256 lines
36 KiB
XML
256 lines
36 KiB
XML
<document id="4B3A5BDE32148980E922411D653D99D4" ID-CLB-Dataset="45679" ID-DOI="10.5281/zenodo.279688" ID-GBIF-Dataset="f3683ee2-9fee-419d-af79-bb0238f4b958" ID-ISSN="1175-5326" ID-Zenodo-Dep="279688" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460426082735" checkinUser="plazi" docAuthor="Konstantinov, Fedor V." docDate="2012" docId="038B87F2CF36D441559DB378C302BD4E" docLanguage="en" docName="zt03157p030.pdf" docOrigin="Zootaxa 3157" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leucodellus Reuter 1906" docType="treatment" docVersion="12" lastPageNumber="20" masterDocId="FFB2FF8ACF37D443550AB13FC078B842" masterDocTitle="Leucodellus Reuter, 1906 (Heteroptera: Miridae: Phylini): key to species, revised species composition and diagnosis" masterLastPageNumber="30" masterPageNumber="18" pageNumber="19" updateTime="1698330704923" updateUser="ExternalLinkService">
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<mods:title id="097F0AFB714A8DF89C698E2DA723CD64">Leucodellus Reuter, 1906 (Heteroptera: Miridae: Phylini): key to species, revised species composition and diagnosis</mods:title>
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<mods:name id="0D0CB1DC2AEAE79BA7F754EEED0196C8" type="personal">
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<mods:namePart id="A8740A719DEADBD37F231A47F22F0A5E">Konstantinov, Fedor V.</mods:namePart>
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<mods:title id="A504B3F70092B9CBD1A4F5279A2C8978">Zootaxa</mods:title>
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<mods:date id="95B73D6996E8ABBE85774ED93365FA54">2012</mods:date>
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<treatment id="038B87F2CF36D441559DB378C302BD4E" ID-DOI="http://doi.org/10.5281/zenodo.6167736" ID-GBIF-Taxon="119428655" ID-Zenodo-Dep="6167736" LSID="urn:lsid:plazi:treatment:038B87F2CF36D441559DB378C302BD4E" httpUri="http://treatment.plazi.org/id/038B87F2CF36D441559DB378C302BD4E" lastPageId="2" lastPageNumber="20" pageId="1" pageNumber="19">
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<subSubSection id="C338656FCF36D442559DB378C1B6BA23" box="[151,462,583,609]" pageId="1" pageNumber="19" type="nomenclature">
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<paragraph id="8B9D36E4CF36D442559DB378C1B6BA23" blockId="1.[151,462,583,609]" box="[151,462,583,609]" pageId="1" pageNumber="19">
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<heading id="D0D58188CF36D442559DB378C1B6BA23" bold="true" box="[151,462,583,609]" fontSize="11" level="1" pageId="1" pageNumber="19" reason="1">
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<taxonomicName id="4C224D67CF36D442559DB378C1B6BA23" ID-CoL="638ZJ" authority="Reuter, 1906" authorityName="Reuter" authorityYear="1906" box="[151,462,583,609]" class="Insecta" family="Miridae" genus="Leucodellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="B956EAF6CF36D442559DB378C1B6BA23" bold="true" box="[151,462,583,609]" pageId="1" pageNumber="19">
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<emphasis id="B956EAF6CF36D442559DB378C152BA23" bold="true" box="[151,298,583,609]" italics="true" pageId="1" pageNumber="19">Leucodellus</emphasis>
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<bibRefCitation id="EFB34B15CF36D442543BB378C1B6BA23" author="Reuter" box="[305,462,583,609]" pageId="1" pageNumber="19" refString="Reuter, O. M. (1906) Capsidae in Prov. Sz'tschwan Chinae a DD. G. Potanin et M. Beresowski Collectae. Ezhegodnik Zoologicheskago Muzeya Imperatorskoi Akademii Nauk, 10, 1 - 81." type="journal article" year="1906">Reuter, 1906</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C338656FCF36D442559DB3B0C5ECBDEC" pageId="1" pageNumber="19" type="materials_examined">
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<paragraph id="8B9D36E4CF36D442559DB3B0C356BAE6" blockId="1.[151,910,654,738]" box="[151,814,654,677]" pageId="1" pageNumber="19">
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<typeStatus id="54998846CF36D442559DB3B0C0B1BAE6" box="[151,201,655,676]" pageId="1" pageNumber="19">Type</typeStatus>
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species by monotypy:
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<taxonomicName id="4C224D67CF36D44254B8B3B0C351BAE6" authority="Reuter 1906: 69" authorityName="Reuter" authorityPageNumber="69" authorityYear="1906" box="[434,809,654,677]" class="Insecta" family="Miridae" genus="Leucodellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="species" species="albidus">
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<emphasis id="B956EAF6CF36D44254B8B3B0C205BAE7" box="[434,637,655,677]" italics="true" pageId="1" pageNumber="19">Leucodellus albidus</emphasis>
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<bibRefCitation id="EFB34B15CF36D442578EB3B0C37BBAE6" author="Reuter" box="[644,771,654,676]" pageId="1" pageNumber="19" refString="Reuter, O. M. (1906) Capsidae in Prov. Sz'tschwan Chinae a DD. G. Potanin et M. Beresowski Collectae. Ezhegodnik Zoologicheskago Muzeya Imperatorskoi Akademii Nauk, 10, 1 - 81." type="journal article" year="1906">Reuter 1906</bibRefCitation>
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: 69
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8B9D36E4CF36D442559DB392C35EBAA3" blockId="1.[151,910,654,738]" pageId="1" pageNumber="19">
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<emphasis id="B956EAF6CF36D442559DB392C145BA81" box="[151,317,685,707]" italics="true" pageId="1" pageNumber="19">Heterochlorillus</emphasis>
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<bibRefCitation id="EFB34B15CF36D442544EB392C19BBA80" author="Putshkov" box="[324,483,685,707]" pageId="1" pageNumber="19" refString="Putshkov, V. G. (1970) Two new monotypic genera of Miridae (Heteroptera) from the West Caucasus. Dopovidi Akademii Nauk Ukrains'koi RSR (B), 1970 (8), 752 - 756. [in Ukrainian, English summary]." type="journal article" year="1970">Putshkov, 1970</bibRefCitation>
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||
: 752; synonymized by
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<bibRefCitation id="EFB34B15CF36D44257C7B392C31BBA80" author="Li" box="[717,867,685,707]" pageId="1" pageNumber="19" refString="Li, X. - M. & Liu, G. - Q. (2007) The genus Leucodellus Reuter, 1906 in China (Hemiptera: Miridae: Phylin). Zootaxa, 1478, 33 - 40." type="journal article" year="2007">Li & Liu 2007</bibRefCitation>
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: 34.
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<emphasis id="B956EAF6CF36D442559DB3F3C16FBAA0" box="[151,279,716,738]" italics="true" pageId="1" pageNumber="19">Leucophylus</emphasis>
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<bibRefCitation id="EFB34B15CF36D4425417B3F4C229BAA3" author="Duwal" box="[285,593,715,738]" pageId="1" pageNumber="19" refString="Duwal, R. K., Yasunaga, T. & Lee, S. (2010) Revision of the plant bug tribe Phylini from Nepal (Heteroptera: Miridae: Phylinae). Entomologica Americana, 116 (1 / 2), 1 - 48." type="journal article" year="2010">Duwal, Yasunaga & Lee 2010</bibRefCitation>
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: 23;
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<emphasis id="B956EAF6CF36D4425788B3F3C35EBAA3" bold="true" box="[642,806,716,737]" pageId="1" pageNumber="19">new synonymy.</emphasis>
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</paragraph>
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<paragraph id="8B9D36E4CF36D442559DB231C33ABCCC" blockId="1.[151,1437,782,2030]" pageId="1" pageNumber="19">
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<emphasis id="B956EAF6CF36D442559DB231C10ABB65" bold="true" box="[151,370,782,807]" pageId="1" pageNumber="19">Revised diagnosis.</emphasis>
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Distinguished by the following characters of the male and female genitalia: male genital segment exceptionally large, 0.6–0.7 of abdomen (
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<figureCitation id="13192A61CF36D44257A2B20DC293BB08" box="[680,747,818,842]" captionStart="FIGURES 1 – 5" captionStartId="4.[151,264,1970,1993]" captionTargetBox="[218,1380,228,1946]" captionTargetId="figure@4.[186,1410,193,1965]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Habitus images, males. 1 — Leucodellus albidus; 2 – 3 — Leucodellus zagdani: 2 — dorsal view, 3 — lateral view; 4 — Compsidolon nathaliae; 5 — Leucodellus amygdali." httpUri="https://zenodo.org/record/279689/files/figure.png" pageId="1" pageNumber="19">Fig. 3</figureCitation>
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), equal to or longer than combined length of head and pronotum; right paramere lanceolate; vesica strongly coiled, with several closely approximating sclerotized ridges running along margin of lateral strap, apically with two characteristically shaped blades, small blade more or less twisted along axis and attenuated laterally, large blade long and thin, gradually curved (
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<figureCitation id="13192A61CF36D4425182B2A1C56CBBF4" box="[1160,1300,926,950]" captionStart="FIGURES 12 – 17" captionStartId="7.[151,264,1975,1998]" captionTargetBox="[209,1392,232,1956]" captionTargetId="figure@7.[184,1408,193,1962]" captionTargetPageId="7" captionText="FIGURES 12 – 17. Vesicae of Leucodellus spp. 12 — L. albidus; 13 — L. amygdali; 14 — L. zagdani; 15 — L. pallescens; 16 — L. variegatus; 17 — L. xizangensis." httpUri="https://zenodo.org/record/279691/files/figure.png" pageId="1" pageNumber="19">Figs. 12–17</figureCitation>
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); secondary gonopore with well developed sculpture and long and distinct gonopore sclerite, far removed from apex of vesica, placed on membranous, dentate lobe; sclerotized rings of female dorsal labiate plate almost triangular, with distinctly tapering anterior angles (
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<figureCitation id="13192A61CF36D442570BB535C22FBC60" box="[513,599,1034,1058]" captionStart="FIGURES 18 – 23" captionStartId="8.[151,264,1706,1729]" captionTargetBox="[186,1406,192,1680]" captionTargetId="figure@8.[151,1436,169,1685]" captionTargetPageId="8" captionText="FIGURES 18 – 23. Genital structures. 18 – 20, vesicae: 18 — Compsidolon nathaliae, 19 — Compsidolon nebulosum; 20 — Compsidolon pumilum; 21 – 22, dorsal labiate plate: 21 — Compsidolon nathaliae, 22 — Leucodellus albidus; 23 — opening of vestibulum in Leucodellus albidus." httpUri="https://zenodo.org/record/279692/files/figure.png" pageId="1" pageNumber="19">Fig. 22</figureCitation>
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). Body form elongate and nearly parallel-sided, length 3.2–4.8 male, 3.0– 5.0 female. Coloration almost completely pale, hemelytra sometimes with pale brown markings, usually with dark markings on femora, brown spot at base of hind tibia and darkened bases of tibial spines. Vestiture with reclining pale simple setae only, rarely somewhat darkened on cuneus.
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</paragraph>
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<paragraph id="8B9D36E4CF36D44255CFB5A5C5ECBDEC" blockId="1.[151,1437,782,2030]" pageId="1" pageNumber="19">
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Similar in size and body proportions, general appearance including dark pattern on legs, and twin bladed vesica with approximating ridges on lateral strap to pale-colored species of
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<taxonomicName id="4C224D67CF36D442569CB581C4B4BC94" authority="Fieber, 1858" authorityName="Fieber" authorityYear="1858" box="[918,1228,1214,1238]" class="Insecta" family="Miridae" genus="Plagiognathus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="B956EAF6CF36D442569CB581C441BC94" box="[918,1081,1214,1238]" italics="true" pageId="1" pageNumber="19">Plagiognathus</emphasis>
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Fieber, 1858
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</taxonomicName>
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(see
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<bibRefCitation id="EFB34B15CF36D442500DB581C5E8BC94" author="Schuh" box="[1287,1424,1214,1238]" pageId="1" pageNumber="19" refString="Schuh, R. T. (2001) Revision of New World Plagiognathus Fieber, with comments on the Palearctic fauna and the description of a new genus (Heteroptera: Miridae: Phylinae). Bulletin of the American Museum of Natural History, 266, 1 - 267." type="journal article" year="2001">Schuh 2001</bibRefCitation>
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),
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<taxonomicName id="4C224D67CF36D442559DB5DCC1DFBCB8" authority="Reuter, 1909" authorityName="Reuter" authorityYear="1909" box="[151,423,1250,1275]" class="Insecta" family="Miridae" genus="Europiella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="B956EAF6CF36D442559DB5DCC176BCB9" box="[151,270,1251,1275]" italics="true" pageId="1" pageNumber="19">Europiella</emphasis>
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Reuter, 1909
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</taxonomicName>
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(see
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<bibRefCitation id="EFB34B15CF36D44254EEB5DDC216BCB8" author="Schuh" box="[484,622,1250,1274]" pageId="1" pageNumber="19" refString="Schuh, R. T. (2004) Revision of Europiella in North America, with the description of a new genus (Heteroptera: Miridae: Phylinae). American Museum Novitates, 3463, 1 - 58." type="journal article" year="2004">Schuh 2004</bibRefCitation>
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),
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<taxonomicName id="4C224D67CF36D4425789B5DCC3D3BCB8" authority="Kerzhner, 1962" authorityName="Kerzhner" authorityYear="1962" box="[643,939,1250,1275]" class="Insecta" family="Miridae" genus="Chlorillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B956EAF6CF36D4425789B5DCC28CBCB9" box="[643,756,1251,1275]" italics="true" pageId="1" pageNumber="19">Chlorillus</emphasis>
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Kerzhner, 1962
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</taxonomicName>
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(see
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<bibRefCitation id="EFB34B15CF36D44256E3B5DCC4FABCB8" author="Wagner" box="[1001,1154,1250,1275]" pageId="1" pageNumber="19" refString="Wagner, E. (1975) Die Miridae Hahn, 183 l, des Mittelmeerraumes und der Makaronesischen Inseln (Hemiptera, Heteroptera). Teil 3. Entomologische Abhandlungen, 40, Suppl., 1 - 483." type="journal article" year="1975">Wagner 1975</bibRefCitation>
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),
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<taxonomicName id="4C224D67CF36D4425192B5DCC0B7BD5C" authority="Kerzhner, 1964" authorityName="Kerzhner" authorityYear="1964" class="Insecta" family="Miridae" genus="Phaeochiton" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B956EAF6CF36D4425192B5DCC55EBCB9" box="[1176,1318,1251,1275]" italics="true" pageId="1" pageNumber="19">Phaeochiton</emphasis>
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Kerzhner, 1964
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||
</taxonomicName>
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(see Konstantinov 2008), and
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<taxonomicName id="4C224D67CF36D442572DB439C2DABD5C" box="[551,674,1286,1310]" class="Magnoliopsida" family="Polygonaceae" genus="Atraphaxis" kingdom="Plantae" order="Polygonales" pageId="1" pageNumber="19" phylum="Tracheophyta" rank="genus">
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||
<emphasis id="B956EAF6CF36D442572DB439C2DABD5C" box="[551,674,1286,1310]" italics="true" pageId="1" pageNumber="19">Atraphaxis</emphasis>
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||
</taxonomicName>
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||
-feeding group of
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<taxonomicName id="4C224D67CF36D4425667B439C4CABD5C" authority="Wagner, 1963" authorityName="Wagner" authorityYear="1963" box="[877,1202,1286,1311]" class="Insecta" family="Miridae" genus="Glaucopterum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B956EAF6CF36D4425667B439C477BD5C" box="[877,1039,1286,1310]" italics="true" pageId="1" pageNumber="19">Glaucopterum</emphasis>
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||
Wagner, 1963
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||
</taxonomicName>
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||
(see
|
||
<bibRefCitation id="EFB34B15CF36D44251E7B439C5EFBD5C" author="Kerzhner" box="[1261,1431,1286,1310]" pageId="1" pageNumber="19" refString="Kerzhner, I. M. (1984) New and little known Heteroptera from Mongolia and from adjacent regions of the USSR. IV. Miridae, I. Nasekomye Mongolii, 9, 35 - 72. [In Russian]." type="journal article" year="1984">Kerzhner 1984</bibRefCitation>
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||
;
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||
<bibRefCitation id="EFB34B15CF36D442559DB415C108BD00" author="Konstantinov" box="[151,368,1322,1346]" pageId="1" pageNumber="19" refString="Konstantinov, F. V. (2006) Two new species of Phylini (Heteroptera, Miridae, Phylinae) from Middle Asia and Caucasus with notes on Compsidolon pumilum (JAKOVLEV, 1876). Denisia, 19, 493 - 502." type="journal article" year="2006">Konstantinov 2006</bibRefCitation>
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||
,
|
||
<bibRefCitation id="EFB34B15CF36D4425474B415C1C2BD00" author="Konstantinov" box="[382,442,1322,1346]" pageId="1" pageNumber="19" refString="Konstantinov, F. V. (2011) New species and new records of plant bugs (Hemiptera: Heteroptera: Miridae) from Northwestern China. Zootaxa, 2836, 27 - 43." type="journal article" year="2011">2011</bibRefCitation>
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||
). Clearly distinguished from these genera by the distinctive curvature of the vesica and the shape of its apical blades, presence of the large gonopore sclerite, and triangular sclerotized rings of dorsal labiate plate. Further differing from
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||
<taxonomicName id="4C224D67CF36D4425702B44CC206BDC9" box="[520,638,1395,1419]" class="Insecta" family="Miridae" genus="Europiella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B956EAF6CF36D4425702B44CC206BDC9" box="[520,638,1395,1419]" italics="true" pageId="1" pageNumber="19">Europiella</emphasis>
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||
</taxonomicName>
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||
and
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||
<taxonomicName id="4C224D67CF36D44257BDB44CC321BDC9" box="[695,857,1395,1419]" class="Insecta" family="Miridae" genus="Glaucopterum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B956EAF6CF36D44257BDB44CC321BDC9" box="[695,857,1395,1419]" italics="true" pageId="1" pageNumber="19">Glaucopterum</emphasis>
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||
</taxonomicName>
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||
in having lanceolate, not truncated right paramere, and from
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||
<emphasis id="B956EAF6CF36D442540EB4A9C279BDEC" box="[260,513,1430,1454]" italics="true" pageId="1" pageNumber="19">
|
||
<taxonomicName id="4C224D67CF36D442540EB4A9C106BDEC" box="[260,382,1430,1454]" class="Insecta" family="Miridae" genus="Europiella" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="genus">Europiella</taxonomicName>
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||
,
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||
<taxonomicName id="4C224D67CF36D4425482B4A9C184BDEC" box="[392,508,1430,1454]" class="Insecta" family="Miridae" genus="Chlorillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="genus">Chlorillus</taxonomicName>
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,
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</emphasis>
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and
|
||
<taxonomicName id="4C224D67CF36D442573DB4A9C2BDBDEC" box="[567,709,1430,1454]" class="Insecta" family="Miridae" genus="Phaeochiton" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B956EAF6CF36D442573DB4A9C2BDBDEC" box="[567,709,1430,1454]" italics="true" pageId="1" pageNumber="19">Phaeochiton</emphasis>
|
||
</taxonomicName>
|
||
in lacking of silvery flattened and dark simple setae on dorsum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338656FCF36D44155CFB485C54CBAAD" lastPageId="2" lastPageNumber="20" pageId="1" pageNumber="19" type="description">
|
||
<paragraph id="8B9D36E4CF36D44255CFB485C44ABD90" blockId="1.[151,1437,782,2030]" box="[197,1074,1466,1491]" pageId="1" pageNumber="19">
|
||
<emphasis id="B956EAF6CF36D44255CFB485C1C5BD91" bold="true" box="[197,445,1466,1491]" pageId="1" pageNumber="19">Redescription. Male.</emphasis>
|
||
Macropterous, with elongate body, total length 3.3–4.7.
|
||
</paragraph>
|
||
<paragraph id="8B9D36E4CF36D44155CFB4E1C536B8B5" blockId="1.[151,1437,782,2030]" lastBlockId="2.[151,1436,151,1293]" lastPageId="2" lastPageNumber="20" pageId="1" pageNumber="19">
|
||
COLORATION: Uniformly pale greenish to pale whitish in life, yellowish to white in collections, dorsum uniformly pale, rarely hemelytron with scarce minute brown spots or diffuse and indistinctly bordered pale brown markings, membrane whitish, semitransparent, typically with oblique wedge-shaped macula at base and variously shaped diffuse paler brown markings behind cells (
|
||
<figureCitation id="13192A61CF36D44257D2B775C31ABE20" box="[728,866,1610,1634]" captionStart="FIGURES 1 – 5" captionStartId="4.[151,264,1970,1993]" captionTargetBox="[218,1380,228,1946]" captionTargetId="figure@4.[186,1410,193,1965]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Habitus images, males. 1 — Leucodellus albidus; 2 – 3 — Leucodellus zagdani: 2 — dorsal view, 3 — lateral view; 4 — Compsidolon nathaliae; 5 — Leucodellus amygdali." httpUri="https://zenodo.org/record/279689/files/figure.png" pageId="1" pageNumber="19">Figs. 1, 2, 5</figureCitation>
|
||
); fore and middle femora with faint minute brown spots apically, rarely uniformly pale, apex of hind femur dorsally and ventrally with distinct dark brown color-pattern; fore and middle tibiae usually, hind tibia always with dark spots at bases of tibial spines and not infrequently with darkened base (
|
||
<figureCitation id="13192A61CF36D442548BB789C184BE8C" box="[385,508,1718,1742]" captionStart="FIGURES 6 – 11" captionStartId="5.[151,264,1415,1438]" captionTargetBox="[169,1428,215,1390]" captionTargetId="figure@5.[151,1436,193,1394]" captionTargetPageId="5" captionText="FIGURES 6 – 11. Color pattern on ventral side of hind femur in males of Leucodellus spp. 6 — L. albidus; 7 — L. amygdali; 8 — L. pallescens; 9 — L. variegatus; 10 — L. xizangensis; 11 — L. zagdani." httpUri="https://zenodo.org/record/279690/files/figure.png" pageId="1" pageNumber="19">Figs. 6–11</figureCitation>
|
||
); venter uniformly pale (
|
||
<figureCitation id="13192A61CF36D442561FB789C321BE8C" box="[789,857,1718,1742]" captionStart="FIGURES 1 – 5" captionStartId="4.[151,264,1970,1993]" captionTargetBox="[218,1380,228,1946]" captionTargetId="figure@4.[186,1410,193,1965]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Habitus images, males. 1 — Leucodellus albidus; 2 – 3 — Leucodellus zagdani: 2 — dorsal view, 3 — lateral view; 4 — Compsidolon nathaliae; 5 — Leucodellus amygdali." httpUri="https://zenodo.org/record/279689/files/figure.png" pageId="1" pageNumber="19">Fig. 3</figureCitation>
|
||
). SURFACE AND VESTITURE: Dorsum smooth, moderately shining; with dense simple uniformly pale setae, rarely somewhat darkened on cuneus, first antennal segment with two pale medial spines, each femur with a few dark or pale spines apically, tibial spines dark brown to whitish. STRUCTURE: Head: Moderately produced anteriorly in dorsal view; eyes relatively large, occupying almost entire height of head in lateral view, posterolateral margins of eyes contiguous with anterolateral margins of pronotum; vertex weakly convex, frons moderately sloping anteriorly in lateral view, clypeus weakly produced, usually not visible in dorsal view; antennae inserted near ventral margin of eye; second antennal segment linear; labium reaching metacoxae. Thorax: Pronotum trapezoidal, 1.9–2.4 times as broad as long, disk weakly convex, calli indistinct, posterior margin nearly straight or weakly concave medially, lateral margins straight, posterolateral angles broadly rounded; mesonotum moderately exposed. Legs: femur elongate, not swollen, usually slightly broader medially, tibia cylindrical, third tarsal segment equal to length of first and second combined, claw elongate, strongly bent at middle, pulvillus small, reaching midpoint of claw, attached to the claw along entire length.
|
||
</paragraph>
|
||
<paragraph id="8B9D36E4CF35D44155CFB03BC560BA55" blockId="2.[151,1436,151,1293]" pageId="2" pageNumber="20">
|
||
GENITALIA: Genital segment large, 0.6–0.7 of abdomen, distinctly longer than basal width, without distinctive ornamentation. Parameres: Right paramere lanceolate, with slender apical process; left paramere with obtuse sensory lobe and straight, gradually tapering apical process. Apex of theca: Shape typical of many phylines, comparatively long and narrow. Vesica: S-shaped, comparatively large, strongly coiled, body of vesica of distinctive curvature and form, with several closely approximating ridges running along margin of lateral strap, apically with two characteristically shaped blades, small blade more or less twisted along axis and attenuated laterally, large blade long and thin, gradually curved (
|
||
<figureCitation id="13192A61CF35D441575EB0E3C298B9B6" box="[596,736,476,500]" captionStart="FIGURES 12 – 17" captionStartId="7.[151,264,1975,1998]" captionTargetBox="[209,1392,232,1956]" captionTargetId="figure@7.[184,1408,193,1962]" captionTargetPageId="7" captionText="FIGURES 12 – 17. Vesicae of Leucodellus spp. 12 — L. albidus; 13 — L. amygdali; 14 — L. zagdani; 15 — L. pallescens; 16 — L. variegatus; 17 — L. xizangensis." httpUri="https://zenodo.org/record/279691/files/figure.png" pageId="2" pageNumber="20">Figs. 12–17</figureCitation>
|
||
); secondary gonopore far removed from apex of vesica, with well developed sculpture and long and distinct gonopore sclerite, placed on membranous, dentate lobe.
|
||
</paragraph>
|
||
<paragraph id="8B9D36E4CF35D44155CFB31BC508BAE5" blockId="2.[151,1436,151,1293]" pageId="2" pageNumber="20">
|
||
<emphasis id="B956EAF6CF35D44155CFB31BC15BBA7E" bold="true" box="[197,291,548,572]" pageId="2" pageNumber="20">Female.</emphasis>
|
||
Macropterous. Similar to male in color, structure, and vestiture, but more strongly ovoid, eyes slightly smaller than in male, with correspondingly broader vertex. GENITALIA: Dorsal labiate plate of bursa copulatrix with almost triangular sclerotized rings, their anterior angles distinctly tapering (
|
||
<figureCitation id="13192A61CF35D441513FB353C4F4BAC6" box="[1077,1164,620,644]" captionStart="FIGURES 18 – 23" captionStartId="8.[151,264,1706,1729]" captionTargetBox="[186,1406,192,1680]" captionTargetId="figure@8.[151,1436,169,1685]" captionTargetPageId="8" captionText="FIGURES 18 – 23. Genital structures. 18 – 20, vesicae: 18 — Compsidolon nathaliae, 19 — Compsidolon nebulosum; 20 — Compsidolon pumilum; 21 – 22, dorsal labiate plate: 21 — Compsidolon nathaliae, 22 — Leucodellus albidus; 23 — opening of vestibulum in Leucodellus albidus." httpUri="https://zenodo.org/record/279692/files/figure.png" pageId="2" pageNumber="20">Fig. 22</figureCitation>
|
||
); posterior wall simple; vestibulum narrow, S-shaped, irregularly sclerotized, vulva with two almost straight sclerites at sides (
|
||
<figureCitation id="13192A61CF35D4415005B3B0C51BBAE5" box="[1295,1379,655,679]" captionStart="FIGURES 18 – 23" captionStartId="8.[151,264,1706,1729]" captionTargetBox="[186,1406,192,1680]" captionTargetId="figure@8.[151,1436,169,1685]" captionTargetPageId="8" captionText="FIGURES 18 – 23. Genital structures. 18 – 20, vesicae: 18 — Compsidolon nathaliae, 19 — Compsidolon nebulosum; 20 — Compsidolon pumilum; 21 – 22, dorsal labiate plate: 21 — Compsidolon nathaliae, 22 — Leucodellus albidus; 23 — opening of vestibulum in Leucodellus albidus." httpUri="https://zenodo.org/record/279692/files/figure.png" pageId="2" pageNumber="20">Fig. 23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9D36E4CF35D44155CFB38BC54CBAAD" blockId="2.[151,1436,151,1293]" pageId="2" pageNumber="20">
|
||
<emphasis id="B956EAF6CF35D44155CFB38BC177BA8E" bold="true" box="[197,271,692,716]" pageId="2" pageNumber="20">Hosts.</emphasis>
|
||
Hosts are documented for three out of six species of the genus, and available data may suggest that either
|
||
<taxonomicName id="4C224D67CF35D44155D7B3E7C11CBAB2" box="[221,356,728,752]" class="Insecta" family="Miridae" genus="Leucodellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B956EAF6CF35D44155D7B3E7C11CBAB2" box="[221,356,728,752]" italics="true" pageId="2" pageNumber="20">Leucodellus</emphasis>
|
||
</taxonomicName>
|
||
spp. feed on a broad range of plant groups or some host data represent a sitting record.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338656FCF35D44155CFB3C4C184BBE6" pageId="2" pageNumber="20" type="distribution">
|
||
<paragraph id="8B9D36E4CF35D44155CFB3C4C184BBE6" blockId="2.[151,1436,151,1293]" pageId="2" pageNumber="20">
|
||
<emphasis id="B956EAF6CF35D44155CFB3C4C126BB56" bold="true" box="[197,350,763,788]" pageId="2" pageNumber="20">Distribution.</emphasis>
|
||
All species of
|
||
<taxonomicName id="4C224D67CF35D4415703B3C3C2E8BB56" box="[521,656,764,788]" class="Insecta" family="Miridae" genus="Leucodellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B956EAF6CF35D4415703B3C3C2E8BB56" box="[521,656,764,788]" italics="true" pageId="2" pageNumber="20">Leucodellus</emphasis>
|
||
</taxonomicName>
|
||
inhabit mountain terrains with moderate elevations. The genus has a highly disjunct distribution, with two closely related species,
|
||
<taxonomicName id="4C224D67CF35D4415646B21EC3ABBB7A" box="[844,979,800,824]" class="Insecta" family="Miridae" genus="Leucodellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="amygdali">
|
||
<emphasis id="B956EAF6CF35D4415646B21EC3ABBB7A" box="[844,979,800,824]" italics="true" pageId="2" pageNumber="20">L. amygdali</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C224D67CF35D4415101B21EC4FABB7A" box="[1035,1154,800,824]" class="Insecta" family="Miridae" genus="Leucodellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="zagdani">
|
||
<emphasis id="B956EAF6CF35D4415101B21EC4FABB7A" box="[1035,1154,800,824]" italics="true" pageId="2" pageNumber="20">L. zagdani</emphasis>
|
||
</taxonomicName>
|
||
, known from the Caucasus, while the rest of the species occurring in
|
||
<collectingCountry id="F3357674CF35D44157A2B27BC296BB1E" box="[680,750,836,860]" name="Nepal" pageId="2" pageNumber="20">Nepal</collectingCountry>
|
||
, Sichuan province and Tibet Autonomous Region of
|
||
<collectingCountry id="F3357674CF35D4415058B27BC5E1BB1E" box="[1362,1433,836,860]" name="China" pageId="2" pageNumber="20">China</collectingCountry>
|
||
. The two ranges are separated by as much as
|
||
<quantity id="4CDA9B01CF35D441578EB258C29EBB3D" box="[644,742,871,895]" metricMagnitude="6" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="20" unit="km" value="3500.0">3500 km</quantity>
|
||
and the genus remains unknown from the relatively well sampled mountains of Central Asia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338656FCF35D44155CFB290C302BD4E" pageId="2" pageNumber="20" type="discussion">
|
||
<paragraph id="8B9D36E4CF35D44155CFB290C098BC1A" blockId="2.[151,1436,151,1293]" pageId="2" pageNumber="20">
|
||
<emphasis id="B956EAF6CF35D44155CFB290C132BB8A" bold="true" box="[197,330,943,968]" pageId="2" pageNumber="20">Discussion.</emphasis>
|
||
A number of features of the male and female genitalia listed in the revised diagnosis unequivocally indicate the monophyly of
|
||
<taxonomicName id="4C224D67CF35D44154DCB2EBC225BBAE" box="[470,605,980,1004]" class="Insecta" family="Miridae" genus="Leucodellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B956EAF6CF35D44154DCB2EBC225BBAE" box="[470,605,980,1004]" italics="true" pageId="2" pageNumber="20">Leucodellus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFB34B15CF35D441577AB2EBC320BBAE" box="[624,856,980,1004]" pageId="2" pageNumber="20" refString="Duwal, R. K., Yasunaga, T. & Lee, S. (2010) Revision of the plant bug tribe Phylini from Nepal (Heteroptera: Miridae: Phylinae). Entomologica Americana, 116 (1 / 2), 1 - 48." type="journal article">Duwal et al. (2010)</bibRefCitation>
|
||
being unaware of the paper by
|
||
<bibRefCitation id="EFB34B15CF35D44151D3B2EBC5E4BBAE" author="Li" box="[1241,1436,980,1004]" pageId="2" pageNumber="20" refString="Li, X. - M. & Liu, G. - Q. (2007) The genus Leucodellus Reuter, 1906 in China (Hemiptera: Miridae: Phylin). Zootaxa, 1478, 33 - 40." type="journal article" year="2007">Li & Liu (2007)</bibRefCitation>
|
||
described the monotypic genus
|
||
<emphasis id="B956EAF6CF35D441570DB2C7C2E4BC52" box="[519,668,1016,1040]" italics="true" pageId="2" pageNumber="20">Leucophyllus</emphasis>
|
||
and correctly pointed out that the new genus is similar in overall appearance and vesica structure to
|
||
<taxonomicName id="4C224D67CF35D441573CB523C322BC76" box="[566,858,1052,1076]" class="Insecta" family="Miridae" genus="Plagiognathus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="pallescens">
|
||
<emphasis id="B956EAF6CF35D441573CB523C322BC76" box="[566,858,1052,1076]" italics="true" pageId="2" pageNumber="20">Plagiognathus pallescens</emphasis>
|
||
</taxonomicName>
|
||
(already transferred to
|
||
<taxonomicName id="4C224D67CF35D441517EB523C483BC76" box="[1140,1275,1052,1076]" class="Insecta" family="Miridae" genus="Leucodellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B956EAF6CF35D441517EB523C483BC76" box="[1140,1275,1052,1076]" italics="true" pageId="2" pageNumber="20">Leucodellus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFB34B15CF35D4415025B523C0ACBC1A" author="Li" pageId="2" pageNumber="20" refString="Li, X. - M. & Liu, G. - Q. (2007) The genus Leucodellus Reuter, 1906 in China (Hemiptera: Miridae: Phylin). Zootaxa, 1478, 33 - 40." type="journal article" year="2007">Li & Liu 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9D36E4CF35D44155CFB55BC302BD4E" blockId="2.[151,1436,151,1293]" pageId="2" pageNumber="20">
|
||
Careful investigation of
|
||
<taxonomicName id="4C224D67CF35D44154DFB55BC2A7BC3E" box="[469,735,1124,1148]" class="Insecta" family="Miridae" genus="Leucophylus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="variegatus">
|
||
<emphasis id="B956EAF6CF35D44154DFB55BC2A7BC3E" box="[469,735,1124,1148]" italics="true" pageId="2" pageNumber="20">Leucophylus variegatus</emphasis>
|
||
</taxonomicName>
|
||
and all species currently assigned to
|
||
<taxonomicName id="4C224D67CF35D4415175B55BC57EBC3E" box="[1151,1286,1124,1148]" class="Insecta" family="Miridae" genus="Leucodellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B956EAF6CF35D4415175B55BC57EBC3E" box="[1151,1286,1124,1148]" italics="true" pageId="2" pageNumber="20">Leucodellus</emphasis>
|
||
</taxonomicName>
|
||
allows me to conclude that they are congeneric. The peculiar vesical structure, large male genital segment and triangular sclerotized rings of the dorsal labiate plate are similar in all species.
|
||
<taxonomicName id="4C224D67CF35D4415658B593C3A1BC86" box="[850,985,1196,1220]" class="Insecta" family="Miridae" genus="Leucodellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B956EAF6CF35D4415658B593C3A1BC86" box="[850,985,1196,1220]" italics="true" pageId="2" pageNumber="20">Leucodellus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B956EAF6CF35D4415105B593C4E4BC86" box="[1039,1180,1196,1220]" italics="true" pageId="2" pageNumber="20">Leucophylus</emphasis>
|
||
share a common color pattern, similar pretarsal structure and vestiture. Based on the above features, the genus
|
||
<emphasis id="B956EAF6CF35D4415165B5EFC485BCAA" box="[1135,1277,1232,1256]" italics="true" pageId="2" pageNumber="20">Leucophylus</emphasis>
|
||
<bibRefCitation id="EFB34B15CF35D4415009B5EFC127BD4E" author="Duwal" pageId="2" pageNumber="20" refString="Duwal, R. K., Yasunaga, T. & Lee, S. (2010) Revision of the plant bug tribe Phylini from Nepal (Heteroptera: Miridae: Phylinae). Entomologica Americana, 116 (1 / 2), 1 - 48." type="journal article" year="2010">Duwal, Yasunaga & Lee, 2010</bibRefCitation>
|
||
is synonymized with
|
||
<taxonomicName id="4C224D67CF35D441575FB5CBC30EBD4E" authority="Reuter, 1906" authorityName="Reuter" authorityYear="1906" box="[597,886,1268,1292]" class="Insecta" family="Miridae" genus="Leucodellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B956EAF6CF35D441575FB5CBC2A5BD4E" box="[597,733,1268,1292]" italics="true" pageId="2" pageNumber="20">Leucodellus</emphasis>
|
||
<bibRefCitation id="EFB34B15CF35D44157EEB5CAC30EBD4E" author="Reuter" box="[740,886,1268,1292]" pageId="2" pageNumber="20" refString="Reuter, O. M. (1906) Capsidae in Prov. Sz'tschwan Chinae a DD. G. Potanin et M. Beresowski Collectae. Ezhegodnik Zoologicheskago Muzeya Imperatorskoi Akademii Nauk, 10, 1 - 81." type="journal article" year="1906">Reuter, 1906</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |