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<document ID-DOI="http://doi.org/10.5281/zenodo.4759324" ID-GBIF-Dataset="66abbd26-0f90-4888-9851-b54dc1f135cb" ID-GBIF-Taxon="182165333" ID-ISSN="1854-0392" ID-Zenodo-Dep="4759324" approvalRequired="2" approvalRequired_for_textStreams="2" checkinTime="1620956958103" checkinUser="felipe" docAuthor="Willett, Michele R. &amp; Stark, Bill P." docDate="2009" docId="038B87E0FFD5FFADFE80FDB0FA7DFEE3" docLanguage="en" docName="Illiesia.5.11.108-127.pdf" docOrigin="Illiesia 5 (11)" docSource="http://illiesia.speciesfile.org/papers/Illiesia05-11.pdf" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Alloperla imbecilla" docType="treatment" docVersion="5" lastPageNumber="123" masterDocId="FFB2FF98FFDAFFA2FFD4FF89FFF1FFF6" masterDocTitle="The Alloperla Leonarda Group Of Eastern North America, With Sem Images Of Four Out-Group Species (Plecoptera: Chloroperlidae)" masterLastPageNumber="127" masterPageNumber="108" pageId="15" pageNumber="123" updateTime="1632022711231" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Alloperla Leonarda Group Of Eastern North America, With Sem Images Of Four Out-Group Species (Plecoptera: Chloroperlidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Willett, Michele R.</mods:namePart>
<mods:affiliation>and Bill P. Stark &amp; Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: michele _ willett @ yahoo. com</mods:affiliation>
<mods:nameIdentifier type="email">michele_willett@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
<mods:affiliation>Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: michele _ willett @ yahoo. com &amp; Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu &amp; and Bill P. Stark &amp; Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: michele _ willett @ yahoo. com</mods:affiliation>
<mods:nameIdentifier type="email">michele_willett@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="volume">
<mods:number>5</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>11</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>108</mods:start>
<mods:end>127</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://illiesia.speciesfile.org/papers/Illiesia05-11.pdf</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4759290</mods:identifier>
<mods:identifier type="GBIF-Dataset">66abbd26-0f90-4888-9851-b54dc1f135cb</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4759290</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4759324" ID-GBIF-Taxon="182165333" ID-Zenodo-Dep="4759324" LSID="urn:lsid:plazi:treatment:038B87E0FFD5FFADFE80FDB0FA7DFEE3" httpUri="http://treatment.plazi.org/id/038B87E0FFD5FFADFE80FDB0FA7DFEE3" lastPageNumber="123" pageId="15" pageNumber="123">
<subSubSection box="[340,646,569,593]" pageId="15" pageNumber="123" type="nomenclature">
<paragraph blockId="15.[340,646,569,593]" box="[340,646,569,593]" pageId="15" pageNumber="123">
<heading bold="true" box="[340,646,569,593]" centered="true" fontSize="10" level="2" pageId="15" pageNumber="123" reason="2">
<taxonomicName authority="(Say)" baseAuthorityName="Say" baseAuthorityYear="1823" box="[340,646,569,593]" class="Insecta" family="Chloroperlidae" genus="Alloperla" kingdom="Animalia" order="Plecoptera" pageId="15" pageNumber="123" phylum="Arthropoda" rank="species" species="imbecilla">
<emphasis bold="true" box="[340,646,569,593]" pageId="15" pageNumber="123">
<emphasis bold="true" box="[340,570,569,593]" italics="true" pageId="15" pageNumber="123">Alloperla imbecilla</emphasis>
(Say)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[422,565,605,629]" pageId="15" pageNumber="123" type="description">
<paragraph blockId="15.[422,565,605,629]" box="[422,565,605,629]" pageId="15" pageNumber="123">
(
