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<document id="10E235686E138CBD5F7C26491172FBB0" ID-DOI="10.5281/zenodo.276189" ID-GBIF-Dataset="d48ca3f0-5056-4e1d-8797-1539d6bd28bc" ID-ISSN="1175-5326" ID-Zenodo-Dep="276189" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460312759491" checkinUser="plazi" docAuthor="González-Moreno, Alejandra, Bordera, Santiago &amp; Delfin, Hugo" docDate="2010" docId="038B87D5FFAF5979FF739BFD3615EF3F" docLanguage="en" docName="zt02648p068.pdf" docOrigin="Zootaxa 2648" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Endasys gibbosus Gonzalez-Moreno et Bordera, sp. nov." docType="treatment" docVersion="6" lastPageNumber="67" masterDocId="FFB2FFADFFAD597FFFE49B6A372FEC48" masterDocTitle="A new species of Endasys Förster (Hymenoptera, Ichneumonidae, Cryptinae) from Mexico" masterLastPageNumber="68" masterPageNumber="61" pageNumber="63" updateTime="1698254448033" updateUser="plazi">
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<mods:title id="F7B0F5B5B9392D8D64AC8317D2E2B704">A new species of Endasys Förster (Hymenoptera, Ichneumonidae, Cryptinae) from Mexico</mods:title>
</mods:titleInfo>
<mods:name id="257E3A6C0C610CF8B4B39E0646EC3103" type="personal">
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<mods:namePart id="EC51387BF09F522D92C21B7CC34E2609">González-Moreno, Alejandra</mods:namePart>
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<mods:namePart id="1EB3ADCB44A12B78D1A5EBDC8DCF059A">Bordera, Santiago</mods:namePart>
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<mods:namePart id="030AEE4A20249F9BE403638116686F8A">Delfin, Hugo</mods:namePart>
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<mods:title id="895B05CE66C65D3DFB6495A49B819DE3">Zootaxa</mods:title>
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<mods:date id="78E2057203BEAB9D481BC4A9C88A637C">2010</mods:date>
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<treatment id="038B87D5FFAF5979FF739BFD3615EF3F" ID-DOI="http://doi.org/10.5281/zenodo.6208959" ID-GBIF-Taxon="119401048" ID-Zenodo-Dep="6208959" LSID="urn:lsid:plazi:treatment:038B87D5FFAF5979FF739BFD3615EF3F" httpUri="http://treatment.plazi.org/id/038B87D5FFAF5979FF739BFD3615EF3F" lastPageId="6" lastPageNumber="67" pageId="2" pageNumber="63">
<subSubSection id="C3386548FFAF597DFF739BFD35A2EC9C" pageId="2" pageNumber="63" type="nomenclature">
<paragraph id="8B9D36C3FFAF597DFF739BFD3461ECF9" blockId="2.[151,846,151,212]" box="[151,846,151,177]" pageId="2" pageNumber="63">
<heading id="D0D581AFFFAF597DFF739BFD3461ECF9" bold="true" box="[151,846,151,177]" fontSize="11" level="1" pageId="2" pageNumber="63" reason="1">
<emphasis id="B956EAD1FFAF597DFF739BFD3461ECF9" bold="true" box="[151,846,151,177]" pageId="2" pageNumber="63">
<taxonomicName id="4C224D40FFAF597DFF739BFD35CCECF9" authority="Gonzalez-Moreno et Bordera" authorityName="Gonzalez-Moreno et Bordera" box="[151,739,151,177]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="gibbosus" status="sp. nov.">
<emphasis id="B956EAD1FFAF597DFF739BFD3642ECF9" bold="true" box="[151,365,151,177]" italics="true" pageId="2" pageNumber="63">Endasys gibbosus</emphasis>
González-Moreno et Bordera
</taxonomicName>
,
<taxonomicNameLabel id="A26557AAFFAF597DFD149BFD3461ECF9" box="[752,846,151,177]" pageId="2" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36C3FFAF597DFF739BD035A2EC9C" blockId="2.[151,846,151,212]" box="[151,653,186,212]" pageId="2" pageNumber="63">
(
<figureCitation id="13192A46FFAF597DFF7B9BD037CAEC9C" box="[159,229,186,212]" captionStart="FIGURE 1" captionStartId="2.[151,257,1522,1546]" captionTargetBox="[151,1436,530,1495]" captionTargetId="figure@2.[151,1436,530,1498]" captionTargetPageId="2" captionText="FIGURE 1. Endasys spp. Habitus. A B, males: A— E. gibbosus holotype, B— E. julianus. C D, females: C — E. gibbosus paratype, D— E. subclavatus." httpUri="https://zenodo.org/record/276190/files/figure.png" pageId="2" pageNumber="63">Figs 1</figureCitation>
A, 1C, 2A2F, 3A, 3C, 3E, 4A, 4C)
</paragraph>
</subSubSection>
<subSubSection id="C3386548FFAF597DFF739A6F3268ED92" pageId="2" pageNumber="63" type="diagnosis">
<paragraph id="8B9D36C3FFAF597DFF739A6F3268ED92" blockId="2.[151,1437,261,474]" pageId="2" pageNumber="63">
<emphasis id="B956EAD1FFAF597DFF739A6F3638ED57" bold="true" box="[151,279,261,287]" pageId="2" pageNumber="63">Diagnosis.</emphasis>
Head and mesosoma black (
<figureCitation id="13192A46FFAF597DFD839A6F3582ED57" box="[615,685,261,287]" captionStart="FIGURE 1" captionStartId="2.[151,257,1522,1546]" captionTargetBox="[151,1436,530,1495]" captionTargetId="figure@2.[151,1436,530,1498]" captionTargetPageId="2" captionText="FIGURE 1. Endasys spp. Habitus. A B, males: A— E. gibbosus holotype, B— E. julianus. C D, females: C — E. gibbosus paratype, D— E. subclavatus." httpUri="https://zenodo.org/record/276190/files/figure.png" pageId="2" pageNumber="63">Figs 1</figureCitation>
A, 1C). Coxa and trochanter orange in female, orange-brown in male. Three or four tyloids in male flagellum (
<figureCitation id="13192A46FFAF597DFD279A403422ED0C" box="[707,781,298,324]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="2" pageNumber="63">Fig. 2</figureCitation>
B). Clypeus black, upturned (
<figureCitation id="13192A46FFAF597DFB8F9A40339CED0C" box="[1131,1203,298,324]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="2" pageNumber="63">Fig. 2</figureCitation>
A), more evident in male, propodeal apophysis as low crest (
<figureCitation id="13192A46FFAF597DFD449A3A35DDED22" box="[672,754,336,362]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="2" pageNumber="63">Figs 2</figureCitation>
D, 3E). Flagellum bicolored dorsally. Median lobe of mesoscutum anteriorly more raised than lateral lobes (
<figureCitation id="13192A46FFAF597DFCC99A1F3458EDC7" box="[813,887,373,399]" captionStart="FIGURE 1" captionStartId="2.[151,257,1522,1546]" captionTargetBox="[151,1436,530,1495]" captionTargetId="figure@2.[151,1436,530,1498]" captionTargetPageId="2" captionText="FIGURE 1. Endasys spp. Habitus. A B, males: A— E. gibbosus holotype, B— E. julianus. C D, females: C — E. gibbosus paratype, D— E. subclavatus." httpUri="https://zenodo.org/record/276190/files/figure.png" pageId="2" pageNumber="63">Figs 1</figureCitation>
A, 1C, 2C, 3C) with coarse dense punctures. Metapleuron aerolate-rugose (
<figureCitation id="13192A46FFAF597DFDE49AF03563EDFC" box="[512,588,410,436]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="2" pageNumber="63">Figs 2</figureCitation>
F, 4C).
<taxonomicName id="4C224D40FFAF597DFD439AF035CEEDFC" box="[679,737,410,436]" class="Insecta" family="Pyralidae" genus="Area" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">Area</taxonomicName>
superomedia with strongly elevated longitudinal wrinkles (
<figureCitation id="13192A46FFAF597DFF7B9AAA37C6ED92" box="[159,233,448,474]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="2" pageNumber="63">Figs 2</figureCitation>
D, 3E). Malar space very reduced in male, about 0.1 times as basal mandibular width (
<figureCitation id="13192A46FFAF597DFB3A9AAA3215ED92" box="[1246,1338,448,474]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="2" pageNumber="63">Fig. 2E</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF5D664BFFAF597DFF739E983519EA61" httpUri="https://zenodo.org/record/276190/files/figure.png" pageId="2" pageNumber="63" targetBox="[151,1436,530,1495]" targetPageId="2">
<paragraph id="8B9D36C3FFAF597DFF739E983519EA61" blockId="2.[151,1437,1522,1577]" pageId="2" pageNumber="63">
<emphasis id="B956EAD1FFAF597DFF739E983632EA42" bold="true" box="[151,285,1522,1546]" pageId="2" pageNumber="63">FIGURE 1.</emphasis>
<taxonomicName id="4C224D40FFAF597DFEC19E9836AEEA41" box="[293,385,1522,1545]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B956EAD1FFAF597DFEC19E9836AEEA41" box="[293,385,1522,1545]" italics="true" pageId="2" pageNumber="63">Endasys</emphasis>
</taxonomicName>
spp. Habitus. AB, males: A—
<taxonomicName id="4C224D40FFAF597DFD009E983449EA41" box="[740,870,1522,1545]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="B956EAD1FFAF597DFD009E983449EA41" box="[740,870,1522,1545]" italics="true" pageId="2" pageNumber="63">E. gibbosus</emphasis>
</taxonomicName>
<emphasis id="B956EAD1FFAF597DFC8A9E9834FCEA42" bold="true" box="[878,979,1522,1546]" pageId="2" pageNumber="63">holotype</emphasis>
, B—
<emphasis id="B956EAD1FFAF597DFBF49E9833BEEA41" box="[1040,1169,1522,1545]" italics="true" pageId="2" pageNumber="63">
<taxonomicName id="4C224D40FFAF597DFBF49E9833A3EA41" box="[1040,1164,1522,1545]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="julianus">E. julianus</taxonomicName>
.
</emphasis>
C D, females: C —
<taxonomicName id="4C224D40FFAF597DFA609E9837D9EA61" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="B956EAD1FFAF597DFA609E9837D9EA61" italics="true" pageId="2" pageNumber="63">E. gibbosus</emphasis>
</taxonomicName>
paratype, D—
<taxonomicName id="4C224D40FFAF597DFE779D78351DEA61" box="[403,562,1554,1577]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="subclavatus">E. subclavatus</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C3386548FFAF597AFF219D3D326BED0F" lastPageId="5" lastPageNumber="66" pageId="2" pageNumber="63" type="description">
<paragraph id="8B9D36C3FFAF597DFF219D3D3530EAF4" blockId="2.[151,1437,1623,2023]" pageId="2" pageNumber="63">
<emphasis id="B956EAD1FFAF597DFF219D3D3672EA39" bold="true" box="[197,349,1623,1649]" pageId="2" pageNumber="63">Description.</emphasis>
MALE (
<specimenCount id="9D24FD4AFFAF597DFE2C9D3D3540EA39" box="[456,623,1623,1649]" pageId="2" pageNumber="63" type="generic">32 specimens</specimenCount>
). Body length 6.08.0 mm; head height 0.91.1; head width 1.61.7; mesosoma length 2.7; mesosoma width (mesoscutum) 1.41.6; fore wing maximum length 4.56.0; first metasomal tergite length 0.60.8.
</paragraph>
<paragraph id="8B9D36C3FFAF597CFF219DA2323FEC9F" blockId="2.[151,1437,1623,2023]" lastBlockId="3.[151,1438,152,215]" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63">
<emphasis id="B956EAD1FFAF597DFF219DA23629EAA9" box="[197,262,1736,1761]" italics="true" pageId="2" pageNumber="63">Head</emphasis>
(
<figureCitation id="13192A46FFAF597DFEF29DAD364CEAA9" box="[278,355,1735,1761]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="2" pageNumber="63">Figs 2</figureCitation>
A2C, 2E). Transverse, 1.81.9 times as wide as long, in dorsal view slightly constricted behind eyes. Gena coarse and densely punctate, in dorsal view 0.60.8 times as long as length of eye. Vertex and frons rugulose punctuate (
<figureCitation id="13192A46FFAF597DFDF29C78354CEB64" box="[534,611,1810,1836]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="2" pageNumber="63">Fig. 2</figureCitation>
C). Posterior ocellus separated from eye by 1.01.2 times its own maximum diameter. Space between hind ocelli 1.61.8 times their diameter. Face strongly convex in central part, 0.6 as high as wide, densely and deeply foveolate. Clypeus about 1.82.0 times as wide as high, densely foveolate, convex, apical margin truncated, smooth, glabrous, sharply upturned (
<figureCitation id="13192A46FFAF597DFBAB9CE833B8EBD4" box="[1103,1175,1922,1948]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="2" pageNumber="63">Fig. 2</figureCitation>
A). Upper mandibular tooth clearly longer than lower tooth. Supraclypeal suture stout, conspicuous; malar space foveolate, 0.1 times as long as basal mandibular width (
<figureCitation id="13192A46FFAF597DFDA79CA7358EEBAF" box="[579,673,1997,2023]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="2" pageNumber="63">Fig. 2E</figureCitation>
); antenna filiform, slender, from base to apex strongly tapered; flagellomeres 122 longer than wide, the first about 2.72.9 times as long as wide; 34 tyloids in flagellomeres 1013 (
<figureCitation id="13192A46FFAE597CFE739BD736F1EC9F" box="[407,478,189,215]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="3" pageNumber="64">Fig. 2</figureCitation>
B). Genal carina joining oral carina barely distad of base of mandible.
</paragraph>
<caption id="DF5D664BFFAE597CFF739DA035A3EB69" httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="3" pageNumber="64" targetBox="[155,1432,269,1710]" targetPageId="3">
<paragraph id="8B9D36C3FFAE597CFF739DA035A3EB69" blockId="3.[151,1437,1738,1825]" pageId="3" pageNumber="64">
<emphasis id="B956EAD1FFAE597CFF739DA0360FEAAA" bold="true" box="[151,288,1738,1762]" pageId="3" pageNumber="64">FIGURE 2.</emphasis>
<taxonomicName id="4C224D40FFAE597CFECC9DA03683EAA9" box="[296,428,1738,1761]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="gibbosus" status="sp. nov.">
<emphasis id="B956EAD1FFAE597CFECC9DA03683EAA9" box="[296,428,1738,1761]" italics="true" pageId="3" pageNumber="64">E. gibbosus</emphasis>
</taxonomicName>
<emphasis id="B956EAD1FFAE597CFE509DA0353FEAAA" bold="true" box="[436,528,1738,1762]" pageId="3" pageNumber="64">
<taxonomicNameLabel id="A26557AAFFAE597CFE509DA0353FEAAA" box="[436,528,1738,1762]" pageId="3" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male
<emphasis id="B956EAD1FFAE597CFDBA9DA035EAEAAA" bold="true" box="[606,709,1738,1762]" pageId="3" pageNumber="64">holotype</emphasis>
: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron.
</paragraph>
</caption>
<paragraph id="8B9D36C3FFAE597BFF219C3A3214EEF4" blockId="3.[151,1436,1871,2009]" lastBlockId="4.[151,1437,152,2007]" lastPageId="4" lastPageNumber="65" pageId="3" pageNumber="64">
<emphasis id="B956EAD1FFAE597CFF219C3A366BEB21" box="[197,324,1872,1897]" italics="true" pageId="3" pageNumber="64">Mesosoma</emphasis>
(
<figureCitation id="13192A46FFAE597CFEB09C25368EEB21" box="[340,417,1871,1897]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="3" pageNumber="64">Figs 2</figureCitation>
C 2F). Propleuron punctate. Upper part of pronotum punctate, lower part ruguloselacunose or longitudinally wrinkled. Epomia strong, evanescent near dorsal margin of pronotum (
<figureCitation id="13192A46FFAE597CFAD59C1F32A0EBC7" box="[1329,1423,1909,1935]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="3" pageNumber="64">Fig. 2E</figureCitation>
). Mesoscutum polished, shiny, coarse, densely punctate. Notaulus strongly impressed, deep in front margin of mesoscutum, reaching tegula level, median lobe of mesoscutum clarly more prominent anteriorly than lateral lobes, with coarse, dense pits. Posterior edge of mesoscutum with trasverse break rather narrow, weak. Prescutellar transverse groove with strong median longitudinal carina. Scutellum flat, without lateral carinae, punctate (
<figureCitation id="13192A46FFA9597BFEEF9B88367DECB4" box="[267,338,226,252]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="4" pageNumber="65">Fig. 2</figureCitation>
C). Mesopleuron punctate, except central
<taxonomicName id="4C224D40FFA9597BFCA69B88345CECB4" box="[834,883,226,252]" class="Insecta" family="Pyralidae" genus="Area" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">area</taxonomicName>
smooth, with very sparse punctures, or almost impunctate. Epicnemial carina stout, its upper end dorsad of subtegular ridge. Sternaulus strongly impressed on anterior half of mesopleuron. Metapleuron very strongly areolate-rugose (
<figureCitation id="13192A46FFA9597BFBCC9A47335FED0F" box="[1064,1136,301,327]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="4" pageNumber="65">Fig. 2</figureCitation>
F). Propodeum carinated, all areas strongly rugose punctate.
<taxonomicName id="4C224D40FFA9597BFDD29A38355FED24" box="[566,624,338,364]" class="Insecta" family="Pyralidae" genus="Area" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">Area</taxonomicName>
superomedia broadly hexagonal with strongly elevated longitudinal wrinkles.
<taxonomicName id="4C224D40FFA9597BFEE89A12366AEDDA" box="[268,325,376,402]" class="Insecta" family="Pyralidae" genus="Area" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">Area</taxonomicName>
basalis transverse very reduced.
<taxonomicName id="4C224D40FFA9597BFD349A123450EDDA" box="[720,895,376,402]" class="Insecta" family="Pyralidae" genus="Area" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="petiolaris">Area petiolaris</taxonomicName>
strongly concave and transversally wrinkled.
<taxonomicName id="4C224D40FFA9597BFF739AF737FEEDFF" box="[151,209,413,439]" class="Insecta" family="Pyralidae" genus="Area" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">Area</taxonomicName>
spiracularis small and triangular.
<taxonomicName id="4C224D40FFA9597BFD829AF7358FEDFF" box="[614,672,413,439]" class="Insecta" family="Pyralidae" genus="Area" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">Area</taxonomicName>
lateralis much longer than wide also triangular. Both transverse carinae of propodeum stout, complete, apical carina forming sublateral lower crest. Propodeal spiracle oval, closer to lateral longitudinal carina than to pleural carina (
<figureCitation id="13192A46FFA9597BFCDC9A823450EE4A" box="[824,895,488,514]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="4" pageNumber="65">Fig. 2</figureCitation>
D). Front wing vein Rs straight; crossvein 1mcu variable, usually absent; marginal cell 0.3 times as long as its maximum width; crossvein 2m-cu vein weakly inclivous, with one wide bulla. Areolet small, pentagonal, strongly convergent towards front side, crossvein 3rs-m usually not pigmented. Vein Cu-a opposite vein Rs&amp;M; abscissa of Cu1 between 1m-cu and Cu1a about 1.82.2 times as long as Cu1b, this one, strongly inclivous.
<collectingCountry id="F3357653FFA9597BFC339917333DEEDF" box="[983,1042,637,663]" name="India" pageId="4" pageNumber="65">Hind</collectingCountry>
wing vein Cu-a clearly inclivous, 0.3 times as long as abcissa of M+Cu between M and Cu1.
<collectingCountry id="F3357653FFA9597BFCAB99C834A5EEF4" box="[847,906,674,700]" name="India" pageId="4" pageNumber="65">Hind</collectingCountry>
femur 3.74.0 times as long as high.
</paragraph>
<paragraph id="8B9D36C3FFA9597BFF2199A236E4EFD4" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="65">
<emphasis id="B956EAD1FFA9597BFF2199A23610EEA9" box="[197,319,712,737]" italics="true" pageId="4" pageNumber="65">Metasoma</emphasis>
(
<figureCitation id="13192A46FFA9597BFEAB99A236BAEEAA" box="[335,405,712,738]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="4" pageNumber="65">Fig. 2</figureCitation>
D). First tergite 1.82.0 as long as its maximum width, smooth and polished, with some sparse setiferous punctures. Median dorsal carinae only present on petiole, becoming obsolescent on the hind half of postpetiole. Postpetiole 0.50.6 times as long as its maximum width, nearly parallel sided (
<figureCitation id="13192A46FFA9597BFACB98783259EF64" box="[1327,1398,786,812]" captionStart="FIGURE 2. E" captionStartId="3.[151,259,1738,1762]" captionTargetBox="[155,1432,269,1710]" captionTargetId="figure@3.[155,1432,269,1717]" captionTargetPageId="3" captionText="FIGURE 2. E. gibbosus sp. nov., male holotype: A—Lower head left, upturned clypeus (arrow). B—Tyloids on flagellum. C—Head and anterior mesosoma dorsal. D—Propodeum dorsal. E—Head lateral and anterior part of mesosoma. F—Mesopleuron and metapleuron." httpUri="https://zenodo.org/record/276191/files/figure.png" pageId="4" pageNumber="65">Fig. 2</figureCitation>
D). Second and rest of tergites smooth or finely coriaceous, with small setiferous punctures, denser in tergites 3 7. Second metasomal tergite 0.6 times as long as its maximum width (dorsally measured). Thyridia circular very small next to margin.
</paragraph>
<paragraph id="8B9D36C3FFA9597BFF2198C236D2E844" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="65">
<emphasis id="B956EAD1FFA9597BFF2198C23626EF89" box="[197,265,936,961]" italics="true" pageId="4" pageNumber="65">Color</emphasis>
. Body black. Scape and pedicel orange, basal flagellomeres dorsally light orange. Mandible dark brown with black teeth. Palpi yellow.
<collectingCountry id="F3357653FFA9597BFD8798A735B0EFAF" box="[611,671,973,999]" name="India" pageId="4" pageNumber="65">Hind</collectingCountry>
leg orange with coxa dorsally brown. Pterostigma dark brown. Wing yellowish. Tegula black.
</paragraph>
<paragraph id="8B9D36C3FFA9597BFF219F723337E834" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="65">
FEMALE (
<specimenCount id="9D24FD4AFFA9597BFEA89F7236CAE87A" box="[332,485,1048,1074]" pageId="4" pageNumber="65" type="generic">2 specimens</specimenCount>
). Body length (without ovipositor) 6.07.0 mm; head maximum height 1.11.2; head width 1.41.6; mesosoma length 2.63.0; mesosoma width (mesoscutum) 1.6; fore wing maximum lenght 6.0; first metasomal tergite length 1.0; ovipositor sheaths length 1.5.
</paragraph>
<paragraph id="8B9D36C3FFA9597BFF219FE23380E9CA" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="65">
<emphasis id="B956EAD1FFA9597BFF219FE2362BE8E9" box="[197,260,1160,1185]" italics="true" pageId="4" pageNumber="65">Head</emphasis>
(
<figureCitation id="13192A46FFA9597BFEF09FE23674E8EA" box="[276,347,1160,1186]" captionStart="FIGURE 1" captionStartId="2.[151,257,1522,1546]" captionTargetBox="[151,1436,530,1495]" captionTargetId="figure@2.[151,1436,530,1498]" captionTargetPageId="2" captionText="FIGURE 1. Endasys spp. Habitus. A B, males: A— E. gibbosus holotype, B— E. julianus. C D, females: C — E. gibbosus paratype, D— E. subclavatus." httpUri="https://zenodo.org/record/276190/files/figure.png" pageId="4" pageNumber="65">Figs 1</figureCitation>
C, 3A, 4A). Transverse, 1.71.9 times as wide as long; in dorsal view not constricted behind eyes. Gena densely foveolate, 1.0 times as long as maximum length of eye (dorsal view). Vertex and frons rugulose-lacunose with very dense coarse punctures. Eye conspicuously pilose (
<figureCitation id="13192A46FFA9597BFB9C9FB833EAE8A4" box="[1144,1221,1234,1260]" captionStart="FIGURE 3" captionStartId="5.[151,255,1853,1877]" captionTargetBox="[151,1436,381,1832]" captionTargetId="figure@5.[151,1436,381,1833]" captionTargetPageId="5" captionText="FIGURE 3. Females: A, C, E: E. gibbosus sp. nov., B, D, F: E. subclavatus. A B, head dorsal. C D, Mesosoma dorsal. E F, Propodeum and petiole dorsal." httpUri="https://zenodo.org/record/276192/files/figure.png" pageId="4" pageNumber="65">Figs 3</figureCitation>
A, 4A). Posterior ocellus separated from eye by 1.41.7 times its own maximum diameter. Space between hind ocelli 2.1 times their diameter. Face 0.5 times as long as wide. Clypeus about 2.12.3 times as wide as long; malar space 0.5 0.7 times as long as basal mandibular width (
<figureCitation id="13192A46FFA9597BFD249E283425E914" box="[704,778,1346,1372]" captionStart="FIGURE 4" captionStartId="6.[151,255,1923,1947]" captionTargetBox="[155,1432,941,1901]" captionTargetId="figure@6.[155,1432,941,1902]" captionTargetPageId="6" captionText="FIGURE 4. Females: A, C: E. gibbosus sp. nov., B, D: E. subclavatus. A B, head and anterior part of mesosoma left. C D, mesosoma left." httpUri="https://zenodo.org/record/276193/files/figure.png" pageId="4" pageNumber="65">Fig. 4</figureCitation>
A); antenna moderately stout, becoming slightly more slender towards apex (
<figureCitation id="13192A46FFA9597BFE7A9E0236CFE9CA" box="[414,480,1384,1410]" captionStart="FIGURE 1" captionStartId="2.[151,257,1522,1546]" captionTargetBox="[151,1436,530,1495]" captionTargetId="figure@2.[151,1436,530,1498]" captionTargetPageId="2" captionText="FIGURE 1. Endasys spp. Habitus. A B, males: A— E. gibbosus holotype, B— E. julianus. C D, females: C — E. gibbosus paratype, D— E. subclavatus." httpUri="https://zenodo.org/record/276190/files/figure.png" pageId="4" pageNumber="65">Fig. 1</figureCitation>
C), with 20 flagellomeres, the first about 1.5 as long as wide.
</paragraph>
<paragraph id="8B9D36C3FFA9597BFF219EE733D5EA2A" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="65">
<emphasis id="B956EAD1FFA9597BFF219EE73669E9EE" box="[197,326,1421,1446]" italics="true" pageId="4" pageNumber="65">Mesosoma</emphasis>
(
<figureCitation id="13192A46FFA9597BFEB29EE7368DE9EF" box="[342,418,1421,1447]" captionStart="FIGURE 3" captionStartId="5.[151,255,1853,1877]" captionTargetBox="[151,1436,381,1832]" captionTargetId="figure@5.[151,1436,381,1833]" captionTargetPageId="5" captionText="FIGURE 3. Females: A, C, E: E. gibbosus sp. nov., B, D, F: E. subclavatus. A B, head dorsal. C D, Mesosoma dorsal. E F, Propodeum and petiole dorsal." httpUri="https://zenodo.org/record/276192/files/figure.png" pageId="4" pageNumber="65">Figs 3</figureCitation>
A, 3C, 3E, 4C). Propleuron foveolate. Pronotum long with a dorsal median circular depression (
<figureCitation id="13192A46FFA9597BFEC09ED83645E984" box="[292,362,1458,1484]" captionStart="FIGURE 3" captionStartId="5.[151,255,1853,1877]" captionTargetBox="[151,1436,381,1832]" captionTargetId="figure@5.[151,1436,381,1833]" captionTargetPageId="5" captionText="FIGURE 3. Females: A, C, E: E. gibbosus sp. nov., B, D, F: E. subclavatus. A B, head dorsal. C D, Mesosoma dorsal. E F, Propodeum and petiole dorsal." httpUri="https://zenodo.org/record/276192/files/figure.png" pageId="4" pageNumber="65">Fig. 3</figureCitation>
A). Mesoscutum as in male, in central part strongly rugulose-lacunose. Notaulus as in male but with a deep concavity next to anterior margin of mesoscutum; reaching tegula level (
<figureCitation id="13192A46FFA9597BFAC29EB2325BE9BA" box="[1318,1396,1496,1522]" captionStart="FIGURE 3" captionStartId="5.[151,255,1853,1877]" captionTargetBox="[151,1436,381,1832]" captionTargetId="figure@5.[151,1436,381,1833]" captionTargetPageId="5" captionText="FIGURE 3. Females: A, C, E: E. gibbosus sp. nov., B, D, F: E. subclavatus. A B, head dorsal. C D, Mesosoma dorsal. E F, Propodeum and petiole dorsal." httpUri="https://zenodo.org/record/276192/files/figure.png" pageId="4" pageNumber="65">Fig. 3</figureCitation>
C). Mesopleuron foveolate, lower two thirds usually longitudinally wrinkled.
<taxonomicName id="4C224D40FFA9597BFBE99E973368EA5F" box="[1037,1095,1533,1559]" class="Insecta" family="Pyralidae" genus="Area" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">Area</taxonomicName>
superomedia confluent with
<taxonomicName id="4C224D40FFA9597BFF739D4837E6EA74" box="[151,201,1570,1596]" class="Insecta" family="Pyralidae" genus="Area" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">area</taxonomicName>
dentipara and lateralis forming a broad inverted U
<taxonomicName id="4C224D40FFA9597BFCDD9D483441EA74" box="[825,878,1570,1596]" class="Insecta" family="Pyralidae" genus="Area" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">area</taxonomicName>
, with strong wrinkles.
<taxonomicName id="4C224D40FFA9597BFB609D483391EA74" box="[1156,1214,1570,1596]" class="Insecta" family="Pyralidae" genus="Area" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">Area</taxonomicName>
externa anteriorly punctate and posteriorly rugulose-lacunose (
<figureCitation id="13192A46FFA9597BFD7A9D2235CBEA2A" box="[670,740,1608,1634]" captionStart="FIGURE 3" captionStartId="5.[151,255,1853,1877]" captionTargetBox="[151,1436,381,1832]" captionTargetId="figure@5.[151,1436,381,1833]" captionTargetPageId="5" captionText="FIGURE 3. Females: A, C, E: E. gibbosus sp. nov., B, D, F: E. subclavatus. A B, head dorsal. C D, Mesosoma dorsal. E F, Propodeum and petiole dorsal." httpUri="https://zenodo.org/record/276192/files/figure.png" pageId="4" pageNumber="65">Fig. 3</figureCitation>
E).
<collectingCountry id="F3357653FFA9597BFCEA9D223466EA2A" box="[782,841,1608,1634]" name="India" pageId="4" pageNumber="65">Hind</collectingCountry>
femur 3.03.1 times as long as high.
</paragraph>
<paragraph id="8B9D36C3FFA9597BFF219D0734C1EABF" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="65">
<emphasis id="B956EAD1FFA9597BFF219D073610EACE" box="[197,319,1645,1670]" italics="true" pageId="4" pageNumber="65">Metasoma</emphasis>
(
<figureCitation id="13192A46FFA9597BFEAB9D0736BBEACF" box="[335,404,1645,1671]" captionStart="FIGURE 3" captionStartId="5.[151,255,1853,1877]" captionTargetBox="[151,1436,381,1832]" captionTargetId="figure@5.[151,1436,381,1833]" captionTargetPageId="5" captionText="FIGURE 3. Females: A, C, E: E. gibbosus sp. nov., B, D, F: E. subclavatus. A B, head dorsal. C D, Mesosoma dorsal. E F, Propodeum and petiole dorsal." httpUri="https://zenodo.org/record/276192/files/figure.png" pageId="4" pageNumber="65">Fig. 3</figureCitation>
E). First tergite 1.41.5 times as long as its maximum width. Postpetiole 0.40.5 times as long as its maximum width, sides slightly divergent posteriorly (
<figureCitation id="13192A46FFA9597BFC5E9DF8332EEAE4" box="[954,1025,1682,1708]" captionStart="FIGURE 3" captionStartId="5.[151,255,1853,1877]" captionTargetBox="[151,1436,381,1832]" captionTargetId="figure@5.[151,1436,381,1833]" captionTargetPageId="5" captionText="FIGURE 3. Females: A, C, E: E. gibbosus sp. nov., B, D, F: E. subclavatus. A B, head dorsal. C D, Mesosoma dorsal. E F, Propodeum and petiole dorsal." httpUri="https://zenodo.org/record/276192/files/figure.png" pageId="4" pageNumber="65">Fig. 3</figureCitation>
E). Second metasomal tergite 0.4 0.5 times as long as its maximum width (dorsally measured). Thyridia indistinct. Ovipositor straight, 0.7 times as long as the hind tibia and 1.1 times as long as basal 4 hind tarsomeres.
</paragraph>
<paragraph id="8B9D36C3FFA9597BFF219C6937F7EB2F" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="65">
<emphasis id="B956EAD1FFA9597BFF219C693625EB54" box="[197,266,1795,1820]" italics="true" pageId="4" pageNumber="65">Color</emphasis>
. Head and mesosoma black. Scape, pedicel, mandible (except teeth) and leg orange. Metasoma orange to reddish brown. Flagellomeres 1011 light yellow, the rest dark brown to black. Other features as in male.
</paragraph>
<paragraph id="8B9D36C3FFA9597BFF219C1836BAEBC4" blockId="4.[151,1437,152,2007]" box="[197,405,1906,1932]" pageId="4" pageNumber="65">
<emphasis id="B956EAD1FFA9597BFF219C18363DEBC4" bold="true" box="[197,274,1906,1932]" pageId="4" pageNumber="65">Hosts.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B9D36C3FFA9597AFF219CFD326BED0F" blockId="4.[151,1437,152,2007]" lastBlockId="5.[151,1437,152,327]" lastPageId="5" lastPageNumber="66" pageId="4" pageNumber="65">
<emphasis id="B956EAD1FFA9597BFF219CFD3603EBF9" bold="true" box="[197,300,1943,1969]" pageId="4" pageNumber="65">Habitat.</emphasis>
Specimens was collected in June and July in a dry forest characterized by trees reaching
<quantity id="4CDA9B26FFA9597BFAB79CF23781EB9F" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="65" unit="m" value="15.0" valueMax="20.0" valueMin="10.0">1020 m</quantity>
in height, 50%75% of which drop their leaves during the dry season, and a shrub and herbaceous layer that develops during the rainy season. Epiphytic plants are abundant (
<collectingCountry id="F3357653FFA8597AFC709BF234F0ECFA" box="[916,991,152,178]" name="Indonesia" pageId="5" pageNumber="66">Flores</collectingCountry>
et Espejel 1994). Common species in this dry forest include
<taxonomicName id="4C224D40FFA8597AFE459BD7349EEC9F" authority="(Schltdl. et Cham.) Benth." authorityName="(Schltdl. et Cham.) Benth." box="[417,945,189,215]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="pennatula">
<emphasis id="B956EAD1FFA8597AFE459BD7355FEC9E" box="[417,624,189,214]" italics="true" pageId="5" pageNumber="66">Acacia pennatula</emphasis>
(Schltdl. et Cham.) Benth.
</taxonomicName>
,
<taxonomicName id="4C224D40FFA8597AFC599BD7320AEC9F" authority="Greenm." authorityName="Greenm." box="[957,1317,189,215]" class="Magnoliopsida" family="Fabaceae" genus="Caesalpinia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="gaumeri">
<emphasis id="B956EAD1FFA8597AFC599BD7339AEC9E" box="[957,1205,189,214]" italics="true" pageId="5" pageNumber="66">Caesalpinia gaumeri</emphasis>
Greenm.
</taxonomicName>
,
<taxonomicName id="4C224D40FFA8597AFAD59BD73602ECB3" class="Magnoliopsida" family="Fabaceae" genus="Lysiloma" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="latisiliquum">
<emphasis id="B956EAD1FFA8597AFAD59BD73602ECB3" italics="true" pageId="5" pageNumber="66">Lysiloma latisiliquum</emphasis>
</taxonomicName>
(L.) Benth.,
<taxonomicName id="4C224D40FFA8597AFE369B8834C3ECB4" authority="(Jacq.) Griseb." authorityName="(Jacq.) Griseb." box="[466,1004,226,252]" class="Magnoliopsida" family="Fabaceae" genus="Enterolobium" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="cyclocarpum">
<emphasis id="B956EAD1FFA8597AFE369B883409ECB3" box="[466,806,226,251]" italics="true" pageId="5" pageNumber="66">Enterolobium cyclocarpum</emphasis>
(Jacq.) Griseb.
</taxonomicName>
,
<taxonomicName id="4C224D40FFA8597AFC1D9B8833F3ECB3" box="[1017,1244,226,251]" class="Magnoliopsida" family="Fabaceae" genus="Piscidia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="piscipula">
<emphasis id="B956EAD1FFA8597AFC1D9B8833F3ECB3" box="[1017,1244,226,251]" italics="true" pageId="5" pageNumber="66">Piscidia piscipula</emphasis>
</taxonomicName>
(L.) Sarg., and
<taxonomicName id="4C224D40FFA8597AFF739A6235B7ED6A" authority="Standl." authorityName="Standl." box="[151,664,264,290]" class="Magnoliopsida" family="Fabaceae" genus="Pithecellobium" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="brownii">
<emphasis id="B956EAD1FFA8597AFF739A623680ED69" box="[151,431,264,289]" italics="true" pageId="5" pageNumber="66">Pithecellobium brownii</emphasis>
Standl. (Fabaceae)
</taxonomicName>
,
<taxonomicName id="4C224D40FFA8597AFD419A623452ED69" box="[677,893,264,289]" class="Magnoliopsida" family="Burseraceae" genus="Bursera" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="simaruba">
<emphasis id="B956EAD1FFA8597AFD419A623452ED69" box="[677,893,264,289]" italics="true" pageId="5" pageNumber="66">Bursera simaruba</emphasis>
</taxonomicName>
(L.) Sarg. (
<taxonomicName id="4C224D40FFA8597AFBEE9A62338DED6A" box="[1034,1186,264,290]" class="Magnoliopsida" family="Burseraceae" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="family">Burseraceae</taxonomicName>
),
<taxonomicName id="4C224D40FFA8597AFB529A6236A2ED0F" authority="Lam." authorityName="Lam." class="Magnoliopsida" family="Malvaceae" genus="Guazuma" kingdom="Plantae" order="Malvales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="ulmifolia">
<emphasis id="B956EAD1FFA8597AFB529A6232B4ED69" box="[1206,1435,264,289]" italics="true" pageId="5" pageNumber="66">Guazuma ulmifolia</emphasis>
Lam. (Sterculiaceae)
</taxonomicName>
,
<taxonomicName id="4C224D40FFA8597AFE7D9A473474ED0F" authority="Greenm." authorityName="Greenm." box="[409,859,301,327]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="gaumeri">
<emphasis id="B956EAD1FFA8597AFE7D9A473513ED0E" box="[409,572,301,326]" italics="true" pageId="5" pageNumber="66">Vitex gaumeri</emphasis>
Greenm. (Verbenaceae)
</taxonomicName>
, and
<taxonomicName id="4C224D40FFA8597AFC7C9A47326EED0F" box="[920,1345,301,327]" class="Magnoliopsida" family="Anacardiaceae" genus="Metopium" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="brownei">
<emphasis id="B956EAD1FFA8597AFC7C9A47335AED0E" box="[920,1141,301,326]" italics="true" pageId="5" pageNumber="66">Metopium brownei</emphasis>
(Anacardiaceae)
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF5D664BFFA8597AFF739C57353BEB3D" httpUri="https://zenodo.org/record/276192/files/figure.png" pageId="5" pageNumber="66" targetBox="[151,1436,381,1832]" targetPageId="5">
<paragraph id="8B9D36C3FFA8597AFF739C57353BEB3D" blockId="5.[151,1437,1853,1909]" pageId="5" pageNumber="66">
<emphasis id="B956EAD1FFA8597AFF739C573636EB1D" bold="true" box="[151,281,1853,1877]" pageId="5" pageNumber="66">FIGURE 3.</emphasis>
Females: A, C, E:
<taxonomicName id="4C224D40FFA8597AFE029C54354CEB1D" box="[486,611,1854,1877]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="gibbosus" status="sp. nov.">
<emphasis id="B956EAD1FFA8597AFE029C54354CEB1D" box="[486,611,1854,1877]" italics="true" pageId="5" pageNumber="66">E. gibbosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557AAFFA8597AFD8D9C573590EB1D" box="[617,703,1853,1877]" pageId="5" pageNumber="66" rank="species">
<emphasis id="B956EAD1FFA8597AFD8D9C573590EB1D" bold="true" box="[617,703,1853,1877]" pageId="5" pageNumber="66">sp. nov.</emphasis>
</taxonomicNameLabel>
, B, D, F:
<emphasis id="B956EAD1FFA8597AFCCD9C5434E3EB1D" box="[809,972,1854,1877]" italics="true" pageId="5" pageNumber="66">
<taxonomicName id="4C224D40FFA8597AFCCD9C5434E7EB1D" box="[809,968,1854,1877]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="subclavatus">E. subclavatus</taxonomicName>
.
</emphasis>
AB, head dorsal. CD, Mesosoma dorsal. EF, Propodeum and petiole dorsal.
</paragraph>
</caption>
<subSubSection id="C3386548FFA85979FF219CC93337EE04" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66" type="discussion">
<paragraph id="8B9D36C3FFA85979FF219CC93337EE04" blockId="5.[151,1437,1955,2019]" lastBlockId="6.[151,1437,152,887]" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66">
<emphasis id="B956EAD1FFA8597AFF219CC93611EBF5" bold="true" box="[197,318,1955,1981]" pageId="5" pageNumber="66">Remarks.</emphasis>
<taxonomicName id="4C224D40FFA8597AFEA29CC93538EBF4" box="[326,535,1955,1980]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="gibbosus" status="sp. nov.">
<emphasis id="B956EAD1FFA8597AFEA29CC93538EBF4" box="[326,535,1955,1980]" italics="true" pageId="5" pageNumber="66">Endasys gibbosus</emphasis>
</taxonomicName>
<emphasis id="B956EAD1FFA8597AFDFB9CC93553EBF5" bold="true" box="[543,636,1955,1981]" pageId="5" pageNumber="66">
<taxonomicNameLabel id="A26557AAFFA8597AFDFB9CC93553EBF5" box="[543,636,1955,1981]" pageId="5" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, can be distinguished from all other Neotropic and Nearctic
<emphasis id="B956EAD1FFA8597AFAA19CC932B4EBF4" box="[1349,1435,1955,1980]" italics="true" pageId="5" pageNumber="66">Endays</emphasis>
species by the combination of characters given in the diagnosis. Nevertheless, two Nearctic species are closely related to
<taxonomicName id="4C224D40FFAB5979FE8F9BF236DAECF9" box="[363,501,152,177]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="B956EAD1FFAB5979FE8F9BF236DAECF9" box="[363,501,152,177]" italics="true" pageId="6" pageNumber="67">E. gibbosus</emphasis>
</taxonomicName>
: males of
<taxonomicName id="4C224D40FFAB5979FD909BF23485ECFA" authority="Luhman, 1990" authorityName="Luhman" authorityYear="1990" box="[628,938,152,178]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="julianus">
<emphasis id="B956EAD1FFAB5979FD909BF235DBECF9" box="[628,756,152,177]" italics="true" pageId="6" pageNumber="67">E. julianus</emphasis>
<bibRefCitation id="EFB34B32FFAB5979FD199BF23485ECFA" author="Luhman" box="[765,938,152,178]" pageId="6" pageNumber="67" refString="Luhman, J. C. (1990) A Taxonomic Revision of Nearctic Endasys Foerster 1868 (Hymenoptera: Ichneumonidae, Gelinae), University of California Press, California, 185 pp." type="book" year="1990">Luhman, 1990</bibRefCitation>
</taxonomicName>
and females of
<taxonomicName id="4C224D40FFAB5979FB8E9BF232B6ECFA" authority="Say, 1836" authorityName="Say" authorityYear="1836" box="[1130,1433,152,178]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="subclavatus">
<emphasis id="B956EAD1FFAB5979FB8E9BF23237ECF9" box="[1130,1304,152,177]" italics="true" pageId="6" pageNumber="67">E. subclavatus</emphasis>
Say, 1836
</taxonomicName>
. The former by having more than 3 tyloids, clypeus strongly upturned, apophysis weak, mesopleuron smooth, postpetiole subquadrate and metasoma partially black, but males of
<taxonomicName id="4C224D40FFAB5979FC539B88336EECB3" box="[951,1089,226,251]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="B956EAD1FFAB5979FC539B88336EECB3" box="[951,1089,226,251]" italics="true" pageId="6" pageNumber="67">E. gibbosus</emphasis>
</taxonomicName>
are clearly distinguishable in having the median lobe of mesoscutum clearly more raised than lateral lobes; metapleuron aerolate-rugose; basal half of antenna light orange or yellow, flagellum strongly tapered from base to apex; postpetiole wider than long, 0.60.7 as long as wide; hind femur, tibia and four basal tarsus entirely orange and metasoma entirely black, except sometimes base of last tergites, which is orange or brown. Females of
<taxonomicName id="4C224D40FFAB5979FB239A12325EEDD9" box="[1223,1393,376,401]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="subclavatus">
<emphasis id="B956EAD1FFAB5979FB239A12325EEDD9" box="[1223,1393,376,401]" italics="true" pageId="6" pageNumber="67">E. subclavatus</emphasis>
</taxonomicName>
are similar to the new species because both have a strong sculpturation in head, gena and frons densely and strongly punctuate and hind femur and tibia entirely orange, but females of
<taxonomicName id="4C224D40FFAB5979FBB69AA833CBED93" box="[1106,1252,450,475]" class="Insecta" family="Ichneumonidae" genus="Endasys" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="B956EAD1FFAB5979FBB69AA833CBED93" box="[1106,1252,450,475]" italics="true" pageId="6" pageNumber="67">E. gibbosus</emphasis>
</taxonomicName>
can be clearly separated by the median lobe of mesoscutum, anteriorly more raised than lateral lobes, with rugulose and coarse dense punctures; gena rugulose-punctate; metapleuron very strongly aerolate-rugose;
<taxonomicName id="4C224D40FFAB5979FB2B9967322FEE6F" box="[1231,1280,525,551]" class="Insecta" family="Pyralidae" genus="Area" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="65" phylum="Arthropoda" rank="genus">area</taxonomicName>
superomedia with strong elevated longitudinal wrinkles and tegula and pterostigma black.
</paragraph>
</subSubSection>
<subSubSection id="C3386548FFAB5979FF21993D3231EE3A" box="[197,1310,599,626]" pageId="6" pageNumber="67" type="etymology">
<paragraph id="8B9D36C3FFAB5979FF21993D3231EE3A" blockId="6.[151,1437,152,887]" box="[197,1310,599,626]" pageId="6" pageNumber="67">
<emphasis id="B956EAD1FFAB5979FF21993D367DEE39" bold="true" box="[197,338,599,625]" pageId="6" pageNumber="67">Etymology.</emphasis>
This species is named referring to the strongly raised median lobe of mesoscutum.
</paragraph>
</subSubSection>
<subSubSection id="C3386548FFAB5979FF219917351CEEDF" box="[197,563,637,663]" pageId="6" pageNumber="67" type="distribution">
<paragraph id="8B9D36C3FFAB5979FF219917351CEEDF" blockId="6.[151,1437,152,887]" box="[197,563,637,663]" pageId="6" pageNumber="67">
<emphasis id="B956EAD1FFAB5979FF21991736E5EEDF" bold="true" box="[197,458,637,663]" pageId="6" pageNumber="67">Distribution records.</emphasis>
<collectingCountry id="F3357653FFAB5979FE369917351FEEDF" box="[466,560,637,663]" name="Mexico" pageId="6" pageNumber="67">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3386548FFAB5979FF2199C83615EF3F" pageId="6" pageNumber="67" type="materials_examined">
<paragraph id="8B9D36C3FFAB5979FF2199C83615EF3F" blockId="6.[151,1437,152,887]" pageId="6" pageNumber="67">
<emphasis id="B956EAD1FFAB5979FF2199C83542EEF4" bold="true" box="[197,621,674,700]" pageId="6" pageNumber="67">
Material examined.
<typeStatus id="54998861FFAB5979FE2199C83542EEF4" box="[453,621,674,700]" pageId="6" pageNumber="67" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
ɗ (
<collectionCode id="ED33AE06FFAB5979FD7E99C835DDEEF4" box="[666,754,674,700]" pageId="6" pageNumber="67">CCBA</collectionCode>
):
<collectingCountry id="F3357653FFAB5979FCE399C8344AEEF4" box="[775,869,674,700]" name="Mexico" pageId="6" pageNumber="67">Mexico</collectingCountry>
, Yucatán State, Ría Lagartos Reserve, Estación Cuyo, dry forest,
<date id="FF9C1003FFAB5979FE8D99A23505EEAA" box="[361,554,712,738]" pageId="6" pageNumber="67" value="2008-07-01" valueMax="2008-07-14" valueMin="2008-07-01">0114.VII.2008</date>
; A González-Moreno leg.; Malaise trap. Dry mounted.
<emphasis id="B956EAD1FFAB5979FB2599AD3242EEA9" bold="true" box="[1217,1389,711,737]" pageId="6" pageNumber="67">
<typeStatus id="54998861FFAB5979FB2599AD3242EEA9" box="[1217,1389,711,737]" pageId="6" pageNumber="67" type="paratype">PARATYPES</typeStatus>
</emphasis>
: 15 ɗɗ (
<collectionCode id="ED33AE06FFAB5979FF299987360BEF4F" box="[205,292,749,775]" pageId="6" pageNumber="67">CCBA</collectionCode>
): same locality and date as
<typeStatus id="54998861FFAB5979FDBB998735E6EF4F" box="[607,713,749,775]" pageId="6" pageNumber="67" type="holotype">holotype</typeStatus>
; 16 ɗɗ(
<collectionCode id="ED33AE06FFAB5979FCCC99873450EF4F" box="[808,895,749,775]" pageId="6" pageNumber="67">CEUA</collectionCode>
): same locality,
<date id="FF9C1003FFAB5979FBDD998733C1EF4F" box="[1081,1262,749,775]" pageId="6" pageNumber="67" value="2009-06-10" valueMax="2009-06-24" valueMin="2009-06-10">1024.VI.2009</date>
; 2 ɗɗ (
<collectionCode id="ED33AE06FFAB5979FAA0998732A0EF4F" LSID="urn:lsid:biocol.org:col:1008" box="[1348,1423,749,775]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1008" name="American Entomological Institute" pageId="6" pageNumber="67">AEIC</collectionCode>
): same locality,
<date id="FF9C1003FFAB5979FEA098783529EF64" box="[324,518,786,812]" pageId="6" pageNumber="67" value="2008-07-01" valueMax="2008-07-14" valueMin="2008-07-01">0114.VII.2008</date>
; 1 ɗ (
<collectionCode id="ED33AE06FFAB5979FDAB987835B3EF64" LSID="urn:lsid:biocol.org:col:1008" box="[591,668,786,812]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1008" name="American Entomological Institute" pageId="6" pageNumber="67">AEIC</collectionCode>
): same locality,
<date id="FF9C1003FFAB5979FCB99878333AEF64" box="[861,1045,786,812]" pageId="6" pageNumber="67" value="2009-06-10" valueMax="2009-06-24" valueMin="2009-06-10">1024.VI.2009</date>
; 1 Ψ(
<collectionCode id="ED33AE06FFAB5979FBB298783381EF64" box="[1110,1198,786,812]" pageId="6" pageNumber="67">CEUA</collectionCode>
): same locality,
<date id="FF9C1003FFAB5979FA8B98783606EF1A" pageId="6" pageNumber="67" value="2008-07-01" valueMax="2008-07-14" valueMin="2008-07-01">01 14.VII.2008</date>
; 1 Ψ (
<collectionCode id="ED33AE06FFAB5979FE8A985236EAEF1A" box="[366,453,824,850]" pageId="6" pageNumber="67">CCBA</collectionCode>
): same locality,
<date id="FF9C1003FFAB5979FD6498523410EF1A" box="[640,831,824,850]" pageId="6" pageNumber="67" value="2009-07-08" valueMax="2009-07-22" valueMin="2009-07-08">0822.VII.2009</date>
; all of them A González-Moreno leg.; Malaise trap. Dry mounted.
</paragraph>
</subSubSection>
</treatment>
</document>