani sp. nov.: A–B, dorsal (A) and front (B) stereo-pair views of holotype ZUTC6881, carapace; C–H, dorsal stereo-pair (C), front stereo-pair (D), left lateral (E), right lateral (F), ventral (G) and posterior (H) views of non-type HUIM195, carapace; I–J, dorsal (I) and front (J) views of non-type HUIM196, carapace; K, dorsal view of paratype HUIM197, carapace; L, dorsal view of paratype HUIM198, carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404978" httpUri="https://zenodo.org/record/6404978/files/figure.png" pageId="6" pageNumber="145">Figs. 5A–D, I–K</figureCitation>
|
||
); other regions of carapace not markedly prominent. Thoracic sternum and male pleon narrow (HUIM199, HUIM202,
|
||
<figureCitation id="13312A56FFD4FFD2FB95F856FB09F85E" box="[1049,1189,1977,2003]" captionStart="FIGURE 6" captionStartId="8.[151,250,1864,1888]" captionTargetBox="[198,1389,181,1840]" captionTargetId="figure-23@8.[198,1389,181,1840]" captionTargetPageId="8" captionText="FIGURE 6. Galene dashtbani sp. nov.: A–C, dorsal (A), right lateral (B) and ventral (C) views of paratype HUIM199, eroded carapace and partial cheliped; D, dorsal view of paratype HUIM200, eroded carapace; E, dorsal view of paratype HUIM201, eroded carapace; F–G, dorsal (F) and ventral (G) views of paratype HUIM202, eroded carapace and fragmentary cheliped; H–I, ventral (H) and left lateral (I) views of paratype HUIM203, cheliped; J–K, ventral (J) and dorsal (K) views of paratype HUIM204, cheliped and partial carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404980" httpUri="https://zenodo.org/record/6404980/files/figure.png" pageId="6" pageNumber="145">Figs. 6C, G</figureCitation>
|
||
). Surface of cheliped manus smooth; palm with posterior tuberculated margin (
|
||
<figureCitation id="13312A56FFD4FFD2FC96F832FCC5F875" box="[794,873,2013,2040]" captionStart="FIGURE 6" captionStartId="8.[151,250,1864,1888]" captionTargetBox="[198,1389,181,1840]" captionTargetId="figure-23@8.[198,1389,181,1840]" captionTargetPageId="8" captionText="FIGURE 6. Galene dashtbani sp. nov.: A–C, dorsal (A), right lateral (B) and ventral (C) views of paratype HUIM199, eroded carapace and partial cheliped; D, dorsal view of paratype HUIM200, eroded carapace; E, dorsal view of paratype HUIM201, eroded carapace; F–G, dorsal (F) and ventral (G) views of paratype HUIM202, eroded carapace and fragmentary cheliped; H–I, ventral (H) and left lateral (I) views of paratype HUIM203, cheliped; J–K, ventral (J) and dorsal (K) views of paratype HUIM204, cheliped and partial carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404980" httpUri="https://zenodo.org/record/6404980/files/figure.png" pageId="6" pageNumber="145">Fig. 6I</figureCitation>
|
||
); dactylus shorter than pollex (
|
||
<figureCitation id="13312A56FFD4FFD2FB48F832FAB2F875" box="[1220,1310,2013,2040]" captionStart="FIGURE 6" captionStartId="8.[151,250,1864,1888]" captionTargetBox="[198,1389,181,1840]" captionTargetId="figure-23@8.[198,1389,181,1840]" captionTargetPageId="8" captionText="FIGURE 6. Galene dashtbani sp. nov.: A–C, dorsal (A), right lateral (B) and ventral (C) views of paratype HUIM199, eroded carapace and partial cheliped; D, dorsal view of paratype HUIM200, eroded carapace; E, dorsal view of paratype HUIM201, eroded carapace; F–G, dorsal (F) and ventral (G) views of paratype HUIM202, eroded carapace and fragmentary cheliped; H–I, ventral (H) and left lateral (I) views of paratype HUIM203, cheliped; J–K, ventral (J) and dorsal (K) views of paratype HUIM204, cheliped and partial carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404980" httpUri="https://zenodo.org/record/6404980/files/figure.png" pageId="6" pageNumber="145">Fig. 6H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF75665BFFD5FFD3FF1BF885FC16F860" ID-DOI="http://doi.org/10.5281/zenodo.6404978" ID-Zenodo-Dep="6404978" httpUri="https://zenodo.org/record/6404978/files/figure.png" pageId="7" pageNumber="146" startId="7.[151,250,1898,1922]" targetBox="[214,1374,181,1874]" targetPageId="7">
|
||
<paragraph id="8BB536D3FFD5FFD3FF1BF885FC16F860" blockId="7.[151,1436,1898,2030]" pageId="7" pageNumber="146">
|
||
<emphasis id="B97EEAC1FFD5FFD3FF1BF885FEBEF80F" bold="true" box="[151,274,1898,1922]" pageId="7" pageNumber="146">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C0A4D50FFD5FFD3FE9BF885FE65F80F" authority="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný, 2022" authorityName="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný" authorityYear="2022" box="[279,457,1898,1922]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="7" pageNumber="146" phylum="Rhodophyta" rank="species" species="dashtbani" status="sp. nov.">
|
||
<emphasis id="B97EEAC1FFD5FFD3FE9BF885FE65F80F" box="[279,457,1898,1922]" italics="true" pageId="7" pageNumber="146">Galene dashtbani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAC1FFD5FFD3FE43F885FDB2F80F" bold="true" box="[463,542,1898,1922]" pageId="7" pageNumber="146">
|
||
<taxonomicNameLabel id="A24D57BAFFD5FFD3FE43F885FDB2F80F" box="[463,542,1898,1922]" pageId="7" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: A–B, dorsal (A) and front (B) stereo-pair views of holotype ZUTC6881, carapace; C–H, dorsal stereo-pair (C), front stereo-pair (D), left lateral (E), right lateral (F), ventral (G) and posterior (H) views of non-type HUIM195, carapace; I–J, dorsal (I) and front (J) views of non-type HUIM196, carapace; K, dorsal view of paratype HUIM197, carapace; L, dorsal view of paratype HUIM198, carapace. Scale bars = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF75665BFFDAFFDCFF1BF8A7FCBDF862" ID-DOI="http://doi.org/10.5281/zenodo.6404980" ID-Zenodo-Dep="6404980" httpUri="https://zenodo.org/record/6404980/files/figure.png" pageId="8" pageNumber="147" startId="8.[151,250,1864,1888]" targetBox="[198,1389,181,1840]" targetPageId="8">
|
||
<paragraph id="8BB536D3FFDAFFDCFF1BF8A7FCBDF862" blockId="8.[151,1437,1864,2032]" pageId="8" pageNumber="147">
|
||
<emphasis id="B97EEAC1FFDAFFDCFF1BF8A7FEBFF8ED" bold="true" box="[151,275,1864,1888]" pageId="8" pageNumber="147">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C0A4D50FFDAFFDCFE95F8A7FE61F8ED" authority="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný, 2022" authorityName="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný" authorityYear="2022" box="[281,461,1864,1888]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="8" pageNumber="147" phylum="Rhodophyta" rank="species" species="dashtbani" status="sp. nov.">
|
||
<emphasis id="B97EEAC1FFDAFFDCFE95F8A7FE61F8ED" box="[281,461,1864,1888]" italics="true" pageId="8" pageNumber="147">Galene dashtbani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAC1FFDAFFDCFE5FF8A7FD81F8ED" bold="true" box="[467,557,1864,1888]" pageId="8" pageNumber="147">
|
||
<taxonomicNameLabel id="A24D57BAFFDAFFDCFE5FF8A7FD84F8ED" box="[467,552,1864,1888]" pageId="8" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
|
||
:
|
||
</emphasis>
|
||
A–C, dorsal (A), right lateral (B) and ventral (C) views of paratype HUIM199, eroded carapace and partial cheliped; D, dorsal view of paratype HUIM200, eroded carapace; E, dorsal view of paratype HUIM201, eroded carapace; F–G, dorsal (F) and ventral (G) views of paratype HUIM202, eroded carapace and fragmentary cheliped; H–I, ventral (H) and left lateral (I) views of paratype HUIM203, cheliped; J–K, ventral (J) and dorsal (K) views of paratype HUIM204, cheliped and partial carapace. Scale bars = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB536D3FFDBFFDDFF4BFF79FB34FF58" blockId="9.[151,1437,150,1041]" pageId="9" pageNumber="148">
|
||
<emphasis id="B97EEAC1FFDBFFDDFF4BFF79FEFEFF3D" bold="true" box="[199,338,150,176]" pageId="9" pageNumber="148">Occurrence</emphasis>
|
||
. The species is known from the Middle Miocene (Langhian) of the Guri Member of the Mishan Formation exposed at Honguyeh and Gohreh sections,
|
||
<collectingRegion id="49CEF831FFDBFFDDFD73FF54FC5CFF58" box="[767,1008,187,213]" country="Iran" name="Hormozgan" pageId="9" pageNumber="148">Hormozgan Province</collectingRegion>
|
||
, southern
|
||
<collectingCountry id="F31D7643FFDBFFDDFBEFFF54FB38FF58" box="[1123,1172,187,213]" name="Iran" pageId="9" pageNumber="148">Iran</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3106558FFDBFFDDFF4BFF30FC72FE90" pageId="9" pageNumber="148" type="etymology">
|
||
<paragraph id="8BB536D3FFDBFFDDFF4BFF30FC72FE90" blockId="9.[151,1437,150,1041]" pageId="9" pageNumber="148">
|
||
<emphasis id="B97EEAC1FFDBFFDDFF4BFF30FEE4FF74" bold="true" box="[199,328,223,249]" pageId="9" pageNumber="148">Etymology</emphasis>
|
||
. This species is named in honor of Mr.Houshang Dashtban; a veteran paleontologist, who contributed deeply into the exploration and the study of Iranian fossils including crabs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3106558FFDBFFDDFF4BFEC8FE0FFB9C" pageId="9" pageNumber="148" type="discussion">
|
||
<paragraph id="8BB536D3FFDBFFDDFF4BFEC8FBB0FD24" blockId="9.[151,1437,150,1041]" pageId="9" pageNumber="148">
|
||
<emphasis id="B97EEAC1FFDBFFDDFF4BFEC8FE98FECC" bold="true" box="[199,308,295,321]" pageId="9" pageNumber="148">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFEC4FEC8FDBFFECC" authority="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný, 2022" authorityName="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný" authorityYear="2022" box="[328,531,295,321]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="9" pageNumber="148" phylum="Rhodophyta" rank="species" species="dashtbani" status="sp. nov.">
|
||
<emphasis id="B97EEAC1FFDBFFDDFEC4FEC8FDBFFECC" box="[328,531,295,321]" italics="true" pageId="9" pageNumber="148">Galene dashtbani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAC1FFDBFFDDFDACFEC8FDD3FECC" bold="true" box="[544,639,295,321]" pageId="9" pageNumber="148">
|
||
<taxonomicNameLabel id="A24D57BAFFDBFFDDFDACFEC8FDD3FECC" box="[544,639,295,321]" pageId="9" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is different from its congeners by having a combination of mixed characteristics. In having less prominent lateral spines, no posterolateral spines and a less protuberant cardiac region (
|
||
<figureCitation id="13312A56FFDBFFDDFF13FE80FECAFE04" box="[159,358,366,393]" captionStart="FIGURE 5" captionStartId="7.[151,250,1898,1922]" captionTargetBox="[214,1374,181,1874]" captionTargetId="figure-19@7.[214,1374,181,1874]" captionTargetPageId="7" captionText="FIGURE 5. Galene dashtbani sp. nov.: A–B, dorsal (A) and front (B) stereo-pair views of holotype ZUTC6881, carapace; C–H, dorsal stereo-pair (C), front stereo-pair (D), left lateral (E), right lateral (F), ventral (G) and posterior (H) views of non-type HUIM195, carapace; I–J, dorsal (I) and front (J) views of non-type HUIM196, carapace; K, dorsal view of paratype HUIM197, carapace; L, dorsal view of paratype HUIM198, carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404978" httpUri="https://zenodo.org/record/6404978/files/figure.png" pageId="9" pageNumber="148">Figs. 5A, C, H, L</figureCitation>
|
||
),
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFEF7FE80FDEDFE04" authority="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný, 2022" authorityName="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný" authorityYear="2022" box="[379,577,367,393]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="9" pageNumber="148" phylum="Rhodophyta" rank="species" species="dashtbani" status="sp. nov.">
|
||
<emphasis id="B97EEAC1FFDBFFDDFEF7FE80FDEDFE04" box="[379,577,367,393]" italics="true" pageId="9" pageNumber="148">Galene dashtbani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAC1FFDBFFDDFDC5FE80FD0EFE04" bold="true" box="[585,674,367,393]" pageId="9" pageNumber="148">
|
||
<taxonomicNameLabel id="A24D57BAFFDBFFDDFDC5FE80FD0EFE04" box="[585,674,367,393]" pageId="9" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is different from
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFCE4FE80FB87FE04" authorityName="Ng et al." authorityYear="2008" baseAuthorityName="Herbst" baseAuthorityYear="1783" box="[872,1067,367,393]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="9" pageNumber="148" phylum="Rhodophyta" rank="species" species="bispinosa">
|
||
<emphasis id="B97EEAC1FFDBFFDDFCE4FE80FB87FE04" box="[872,1067,367,393]" italics="true" pageId="9" pageNumber="148">Galene bispinosa</emphasis>
|
||
</taxonomicName>
|
||
, including specimens previously documented as
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFECDFE7CFDEDFE21" authorityName="Desmarest" authorityYear="1822" box="[321,577,403,429]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="148" phylum="Arthropoda" rank="species" species="trispinosus">
|
||
<emphasis id="B97EEAC1FFDBFFDDFECDFE7CFDEDFE21" box="[321,577,403,429]" italics="true" pageId="9" pageNumber="148">Gecarcinus trispinosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFDFFFE7CFCF1FE20" authorityName="McCoy" authorityYear="1849" box="[627,861,403,429]" class="Florideophyceae" family="Halymeniaceae" genus="Podopilumnus" kingdom="Plantae" order="Halymeniales" pageId="9" pageNumber="148" phylum="Rhodophyta" rank="species" species="fittoni">
|
||
<emphasis id="B97EEAC1FFDBFFDDFDFFFE7CFCF1FE20" box="[627,861,403,429]" italics="true" pageId="9" pageNumber="148">Podopilumnus fittoni</emphasis>
|
||
</taxonomicName>
|
||
(also see
|
||
<bibRefCitation id="EF9B4B22FFDBFFDDFC49FE7CFABCFE20" author="Etheridge, R. E. & McCulloch, A. R." box="[965,1296,403,429]" pageId="9" pageNumber="148" pagination="1 - 14" refId="ref6561" refString="Etheridge, R. E. & McCulloch, A. R. (1916) Sub-fossil crustaceans from the coasts of Australia. Records of the Australian Museum, 11, 1 - 14, pls. 1 - 7. https: // doi. org / 10.3853 / j. 0067 - 1975.11.1916.906" type="journal article" year="1916">Etheridge & McCulloch 1916</bibRefCitation>
|
||
: pl. III fig. 3;
|
||
<bibRefCitation id="EF9B4B22FFDBFFDDFF1BFE58FE22FE5C" author="Valinassab, T. & Fatemi, S. & Ghotbeddin, N." box="[151,398,439,465]" pageId="9" pageNumber="148" pagination="439 - 441" refId="ref10023" refString="Valinassab, T., Fatemi, S. & Ghotbeddin, N. (2012) First record of Galene bispinosa Herbst, 1783 [Decapoda; Brachyura; Galenidae] from the Gulf of Oman, Iran. World Applied Sciences Journal, 17 (4), 439 - 441." type="journal article" year="2012">
|
||
Valinassab
|
||
<emphasis id="B97EEAC1FFDBFFDDFE99FE58FEE0FE5C" box="[277,332,439,465]" italics="true" pageId="9" pageNumber="148">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
: fig. 2a;
|
||
<bibRefCitation id="EF9B4B22FFDBFFDDFE61FE58FD22FE5C" author="Ng, P. K. L. & Shih, H. T. & Ho, P. H. & Wang, C. H." box="[493,654,439,465]" pageId="9" pageNumber="148" pagination="1 - 185" refId="ref9048" refString="Ng, P. K. L., Shih, H. T., Ho, P. H. & Wang, C. H. (2017) An updated annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Journal of the National Taiwan Museum, 70 (3 & 4), 1 - 185." type="journal article" year="2017">
|
||
Ng
|
||
<emphasis id="B97EEAC1FFDBFFDDFD99FE58FDE0FE5C" box="[533,588,439,465]" italics="true" pageId="9" pageNumber="148">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
: fig. 5h),
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFD7AFE58FB08FE5C" authority="Morris & Collins, 1991" authorityName="Morris & Collins" authorityYear="1991" box="[758,1188,439,465]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="9" pageNumber="148" phylum="Rhodophyta" rank="species" species="stipata">
|
||
<emphasis id="B97EEAC1FFDBFFDDFD7AFE58FC35FE5D" box="[758,921,439,465]" italics="true" pageId="9" pageNumber="148">Galene stipata</emphasis>
|
||
<bibRefCitation id="EF9B4B22FFDBFFDDFC2CFE58FB08FE5C" author="Morris, S. F. & Collins, J. S." box="[928,1188,439,465]" pageId="9" pageNumber="148" pagination="1 - 33" refId="ref8616" refString="Morris, S. F. & Collins, J. S. (1991) Neogene crabs from Brunei, Sabah and Sarawak. Bulletin of the British Museum (Natural History) Geology, 47 (1), 1 - 33." type="journal article" year="1991">Morris & Collins, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF9B4B22FFDBFFDDFB51FE58FF42FE78" author="Morris, S. F. & Collins, J. S." pageId="9" pageNumber="148" pagination="1 - 33" refId="ref8616" refString="Morris, S. F. & Collins, J. S. (1991) Neogene crabs from Brunei, Sabah and Sarawak. Bulletin of the British Museum (Natural History) Geology, 47 (1), 1 - 33." type="journal article" year="1991">Morris & Collins 1991: 8</bibRefCitation>
|
||
, fig. 54a),
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFEE0FE34FC93FE78" authority="A. Milne-Edwards, 1865" authorityName="A. Milne-Edwards" authorityYear="1865" box="[364,831,474,501]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="9" pageNumber="148" phylum="Rhodophyta" rank="species" species="obscura">
|
||
<emphasis id="B97EEAC1FFDBFFDDFEE0FE34FDB3FE78" box="[364,543,475,501]" italics="true" pageId="9" pageNumber="148">Galene obscura</emphasis>
|
||
A.
|
||
<bibRefCitation id="EF9B4B22FFDBFFDDFDC5FE34FC93FE78" author="Milne-Edwards, A." box="[585,831,474,501]" pageId="9" pageNumber="148" pagination="297 - 351" refId="ref8583" refString="Milne-Edwards, A. (1865) Monographie des crustaces fossiles de la famille des canceriens. Annales des Sciences naturelles, Zoologie, 5 (3), 297 - 351." type="journal article" year="1865">Milne-Edwards, 1865</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF9B4B22FFDBFFDDFCF1FE34FBFFFE78" author="Collins, J. S. & Lee, C. & Noad, J." box="[893,1107,475,501]" pageId="9" pageNumber="148" pagination="187 - 226" refId="ref5743" refString="Collins, J. S., Lee, C. & Noad, J. (2003) Miocene and Pleistocene crabs (Crustacea, Decapoda) from Sabah and Sarawak. Journal of Systematic Paleontology, 1 (3), 187 - 226. https: // doi. org / 10.1017 / S 1477201903001068" type="journal article" year="2003">
|
||
Collins
|
||
<emphasis id="B97EEAC1FFDBFFDDFC5AFE34FBA3FE78" box="[982,1039,475,501]" italics="true" pageId="9" pageNumber="148">et al.</emphasis>
|
||
2003
|
||
</bibRefCitation>
|
||
: fig. 6a) and
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFB66FE34FE65FD94" authority="Collins, Lee & Noad, 2003" authorityName="Collins, Lee & Noad" authorityYear="2003" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="9" pageNumber="148" phylum="Rhodophyta" rank="species" species="litoralis">
|
||
<emphasis id="B97EEAC1FFDBFFDDFB66FE34FA31FE78" box="[1258,1437,475,501]" italics="true" pageId="9" pageNumber="148">Galene litoralis</emphasis>
|
||
<bibRefCitation id="EF9B4B22FFDBFFDDFF1BFE10FE65FD94" author="Collins, J. S. & Lee, C. & Noad, J." box="[151,457,511,537]" pageId="9" pageNumber="148" pagination="187 - 226" refId="ref5743" refString="Collins, J. S., Lee, C. & Noad, J. (2003) Miocene and Pleistocene crabs (Crustacea, Decapoda) from Sabah and Sarawak. Journal of Systematic Paleontology, 1 (3), 187 - 226. https: // doi. org / 10.1017 / S 1477201903001068" type="journal article" year="2003">Collins, Lee & Noad, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF9B4B22FFDBFFDDFD89FE10FCBDFD94" author="Collins, J. S. & Lee, C. & Noad, J." box="[517,785,511,537]" pageId="9" pageNumber="148" pagination="187 - 226" refId="ref5743" refString="Collins, J. S., Lee, C. & Noad, J. (2003) Miocene and Pleistocene crabs (Crustacea, Decapoda) from Sabah and Sarawak. Journal of Systematic Paleontology, 1 (3), 187 - 226. https: // doi. org / 10.1017 / S 1477201903001068" type="journal article" year="2003">
|
||
Collins
|
||
<emphasis id="B97EEAC1FFDBFFDDFDD2FE10FD3BFD94" box="[606,663,511,537]" italics="true" pageId="9" pageNumber="148">et al.</emphasis>
|
||
2003: 221
|
||
</bibRefCitation>
|
||
, fig. 7a). The carapace frontal margin of
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFB6EFE10FADFFD94" authority="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný, 2022" authorityName="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný" authorityYear="2022" box="[1250,1395,511,537]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="9" pageNumber="148" phylum="Rhodophyta" rank="species" species="dashtbani" status="sp. nov.">
|
||
<emphasis id="B97EEAC1FFDBFFDDFB6EFE10FADFFD94" box="[1250,1395,511,537]" italics="true" pageId="9" pageNumber="148">G. dashtbani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAC1FFDBFFDDFAF7FE10FF64FDB0" bold="true" pageId="9" pageNumber="148">
|
||
<taxonomicNameLabel id="A24D57BAFFDBFFDDFAF7FE10FF64FDB0" pageId="9" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
comprises half the orbitofrontal margin, compared to less than half in
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFC6CFDCCFA87FDB0" authority="Lin, 1947" authorityName="Lin" authorityYear="1947" box="[992,1323,547,573]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="9" pageNumber="148" phylum="Rhodophyta" rank="species" species="granulifera">
|
||
<emphasis id="B97EEAC1FFDBFFDDFC6CFDCCFB14FDB0" box="[992,1208,547,573]" italics="true" pageId="9" pageNumber="148">Galene granulifera</emphasis>
|
||
<bibRefCitation id="EF9B4B22FFDBFFDDFB32FDCCFA87FDB0" author="Lin, C." box="[1214,1323,547,573]" pageId="9" pageNumber="148" pagination="139 - 154" refId="ref8461" refString="Lin, C. (1947) On a new xanthid crab from the Neogene Formation of Taiwan. Acta Geologica Taiwanica, 1 (2), 139 - 154." type="journal article" year="1947">Lin, 1947</bibRefCitation>
|
||
</taxonomicName>
|
||
(compare
|
||
<bibRefCitation id="EF9B4B22FFDBFFDDFF1BFDA8FEEEFDEC" author="Hu, C. H. & Tao, H. J." box="[151,322,583,609]" pageId="9" pageNumber="148" pagination="537 - 555" refId="ref7824" refString="Hu, C. H. & Tao, H. J. (2004) Studeis on the Neogene crabs from South-Western foot hills of Taiwan. Acta Palaeontologica Sinica, 43 (4), 537 - 555." type="journal article" year="2004">Hu & Tao 2004</bibRefCitation>
|
||
L pl. V figs. 4,7) and
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFDA1FDA8FD05FDEC" authorityName="A. Milne-Edwards" authorityYear="1865" box="[557,681,583,609]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="9" pageNumber="148" phylum="Rhodophyta" rank="species" species="obscura">
|
||
<emphasis id="B97EEAC1FFDBFFDDFDA1FDA8FD05FDEC" box="[557,681,583,609]" italics="true" pageId="9" pageNumber="148">G. obscura</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF9B4B22FFDBFFDDFD6EFDA8FC1FFDEC" author="Collins, J. S. & Lee, C. & Noad, J." box="[738,947,583,609]" pageId="9" pageNumber="148" pagination="187 - 226" refId="ref5743" refString="Collins, J. S., Lee, C. & Noad, J. (2003) Miocene and Pleistocene crabs (Crustacea, Decapoda) from Sabah and Sarawak. Journal of Systematic Paleontology, 1 (3), 187 - 226. https: // doi. org / 10.1017 / S 1477201903001068" type="journal article" year="2003">
|
||
Collins
|
||
<emphasis id="B97EEAC1FFDBFFDDFCB6FDA8FCDDFDEC" box="[826,881,583,609]" italics="true" pageId="9" pageNumber="148">et al.</emphasis>
|
||
2003
|
||
</bibRefCitation>
|
||
: figs. 6a, 6c).
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFBC0FDA8FABDFDEC" authority="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný, 2022" authorityName="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný" authorityYear="2022" box="[1100,1297,583,609]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="9" pageNumber="148" phylum="Rhodophyta" rank="species" species="dashtbani" status="sp. nov.">
|
||
<emphasis id="B97EEAC1FFDBFFDDFBC0FDA8FABDFDEC" box="[1100,1297,583,609]" italics="true" pageId="9" pageNumber="148">Galene dashtbani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAC1FFDBFFDDFA94FDA8FADDFDEC" bold="true" box="[1304,1393,583,609]" pageId="9" pageNumber="148">
|
||
<taxonomicNameLabel id="A24D57BAFFDBFFDDFA94FDA8FADDFDEC" box="[1304,1393,583,609]" pageId="9" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a smoother carapace surface, both in smaller and larger specimens (
|
||
<figureCitation id="13312A56FFDBFFDDFC18FD84FBFAFD08" box="[916,1110,618,645]" captionStart="FIGURE 5" captionStartId="7.[151,250,1898,1922]" captionTargetBox="[214,1374,181,1874]" captionTargetId="figure-19@7.[214,1374,181,1874]" captionTargetPageId="7" captionText="FIGURE 5. Galene dashtbani sp. nov.: A–B, dorsal (A) and front (B) stereo-pair views of holotype ZUTC6881, carapace; C–H, dorsal stereo-pair (C), front stereo-pair (D), left lateral (E), right lateral (F), ventral (G) and posterior (H) views of non-type HUIM195, carapace; I–J, dorsal (I) and front (J) views of non-type HUIM196, carapace; K, dorsal view of paratype HUIM197, carapace; L, dorsal view of paratype HUIM198, carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404978" httpUri="https://zenodo.org/record/6404978/files/figure.png" pageId="9" pageNumber="148">Figs. 5A, C, I, K</figureCitation>
|
||
), in contrast to
|
||
<emphasis id="B97EEAC1FFDBFFDDFA8BFD84FA30FD08" box="[1287,1436,619,645]" italics="true" pageId="9" pageNumber="148">
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFA8BFD84FA34FD08" authorityName="Ng et al." authorityYear="2008" baseAuthorityName="Herbst" baseAuthorityYear="1783" box="[1287,1432,619,645]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="9" pageNumber="148" phylum="Rhodophyta" rank="species" species="bispinosa">G. bispinosa</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
which bears a more granular carapace in smaller specimens (
|
||
<bibRefCitation id="EF9B4B22FFDBFFDDFCCCFD60FBA3FD24" author="Ng, P. K. L. & Wang, C. H. & Ho, P. H. & Shih, H. T." box="[832,1039,655,681]" pageId="9" pageNumber="148" pagination="86" refId="ref9113" refString="Ng, P. K. L., Wang, C. H., Ho, P. H. & Shih, H. T. (2001) An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum Special Publication Series no. 11, 86." type="journal article" year="2001">
|
||
Ng
|
||
<emphasis id="B97EEAC1FFDBFFDDFCE6FD60FC0EFD24" box="[874,930,655,681]" italics="true" pageId="9" pageNumber="148">et al.</emphasis>
|
||
2001: 31
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB536D3FFDBFFDDFF4BFD5CFACEFCB4" blockId="9.[151,1437,150,1041]" pageId="9" pageNumber="148">
|
||
The most prominent character of
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFDCAFD5CFCA2FD40" authority="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný, 2022" authorityName="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný" authorityYear="2022" box="[582,782,691,717]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="9" pageNumber="148" phylum="Rhodophyta" rank="species" species="dashtbani" status="sp. nov.">
|
||
<emphasis id="B97EEAC1FFDBFFDDFDCAFD5CFCA2FD40" box="[582,782,691,717]" italics="true" pageId="9" pageNumber="148">Galene dashtbani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAC1FFDBFFDDFC9BFD5CFCDFFD40" bold="true" box="[791,883,691,717]" pageId="9" pageNumber="148">
|
||
<taxonomicNameLabel id="A24D57BAFFDBFFDDFC9BFD5CFCDFFD40" box="[791,883,691,717]" pageId="9" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the spineless and globular form of the frontal margin and superior margin of the orbit, at the same time, bearing swollen tuberosities on these regions (
|
||
<figureCitation id="13312A56FFDBFFDDFAB9FD38FF61FC98" captionStart="FIGURE 5" captionStartId="7.[151,250,1898,1922]" captionTargetBox="[214,1374,181,1874]" captionTargetId="figure-19@7.[214,1374,181,1874]" captionTargetPageId="7" captionText="FIGURE 5. Galene dashtbani sp. nov.: A–B, dorsal (A) and front (B) stereo-pair views of holotype ZUTC6881, carapace; C–H, dorsal stereo-pair (C), front stereo-pair (D), left lateral (E), right lateral (F), ventral (G) and posterior (H) views of non-type HUIM195, carapace; I–J, dorsal (I) and front (J) views of non-type HUIM196, carapace; K, dorsal view of paratype HUIM197, carapace; L, dorsal view of paratype HUIM198, carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404978" httpUri="https://zenodo.org/record/6404978/files/figure.png" pageId="9" pageNumber="148">Figs. 5B, D, E</figureCitation>
|
||
). Despite the size, this feature is prominent in all studied specimens with a well-preserved frontal margin. Even in the smallest specimen (HUIM195,
|
||
<figureCitation id="13312A56FFDBFFDDFDB1FCF0FD11FCB4" box="[573,701,798,825]" captionStart="FIGURE 5" captionStartId="7.[151,250,1898,1922]" captionTargetBox="[214,1374,181,1874]" captionTargetId="figure-19@7.[214,1374,181,1874]" captionTargetPageId="7" captionText="FIGURE 5. Galene dashtbani sp. nov.: A–B, dorsal (A) and front (B) stereo-pair views of holotype ZUTC6881, carapace; C–H, dorsal stereo-pair (C), front stereo-pair (D), left lateral (E), right lateral (F), ventral (G) and posterior (H) views of non-type HUIM195, carapace; I–J, dorsal (I) and front (J) views of non-type HUIM196, carapace; K, dorsal view of paratype HUIM197, carapace; L, dorsal view of paratype HUIM198, carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404978" httpUri="https://zenodo.org/record/6404978/files/figure.png" pageId="9" pageNumber="148">Figs. 5C–F</figureCitation>
|
||
), the globular form of frontal and orbital lobes is prominent.
|
||
</paragraph>
|
||
<paragraph id="8BB536D3FFDBFFDDFF4BFCACFE0FFB9C" blockId="9.[151,1437,150,1041]" pageId="9" pageNumber="148">
|
||
Two small specimens (HUIM196, HUIM197) of
|
||
<taxonomicName id="4C0A4D50FFDBFFDDFD6CFCACFC0FFCD0" authority="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný, 2022" authorityName="Khosravi & Sari & Mirziee-Ataabadi & Gholamalian & Hyžný" authorityYear="2022" box="[736,931,835,861]" class="Florideophyceae" family="Halymeniaceae" genus="Galene" kingdom="Plantae" order="Halymeniales" pageId="9" pageNumber="148" phylum="Rhodophyta" rank="species" species="dashtbani" status="sp. nov.">
|
||
<emphasis id="B97EEAC1FFDBFFDDFD6CFCACFC0FFCD0" box="[736,931,835,861]" italics="true" pageId="9" pageNumber="148">Galene dashtbani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97EEAC1FFDBFFDDFC24FCACFC53FCD0" bold="true" box="[936,1023,835,861]" pageId="9" pageNumber="148">
|
||
<taxonomicNameLabel id="A24D57BAFFDBFFDDFC24FCACFC53FCD0" box="[936,1023,835,861]" pageId="9" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
appear to have a proportionally wider carapace (
|
||
<figureCitation id="13312A56FFDBFFDDFE86FC88FE25FC0C" box="[266,393,870,897]" captionStart="FIGURE 5" captionStartId="7.[151,250,1898,1922]" captionTargetBox="[214,1374,181,1874]" captionTargetId="figure-19@7.[214,1374,181,1874]" captionTargetPageId="7" captionText="FIGURE 5. Galene dashtbani sp. nov.: A–B, dorsal (A) and front (B) stereo-pair views of holotype ZUTC6881, carapace; C–H, dorsal stereo-pair (C), front stereo-pair (D), left lateral (E), right lateral (F), ventral (G) and posterior (H) views of non-type HUIM195, carapace; I–J, dorsal (I) and front (J) views of non-type HUIM196, carapace; K, dorsal view of paratype HUIM197, carapace; L, dorsal view of paratype HUIM198, carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404978" httpUri="https://zenodo.org/record/6404978/files/figure.png" pageId="9" pageNumber="148">Figs. 5I, K</figureCitation>
|
||
). These specimens have less salient frontal and orbital margins and appear morphologically different from the others. These possess a flatter carapace, and deeper grooves between the mesogastric, cardiac and branchial lobes (
|
||
<figureCitation id="13312A56FFDBFFDDFEDFFC40FE7DFC44" box="[339,465,942,969]" captionStart="FIGURE 5" captionStartId="7.[151,250,1898,1922]" captionTargetBox="[214,1374,181,1874]" captionTargetId="figure-19@7.[214,1374,181,1874]" captionTargetPageId="7" captionText="FIGURE 5. Galene dashtbani sp. nov.: A–B, dorsal (A) and front (B) stereo-pair views of holotype ZUTC6881, carapace; C–H, dorsal stereo-pair (C), front stereo-pair (D), left lateral (E), right lateral (F), ventral (G) and posterior (H) views of non-type HUIM195, carapace; I–J, dorsal (I) and front (J) views of non-type HUIM196, carapace; K, dorsal view of paratype HUIM197, carapace; L, dorsal view of paratype HUIM198, carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404978" httpUri="https://zenodo.org/record/6404978/files/figure.png" pageId="9" pageNumber="148">Figs. 5I–K</figureCitation>
|
||
). In these specimens, the widest part of the mesogastric region is at the level between two lateral tuberosities of the carapace. The width of the posterior margin of the intestinal region equals the width of the front (
|
||
<figureCitation id="13312A56FFDBFFDDFEA8FC18FE3AFB9C" box="[292,406,1014,1041]" captionStart="FIGURE 5" captionStartId="7.[151,250,1898,1922]" captionTargetBox="[214,1374,181,1874]" captionTargetId="figure-19@7.[214,1374,181,1874]" captionTargetPageId="7" captionText="FIGURE 5. Galene dashtbani sp. nov.: A–B, dorsal (A) and front (B) stereo-pair views of holotype ZUTC6881, carapace; C–H, dorsal stereo-pair (C), front stereo-pair (D), left lateral (E), right lateral (F), ventral (G) and posterior (H) views of non-type HUIM195, carapace; I–J, dorsal (I) and front (J) views of non-type HUIM196, carapace; K, dorsal view of paratype HUIM197, carapace; L, dorsal view of paratype HUIM198, carapace. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6404978" httpUri="https://zenodo.org/record/6404978/files/figure.png" pageId="9" pageNumber="148">Fig. 5I, K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |