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<document id="457945286848B5F6BA2C024A7C6727B4" ID="10.11646/zootaxa.4267.1.1SLASH11021" ID-CLB-Dataset="33061" ID-DOI="10.5281/zenodo.575641" ID-GBIF-Dataset="810b55e4-b8ba-4368-bf9c-8517ee33e7d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="575641" ID-ZooBank="8920893E-E5F8-482A-A60D-7A248D2B0DCB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494577389872" checkinUser="plazi" docAuthor="Skelley, Paul, Xu, Guang, Tang, William, Lindström, Anders J., Marler, Thomas, Khuraijam, Jibankumar Singh, Singh, Rita &amp; Rich, Stephen" docDate="2017" docId="03A38781FFEDFF81FF70FCF3FB32C75C" docLanguage="en" docName="zootaxa.4267.1.1SLASH11021.pdf" docOrigin="Zootaxa 4267 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cycadophila (Cycadophila) Xu, Tang &amp; Skelley" docType="treatment" docVersion="8" lastPageNumber="15" masterDocId="FF9AFFF9FFE6FF8FFFE7FFEBFFD5C639" masterDocTitle="Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species" masterLastPageNumber="63" masterPageNumber="1" pageNumber="12" updateTime="1698439510239" updateUser="ExternalLinkService">
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<mods:title id="1C532580E6196C14296D170DB874386A">Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species</mods:title>
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<mods:namePart id="1EA534823C15974FDE336E9EDDCC1D16">Skelley, Paul</mods:namePart>
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<mods:namePart id="8A93850BADC686FFEA563929B484BA38">Xu, Guang</mods:namePart>
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<mods:namePart id="C39A87AECA111538D56A1B8F44EB4676">Tang, William</mods:namePart>
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<mods:namePart id="587B2B13385C148EB79542050CF55E0D">Lindström, Anders J.</mods:namePart>
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<mods:namePart id="CAF5CEC19490C8F58A5C75865A57EF4E">Marler, Thomas</mods:namePart>
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<mods:namePart id="187162E5741AD5944A7E2E335DAEEC13">Khuraijam, Jibankumar Singh</mods:namePart>
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<mods:namePart id="1FA9EA402F83264A17B4D4EF39008D32">Rich, Stephen</mods:namePart>
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<mods:date id="27B113551A2D969F26E9143FFF90906C">2017</mods:date>
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<treatment id="03A38781FFEDFF81FF70FCF3FB32C75C" ID-DOI="http://doi.org/10.5281/zenodo.5999173" ID-GBIF-Taxon="130809492" ID-Zenodo-Dep="5999173" LSID="urn:lsid:plazi:treatment:03A38781FFEDFF81FF70FCF3FB32C75C" httpUri="http://treatment.plazi.org/id/03A38781FFEDFF81FF70FCF3FB32C75C" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
<subSubSection id="C310651CFFEDFF84FF70FCF3FC0DC50A" box="[151,984,792,819]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BB53697FFEDFF84FF70FCF3FC0DC50A" blockId="11.[151,984,792,819]" box="[151,984,792,819]" pageId="11" pageNumber="12">
<heading id="D0FD81FBFFEDFF84FF70FCF3FC0DC50A" bold="true" box="[151,984,792,819]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B97EEA85FFEDFF84FF70FCF3FC0DC50A" bold="true" box="[151,984,792,819]" pageId="11" pageNumber="12">
<taxonomicName id="4C0A4D14FFEDFF84FF70FCF3FD0EC50A" authority="Xu, Tang &amp; Skelley" authorityName="Xu, Tang &amp; Skelley" box="[151,731,792,819]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subGenus" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEDFF84FF70FCF3FEFAC50B" bold="true" box="[151,303,792,818]" italics="true" pageId="11" pageNumber="12">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFEDFF84FED8FCF3FE03C50B" bold="true" box="[319,470,792,818]" italics="true" pageId="11" pageNumber="12">Cycadophila</emphasis>
) Xu, Tang &amp; Skelley
</taxonomicName>
, nominate subgenus
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFEDFF84FF70FCB2FB53C54B" box="[151,1158,857,882]" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BB53697FFEDFF84FF70FCB2FB53C54B" blockId="11.[151,1437,857,2034]" box="[151,1158,857,882]" pageId="11" pageNumber="12">
<emphasis id="B97EEA85FFEDFF84FF70FCB2FEFEC54B" bold="true" box="[151,299,857,882]" pageId="11" pageNumber="12">Type species</emphasis>
.
<taxonomicName id="4C0A4D14FFEDFF84FEDEFCB2FCA5C54B" authority="Xu, Tang &amp; Skelley, 2015" authorityName="Xu, Tang &amp; Skelley" authorityYear="2015" box="[313,880,857,882]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="debaonica">
<emphasis id="B97EEA85FFEDFF84FEDEFCB2FD91C54B" box="[313,580,857,882]" italics="true" pageId="11" pageNumber="12">Cycadophila debaonica</emphasis>
<bibRefCitation id="EF9B4B66FFEDFF84FDACFCB1FCA5C54B" author="Xu" box="[587,880,857,882]" pageId="11" pageNumber="12" refString="Xu, G., Tang, W., Skelley, P., Liu, N., &amp; Rich, S. (2015) Cycadophila, a new genus (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas debaoensis (Cycadaceae) in Asia. Zootaxa, 3986 (3), 251 - 278." type="journal article" year="2015">Xu, Tang &amp; Skelley, 2015</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFEDFF84FF20FC97FB5AC1CB" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8BB53697FFEDFF84FF20FC97FD8CC5C4" blockId="11.[151,1437,857,2034]" pageId="11" pageNumber="12">
<emphasis id="B97EEA85FFEDFF84FF20FC97FE56C5AC" bold="true" box="[199,387,892,917]" pageId="11" pageNumber="12">Adult diagnosis</emphasis>
.
<taxonomicName id="4C0A4D14FFEDFF84FE74FC97FD19C5AD" box="[403,716,892,917]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFEDFF84FE74FC97FDF7C5AC" box="[403,546,892,917]" italics="true" pageId="11" pageNumber="12">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFEDFF84FDD3FC97FD16C5AC" box="[564,707,892,917]" italics="true" pageId="11" pageNumber="12">Cycadophila</emphasis>
)
</taxonomicName>
is distinguished externally from the subgenus
<emphasis id="B97EEA85FFEDFF84FB02FC97FAA2C5AC" box="[1253,1399,892,917]" italics="true" pageId="11" pageNumber="12">Strobilophila</emphasis>
by the thin lateral pronotal carinae in lateral view with single row of punctation, by the lack of stout setae on the lateral margin of the protibia or with only a few setae at the extreme apex and with sexual dimorphism present or not, when present appearing on various legs.
</paragraph>
<paragraph id="8BB53697FFEDFF84FF20FBECFB5AC1CB" blockId="11.[151,1437,857,2034]" pageId="11" pageNumber="12">
<emphasis id="B97EEA85FFEDFF84FF20FBECFE4FC219" bold="true" box="[199,410,1031,1056]" pageId="11" pageNumber="12">Adult description</emphasis>
. Length
<quantity id="4CF29B72FFEDFF84FDE4FBECFD7EC226" box="[515,683,1031,1056]" metricMagnitude="-3" metricUnit="m" metricValue="4.995" metricValueMax="6.95" metricValueMin="3.04" pageId="11" pageNumber="12" unit="mm" value="4.995" valueMax="6.95" valueMin="3.04">3.046.95 mm</quantity>
, width
<quantity id="4CF29B72FFEDFF84FCE5FBECFC7FC226" box="[770,938,1031,1056]" metricMagnitude="-3" metricUnit="m" metricValue="1.9749999999999999" metricValueMax="2.8" metricValueMin="1.15" pageId="11" pageNumber="12" unit="mm" value="1.9749999999999999" valueMax="2.8" valueMin="1.15">1.152.80 mm</quantity>
.
<emphasis id="B97EEA85FFEDFF84FC50FBECFC20C219" bold="true" box="[951,1013,1031,1056]" pageId="11" pageNumber="12">Body</emphasis>
: form elongate, not strongly parallel sided; dorsoventrally flattened to weakly convex; widest at middle of elytra; color reddish-brown or dark brown, with or without color pattern; dorsal punctation moderately coarse, evenly distributed, dense.
<emphasis id="B97EEA85FFEDFF84FB56FBA6FB25C25F" bold="true" box="[1201,1264,1101,1126]" pageId="11" pageNumber="12">Head</emphasis>
: dorsal surface of head relatively flat to convex in profile; convex between eyes, clypeus flat to convex apically; some with vague depressions basally; clypeus truncate anteriorly; supraocular striae length variable; transverse occipital ridge present (vertexal line); stridulatory file on occipital region of head present, one on each side, widely separated, parallel; eyes prominent, coarsely faceted; usually lacking lateral temple behind eye.
<emphasis id="B97EEA85FFEDFF84FBBEFB32FB6AC2CB" bold="true" box="[1113,1215,1241,1266]" pageId="11" pageNumber="12">Antenna</emphasis>
: antennomere IVI length subequal to width, moniliform in most; antenna with 3-segmented club.
<emphasis id="B97EEA85FFEDFF84FBCEFB17FB6DC32C" bold="true" box="[1065,1208,1276,1301]" pageId="11" pageNumber="12">Mouthparts</emphasis>
: terminal maxillary palpomere cylindrical, acuminate at apex, elongate, 23× longer than preceding palpomere; terminal labial palpomere flattened, roughly oval in shape, 3/4 length of terminal maxillary palpomere; mentum with large triangular base defined by carina; lateral pockets in front of carina evident or not; submentum and gula relatively flat, suture not obvious, medially with weak to distinct depression bearing setose punctures.
<emphasis id="B97EEA85FFEDFF84FB52FA6CFAFBC399" bold="true" box="[1205,1326,1415,1440]" pageId="11" pageNumber="12">Pronotum</emphasis>
: pronotal anterior marginal bead complete, fine; pronotal anterior margin near eyes emarginate, anterior angles rounded; pronotal basal pits present, associated longitudinal groove distinct, groove approximately 1/4 pronotal length; secondary transverse groove between pits present, broad, curving along posterior margin except medially where slightly separated from margin; pronotal lateral carinae evenly arcuate, lacking modifications; pronotal lateral carinal bead narrow.
<emphasis id="B97EEA85FFEDFF84FE6AF9DDFD9FC076" bold="true" box="[397,586,1590,1615]" pageId="11" pageNumber="12">Scutellar shield</emphasis>
: broadly pentagonal.
<emphasis id="B97EEA85FFEDFF84FCAEF9DDFC40C076" bold="true" box="[841,917,1590,1615]" pageId="11" pageNumber="12">Elytra</emphasis>
: finely setose, indistinct; with coarse, dense interval punctures; elytra with marginal bead basally.
<emphasis id="B97EEA85FFEDFF84FD15F9B2FC55C04B" bold="true" box="[754,896,1625,1650]" pageId="11" pageNumber="12">Prosternum</emphasis>
: mostly flat, depressed or not posterior of coxa; procoxal cavities narrowly open; pronotal hypomeron with longitudinal striations, weak in some.
<emphasis id="B97EEA85FFEDFF84FB1CF997FAE6C0AC" bold="true" box="[1275,1331,1660,1685]" pageId="11" pageNumber="12">Legs</emphasis>
: protibia dilated distally or not, angled laterally but without apical tooth; ventral apical margin with or without fringe of stout ventrally directed, tooth-like spinules, some present on disto-lateral margin; meso- and metafemora elongate oval, meso- and metatibiae moderately angularly dilated at apex.
<emphasis id="B97EEA85FFEDFF84FCDCF90EFC7AC0C7" bold="true" box="[827,943,1765,1790]" pageId="11" pageNumber="12">Abdomen</emphasis>
: ventrites IIV each with pair of long erect sensory hairs located adjacent to midline (often abraded).
<emphasis id="B97EEA85FFEDFF84FCC5F8E3FBDEC118" bold="true" box="[802,1035,1800,1825]" pageId="11" pageNumber="12">Sexual dimorphism</emphasis>
: when evident, present in variously developed legs of males, being more robust, dilated distally or variously curved.
<emphasis id="B97EEA85FFEDFF84FBC0F8C0FB06C17D" bold="true" box="[1063,1235,1835,1860]" pageId="11" pageNumber="12">Male genitalia</emphasis>
: median lobe and tegmen flattened laterally, weakly twisted basally; penile struts not fused, separate for entire length, each 56× longer than median lobe; flagellum shorter than penile strut, not coiled; spiculum gastrale asymmetrical.
<emphasis id="B97EEA85FFEDFF84FAA2F89AFF2BC194" bold="true" pageId="11" pageNumber="12">Female genitalia</emphasis>
: gonocoxites triangular, gradually narrowed posteriorly; apices of gonocoxites laterally with concave impression and setae. Gonostylus small, inserted at the concave impression of the gonocoxite, with several short setae and one long seta. Valvifers expanded posteriorly. Spermatheca elongated, C-shaped.
</paragraph>
</subSubSection>
<caption id="DF75661FFFEAFF83FF70F8DDFA49C1FF" httpUri="https://zenodo.org/record/575643/files/figure.png" pageId="12" pageNumber="13" targetBox="[170,1433,205,1803]" targetPageId="12">
<paragraph id="8BB53697FFEAFF83FF70F8DDFA49C1FF" blockId="12.[151,1436,1846,1990]" pageId="12" pageNumber="13">
<emphasis id="B97EEA85FFEAFF83FF70F8DDFEC7C175" bold="true" box="[151,274,1846,1868]" pageId="12" pageNumber="13">FIGURE 2.</emphasis>
Key characters of adults: photographs of lateral head and pronotal margin: A)
<taxonomicName id="4C0A4D14FFEAFF83FBC8F8DDFA82C175" box="[1071,1367,1846,1868]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="fupingensis" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEAFF83FBC8F8DDFB65C175" box="[1071,1200,1846,1868]" italics="true" pageId="12" pageNumber="13">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FB58F8DCFB03C175" box="[1215,1238,1847,1868]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FB03F8DDFA82C175" box="[1252,1367,1846,1868]" italics="true" pageId="12" pageNumber="13">fupingensis</emphasis>
</taxonomicName>
; B)
<taxonomicName id="4C0A4D14FFEAFF83FA63F8DCFE41C153" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tansachai" subGenus="Strobilophila">
<emphasis id="B97EEA85FFEAFF83FA63F8DCFA49C175" box="[1412,1436,1847,1868]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FF78F8BFFEF6C153" box="[159,291,1876,1898]" italics="true" pageId="12" pageNumber="13">Strobilophila</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FED5F8BFFE41C153" box="[306,404,1876,1898]" italics="true" pageId="12" pageNumber="13">tansachai</emphasis>
</taxonomicName>
; C)
<taxonomicName id="4C0A4D14FFEAFF83FE26F8BEFD5FC153" box="[449,650,1876,1898]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="yunnanensis" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEAFF83FE26F8BEFE07C153" box="[449,466,1877,1898]" italics="true" pageId="12" pageNumber="13">C</emphasis>
. (
<emphasis id="B97EEA85FFEAFF83FE00F8BEFE2AC153" box="[487,511,1877,1898]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FDEAF8BEFD5FC153" box="[525,650,1877,1898]" italics="true" pageId="12" pageNumber="13">yunnanensis</emphasis>
</taxonomicName>
; SEM of protibia, ventral view: D)
<taxonomicName id="4C0A4D14FFEAFF83FC12F8BEFB61C153" box="[1013,1204,1876,1898]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="fupingensis" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEAFF83FC12F8BEFBD8C153" box="[1013,1037,1877,1898]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FBFCF8BEFBE7C153" box="[1051,1074,1877,1898]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FBA6F8BFFB61C153" box="[1089,1204,1876,1898]" italics="true" pageId="12" pageNumber="13">fupingensis</emphasis>
</taxonomicName>
; E)
<taxonomicName id="4C0A4D14FFEAFF83FB38F8BEFA40C153" box="[1247,1429,1876,1898]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="debaonica" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEAFF83FB38F8BEFB22C153" box="[1247,1271,1877,1898]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FAE2F8BEFAC8C153" box="[1285,1309,1877,1898]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FACCF8BFFA40C153" box="[1323,1429,1876,1898]" italics="true" pageId="12" pageNumber="13">debaonica</emphasis>
</taxonomicName>
; F)
<taxonomicName id="4C0A4D14FFEAFF83FF5FF89FFEB9C1B0" box="[184,364,1907,1929]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tansachai" subGenus="Strobilophila">
<emphasis id="B97EEA85FFEAFF83FF5FF89FFF05C1B0" box="[184,208,1908,1929]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FF04F89FFF23C1B0" box="[227,246,1908,1929]" italics="true" pageId="12" pageNumber="13">S.</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FEEEF898FEB9C1B0" box="[265,364,1907,1929]" italics="true" pageId="12" pageNumber="13">tansachai</emphasis>
</taxonomicName>
; G)
<taxonomicName id="4C0A4D14FFEAFF83FE43F89FFDF9C1B0" box="[420,556,1907,1929]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="yangi" subGenus="Strobilophila">
<emphasis id="B97EEA85FFEAFF83FE43F89FFE6EC1B0" box="[420,443,1908,1929]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FE29F89FFE37C1B0" box="[462,482,1908,1929]" italics="true" pageId="12" pageNumber="13">S.</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FE13F89FFDF9C1B0" box="[500,556,1908,1929]" italics="true" pageId="12" pageNumber="13">yangi</emphasis>
</taxonomicName>
; photographs of mesotibia, ventral view: H)
<taxonomicName id="4C0A4D14FFEAFF83FBF2F89FFB01C1B0" box="[1045,1236,1907,1929]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="debaonica" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEAFF83FBF2F89FFBF8C1B0" box="[1045,1069,1908,1929]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FBA7F89FFB8DC1B0" box="[1088,1112,1908,1929]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FB8DF898FB01C1B0" box="[1130,1236,1907,1929]" italics="true" pageId="12" pageNumber="13">debaonica</emphasis>
</taxonomicName>
female; I)
<taxonomicName id="4C0A4D14FFEAFF83FAB5F89FFED4C191" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="debaonica" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEAFF83FAB5F89FFABCC1B0" box="[1362,1385,1908,1929]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FA9BF89FFA41C1B0" box="[1404,1428,1908,1929]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FF70F879FED4C191" box="[151,257,1938,1960]" italics="true" pageId="12" pageNumber="13">debaonica</emphasis>
</taxonomicName>
male; J)
<taxonomicName id="4C0A4D14FFEAFF83FEBAF879FE05C191" box="[349,464,1938,1960]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="lata" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEAFF83FEBAF879FEA0C19E" box="[349,373,1938,1959]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FE64F879FE4EC19E" box="[387,411,1938,1959]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FE4EF879FE05C191" box="[425,464,1938,1960]" italics="true" pageId="12" pageNumber="13">lata</emphasis>
</taxonomicName>
male; photographs of left half of pronotum: K)
<taxonomicName id="4C0A4D14FFEAFF83FC52F879FB92C191" box="[949,1095,1938,1960]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="collina" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEAFF83FC52F879FC18C19E" box="[949,973,1938,1959]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FC3CF879FC27C19E" box="[987,1010,1938,1959]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FBE6F879FB92C191" box="[1025,1095,1938,1960]" italics="true" pageId="12" pageNumber="13">collina</emphasis>
</taxonomicName>
; L)
<taxonomicName id="4C0A4D14FFEAFF83FB94F879FAFDC191" box="[1139,1320,1938,1960]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="debaonica" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEAFF83FB94F879FB5EC19E" box="[1139,1163,1938,1959]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FB7EF879FB65C19E" box="[1177,1200,1938,1959]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FB58F879FAFDC191" box="[1215,1320,1938,1960]" italics="true" pageId="12" pageNumber="13">debaonica</emphasis>
</taxonomicName>
; M)
<taxonomicName id="4C0A4D14FFEAFF83FABCF879FF2FC1FF" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="kwaiensis" subGenus="Strobilophila">
<emphasis id="B97EEA85FFEAFF83FABCF879FAA7C19E" box="[1371,1394,1938,1959]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FA67F879FA41C19E" box="[1408,1428,1938,1959]" italics="true" pageId="12" pageNumber="13">S.</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FF70F85BFF2FC1FF" box="[151,250,1968,1990]" italics="true" pageId="12" pageNumber="13">kwaiensis</emphasis>
</taxonomicName>
; N)
<taxonomicName id="4C0A4D14FFEAFF83FEC0F85AFE05C1FF" box="[295,464,1968,1990]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tansachai" subGenus="Strobilophila">
<emphasis id="B97EEA85FFEAFF83FEC0F85AFEEBC1FF" box="[295,318,1969,1990]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FEABF85AFEB5C1FF" box="[332,352,1969,1990]" italics="true" pageId="12" pageNumber="13">S.</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FE8AF85BFE05C1FF" box="[365,464,1968,1990]" italics="true" pageId="12" pageNumber="13">tansachai</emphasis>
</taxonomicName>
; photographs of mesofemur of major male: O)
<taxonomicName id="4C0A4D14FFEAFF83FC4AF85AFBEBC1FF" box="[941,1086,1968,1990]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="collina" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEAFF83FC4AF85AFC10C1FF" box="[941,965,1969,1990]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FC34F85AFC3FC1FF" box="[979,1002,1969,1990]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
)
<emphasis id="B97EEA85FFEAFF83FC1FF85BFBEBC1FF" box="[1016,1086,1968,1990]" italics="true" pageId="12" pageNumber="13">collina</emphasis>
</taxonomicName>
ex Vietnam; P)
<taxonomicName id="4C0A4D14FFEAFF83FB06F85AFA4DC1FF" box="[1249,1432,1968,1990]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="debaonica" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEAFF83FB06F85AFB2CC1FF" box="[1249,1273,1969,1990]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
(
<emphasis id="B97EEA85FFEAFF83FAE1F85AFACBC1FF" box="[1286,1310,1969,1990]" italics="true" pageId="12" pageNumber="13">C.</emphasis>
) debaonica
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF75661FFFEBFF82FF70F87FFA4EC1D6" httpUri="https://zenodo.org/record/575644/files/figure.png" pageId="13" pageNumber="14" targetBox="[201,1384,180,1919]" targetPageId="13">
<paragraph id="8BB53697FFEBFF82FF70F87FFA4EC1D6" blockId="13.[151,1436,1940,2031]" pageId="13" pageNumber="14">
<emphasis id="B97EEA85FFEBFF82FF70F87FFEC1C193" bold="true" box="[151,276,1940,1962]" pageId="13" pageNumber="14">FIGURE 3.</emphasis>
Photographs of key characters of larvae: terminal segments of abdomen, lateral and dorsal view (t1t3 indicate basal tubercles of urogomphi): AB)
<taxonomicName id="4C0A4D14FFEBFF82FDFBF85DFCC6C1F5" box="[540,787,1974,1996]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="floridana">
<emphasis id="B97EEA85FFEBFF82FDFBF85DFCC6C1F5" box="[540,787,1974,1996]" italics="true" pageId="13" pageNumber="14">Pharaxonotha floridana</emphasis>
</taxonomicName>
; CD)
<taxonomicName id="4C0A4D14FFEBFF82FC83F85DFB52C1F5" box="[868,1159,1974,1996]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="debaonica" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEBFF82FC83F85DFC30C1F5" box="[868,997,1974,1996]" italics="true" pageId="13" pageNumber="14">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFEBFF82FC12F85CFBD8C1F5" box="[1013,1037,1975,1996]" italics="true" pageId="13" pageNumber="14">C.</emphasis>
)
<emphasis id="B97EEA85FFEBFF82FBFAF85DFB52C1F5" box="[1053,1159,1974,1996]" italics="true" pageId="13" pageNumber="14">debaonica</emphasis>
</taxonomicName>
; EF)
<taxonomicName id="4C0A4D14FFEBFF82FB36F85CFA40C1F5" box="[1233,1429,1974,1996]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fupingensis" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEBFF82FB36F85CFB3CC1F5" box="[1233,1257,1975,1996]" italics="true" pageId="13" pageNumber="14">C.</emphasis>
(
<emphasis id="B97EEA85FFEBFF82FB1DF85CFAC4C1F5" box="[1274,1297,1975,1996]" italics="true" pageId="13" pageNumber="14">C.</emphasis>
)
<emphasis id="B97EEA85FFEBFF82FAC5F85DFA40C1F5" box="[1314,1429,1974,1996]" italics="true" pageId="13" pageNumber="14">fupingensis</emphasis>
</taxonomicName>
; head, ventral view: G)
<taxonomicName id="4C0A4D14FFEBFF82FE99F831FDE9C1D6" box="[382,572,2009,2031]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fupingensis" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEBFF82FE99F831FE43C1D6" box="[382,406,2010,2031]" italics="true" pageId="13" pageNumber="14">C.</emphasis>
(
<emphasis id="B97EEA85FFEBFF82FE43F831FE6EC1D6" box="[420,443,2010,2031]" italics="true" pageId="13" pageNumber="14">C.</emphasis>
)
<emphasis id="B97EEA85FFEBFF82FE2EF832FDE9C1D6" box="[457,572,2009,2031]" italics="true" pageId="13" pageNumber="14">fupingensis</emphasis>
</taxonomicName>
; right mandible, dissected, ventral view: H)
<taxonomicName id="4C0A4D14FFEBFF82FC19F831FB68C1D6" box="[1022,1213,2009,2031]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fupingensis" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEBFF82FC19F831FBC3C1D6" box="[1022,1046,2010,2031]" italics="true" pageId="13" pageNumber="14">C.</emphasis>
(
<emphasis id="B97EEA85FFEBFF82FBC3F831FBEEC1D6" box="[1060,1083,2010,2031]" italics="true" pageId="13" pageNumber="14">C.</emphasis>
)
<emphasis id="B97EEA85FFEBFF82FBADF832FB68C1D6" box="[1098,1213,2009,2031]" italics="true" pageId="13" pageNumber="14">fupingensis</emphasis>
</taxonomicName>
; I)
<taxonomicName id="4C0A4D14FFEBFF82FB06F831FA40C1D6" box="[1249,1429,2009,2031]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="debaonica" subGenus="Cycadophila">
<emphasis id="B97EEA85FFEBFF82FB06F831FB2DC1D6" box="[1249,1272,2010,2031]" italics="true" pageId="13" pageNumber="14">C.</emphasis>
(
<emphasis id="B97EEA85FFEBFF82FAE1F831FACBC1D6" box="[1286,1310,2010,2031]" italics="true" pageId="13" pageNumber="14">C.</emphasis>
)
<emphasis id="B97EEA85FFEBFF82FACBF832FA40C1D6" box="[1324,1429,2009,2031]" italics="true" pageId="13" pageNumber="14">debaonica</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C310651CFFE8FF81FF20FF7CFB32C75C" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BB53697FFE8FF81FF20FF7CFB32C75C" blockId="14.[151,1436,151,357]" pageId="14" pageNumber="15">
<emphasis id="B97EEA85FFE8FF81FF20FF7CFEE1C689" bold="true" box="[199,308,151,176]" pageId="14" pageNumber="15">Remarks</emphasis>
.
<taxonomicName id="4C0A4D14FFE8FF81FEA3FF7CFDABC689" box="[324,638,151,176]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFE8FF81FEA3FF7CFE06C689" box="[324,467,151,176]" italics="true" pageId="14" pageNumber="15">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFE8FF81FE02FF7CFDA1C689" box="[485,628,151,176]" italics="true" pageId="14" pageNumber="15">Cycadophila</emphasis>
)
</taxonomicName>
presently comprises four species groups as diagnosed below: the
<taxonomicName id="4C0A4D14FFE8FF81FA93FF7CFA49C689" box="[1396,1436,151,176]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="lata">lata</taxonomicName>
group,
<taxonomicName id="4C0A4D14FFE8FF81FF01FF57FEB2C6EC" box="[230,359,188,213]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="fupingensis">fupingensis</taxonomicName>
group,
<taxonomicName id="4C0A4D14FFE8FF81FE5AFF57FE22C6EC" box="[445,503,188,213]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nigra">nigra</taxonomicName>
group and
<taxonomicName id="4C0A4D14FFE8FF81FD91FF56FD6CC6ED" box="[630,697,189,212]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="papua">papua</taxonomicName>
group. Preliminary morphological and molecular data show these to be monophyletic groups, but further data and analyses are needed to substantiate relationships amongst them before they can be confidently assigned into separate subgenera. They are readily distinguished from each other by characters in the key and by diagnostic characters presented in the following accounts. The external morphological diversity of these groups makes a simple diagnosis of
<emphasis id="B97EEA85FFE8FF81FD13FEA6FCD2C75D" box="[756,775,333,356]" italics="true" pageId="14" pageNumber="15">C</emphasis>
. (
<taxonomicName id="4C0A4D14FFE8FF81FCF9FEA7FC78C75C" box="[798,941,332,357]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFE8FF81FCF9FEA7FC78C75C" box="[798,941,332,357]" italics="true" pageId="14" pageNumber="15">Cycadophila</emphasis>
</taxonomicName>
) as presented here difficult.
</paragraph>
</subSubSection>
</treatment>
</document>