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<document id="595046EF04596B82349D503F2584D1AA" ID="10.11646/zootaxa.4267.1.1SLASH11021" ID-CLB-Dataset="33061" ID-DOI="10.5281/zenodo.575641" ID-GBIF-Dataset="810b55e4-b8ba-4368-bf9c-8517ee33e7d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="575641" ID-ZooBank="8920893E-E5F8-482A-A60D-7A248D2B0DCB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494577389872" checkinUser="plazi" docAuthor="Skelley, Paul, Xu, Guang, Tang, William, Lindström, Anders J., Marler, Thomas, Khuraijam, Jibankumar Singh, Singh, Rita &amp; Rich, Stephen" docDate="2017" docId="03A38781FFFCFF93FF70FB26FBD1C06A" docLanguage="en" docName="zootaxa.4267.1.1SLASH11021.pdf" docOrigin="Zootaxa 4267 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cycadophila (Cycadophila) cyclochasma Skelley, Xu &amp; Tang, new species" docType="treatment" docVersion="8" lastPageNumber="29" masterDocId="FF9AFFF9FFE6FF8FFFE7FFEBFFD5C639" masterDocTitle="Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species" masterLastPageNumber="63" masterPageNumber="1" pageNumber="27" updateTime="1698439510239" updateUser="ExternalLinkService">
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<mods:title id="F65D484A570A9F0FCF9B75995373F5C1">Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species</mods:title>
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<mods:namePart id="027031AF316689DBECF18FF2445F53CE">Skelley, Paul</mods:namePart>
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<mods:namePart id="940FA6853BA639EBB076A94DA64DC4D4">Xu, Guang</mods:namePart>
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<mods:namePart id="BD17B90B8F96266BD0E26B864D8A0224">Tang, William</mods:namePart>
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<mods:namePart id="33983E00164B0220437C7774F1A657B5">Lindström, Anders J.</mods:namePart>
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<mods:namePart id="4C47DB2BB4E116042BB09212244FCE63">Marler, Thomas</mods:namePart>
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<mods:namePart id="15EF22E8023EF0DEFBA2AEAA613CFB21">Khuraijam, Jibankumar Singh</mods:namePart>
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<mods:namePart id="15E33CAC05D3EBDE45BE55FCB705F33A">Singh, Rita</mods:namePart>
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<mods:namePart id="256F9A977F610436BF4AEA6E890FE588">Rich, Stephen</mods:namePart>
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<mods:date id="7E2F0C58959F8D7B95362092F6FC3B1D">2017</mods:date>
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<mods:number id="9415FCDADA76225A2A7C486311A30E50">4267</mods:number>
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<mods:identifier id="B9C6D52AD55B3CDE11275D8EA43D4516" type="DOI">10.5281/zenodo.575641</mods:identifier>
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<treatment id="03A38781FFFCFF93FF70FB26FBD1C06A" ID-DOI="http://doi.org/10.5281/zenodo.5999196" ID-GBIF-Taxon="130809481" ID-Zenodo-Dep="5999196" LSID="urn:lsid:plazi:treatment:03A38781FFFCFF93FF70FB26FBD1C06A" httpUri="http://treatment.plazi.org/id/03A38781FFFCFF93FF70FB26FBD1C06A" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">
<subSubSection id="C310651CFFFCFF95FF70FB26FEC3C331" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8BB53697FFFCFF95FF70FB26FBCFC2DE" blockId="26.[151,1050,1229,1289]" box="[151,1050,1229,1255]" pageId="26" pageNumber="27">
<heading id="D0FD81FBFFFCFF95FF70FB26FBCFC2DE" bold="true" box="[151,1050,1229,1255]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="B97EEA85FFFCFF95FF70FB26FBCFC2DE" bold="true" box="[151,1050,1229,1255]" pageId="26" pageNumber="27">
<taxonomicName id="4C0A4D14FFFCFF95FF70FB26FCA8C2DE" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[151,893,1229,1255]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="cyclochasma" status="sp. nov." subGenus="Cycadophila">
<emphasis id="B97EEA85FFFCFF95FF70FB26FEFAC2DE" bold="true" box="[151,303,1229,1255]" italics="true" pageId="26" pageNumber="27">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFFCFF95FED8FB26FE03C2DE" bold="true" box="[319,470,1229,1255]" italics="true" pageId="26" pageNumber="27">Cycadophila</emphasis>
)
<emphasis id="B97EEA85FFFCFF95FE0FFB26FD56C2DE" bold="true" box="[488,643,1229,1255]" italics="true" pageId="26" pageNumber="27">cyclochasma</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFFCFF95FC6DFB26FBCFC2DE" box="[906,1050,1229,1255]" pageId="26" pageNumber="27" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB53697FFFCFF95FF70FB1BFEC3C331" blockId="26.[151,1050,1229,1289]" box="[151,278,1264,1289]" pageId="26" pageNumber="27">
<figureCitation id="13312A12FFFCFF95FF70FB1BFF34C331" box="[151,225,1264,1289]" captionStart="FIGURE 8" captionStartId="27.[151,250,1563,1585]" captionTargetBox="[150,1436,409,1541]" captionTargetId="figure@27.[150,1436,409,1541]" captionTargetPageId="27" captionText="FIGURE 8. Phorographs of Cycadophila (Cycadophila) cyclochasma, holotype: A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia." httpUri="https://zenodo.org/record/575649/files/figure.png" pageId="26" pageNumber="27">Figs. 8</figureCitation>
A–E
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFFCFF95FF70FAD3FD2DC075" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8BB53697FFFCFF95FF70FAD3FEABC3A0" blockId="26.[151,1437,1336,2009]" pageId="26" pageNumber="27">
<emphasis id="B97EEA85FFFCFF95FF70FAD3FE8AC368" bold="true" box="[151,351,1336,1361]" pageId="26" pageNumber="27">Adult diagnosis.</emphasis>
A member of
<taxonomicName id="4C0A4D14FFFCFF95FDFAFAD3FC89C369" box="[541,860,1336,1361]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFFCFF95FDFAFAD3FD79C368" box="[541,684,1336,1361]" italics="true" pageId="26" pageNumber="27">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFFCFF95FD23FAD3FC86C368" box="[708,851,1336,1361]" italics="true" pageId="26" pageNumber="27">Cycadophila</emphasis>
)
</taxonomicName>
<taxonomicName id="4C0A4D14FFFCFF95FC8DFAD3FC76C368" box="[874,931,1336,1361]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="nigra">nigra</taxonomicName>
species group as discussed above readily distinguished from all others by its large body size, large deep submental-gular pit, and elongate pronotum with nearly straight sides.
</paragraph>
<paragraph id="8BB53697FFFCFF95FF20FA4FFD2DC075" blockId="26.[151,1437,1336,2009]" pageId="26" pageNumber="27">
<emphasis id="B97EEA85FFFCFF95FF20FA4FFE74C384" bold="true" box="[199,417,1444,1469]" pageId="26" pageNumber="27">Adult description.</emphasis>
Length
<quantity id="4CF29B72FFFCFF95FDE5FA4FFDBFC385" box="[514,618,1444,1469]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="26" pageNumber="27" unit="mm" value="6.9">6.90 mm</quantity>
, width
<quantity id="4CF29B72FFFCFF95FD26FA4FFCFFC385" box="[705,810,1444,1469]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="27" unit="mm" value="2.5">2.50 mm</quantity>
. Body in dorsal view elongate, sides gradually arched, almost parallel-sided, greatest width at middle of elytra; in lateral view convex dorsally. General body color entirely dark reddish-brown, appendages and lateral margins lighter; dorsal surface distinctly punctate, surface appearing slightly shiny, dorsal punctation with short procumbent hairs barely protruding out of punctures, ventrally mostly covered with short procumbent setae.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFFCFF93FF20F9B3FEAEC566" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="description">
<paragraph id="8BB53697FFFCFF95FF20F9B3FA48C1E0" blockId="26.[151,1437,1336,2009]" pageId="26" pageNumber="27">
<emphasis id="B97EEA85FFFCFF95FF20F9B3FED0C048" bold="true" box="[199,261,1624,1649]" italics="true" pageId="26" pageNumber="27">Head</emphasis>
in dorsal view conical, gradually narrowed apically, surface convex, appearing swollen above eye, distinctly coarsely punctured; width
<quantity id="4CF29B72FFFCFF95FDD2F997FD4BC0AD" box="[565,670,1660,1685]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="26" pageNumber="27" unit="mm" value="1.05">1.05 mm</quantity>
; dorsal interocular distance
<quantity id="4CF29B72FFFCFF95FC3AF997FB91C0AD" box="[989,1092,1660,1685]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="26" pageNumber="27" unit="mm" value="0.67">0.67 mm</quantity>
, head width/dorsal interocular distance ratio 1.57, ventral interocular distance
<quantity id="4CF29B72FFFCFF95FD5FF94BFCF4C081" box="[696,801,1696,1721]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="26" pageNumber="27" unit="mm" value="0.55">0.55 mm</quantity>
, head width/ventral interocular distance ratio 1.91. Eye globular; bordered dorsally by swollen head, so that dorsal margin of eye with deep groove, supraocular stria less than half length of eye but coarsely punctured; projected laterally, with large black facets and short, inconspicuous interfacetal setae. Frons with short, dark, subcuticular suture on each side, approximately 1/4 width of frons in length, extending obliquely to lateral margin above and anterior to antennal insertion. Antennal length about 3/4 pronotal width, and slightly longer than head width, antennomere relative lengths from base to end approximately 16:11:13:10:10:10:10:11:16:16:17; antennomere I (scape) fairly large, slightly elongate, antennomere II slightly shorter than III; IV–VIII small, width equals length; club fairly large, IX–XI similar in length, XI globular. Clypeus rounded apically, anterior margin with long setae; densely punctate; somewhat emarginate, with narrow margin.
</paragraph>
<paragraph id="8BB53697FFFDFF94FF70FF7CFE15C75C" blockId="27.[151,1436,151,357]" pageId="27" pageNumber="28">
Maxillary palpomeres II, III slightly longer than wide; IV elongate, apex densely papillate; relative lengths of II–IV approximately 4:2:6. Labium with mentum pentagonal with carina forming triangular plate projecting medially, with distinct lateral pockets; labial palpomere III elongate-oval.
<collectingRegion id="49CEF875FFFDFF94FC85FF0BFC14C6CE" box="[866,961,224,247]" country="Myanmar" name="Chin" pageId="27" pageNumber="28">Mentum</collectingRegion>
and submentum with moderate punctation and setation, setae short. Gular area smooth, without setae or punctation, except at suture with submentum where a deep circular depression surrounds a central area with setose punctures; gula with deep groove surrounding medial and posterior eye margins.
</paragraph>
<caption id="DF75661FFFFDFF94FF70F9F0FCE8C06A" httpUri="https://zenodo.org/record/575649/files/figure.png" pageId="27" pageNumber="28" targetBox="[150,1436,409,1541]" targetPageId="27">
<paragraph id="8BB53697FFFDFF94FF70F9F0FCE8C06A" blockId="27.[151,1435,1563,1619]" pageId="27" pageNumber="28">
<emphasis id="B97EEA85FFFDFF94FF70F9F0FEC3C008" bold="true" box="[151,278,1563,1585]" pageId="27" pageNumber="28">FIGURE 8.</emphasis>
Phorographs of
<taxonomicName id="4C0A4D14FFFDFF94FE2EF9F0FCA1C008" box="[457,884,1563,1585]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="cyclochasma" subGenus="Cycadophila">
<emphasis id="B97EEA85FFFDFF94FE2EF9F0FD9EC008" box="[457,587,1563,1585]" italics="true" pageId="27" pageNumber="28">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFFDFF94FDBAF9F0FD0BC008" box="[605,734,1563,1585]" italics="true" pageId="27" pageNumber="28">Cycadophila</emphasis>
)
<emphasis id="B97EEA85FFFDFF94FD16F9F0FCA1C008" box="[753,884,1563,1585]" italics="true" pageId="27" pageNumber="28">cyclochasma</emphasis>
</taxonomicName>
, holotype: A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia.
</paragraph>
</caption>
<paragraph id="8BB53697FFFDFF93FF20F96FFDE7C7B1" blockId="27.[151,1437,1668,2020]" lastBlockId="28.[151,1437,151,1619]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="B97EEA85FFFDFF94FF20F96FFEC8C0A4" bold="true" box="[199,285,1668,1693]" italics="true" pageId="27" pageNumber="28">Thorax</emphasis>
with pronotum transverse in dorsal view, with marginal beads on all sides; length/width ratio 0.82, convex, nearly parallel-sided, vaguely narrowing anteriorly; anterior angles projecting, weakly angulate; lateral carinae nearly straight, with thin bead, bead bearing row of setose punctures; posterior angles sharp, nearly 90o; posterior margin projecting medially; with narrow, short longitudinal furrow posterolaterally, beginning at pore on posterior margin, extending forward and ending abruptly, length of furrows relative to pronotal length 0.25. Prosternum with anterior margin slightly emarginate, finely denticulate with fringe of long, anteriorly directed setae; posterior prosternal process truncate, not turning inward at apex. Hypomeron with coarse punctures laterally, medially without longitudinal striations. Scutellar shield transverse, posterior margin roundly projected, pentagonal. Elytra elongate with sides gradually arched, convex; length/width 3.52, greatest width near midlength; with marginal bead basally, scutellary striole with 9 punctures; intervals of striae with distinct punctures. All elytral punctures bearing a single short seta; seta length just reaching outside of punctures. Punctation on meso- and metaventrite distinct. Metaventrite long, convex laterally, slightly impressed medially, metathoracic discrimen extending slightly over half metaventrite length; line behind mesocoxae with distinct, coarse punctures. Legs narrow, similar in length; femora compressed laterally; male with metafemur swollen basally, posterior margin curved; tibiae shorter than femora, weakly dilated to narrow apex. Protibia not dilated at apex, with fine setal fringe on ventral, lateral margin smooth without stout spinules; few small stout spinules at disto-lateral angle. Meso- and metatibia narrow, similar to protibia.
</paragraph>
<paragraph id="8BB53697FFFAFF93FF20FE7FFD7DC466" blockId="28.[151,1437,151,1619]" pageId="28" pageNumber="29">
<emphasis id="B97EEA85FFFAFF93FF20FE7FFEE0C794" bold="true" box="[199,309,404,429]" italics="true" pageId="28" pageNumber="29">Abdomen</emphasis>
with 5 ventrites bearing fine punctures; anterior margin with intercoxal process narrow, with triangular point anteromedially, lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I lacking subcoxal line, slightly longer medially than II; I–IV lacking pair of erect sensory hairs located on either side of the middle; II–IV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex bearing row of short appressed, densely arranged setae.
</paragraph>
<paragraph id="8BB53697FFFAFF93FF20FD85FEAEC566" blockId="28.[151,1437,151,1619]" pageId="28" pageNumber="29">
<emphasis id="B97EEA85FFFAFF93FF20FD85FEBBC4BE" bold="true" box="[199,366,622,647]" italics="true" pageId="28" pageNumber="29">Male genitalia</emphasis>
(tegmen and median lobe of aedeagus) twisted approximately 45o toward the left side of animal and resting on side (
<figureCitation id="13312A12FFFAFF93FE74FD79FE0EC492" box="[403,475,658,683]" captionStart="FIGURE 8" captionStartId="27.[151,250,1563,1585]" captionTargetBox="[150,1436,409,1541]" captionTargetId="figure@27.[150,1436,409,1541]" captionTargetPageId="27" captionText="FIGURE 8. Phorographs of Cycadophila (Cycadophila) cyclochasma, holotype: A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia." httpUri="https://zenodo.org/record/575649/files/figure.png" pageId="28" pageNumber="29">Fig. 8</figureCitation>
G). Tegmen sclerotized, triangular, gently twisted; with anterior region ring-like, posterior region sheath-like, posterior margin slightly arcuate; lateral margins gradually converging posteriorly; dorsally with 2 large, elongate, somewhat spatula-shaped parameres. Parameres with long setae apically; length/ width ratio 2.00. Aedeagus with median lobe short, slender, highly sclerotized, apex long and acuminate. Ratio of median lobe length to penile strut length 1:5.
<emphasis id="B97EEA85FFFAFF93FD78FCC9FD22C502" bold="true" box="[671,759,802,827]" italics="true" pageId="28" pageNumber="29">Female</emphasis>
unknown, but as with
<taxonomicName id="4C0A4D14FFFAFF93FC1BFCC8FBBFC502" box="[1020,1130,802,827]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="abyssa">
<emphasis id="B97EEA85FFFAFF93FC1BFCC8FBBFC502" box="[1020,1130,802,827]" italics="true" pageId="28" pageNumber="29">C. abyssa</emphasis>
</taxonomicName>
, they are expected to have narrow metafemora.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFFAFF93FF20FC81FD75C20E" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BB53697FFFAFF93FF20FC81FD15C5BA" blockId="28.[151,1437,151,1619]" box="[199,704,874,899]" pageId="28" pageNumber="29">
<materialsCitation id="3B623CCAFFFAFF93FF20FC81FD15C5BA" ID-GBIF-Occurrence="1502685085" box="[199,704,874,899]" country="Vietnam" location="Vinh Phuc Province" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Vinh Phuc Province" typeStatus="holotype">
<emphasis id="B97EEA85FFFAFF93FF20FC81FEB0C5BA" bold="true" box="[199,357,874,899]" pageId="28" pageNumber="29">
<typeStatus id="54B18835FFFAFF93FF20FC81FED7C5BA" box="[199,258,874,899]" pageId="28" pageNumber="29">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F31D7607FFFAFF93FE8BFC81FE1BC5BA" box="[364,462,874,899]" name="Vietnam" pageId="28" pageNumber="29">Vietnam</collectingCountry>
,
<collectingRegion id="49CEF875FFFAFF93FE3EFC81FD69C5BA" box="[473,700,874,899]" country="Vietnam" name="Vinh Phuc" pageId="28" pageNumber="29">
Vinh
<collectingRegion id="49CEF875FFFAFF93FDF1FC81FD69C5BA" box="[534,700,874,899]" inferredBySuffix="true" pageId="28" pageNumber="29">Phuc Province</collectingRegion>
</collectingRegion>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB53697FFFAFF93FF20FC65FDA0C59E" blockId="28.[151,1437,151,1619]" box="[199,629,910,935]" pageId="28" pageNumber="29">
<materialsCitation id="3B623CCAFFFAFF93FF20FC65FDA0C59E" ID-GBIF-Occurrence="1502685086" box="[199,629,910,935]" country="Vietnam" location="Vinh Phuc Province" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Vinh Phuc Province">
<emphasis id="B97EEA85FFFAFF93FF20FC65FECFC59E" bold="true" box="[199,282,910,935]" pageId="28" pageNumber="29">Range.</emphasis>
<collectingCountry id="F31D7607FFFAFF93FEC6FC65FE56C59E" box="[289,387,910,935]" name="Vietnam" pageId="28" pageNumber="29">Vietnam</collectingCountry>
,
<collectingRegion id="49CEF875FFFAFF93FE69FC65FDA4C59E" box="[398,625,910,935]" country="Vietnam" name="Vinh Phuc" pageId="28" pageNumber="29">
Vinh
<collectingRegion id="49CEF875FFFAFF93FE2CFC65FDA4C59E" box="[459,625,910,935]" inferredBySuffix="true" pageId="28" pageNumber="29">Phuc Province</collectingRegion>
</collectingRegion>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB53697FFFAFF93FF20FC59FD75C20E" blockId="28.[151,1437,151,1619]" pageId="28" pageNumber="29">
<materialsCitation id="3B623CCAFFFAFF93FF20FC59FB27C5D6" ID-GBIF-Occurrence="1502685087" collectingDate="2013-09-27" collectingDateMax="2013-09-29" collectingDateMin="2013-09-27" collectorName="Material" country="Vietnam" location="Dai Lai Lake" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Vinh Phuc" typeStatus="holotype">
<emphasis id="B97EEA85FFFAFF93FF20FC59FDF6C5F2" bold="true" box="[199,547,946,971]" pageId="28" pageNumber="29">
<collectorName id="26FF5341FFFAFF93FF20FC59FEFAC5F2" box="[199,303,946,971]" pageId="28" pageNumber="29">Material</collectorName>
examined.
<typeStatus id="54B18835FFFAFF93FE50FC59FDF6C5F2" box="[439,547,946,971]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
(by designation) male with the following labels: 1) [rectangular; white; printed in black ink] “
<collectingCountry id="F31D7607FFFAFF93FED9FC3CFE13C5D6" box="[318,454,983,1007]" name="Vietnam" pageId="28" pageNumber="29">VIETNAM</collectingCountry>
:
<collectingRegion id="49CEF875FFFAFF93FE35FC3DFD9FC5D6" box="[466,586,982,1007]" country="Vietnam" name="Vinh Phuc" pageId="28" pageNumber="29">Vinh Phuc</collectingRegion>
Prov., Me Linh Biodiv. Sta.;
<location id="8ED5604CFFFAFF93FC44FC3DFB95C5D6" LSID="urn:lsid:plazi:treatment:03A38781FFFCFF93FF70FB26FBD1C06A:8ED5604CFFFAFF93FC44FC3DFB95C5D6" box="[931,1088,982,1007]" country="Vietnam" name="Dai Lai Lake" pageId="28" pageNumber="29" stateProvince="Vinh Phuc">Dai Lai Lake</location>
,
<date id="FFB41057FFFAFF93FBAAFC3DFB27C5D6" box="[1101,1266,982,1007]" pageId="28" pageNumber="29" value="2013-09-27" valueMax="2013-09-29" valueMin="2013-09-27">
<collectingDate id="EFF0E9BFFFFAFF93FBAAFC3DFB27C5D6" box="[1101,1266,982,1007]" pageId="28" pageNumber="29" value="2013-09-27" valueMax="2013-09-29" valueMin="2013-09-27">27–29.ix.2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B623CCAFFFAFF93FAE6FC3CFD75C20E" ID-GBIF-Occurrence="1502685080" collectionCode="FSCA" collectorName="Col. J. B." country="Vietnam" elevation="100" location="Deposited" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phuc" typeStatus="holotype">
<quantity id="4CF29B72FFFAFF93FAE6FC3CFA96C5D6" box="[1281,1347,983,1007]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="m" value="100.0">
<elevation id="0027D1A4FFFAFF93FAE6FC3CFA96C5D6" box="[1281,1347,983,1007]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="m" value="100.0">100m</elevation>
</quantity>
,
<collectorName id="26FF5341FFFAFF93FAB6FC3DFF79C22A" pageId="28" pageNumber="29">Col. J. B.</collectorName>
Heppner”; 2) [rectangular; red, printed in black ink] “
<typeStatus id="54B18835FFFAFF93FCF0FC10FC78C22A" box="[791,941,1019,1043]" pageId="28" pageNumber="29" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D0CFD1EFFFAFF93FC51FC10FC1EC22B" box="[950,971,1019,1042]" pageId="28" pageNumber="29" type="male">♂</specimenCount>
<taxonomicName id="4C0A4D14FFFAFF93FC33FC11FEABC20E" authority="P. Skelley, G. Xu &amp; W. Tang 2017" authorityName="P. Skelley, G. Xu &amp; W. Tang" authorityYear="2017" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="cyclochasma">
<emphasis id="B97EEA85FFFAFF93FC33FC11FB28C22A" box="[980,1277,1018,1043]" italics="true" pageId="28" pageNumber="29">Cycadophila cyclochasma</emphasis>
P. Skelley, G. Xu &amp; W. Tang 2017
</taxonomicName>
”.
<location id="8ED5604CFFFAFF93FE74FBF5FDD0C20E" LSID="urn:lsid:plazi:treatment:03A38781FFFCFF93FF70FB26FBD1C06A:8ED5604CFFFAFF93FE74FBF5FDD0C20E" box="[403,517,1054,1079]" country="Vietnam" name="Deposited" pageId="28" pageNumber="29" stateProvince="Vinh Phuc">Deposited</location>
in the
<collectionCode id="ED1BAE52FFFAFF93FDB5FBF4FD4EC20E" box="[594,667,1055,1079]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="28" pageNumber="29">FSCA</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFFAFF93FF20FBA9FC3BC246" pageId="28" pageNumber="29" type="etymology">
<paragraph id="8BB53697FFFAFF93FF20FBA9FC3BC246" blockId="28.[151,1437,151,1619]" pageId="28" pageNumber="29">
<emphasis id="B97EEA85FFFAFF93FF20FBA9FE9BC262" bold="true" box="[199,334,1090,1115]" pageId="28" pageNumber="29">Etymology.</emphasis>
The species epithet comes from the Greek, ‘cyclo’- for circular, and ‘khasma’—for deep void, pit, in combination referring to the ‘circular pit’ at the submental-gular juncture.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFFAFF93FF20FB61FBD1C06A" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8BB53697FFFAFF93FF20FB61FBD1C06A" blockId="28.[151,1437,151,1619]" pageId="28" pageNumber="29">
<materialsCitation id="3B623CCAFFFAFF93FF20FB61FEB2C36E" ID-GBIF-Occurrence="1502685083" location="northern Vietnam" municipality="southeastern China" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Vinh Phuc province" typeStatus="holotype">
<emphasis id="B97EEA85FFFAFF93FF20FB61FEEEC29A" bold="true" box="[199,315,1162,1187]" pageId="28" pageNumber="29">Remarks.</emphasis>
<taxonomicName id="4C0A4D14FFFAFF93FEA2FB61FDBAC29A" box="[325,623,1162,1187]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="cyclochasma">
<emphasis id="B97EEA85FFFAFF93FEA2FB61FDBAC29A" box="[325,623,1162,1187]" italics="true" pageId="28" pageNumber="29">Cycadophila cyclochasma</emphasis>
</taxonomicName>
is most similar in all morphological characters to
<taxonomicName id="4C0A4D14FFFAFF93FB5AFB60FAF8C29A" box="[1213,1325,1162,1187]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="abyssa">
<emphasis id="B97EEA85FFFAFF93FB5AFB60FAF8C29A" box="[1213,1325,1162,1187]" italics="true" pageId="28" pageNumber="29">C. abyssa</emphasis>
</taxonomicName>
. The key differences are subtle, but these differences are consistent in available specimens. Over 15 species of
<taxonomicName id="4C0A4D14FFFAFF93FAF6FB44FA80C2FF" box="[1297,1365,1199,1222]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEA85FFFAFF93FAF6FB44FA80C2FF" box="[1297,1365,1199,1222]" italics="true" pageId="28" pageNumber="29">Cycas</emphasis>
</taxonomicName>
occur in the general region of
<location id="8ED5604CFFFAFF93FE4AFB39FDA1C2D2" LSID="urn:lsid:plazi:treatment:03A38781FFFCFF93FF70FB26FBD1C06A:8ED5604CFFFAFF93FE4AFB39FDA1C2D2" box="[429,628,1234,1259]" municipality="southeastern China" name="northern Vietnam" pageId="28" pageNumber="29" stateProvince="Vinh Phuc province">northern Vietnam</location>
and adjacent regions of
<collectingMunicipality id="6BD1ACEDFFFAFF93FC76FB39FBBCC2D2" box="[913,1129,1234,1259]" pageId="28" pageNumber="29">southeastern China</collectingMunicipality>
(
<bibRefCitation id="EF9B4B66FFFAFF93FB9DFB39FAB1C2D2" author="Osborne" box="[1146,1380,1234,1259]" pageId="28" pageNumber="29" refString="Osborne, R., Hill, K. D., Nguyen, H. T. &amp; Ke, L. P. (2007) Cycads of Fietnam. Tien Wah Press (Pte) Ltd, Singapore, 116 pp." type="book" year="2007">
Osborne
<emphasis id="B97EEA85FFFAFF93FB03FB3FFACBC2D2" box="[1252,1310,1234,1259]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
2007
</bibRefCitation>
). All these
<taxonomicName id="4C0A4D14FFFAFF93FF3FFB1CFEC8C337" box="[216,285,1271,1294]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEA85FFFAFF93FF3FFB1CFEC8C337" box="[216,285,1271,1294]" italics="true" pageId="28" pageNumber="29">Cycas</emphasis>
</taxonomicName>
belong in the Section
<emphasis id="B97EEA85FFFAFF93FDFEFB1DFD67C336" box="[537,690,1270,1295]" italics="true" pageId="28" pageNumber="29">Stangerioides</emphasis>
. Only one species of
<taxonomicName id="4C0A4D14FFFAFF93FC42FB1CFC3CC337" box="[933,1001,1271,1294]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEA85FFFAFF93FC42FB1CFC3CC337" box="[933,1001,1271,1294]" italics="true" pageId="28" pageNumber="29">Cycas</emphasis>
</taxonomicName>
,
<emphasis id="B97EEA85FFFAFF93FC1FFB1CFB50C336" box="[1016,1157,1270,1295]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="4C0A4D14FFFAFF93FC1FFB1CFB54C336" box="[1016,1153,1270,1295]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="balansae">C. balansae</taxonomicName>
,
</emphasis>
belonging in Subsection
<emphasis id="B97EEA85FFFAFF93FF70FAF1FEE5C30A" box="[151,304,1306,1331]" italics="true" pageId="28" pageNumber="29">Stangerioides</emphasis>
, is currently known from the
<typeStatus id="54B18835FFFAFF93FD62FAF0FD60C30A" box="[645,693,1307,1331]" pageId="28" pageNumber="29">type</typeStatus>
locality of
<collectingRegion id="49CEF875FFFAFF93FCDEFAF1FBCCC30A" box="[825,1049,1306,1331]" country="Vietnam" name="Vinh Phuc" pageId="28" pageNumber="29">Vinh Phuc province</collectingRegion>
and this species ranges north into
<collectingRegion id="49CEF875FFFAFF93FF70FAD5FF2DC36E" box="[151,248,1342,1367]" country="China" name="Guangxi" pageId="28" pageNumber="29">Guangxi</collectingRegion>
province
</materialsCitation>
, China.
<materialsCitation id="3B623CCAFFFAFF93FE2EFAD5FC80C342" ID-GBIF-Occurrence="1502685088" country="Iceland" location="Hainan" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Hainan" typeStatus="holotype">
The closely related
<taxonomicName id="4C0A4D14FFFAFF93FD48FAD5FC43C36E" box="[687,918,1342,1367]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="abyssa">
<emphasis id="B97EEA85FFFAFF93FD48FAD5FC43C36E" box="[687,918,1342,1367]" italics="true" pageId="28" pageNumber="29">Cycadophila abyssa</emphasis>
</taxonomicName>
is known from
<taxonomicName id="4C0A4D14FFFAFF93FBB3FAD4FACFC36E" box="[1108,1306,1342,1367]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="hainanesis">
<emphasis id="B97EEA85FFFAFF93FBB3FAD4FACFC36E" box="[1108,1306,1342,1367]" italics="true" pageId="28" pageNumber="29">Cycas hainanesis</emphasis>
</taxonomicName>
on
<collectingRegion id="49CEF875FFFAFF93FAACFAD5FA49C36E" box="[1355,1436,1342,1367]" country="China" name="Hainan" pageId="28" pageNumber="29">Hainan</collectingRegion>
<collectingCountry id="F31D7607FFFAFF93FF70FA89FF0EC342" box="[151,219,1378,1403]" name="Iceland" pageId="28" pageNumber="29">Island</collectingCountry>
and habitat of
<taxonomicName id="4C0A4D14FFFAFF93FE7DFA88FDE7C342" box="[410,562,1378,1403]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="fairylakea">
<emphasis id="B97EEA85FFFAFF93FE7DFA88FDE7C342" box="[410,562,1378,1403]" italics="true" pageId="28" pageNumber="29">C. fairylakea</emphasis>
</taxonomicName>
in
<collectingRegion id="49CEF875FFFAFF93FD87FA89FC80C342" box="[608,853,1378,1403]" country="China" name="Guangdong" pageId="28" pageNumber="29">Guangdong province</collectingRegion>
</materialsCitation>
, China (both belonging to Section
<emphasis id="B97EEA85FFFAFF93FB1BFA89FA40C342" box="[1276,1429,1378,1403]" italics="true" pageId="28" pageNumber="29">Stangerioides</emphasis>
, Subsection
<emphasis id="B97EEA85FFFAFF93FEFBFA63FE4BC3A6" box="[284,414,1416,1439]" italics="true" pageId="28" pageNumber="29">Taiwaninae</emphasis>
), farther to the east. Based on these differences in geographical distributions, available hosts and morphology we consider these beetles to be separate species. Although molecular evidence is not currently available to confirm this conclusion, describing the species officially recognizes these differences and provides a name for future research.
<taxonomicName id="4C0A4D14FFFAFF93FDD3FA19FC8EC032" box="[564,859,1522,1547]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="cyclochasma">
<emphasis id="B97EEA85FFFAFF93FDD3FA19FC8EC032" box="[564,859,1522,1547]" italics="true" pageId="28" pageNumber="29">Cycadophila cyclochasma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0A4D14FFFAFF93FC75FA18FC2AC032" box="[914,1023,1522,1547]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="abyssa">
<emphasis id="B97EEA85FFFAFF93FC75FA18FC2AC032" box="[914,1023,1522,1547]" italics="true" pageId="28" pageNumber="29">C. abyssa</emphasis>
</taxonomicName>
differ from all other members of the genus by sharing a number of unique features on the head, most notably the deep pit at the submental-gular juncture, but also the swollen head that creates a deep groove around the eyes.
</paragraph>
</subSubSection>
</treatment>
</document>
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