<figureCitation box="[430,557,605,629]" captionStart="Figs" captionStartId="16.[189,241,1816,1840]" captionTargetBox="[249,1383,314,1739]" captionTargetId="figure-54@16.[249,1383,314,1739]" captionTargetPageId="16" captionText="Figs. 61-66. Alloperla imbecilla epiproct structures, Devils Fork, Scott Co., VA. 61. Male terminalia, dorsal aspect. 62. Epiproct and cowl, dorsal aspect. 63. Apex of epiproct, dorsal aspect. 64. Apex of epiproct, lateral aspect. 65. Apex of epiproct, frontal aspect. 66. Detail of basal epiproct surface." figureDoi="http://doi.org/10.5281/zenodo.4759328" httpUri="https://zenodo.org/record/4759328/files/figure.png" pageId="15" pageNumber="123">Figs. 61-66</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="123" type="reference_group">
<paragraph blockId="15.[227,797,673,890]" pageId="15" pageNumber="123">
<treatmentCitationGroup pageId="15" pageNumber="123">
<treatmentCitation author="Say, T." box="[227,537,673,695]" page="165" pageId="15" pageNumber="123" year="1823">
<taxonomicName authority="Say, 1823: 165" authorityName="Say" authorityPageNumber="165" authorityYear="1823" box="[227,537,673,695]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="15" pageNumber="123" phylum="Arthropoda" rank="species" species="imbecilla">
<emphasis box="[227,377,673,694]" italics="true" pageId="15" pageNumber="123">Sialis imbecilla</emphasis>
<bibRefCitation author="Say, T." box="[389,537,673,695]" pageId="15" pageNumber="123" pagination="170 - 175" refId="ref7485" refString="Say, T. 1823. Description of insects belonging to the order Neuroptera Lin., Latr. collected by the expedition authorized by J. C. Calhoun, Secretary of War, under the command of Major S. H. Long. Western Quarterly Reporter, 2: 160 - 165, (reprinted 1883, pp. 170 - 175)." type="journal article" year="1823">Say, 1823:165</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus box="[551,605,674,695]" pageId="15" pageNumber="123">Type</typeStatus>
lost,
<collectingRegion box="[670,723,673,695]" country="United States of America" name="Ohio" pageId="15" pageNumber="123">Ohio</collectingRegion>
River, Cincinnati,
<collectingRegion box="[350,403,705,727]" country="United States of America" name="Ohio" pageId="15" pageNumber="123">Ohio</collectingRegion>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="15.[227,797,673,890]" box="[227,659,738,760]" pageId="15" pageNumber="123">
<treatmentCitationGroup box="[227,659,738,760]" pageId="15" pageNumber="123">
<treatmentCitation author="Baumann, R. W." box="[227,655,738,760]" page="258" pageId="15" pageNumber="123" year="1974">
<taxonomicName authority=": Baumann, 1974: 258" authorityName="Baumann" authorityPageNumber="258" authorityYear="1974" box="[227,655,738,760]" class="Insecta" family="Chloroperlidae" genus="Alloperla" kingdom="Animalia" order="Plecoptera" pageId="15" pageNumber="123" phylum="Arthropoda" rank="species" species="imbecilla">
<emphasis box="[227,417,738,759]" italics="true" pageId="15" pageNumber="123">Alloperla imbecilla</emphasis>
:
<bibRefCitation author="Baumann, R. W." box="[438,655,738,760]" pageId="15" pageNumber="123" pagination="257 - 264" refId="ref6895" refString="Baumann, R. W. 1974. What is Alloperla imbecilla (Say)? Designation of a neotype, and a new Alloperla from eastern North America (Plecoptera: Chloroperlidae). Proceedings of the Biological Society of Washington, 87: 257 - 264." type="journal article" year="1974">Baumann, 1974:258</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="15.[227,797,673,890]" pageId="15" pageNumber="123">
<materialsCitation ID-GBIF-Occurrence="3112004331" county="Adams Co." location="Vastine Run" pageId="15" pageNumber="123" specimenCount="1" specimenCount-male="1" stateProvince="Ohio" typeStatus="neotype">
<typeStatus box="[673,764,739,760]" pageId="15" pageNumber="123">Neotype</typeStatus>
<specimenCount box="[779,797,738,760]" pageId="15" pageNumber="123" type="male">♂</specimenCount>
(United States National Museum),
<location LSID="urn:lsid:plazi:treatment:038B87E0FFD5FFADFE80FDB0FA7DFEE3:8EFD602DFFD5FFADFD9BFC8AFD20FCEE" box="[591,721,771,793]" country="United States of America" county="Adams Co." municipality="National Museum" name="Vastine Run" pageId="15" pageNumber="123" stateProvince="Ohio">Vastine Run</location>
,
<location LSID="urn:lsid:plazi:treatment:038B87E0FFD5FFADFE80FDB0FA7DFEE3:8EFD602DFFD5FFADFD0EFC8AFE93FCCF" country="United States of America" county="Adams Co." municipality="National Museum" name="Lower Twin Creek" pageId="15" pageNumber="123" stateProvince="Ohio">Lower Twin Creek</location>
,
<collectingCounty box="[364,485,803,825]" pageId="15" pageNumber="123">Adams Co.</collectingCounty>
,
<collectingRegion box="[495,548,803,825]" country="United States of America" name="Ohio" pageId="15" pageNumber="123">Ohio</collectingRegion>
</materialsCitation>
</paragraph>
<paragraph blockId="15.[227,797,673,890]" pageId="15" pageNumber="123">
<treatmentCitationGroup pageId="15" pageNumber="123">
<treatmentCitation author="Surdick, R. F." box="[227,595,836,858]" page="17" pageId="15" pageNumber="123" year="2004">
<taxonomicName authority=": Surdick, 2004: 17" authorityName=": Surdick" authorityPageNumber="17" authorityYear="2004" box="[227,595,836,858]" class="Insecta" family="Chloroperlidae" genus="Alloperla" kingdom="Animalia" order="Plecoptera" pageId="15" pageNumber="123" phylum="Arthropoda" rank="species" species="imbecilla">
<emphasis box="[227,408,836,857]" italics="true" pageId="15" pageNumber="123">Alloperla imbecilla</emphasis>
:
<bibRefCitation author="Surdick, R. F." box="[420,595,836,858]" pageId="15" pageNumber="123" refId="ref7688" refString="Surdick, R. F. 2004. Chloroperlidae (The Sallflies). Pp. 1 - 60 in. B. P. Stark &amp; B. J. Armitage (eds.). Stoneflies (Plecoptera) of eastern North America. Volume II. Chloroperlidae, Perlidae, and Perlodidae (Perlodinae). Bulletin of the Ohio Biological Survey, New Series, Volume 14. Ohio Biological Survey, Columbus. 192 pp." type="book" year="2004">Surdick, 2004:17</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Redescription and distribution update
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="123" type="materials_examined">
<paragraph blockId="15.[189,798,933,1918]" pageId="15" pageNumber="123">
<emphasis bold="true" box="[189,427,933,957]" pageId="15" pageNumber="123">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3112004328" collectingDate="1997-05-17" collectionCode="BPS" collectorName="Devils Fork, FR &amp; B. Stark" county="Scott Co." location="Scott Co." pageId="15" pageNumber="123" specimenCount="16" specimenCount-female="8" specimenCount-male="8">
VA:
<collectingCounty box="[491,599,934,958]" pageId="15" pageNumber="123">Scott Co.</collectingCounty>
,
<collectorName box="[612,796,934,958]" pageId="15" pageNumber="123">Devils Fork, FR</collectorName>
619,
<date box="[243,391,970,994]" pageId="15" pageNumber="123" value="1997-05-17">
<collectingDate box="[243,391,970,994]" pageId="15" pageNumber="123" value="1997-05-17">17 May 1997</collectingDate>
</date>
,
<collectorName box="[402,495,969,993]" pageId="15" pageNumber="123">B. Stark</collectorName>
,
<specimenCount box="[505,545,969,994]" pageId="15" pageNumber="123" type="male">8 ♂</specimenCount>
,
<specimenCount box="[558,597,969,994]" pageId="15" pageNumber="123" type="female">8 ♀</specimenCount>
(
<collectionCode box="[611,663,970,994]" country="USA" lsid="urn:lsid:biocol.org:col:14922" name="California Department of Food and Agriculture" pageId="15" pageNumber="123" type="Herbarium">BPS</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3112004303" collectingDate="2009-05-19" collectionCode="BPS" collectorName="B. Stark" location="Same" pageId="15" pageNumber="123" specimenCount="16" specimenCount-female="6" specimenCount-male="10">
<location LSID="urn:lsid:plazi:treatment:038B87E0FFD5FFADFE80FDB0FA7DFEE3:8EFD602DFFD5FFADFD7CFC43FD16FC14" box="[680,743,970,994]" name="Same" pageId="15" pageNumber="123">Same</location>
site,
<date box="[189,338,1005,1030]" pageId="15" pageNumber="123" value="2009-05-19">
<collectingDate box="[189,338,1005,1030]" pageId="15" pageNumber="123" value="2009-05-19">19 May 2009</collectingDate>
</date>
,
<collectorName box="[349,442,1005,1029]" pageId="15" pageNumber="123">B. Stark</collectorName>
,
<specimenCount box="[452,506,1005,1029]" pageId="15" pageNumber="123" type="male">10 ♂</specimenCount>
,
<specimenCount box="[518,558,1005,1029]" pageId="15" pageNumber="123" type="female">6 ♀</specimenCount>
(
<collectionCode box="[572,624,1005,1029]" country="USA" lsid="urn:lsid:biocol.org:col:14922" name="California Department of Food and Agriculture" pageId="15" pageNumber="123" type="Herbarium">BPS</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3112004313" collectingDate="2009-05-19" collectionCode="BPS" collectorName="Big Stony Creek, FR &amp; B. Stark" county="Scott Co." location="Big Stony Creek" pageId="15" pageNumber="123" specimenCount="1" specimenCount-male="1">
<collectingCounty box="[641,747,1005,1029]" pageId="15" pageNumber="123">Scott Co.</collectingCounty>
,
<collectorName pageId="15" pageNumber="123">
<location LSID="urn:lsid:plazi:treatment:038B87E0FFD5FFADFE80FDB0FA7DFEE3:8EFD602DFFD5FFADFD22FC64FEA4FBDE" county="Scott Co." name="Big Stony Creek" pageId="15" pageNumber="123">Big Stony Creek</location>
, FR
</collectorName>
657,
<date box="[461,621,1041,1065]" pageId="15" pageNumber="123" value="2009-05-19">
<collectingDate box="[461,621,1041,1065]" pageId="15" pageNumber="123" value="2009-05-19">19 May 2009</collectingDate>
</date>
,
<collectorName box="[637,736,1040,1064]" pageId="15" pageNumber="123">B. Stark</collectorName>
,
<specimenCount box="[751,797,1040,1065]" pageId="15" pageNumber="123" type="male">1 ♂</specimenCount>
(
<collectionCode box="[197,249,1076,1100]" country="USA" lsid="urn:lsid:biocol.org:col:14922" name="California Department of Food and Agriculture" pageId="15" pageNumber="123" type="Herbarium">BPS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="15.[189,798,933,1918]" pageId="15" pageNumber="123">
<emphasis bold="true" box="[189,299,1111,1135]" pageId="15" pageNumber="123">Epiproct.</emphasis>
Length from base to tip, ca. 266-275 μm. Greatest width beyond base, ca. 39-46 μm. Greatest apical thickness, ca. 43 μm. Epiproct cowl without enlarged lobes (
<figureCitation box="[377,547,1218,1242]" captionStart="Figs" captionStartId="16.[189,241,1816,1840]" captionTargetBox="[249,1383,314,1739]" captionTargetId="figure-54@16.[249,1383,314,1739]" captionTargetPageId="16" captionText="Figs. 61-66. Alloperla imbecilla epiproct structures, Devils Fork, Scott Co., VA. 61. Male terminalia, dorsal aspect. 62. Epiproct and cowl, dorsal aspect. 63. Apex of epiproct, dorsal aspect. 64. Apex of epiproct, lateral aspect. 65. Apex of epiproct, frontal aspect. 66. Detail of basal epiproct surface." figureDoi="http://doi.org/10.5281/zenodo.4759328" httpUri="https://zenodo.org/record/4759328/files/figure.png" pageId="15" pageNumber="123">Figs. 61-62, 66</figureCitation>
); densely setose over most of cowl surface but bare near base of epiproct. Epiproct widest at base (ca. 61 μm) and on subapical cap (ca. 39 μm), but similar in width through most of length (
<figureCitation box="[285,374,1360,1385]" captionStart="Figs" captionStartId="16.[189,241,1816,1840]" captionTargetBox="[249,1383,314,1739]" captionTargetId="figure-54@16.[249,1383,314,1739]" captionTargetPageId="16" captionText="Figs. 61-66. Alloperla imbecilla epiproct structures, Devils Fork, Scott Co., VA. 61. Male terminalia, dorsal aspect. 62. Epiproct and cowl, dorsal aspect. 63. Apex of epiproct, dorsal aspect. 64. Apex of epiproct, lateral aspect. 65. Apex of epiproct, frontal aspect. 66. Detail of basal epiproct surface." figureDoi="http://doi.org/10.5281/zenodo.4759328" httpUri="https://zenodo.org/record/4759328/files/figure.png" pageId="15" pageNumber="123">Fig. 62</figureCitation>
). Most of dorsum covered with a closely appressed, densely setose mat (
<figureCitation captionStart="Figs" captionStartId="16.[189,241,1816,1840]" captionTargetBox="[249,1383,314,1739]" captionTargetId="figure-54@16.[249,1383,314,1739]" captionTargetPageId="16" captionText="Figs. 61-66. Alloperla imbecilla epiproct structures, Devils Fork, Scott Co., VA. 61. Male terminalia, dorsal aspect. 62. Epiproct and cowl, dorsal aspect. 63. Apex of epiproct, dorsal aspect. 64. Apex of epiproct, lateral aspect. 65. Apex of epiproct, frontal aspect. 66. Detail of basal epiproct surface." figureDoi="http://doi.org/10.5281/zenodo.4759328" httpUri="https://zenodo.org/record/4759328/files/figure.png" pageId="15" pageNumber="123">Figs. 62-63, 66</figureCitation>
), but small basolateral and apical areas, and entire ventral surface bare; apex with a distinctive ventral row of long setae (
<figureCitation box="[451,589,1502,1527]" captionStart="Figs" captionStartId="16.[189,241,1816,1840]" captionTargetBox="[249,1383,314,1739]" captionTargetId="figure-54@16.[249,1383,314,1739]" captionTargetPageId="16" captionText="Figs. 61-66. Alloperla imbecilla epiproct structures, Devils Fork, Scott Co., VA. 61. Male terminalia, dorsal aspect. 62. Epiproct and cowl, dorsal aspect. 63. Apex of epiproct, dorsal aspect. 64. Apex of epiproct, lateral aspect. 65. Apex of epiproct, frontal aspect. 66. Detail of basal epiproct surface." figureDoi="http://doi.org/10.5281/zenodo.4759328" httpUri="https://zenodo.org/record/4759328/files/figure.png" pageId="15" pageNumber="123">Figs. 63-65</figureCitation>
) and a median longitudinal patch of additional setae extends onto spiny process laterally or basolaterally (
<figureCitation box="[648,726,1574,1598]" captionStart="Figs" captionStartId="16.[189,241,1816,1840]" captionTargetBox="[249,1383,314,1739]" captionTargetId="figure-54@16.[249,1383,314,1739]" captionTargetPageId="16" captionText="Figs. 61-66. Alloperla imbecilla epiproct structures, Devils Fork, Scott Co., VA. 61. Male terminalia, dorsal aspect. 62. Epiproct and cowl, dorsal aspect. 63. Apex of epiproct, dorsal aspect. 64. Apex of epiproct, lateral aspect. 65. Apex of epiproct, frontal aspect. 66. Detail of basal epiproct surface." figureDoi="http://doi.org/10.5281/zenodo.4759328" httpUri="https://zenodo.org/record/4759328/files/figure.png" pageId="15" pageNumber="123">Fig. 63</figureCitation>
); apex bearing a slightly thickened, multispined process (
<figureCitation box="[197,326,1645,1669]" captionStart="Figs" captionStartId="16.[189,241,1816,1840]" captionTargetBox="[249,1383,314,1739]" captionTargetId="figure-54@16.[249,1383,314,1739]" captionTargetPageId="16" captionText="Figs. 61-66. Alloperla imbecilla epiproct structures, Devils Fork, Scott Co., VA. 61. Male terminalia, dorsal aspect. 62. Epiproct and cowl, dorsal aspect. 63. Apex of epiproct, dorsal aspect. 64. Apex of epiproct, lateral aspect. 65. Apex of epiproct, frontal aspect. 66. Detail of basal epiproct surface." figureDoi="http://doi.org/10.5281/zenodo.4759328" httpUri="https://zenodo.org/record/4759328/files/figure.png" pageId="15" pageNumber="123">Figs. 63-64</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="123" type="discussion">
<paragraph blockId="15.[189,798,933,1918]" lastBlockId="15.[835,1420,253,277]" pageId="15" pageNumber="123">
<emphasis bold="true" box="[189,304,1680,1704]" pageId="15" pageNumber="123">Remarks.</emphasis>
The specimens examined in this study represent the southern most population known for this species, and they exhibit subtle variations from the figures presented by
<bibRefCitation author="Surdick, R. F." box="[544,729,1787,1811]" pageId="15" pageNumber="123" refId="ref7688" refString="Surdick, R. F. 2004. Chloroperlidae (The Sallflies). Pp. 1 - 60 in. B. P. Stark &amp; B. J. Armitage (eds.). Stoneflies (Plecoptera) of eastern North America. Volume II. Chloroperlidae, Perlidae, and Perlodidae (Perlodinae). Bulletin of the Ohio Biological Survey, New Series, Volume 14. Ohio Biological Survey, Columbus. 192 pp." type="book" year="2004">Surdick (2004)</bibRefCitation>
and
<bibRefCitation author="Baumann, R. W." box="[189,394,1823,1847]" pageId="15" pageNumber="123" pagination="257 - 264" refId="ref6895" refString="Baumann, R. W. 1974. What is Alloperla imbecilla (Say)? Designation of a neotype, and a new Alloperla from eastern North America (Plecoptera: Chloroperlidae). Proceedings of the Biological Society of Washington, 87: 257 - 264." type="journal article" year="1974">Baumann (1974)</bibRefCitation>
. A comparative SEM study is needed for several populations to evaluate the extent of variation in the shape, setation and number of serrae for the apical spiny portion of the epiproct.
</paragraph>
</subSubSection>
</treatment>
</document>
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