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<document id="09009030E3B1197E43778BF158503DA1" ID-CLB-Dataset="7001" ID-DOI="10.11646/zootaxa.4965.3.1" ID-GBIF-Dataset="6cadad22-abf2-4abe-9165-156d5c63ece3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4752531" ID-ZooBank="94FB89CF-2083-4FAC-AE60-B8CCF1D5FE8E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619690781738" checkinUser="plazi" docAuthor="Bennett, Robb, Copley, Claudia &amp; Copley, Darren" docDate="2021" docId="038B87C31901BE2CFF3A6CE2FA589E3E" docLanguage="en" docName="zootaxa.4965.3.1.pdf" docOrigin="Zootaxa 4965 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cybaeus consocius Chamberlin &amp; Ivie 1932" docType="key" docVersion="7" lastPageNumber="404" masterDocId="FFB2FFBB1903BE2FFFAD6F15FFC49C5B" masterDocTitle="Cybaeus (Araneae: Cybaeidae): the consocius species group of the Californian clade" masterLastPageNumber="436" masterPageNumber="401" pageNumber="403" updateTime="1698937978772" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="80AABB9058DBC01E164EA588E06A45DA">Cybaeus (Araneae: Cybaeidae): the consocius species group of the Californian clade</mods:title>
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<mods:namePart id="FE21A0C964C8C32103D3F2615DAC0F38">Bennett, Robb</mods:namePart>
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<mods:nameIdentifier id="3EABD280FBC691810ADB22AC4289519B" type="email">ennett@shaw.ca</mods:nameIdentifier>
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<mods:namePart id="4E291B59363FC1B66B9D7421588F6191">Copley, Claudia</mods:namePart>
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<mods:affiliation id="7FA864FFF249D46A7E0797329701495A">ccopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 8184 - 5819</mods:affiliation>
<mods:nameIdentifier id="52369C2C2AD893DEAA118F3D26A60D45" type="email">ccopley@royalbcmuseum.bc.ca</mods:nameIdentifier>
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<mods:namePart id="22C7ADD82B5FDD2D16392CDC34260CFC">Copley, Darren</mods:namePart>
<mods:nameIdentifier id="B3EEFADA85061F531B131144CE5F0A53" type="ORCID">0000-0002-1944-4272</mods:nameIdentifier>
<mods:affiliation id="0A23ACDF4F5CBC36FE8EF4DEB4963147">dcopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 1944 - 4272</mods:affiliation>
<mods:nameIdentifier id="FAA7E2829924895FE85215F90848F91B" type="email">dcopley@royalbcmuseum.bc.ca</mods:nameIdentifier>
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<treatment id="038B87C31901BE2CFF3A6CE2FA589E3E" ID-DOI="http://doi.org/10.5281/zenodo.4727166" ID-GBIF-Taxon="181983468" ID-Zenodo-Dep="4727166" LSID="urn:lsid:plazi:treatment:038B87C31901BE2CFF3A6CE2FA589E3E" httpUri="http://treatment.plazi.org/id/038B87C31901BE2CFF3A6CE2FA589E3E" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403">
<subSubSection id="C338655E1901BE2DFF3A6CE2FCC5984A" box="[151,769,1014,1041]" pageId="2" pageNumber="403" type="nomenclature">
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<heading id="D0D581B91901BE2DFF3A6CE2FCC5984A" bold="true" box="[151,769,1014,1041]" fontSize="11" level="1" pageId="2" pageNumber="403" reason="1">
<emphasis id="B956EAC71901BE2DFF3A6CE2FCC5984A" bold="true" box="[151,769,1014,1041]" pageId="2" pageNumber="403">
Key to the species of the
<taxonomicName id="4C224D561901BE2DFE666CE3FD74984A" ID-CoL="6C55P" authorityName="Chamberlin &amp; Ivie" authorityYear="1932" box="[459,688,1014,1041]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="consocius">
<emphasis id="B956EAC71901BE2DFE666CE3FD74984A" bold="true" box="[459,688,1014,1041]" italics="true" pageId="2" pageNumber="403">Cybaeus consocius</emphasis>
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group
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</paragraph>
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<keyStep id="30D62E701901BE2DFF3A6B29FA58995A" pageId="2" pageNumber="403">
<paragraph id="8B9D36D51901BE2DFF3A6B29FA5898D0" blockId="2.[151,1437,1084,2015]" pageId="2" pageNumber="403">
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1. Proximal arm of tegular apophysis in ventral view with tip forked (
<figureCitation id="13192A501901BE2DFCC66B29FC61980A" box="[875,933,1084,1106]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1863,1887]" captionTargetBox="[252,1338,241,1831]" captionTargetId="figure-62@4.[246,1341,234,1837]" captionTargetPageId="4" captionText="FIGURES 1–4. Cybaeus spp., males, left palpus (1 C. simplex from Ashland watershed, Oregon; 2–4 C. consocius from Muir Woods, California). 1 Patella, tibia, and genital bulb, ventral. 2 Patella and tibia, dorsal. 3–4 Genital bulb (3 ventral, 4 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752533" httpUri="https://zenodo.org/record/4752533/files/figure.png" pageId="2" pageNumber="403">Fig. 3</figureCitation>
,
<figureCitation id="13192A501901BE2DFC1D6B29FC0F9809" box="[944,971,1084,1106]" captionStart="FIGURES 20–22" captionStartId="10.[151,264,714,738]" captionTargetBox="[151,1436,181,690]" captionTargetId="figure-27@10.[151,1436,181,690]" captionTargetPageId="10" captionText="FIGURES 20–22. Cybaeus spp., males from California (20 C. consocius from Muir Woods, 21 C. hesper from Skyline Boulevard at King’s Mountain, 22 C. hummeli spec. nov. from Skyline Boulevard at King’s Mountain). 20, 22 Left palpus, proximal tip of tegular apophysis, ventral. 21 Right palpus, proximal tip of tegular apophysis, ventral. Unlabelled arrows indicate diagnostic characters (20 right angle forked tip, 21 inconspicuous bifid tip, 23 small acuminate tip and central longitudinal concavity)." figureDoi="http://doi.org/10.5281/zenodo.4752560" httpUri="https://zenodo.org/record/4752560/files/figure.png" pageId="2" pageNumber="403">20</figureCitation>
), bifid (
<figureCitation id="13192A501901BE2DFBBA6B29FBA5980A" box="[1047,1121,1084,1106]" captionStart="FIGURES 10–12" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[202,1388,192,1918]" captionTargetId="figure-17@7.[200,1388,179,1918]" captionTargetPageId="7" captionText="FIGURES 10–12. Cybaeus hesper, male from Skyline Boulevard at King’s Mountain, California, right palpus. 10 Patella and tibia, dorsal. 11–12 Genital bulb (11 ventral, 12 retrolateral). PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752546" httpUri="https://zenodo.org/record/4752546/files/figure.png" pageId="2" pageNumber="403">Figs 11</figureCitation>
,
<figureCitation id="13192A501901BE2DFBC16B29FB429809" box="[1132,1158,1084,1106]" captionStart="FIGURES 20–22" captionStartId="10.[151,264,714,738]" captionTargetBox="[151,1436,181,690]" captionTargetId="figure-27@10.[151,1436,181,690]" captionTargetPageId="10" captionText="FIGURES 20–22. Cybaeus spp., males from California (20 C. consocius from Muir Woods, 21 C. hesper from Skyline Boulevard at King’s Mountain, 22 C. hummeli spec. nov. from Skyline Boulevard at King’s Mountain). 20, 22 Left palpus, proximal tip of tegular apophysis, ventral. 21 Right palpus, proximal tip of tegular apophysis, ventral. Unlabelled arrows indicate diagnostic characters (20 right angle forked tip, 21 inconspicuous bifid tip, 23 small acuminate tip and central longitudinal concavity)." figureDoi="http://doi.org/10.5281/zenodo.4752560" httpUri="https://zenodo.org/record/4752560/files/figure.png" pageId="2" pageNumber="403">21</figureCitation>
), or a broad trough (
<figureCitation id="13192A501901BE2DFAE06B29FA5C980A" box="[1357,1432,1084,1106]" captionStart="FIGURES 18–19" captionStartId="9.[151,264,1430,1454]" captionTargetBox="[154,1436,573,1404]" captionTargetId="figure-158@9.[151,1436,562,1407]" captionTargetPageId="9" captionText="FIGURES 18–19. Cybaeus hummeli spec. nov., male from Skyline Boulevard at King’s Mountain, California, left palpus, genital bulb (18 ventral, 19 retrolateral). E—embolus, PA—proximal arm of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752554" httpUri="https://zenodo.org/record/4752554/files/figure.png" pageId="2" pageNumber="403">Figs 18</figureCitation>
,
<figureCitation id="13192A501901BE2DFF4F6B4CFF399834" box="[226,253,1113,1135]" captionStart="FIGURES 20–22" captionStartId="10.[151,264,714,738]" captionTargetBox="[151,1436,181,690]" captionTargetId="figure-27@10.[151,1436,181,690]" captionTargetPageId="10" captionText="FIGURES 20–22. Cybaeus spp., males from California (20 C. consocius from Muir Woods, 21 C. hesper from Skyline Boulevard at King’s Mountain, 22 C. hummeli spec. nov. from Skyline Boulevard at King’s Mountain). 20, 22 Left palpus, proximal tip of tegular apophysis, ventral. 21 Right palpus, proximal tip of tegular apophysis, ventral. Unlabelled arrows indicate diagnostic characters (20 right angle forked tip, 21 inconspicuous bifid tip, 23 small acuminate tip and central longitudinal concavity)." figureDoi="http://doi.org/10.5281/zenodo.4752560" httpUri="https://zenodo.org/record/4752560/files/figure.png" pageId="2" pageNumber="403">22</figureCitation>
); in retrolateral view with smoothly, simply curved dorsal margin (
<figureCitation id="13192A501901BE2DFC296B4CFC079835" box="[900,963,1113,1135]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1863,1887]" captionTargetBox="[252,1338,241,1831]" captionTargetId="figure-62@4.[246,1341,234,1837]" captionTargetPageId="4" captionText="FIGURES 1–4. Cybaeus spp., males, left palpus (1 C. simplex from Ashland watershed, Oregon; 2–4 C. consocius from Muir Woods, California). 1 Patella, tibia, and genital bulb, ventral. 2 Patella and tibia, dorsal. 3–4 Genital bulb (3 ventral, 4 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752533" httpUri="https://zenodo.org/record/4752533/files/figure.png" pageId="2" pageNumber="403">Figs 4</figureCitation>
,
<figureCitation id="13192A501901BE2DFC636B4CFC2C9835" box="[974,1000,1113,1134]" captionStart="FIGURES 10–12" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[202,1388,192,1918]" captionTargetId="figure-17@7.[200,1388,179,1918]" captionTargetPageId="7" captionText="FIGURES 10–12. Cybaeus hesper, male from Skyline Boulevard at King’s Mountain, California, right palpus. 10 Patella and tibia, dorsal. 11–12 Genital bulb (11 ventral, 12 retrolateral). PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752546" httpUri="https://zenodo.org/record/4752546/files/figure.png" pageId="2" pageNumber="403">12</figureCitation>
,
<figureCitation id="13192A501901BE2DFC5F6B4CFBC99834" box="[1010,1037,1113,1135]" captionStart="FIGURES 18–19" captionStartId="9.[151,264,1430,1454]" captionTargetBox="[154,1436,573,1404]" captionTargetId="figure-158@9.[151,1436,562,1407]" captionTargetPageId="9" captionText="FIGURES 18–19. Cybaeus hummeli spec. nov., male from Skyline Boulevard at King’s Mountain, California, left palpus, genital bulb (18 ventral, 19 retrolateral). E—embolus, PA—proximal arm of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752554" httpUri="https://zenodo.org/record/4752554/files/figure.png" pageId="2" pageNumber="403">19</figureCitation>
). Proximal loops of copulatory ducts distinctly U-shaped in dorsal view of vulva (
<figureCitation id="13192A501901BE2DFDDC6B63FD7498D7" box="[625,688,1142,1164]" captionStart="FIGURES 5–9" captionStartId="5.[151,264,1417,1441]" captionTargetBox="[156,1436,180,1384]" captionTargetId="figure-17@5.[151,1436,179,1391]" captionTargetPageId="5" captionText="FIGURES 5–9. Cybaeus consocius, females from California, copulatory organ (5–7 holotype, 8 from Mount Tamalpais, 9 from near Oakville). 5 Epigynum, ventral. 6–8 Vulva (6, 8 ventral, 7 dorsal). 9 Atria, ventral. BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate anterior margin of atrium." figureDoi="http://doi.org/10.5281/zenodo.4752539" httpUri="https://zenodo.org/record/4752539/files/figure.png" pageId="2" pageNumber="403">Figs 7</figureCitation>
,
<figureCitation id="13192A501901BE2DFD176B63FD1098D7" box="[698,724,1142,1164]" captionStart="FIGURES 13–17" captionStartId="8.[151,264,1664,1688]" captionTargetBox="[164,1435,184,1637]" captionTargetId="figure-27@8.[151,1436,181,1640]" captionTargetPageId="8" captionText="FIGURES 13–17. Cybaeus spp., females, copulatory organ (13–15 C. hesper, syntype of C. minor; 16–17 C. consocius, holotype of C. marinensis). 13, 16 Epigynum, ventral; 14–15, 17 Vulva (14, 17 ventral, 15 dorsal). CD—copulatory duct. Unlabelled arrows indicate anterior margin of atrium." figureDoi="http://doi.org/10.5281/zenodo.4752550" httpUri="https://zenodo.org/record/4752550/files/figure.png" pageId="2" pageNumber="403">15</figureCitation>
,
<figureCitation id="13192A501901BE2DFD736B63FD3398D7" box="[734,759,1142,1164]" captionStart="FIGURES 23–26" captionStartId="10.[151,264,1921,1945]" captionTargetBox="[151,1436,906,1894]" captionTargetId="figure-111@10.[151,1436,906,1898]" captionTargetPageId="10" captionText="FIGURES 23–26. Cybaeus hummeli spec. nov., females from California, copulatory organ (23–25 from Miramar, 26 from San Francisco). 23, 26 Epigynum, ventral. 24–25 Vulva (24 ventral, 25 dorsal). CD—copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.4752564" httpUri="https://zenodo.org/record/4752564/files/figure.png" pageId="2" pageNumber="403">25</figureCitation>
)...................................................... 2
</keyLead>
</paragraph>
<paragraph id="8B9D36D51901BE2DFF3A6B81FA58995A" blockId="2.[151,1437,1084,2015]" pageId="2" pageNumber="403">
<keyLead id="30D395E01901BE2DFF3A6B81FA58995A" pageId="2" pageNumber="403">
- Proximal arm of tegular apophysis in ventral view with tip acuminate and/or spirally twisted (
<figureCitation id="13192A501901BE2DFBF36B81FB6C98F2" box="[1118,1192,1172,1194]" captionStart="FIGURES 27–29" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[162,1415,184,1917]" captionTargetId="figure-27@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURES 27–29. Cybaeus opulentus spec. nov., male from Riverton, California, left palpus. 27 Patella and tibia, dorsal. 28–29 Genital bulb (28 ventral, 29 retrolateral). PA—proximal arm of tegular apophysis. Unlabelled arrows indicate compound curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752568" httpUri="https://zenodo.org/record/4752568/files/figure.png" pageId="2" pageNumber="403">Figs 28</figureCitation>
,
<figureCitation id="13192A501901BE2DFB1C6B86FB0F98F2" box="[1201,1227,1171,1193]" captionStart="FIGURES 33–36" captionStartId="14.[151,264,1933,1957]" captionTargetBox="[207,1372,273,1901]" captionTargetId="figure-71@14.[207,1381,273,1909]" captionTargetPageId="14" captionText="FIGURES 33–36. Cybaeus pan spec. nov., male from Calpine, California, left palpus. 33–34 Patella and tibia (33 retrolateral, 34 dorsal). 35–36 Genital bulb (35 ventral, 36 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752576" httpUri="https://zenodo.org/record/4752576/files/figure.png" pageId="2" pageNumber="403">35</figureCitation>
,
<figureCitation id="13192A501901BE2DFB786B81FB2B98F2" box="[1237,1263,1172,1193]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1806,1830]" captionTargetBox="[151,1419,155,1775]" captionTargetId="figure-17@17.[151,1436,151,1783]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus penedentatus, male from Holy City, California, left palpus. 42–43 Patella and tibia (42 retrolateral, 43 dorsal). 44–45 Genital bulb (44 ventral, 45 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate ventral keel of proximal arm of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752582" httpUri="https://zenodo.org/record/4752582/files/figure.png" pageId="2" pageNumber="403">44</figureCitation>
,
<figureCitation id="13192A501901BE2DFB556B86FAF298F2" box="[1272,1334,1171,1193]" captionStart="FIGURES 55–58" captionStartId="20.[151,264,1895,1919]" captionTargetBox="[177,1409,381,1872]" captionTargetId="figure-139@20.[177,1411,381,1872]" captionTargetPageId="20" captionText="FIGURES 55–58. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis (55 C. opulentus spec. nov. from Riverton, 56 C. penedentatus from Holy City, 57 C. vulpinus from Columbia, 58 C. pan from Calpine) (55–56 ventral, 57–58 prolateral). Arrows indicate diagnostic characters (55 slightly twisted tip and prominent basal protrusion; 56 acuminate slightly twisted tip and longitudinal ventral keel; 57 smoothly curved, twisted tip; 58 triangular keel dorsally on tip)." figureDoi="http://doi.org/10.5281/zenodo.4752596" httpUri="https://zenodo.org/record/4752596/files/figure.png" pageId="2" pageNumber="403">55–56</figureCitation>
,
<figureCitation id="13192A501901BE2DFA926B81FA9D98F1" box="[1343,1369,1172,1194]" captionStart="FIGURES 59–63" captionStartId="21.[151,264,1872,1896]" captionTargetBox="[176,1407,183,1835]" captionTargetId="figure-17@21.[176,1410,181,1849]" captionTargetPageId="21" captionText="FIGURES 59–63. Cybaeus simplex, males from Oregon, left palpus (59 from Ashland watershed, 60–63 from H.J. Andrews Experimental Forest). 59 Tibia, retrolateral. 60 Patella and tibia, dorsal. 61–62 Genital bulb (61 ventral, 62 retrolateral). 63 Tip of proximal arm of tegular apophysis, ventral. Arrows indicate diagnostic characters (59 distal extension of retrolateral tibial apophysis, 61 strongly twisted tip and prominent basal projection of proximal arm of tegular apophysis, 62 compound curve of dorsal margin of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.4752600" httpUri="https://zenodo.org/record/4752600/files/figure.png" pageId="2" pageNumber="403">61</figureCitation>
,
<figureCitation id="13192A501901BE2DFACE6B81FAB898F1" box="[1379,1404,1172,1194]" captionStart="FIGURES 80–83" captionStartId="27.[151,264,1843,1867]" captionTargetBox="[151,1420,199,1819]" captionTargetId="figure-17@27.[151,1436,181,1819]" captionTargetPageId="27" captionText="FIGURES 80–83. Cybaeus vulpinus, male from Columbia, California, left palpus. 80–81 Patella and tibia (80 retrolateral, 81 dorsal). 82–83 Genital bulb (82 ventral, 83 retrolateral). Arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752620" httpUri="https://zenodo.org/record/4752620/files/figure.png" pageId="2" pageNumber="403">82</figureCitation>
) or blunt with long, narrow ventral trough (
<figureCitation id="13192A501901BE2DFDF26BA4FD6E989D" box="[607,682,1201,1223]" captionStart="FIGURES 68–71" captionStartId="24.[151,264,1762,1786]" captionTargetBox="[151,1418,186,1738]" captionTargetId="figure-27@24.[151,1436,181,1738]" captionTargetPageId="24" captionText="FIGURES 68–71. Cybaeus ubicki spec. nov., males from California, left palpus (68–69, 71 from Novato, 70 from Anchor Bay). 68–69 Patella and tibia (68 retrolateral, 69 dorsal). 70–71 Genital bulb (70 ventral, 71 retrolateral). E—embolus, PA—proximal arm of tegular apophysis. RTA—retrolateral tibial apophysis. Unlabelled arrows indicate dorsal tibial ridge and dorsal prominence of patellar apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752607" httpUri="https://zenodo.org/record/4752607/files/figure.png" pageId="2" pageNumber="403">Figs 70</figureCitation>
,
<figureCitation id="13192A501901BE2DFD196BA4FD0B989C" box="[692,719,1201,1223]" captionStart="FIGURES 72–73" captionStartId="25.[151,264,977,1001]" captionTargetBox="[151,1436,181,953]" captionTargetId="figure-17@25.[151,1436,181,953]" captionTargetPageId="25" captionText="FIGURES 72–73. Cybaeus ubicki spec. nov., male from Novato, California, left palpus. 72 Patellar apophysis, ventral. 73 Proximal arm of tegular apophysis, ventral. Arrows indicate diagnostic characters (72 dorsal prominence of patellar apophysis, 73 ventral longitudinal groove of proximal arm of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.4752611" httpUri="https://zenodo.org/record/4752611/files/figure.png" pageId="2" pageNumber="403">73</figureCitation>
); in retrolateral view, dorsal margin a simple curve (
<figureCitation id="13192A501901BE2DFB686BA4FAD4989C" box="[1221,1296,1201,1223]" captionStart="FIGURES 33–36" captionStartId="14.[151,264,1933,1957]" captionTargetBox="[207,1372,273,1901]" captionTargetId="figure-71@14.[207,1381,273,1909]" captionTargetPageId="14" captionText="FIGURES 33–36. Cybaeus pan spec. nov., male from Calpine, California, left palpus. 33–34 Patella and tibia (33 retrolateral, 34 dorsal). 35–36 Genital bulb (35 ventral, 36 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752576" httpUri="https://zenodo.org/record/4752576/files/figure.png" pageId="2" pageNumber="403">Figs 36</figureCitation>
,
<figureCitation id="13192A501901BE2DFAB76BA4FAF7989C" box="[1306,1331,1201,1223]" captionStart="FIGURES 80–83" captionStartId="27.[151,264,1843,1867]" captionTargetBox="[151,1420,199,1819]" captionTargetId="figure-17@27.[151,1436,181,1819]" captionTargetPageId="27" captionText="FIGURES 80–83. Cybaeus vulpinus, male from Columbia, California, left palpus. 80–81 Patella and tibia (80 retrolateral, 81 dorsal). 82–83 Genital bulb (82 ventral, 83 retrolateral). Arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752620" httpUri="https://zenodo.org/record/4752620/files/figure.png" pageId="2" pageNumber="403">83</figureCitation>
) or a more complex compound curve (
<figureCitation id="13192A501901BE2DFE486BDBFDEB98B8" box="[485,559,1230,1252]" captionStart="FIGURES 27–29" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[162,1415,184,1917]" captionTargetId="figure-27@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURES 27–29. Cybaeus opulentus spec. nov., male from Riverton, California, left palpus. 27 Patella and tibia, dorsal. 28–29 Genital bulb (28 ventral, 29 retrolateral). PA—proximal arm of tegular apophysis. Unlabelled arrows indicate compound curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752568" httpUri="https://zenodo.org/record/4752568/files/figure.png" pageId="2" pageNumber="403">Figs 29</figureCitation>
,
<figureCitation id="13192A501901BE2DFD956BDBFD9698BF" box="[568,594,1230,1252]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1806,1830]" captionTargetBox="[151,1419,155,1775]" captionTargetId="figure-17@17.[151,1436,151,1783]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus penedentatus, male from Holy City, California, left palpus. 42–43 Patella and tibia (42 retrolateral, 43 dorsal). 44–45 Genital bulb (44 ventral, 45 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate ventral keel of proximal arm of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752582" httpUri="https://zenodo.org/record/4752582/files/figure.png" pageId="2" pageNumber="403">45</figureCitation>
,
<figureCitation id="13192A501901BE2DFDF66BDBFDB298BF" box="[603,630,1230,1252]" captionStart="FIGURES 59–63" captionStartId="21.[151,264,1872,1896]" captionTargetBox="[176,1407,183,1835]" captionTargetId="figure-17@21.[176,1410,181,1849]" captionTargetPageId="21" captionText="FIGURES 59–63. Cybaeus simplex, males from Oregon, left palpus (59 from Ashland watershed, 60–63 from H.J. Andrews Experimental Forest). 59 Tibia, retrolateral. 60 Patella and tibia, dorsal. 61–62 Genital bulb (61 ventral, 62 retrolateral). 63 Tip of proximal arm of tegular apophysis, ventral. Arrows indicate diagnostic characters (59 distal extension of retrolateral tibial apophysis, 61 strongly twisted tip and prominent basal projection of proximal arm of tegular apophysis, 62 compound curve of dorsal margin of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.4752600" httpUri="https://zenodo.org/record/4752600/files/figure.png" pageId="2" pageNumber="403">62</figureCitation>
). Proximal loops of copulatory ducts not distinctly U-shaped in dorsal view (
<figureCitation id="13192A501901BE2DFAE36BDBFA5C98B8" box="[1358,1432,1230,1252]" captionStart="FIGURES 30–32" captionStartId="13.[151,264,1111,1135]" captionTargetBox="[156,1436,186,1087]" captionTargetId="figure-17@13.[151,1436,181,1087]" captionTargetPageId="13" captionText="FIGURES 30–32. Cybaeus opulentus spec. nov., female from Riverton, California, copulatory organ. 30 Epigynum, ventral. 31–32 Vulva (31 ventral, 32 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4752572" httpUri="https://zenodo.org/record/4752572/files/figure.png" pageId="2" pageNumber="403">Figs 32</figureCitation>
,
<figureCitation id="13192A501901BE2DFF4F6BF9FF38995A" box="[226,252,1260,1281]" captionStart="FIGURES 37–41" captionStartId="15.[151,264,1667,1691]" captionTargetBox="[151,1431,185,1639]" captionTargetId="figure-17@15.[151,1436,181,1643]" captionTargetPageId="15" captionText="FIGURES 37–41. Cybaeus spp., females from California, copulatory organ (37–38 C. pan spec. nov. from near Calpine, 39 C. pan spec. nov. from Sierra City, 40–41 C. vulpinus from Big Trees State Park). 37 Epigynum, ventral. 38–41 Vulva (38, 40 ventral; 39, 41 dorsal). CD—copulatory duct, SS—spermathecal stalk. Unlabelled arrow indicates epigynal notch." figureDoi="http://doi.org/10.5281/zenodo.4752580" httpUri="https://zenodo.org/record/4752580/files/figure.png" pageId="2" pageNumber="403">41</figureCitation>
,
<figureCitation id="13192A501901BE2DFEAB6BF9FEE4995A" box="[262,288,1260,1281]" captionStart="FIGURES 46–51" captionStartId="18.[151,264,1787,1811]" captionTargetBox="[151,1433,181,1759]" captionTargetId="figure-27@18.[151,1437,181,1763]" captionTargetPageId="18" captionText="FIGURES 46–51. Cybaeus penedentatus, females from California, copulatory organ (46–48 from Ben Lomond, 49 from Pescadero Creek, 50–51 from Mount Madonna). 46 Epigynum, ventral. 47–51 Vulva (47, 49–50 ventral; 48, 51 dorsal). AT—atrium." figureDoi="http://doi.org/10.5281/zenodo.4752586" httpUri="https://zenodo.org/record/4752586/files/figure.png" pageId="2" pageNumber="403">48</figureCitation>
,
<figureCitation id="13192A501901BE2DFE876BF9FE809959" box="[298,324,1260,1282]" captionStart="FIGURES 64–67" captionStartId="22.[151,264,1249,1273]" captionTargetBox="[153,1436,184,1225]" captionTargetId="figure-27@22.[151,1436,181,1226]" captionTargetPageId="22" captionText="FIGURES 64–67. Cybaeus simplex, females from Cascadia, Oregon, copulatory organ. 64 Epigynum, ventral. 65–67 Vulva (65 ventral, 66–67 dorsal). CD—copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.4752604" httpUri="https://zenodo.org/record/4752604/files/figure.png" pageId="2" pageNumber="403">66</figureCitation>
,
<figureCitation id="13192A501901BE2DFEE36BF9FEA39959" box="[334,359,1260,1282]" captionStart="FIGURES 74–79" captionStartId="26.[151,264,1772,1796]" captionTargetBox="[162,1436,186,1747]" captionTargetId="figure-27@26.[151,1436,181,1748]" captionTargetPageId="26" captionText="FIGURES 74–79. Cybaeus ubicki spec. nov., females from California, copulatory organ (74–76 from Point Reyes Station, 77– 79 from Anchor Bay). 74, 77 Epigynum, ventral. 75–76, 78–79 Vulva (75, 78, ventral, 76, 79 dorsal). CD—copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.4752616" httpUri="https://zenodo.org/record/4752616/files/figure.png" pageId="2" pageNumber="403">79</figureCitation>
)........................................................................................ 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D62E701901BE2DFF3A6A1CFA58992D" pageId="2" pageNumber="403">
<paragraph id="8B9D36D51901BE2DFF3A6A1CFA589960" blockId="2.[151,1437,1084,2015]" pageId="2" pageNumber="403">
<keyLead id="30D395E01901BE2DFF3A6A1CFA589960" pageId="2" pageNumber="403">
2(1). Proximal arm of tegular apophysis in ventral view with broad, ventral trough terminating in a small point (
<figureCitation id="13192A501901BE2DFB746A1CFAE79945" box="[1241,1315,1289,1311]" captionStart="FIGURES 18–19" captionStartId="9.[151,264,1430,1454]" captionTargetBox="[154,1436,573,1404]" captionTargetId="figure-158@9.[151,1436,562,1407]" captionTargetPageId="9" captionText="FIGURES 18–19. Cybaeus hummeli spec. nov., male from Skyline Boulevard at King’s Mountain, California, left palpus, genital bulb (18 ventral, 19 retrolateral). E—embolus, PA—proximal arm of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752554" httpUri="https://zenodo.org/record/4752554/files/figure.png" pageId="2" pageNumber="403">Figs 18</figureCitation>
,
<figureCitation id="13192A501901BE2DFA806A1CFA8C9944" box="[1325,1352,1289,1311]" captionStart="FIGURES 20–22" captionStartId="10.[151,264,714,738]" captionTargetBox="[151,1436,181,690]" captionTargetId="figure-27@10.[151,1436,181,690]" captionTargetPageId="10" captionText="FIGURES 20–22. Cybaeus spp., males from California (20 C. consocius from Muir Woods, 21 C. hesper from Skyline Boulevard at King’s Mountain, 22 C. hummeli spec. nov. from Skyline Boulevard at King’s Mountain). 20, 22 Left palpus, proximal tip of tegular apophysis, ventral. 21 Right palpus, proximal tip of tegular apophysis, ventral. Unlabelled arrows indicate diagnostic characters (20 right angle forked tip, 21 inconspicuous bifid tip, 23 small acuminate tip and central longitudinal concavity)." figureDoi="http://doi.org/10.5281/zenodo.4752560" httpUri="https://zenodo.org/record/4752560/files/figure.png" pageId="2" pageNumber="403">22</figureCitation>
). Atrium divided (
<figureCitation id="13192A501901BE2DFE9A6A33FE609967" box="[311,420,1318,1340]" captionStart="FIGURES 23–26" captionStartId="10.[151,264,1921,1945]" captionTargetBox="[151,1436,906,1894]" captionTargetId="figure-111@10.[151,1436,906,1898]" captionTargetPageId="10" captionText="FIGURES 23–26. Cybaeus hummeli spec. nov., females from California, copulatory organ (23–25 from Miramar, 26 from San Francisco). 23, 26 Epigynum, ventral. 24–25 Vulva (24 ventral, 25 dorsal). CD—copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.4752564" httpUri="https://zenodo.org/record/4752564/files/figure.png" pageId="2" pageNumber="403">Figs 23, 26</figureCitation>
).............................................................
<taxonomicName id="4C224D561901BE2DFB3C6A33FAF19967" ID-CoL="98YZV" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1169,1333,1318,1340]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="hummeli" status="sp. nov.">
<emphasis id="B956EAC71901BE2DFB3C6A33FB209960" box="[1169,1252,1318,1339]" italics="true" pageId="2" pageNumber="403">hummeli</emphasis>
Bennett
</taxonomicName>
<emphasis id="B956EAC71901BE2DFA976A33FA589960" bold="true" box="[1338,1436,1318,1339]" pageId="2" pageNumber="403">
<taxonomicNameLabel id="A26557BC1901BE2DFA976A33FA589960" box="[1338,1436,1318,1339]" pageId="2" pageNumber="403" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9D36D51901BE2DFF3A6A51FA58992D" blockId="2.[151,1437,1084,2015]" pageId="2" pageNumber="403">
<keyLead id="30D395E01901BE2DFF3A6A51FA58992D" pageId="2" pageNumber="403">
- Proximal arm of tegular apophysis in ventral view conspicuously forked (
<figureCitation id="13192A501901BE2DFC1F6A51FC289901" box="[946,1004,1348,1370]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1863,1887]" captionTargetBox="[252,1338,241,1831]" captionTargetId="figure-62@4.[246,1341,234,1837]" captionTargetPageId="4" captionText="FIGURES 1–4. Cybaeus spp., males, left palpus (1 C. simplex from Ashland watershed, Oregon; 2–4 C. consocius from Muir Woods, California). 1 Patella, tibia, and genital bulb, ventral. 2 Patella and tibia, dorsal. 3–4 Genital bulb (3 ventral, 4 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752533" httpUri="https://zenodo.org/record/4752533/files/figure.png" pageId="2" pageNumber="403">Fig. 3</figureCitation>
) or with inconspicuously bifid tip (
<figureCitation id="13192A501901BE2DFAE46A51FA549901" box="[1353,1424,1348,1370]" captionStart="FIGURES 10–12" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[202,1388,192,1918]" captionTargetId="figure-17@7.[200,1388,179,1918]" captionTargetPageId="7" captionText="FIGURES 10–12. Cybaeus hesper, male from Skyline Boulevard at King’s Mountain, California, right palpus. 10 Patella and tibia, dorsal. 11–12 Genital bulb (11 ventral, 12 retrolateral). PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752546" httpUri="https://zenodo.org/record/4752546/files/figure.png" pageId="2" pageNumber="403">Fig. 11</figureCitation>
). Atrium undivided, inverted U-shaped (
<figureCitation id="13192A501901BE2DFDFA6A74FD69992D" box="[599,685,1377,1399]" captionStart="FIGURES 5–9" captionStartId="5.[151,264,1417,1441]" captionTargetBox="[156,1436,180,1384]" captionTargetId="figure-17@5.[151,1436,179,1391]" captionTargetPageId="5" captionText="FIGURES 5–9. Cybaeus consocius, females from California, copulatory organ (5–7 holotype, 8 from Mount Tamalpais, 9 from near Oakville). 5 Epigynum, ventral. 6–8 Vulva (6, 8 ventral, 7 dorsal). 9 Atria, ventral. BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate anterior margin of atrium." figureDoi="http://doi.org/10.5281/zenodo.4752539" httpUri="https://zenodo.org/record/4752539/files/figure.png" pageId="2" pageNumber="403">Figs 5, 9</figureCitation>
,
<figureCitation id="13192A501901BE2DFD1A6A74FD15992C" box="[695,721,1377,1399]" captionStart="FIGURES 13–17" captionStartId="8.[151,264,1664,1688]" captionTargetBox="[164,1435,184,1637]" captionTargetId="figure-27@8.[151,1436,181,1640]" captionTargetPageId="8" captionText="FIGURES 13–17. Cybaeus spp., females, copulatory organ (13–15 C. hesper, syntype of C. minor; 16–17 C. consocius, holotype of C. marinensis). 13, 16 Epigynum, ventral; 14–15, 17 Vulva (14, 17 ventral, 15 dorsal). CD—copulatory duct. Unlabelled arrows indicate anterior margin of atrium." figureDoi="http://doi.org/10.5281/zenodo.4752550" httpUri="https://zenodo.org/record/4752550/files/figure.png" pageId="2" pageNumber="403">13</figureCitation>
).......................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D62E701901BE2DFF3A6A6BFA5F9A51" pageId="2" pageNumber="403">
<paragraph id="8B9D36D51901BE2DFF3A6A6BFA589994" blockId="2.[151,1437,1084,2015]" pageId="2" pageNumber="403">
<keyLead id="30D395E01901BE2DFF3A6A6BFA589994" pageId="2" pageNumber="403">
3(2). Proximal arm of tegular apophysis with inconspicuously bifid tip (
<figureCitation id="13192A501901BE2DFCFB6A6BFC5F99CF" box="[854,923,1406,1428]" captionStart="FIGURES 10–12" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[202,1388,192,1918]" captionTargetId="figure-17@7.[200,1388,179,1918]" captionTargetPageId="7" captionText="FIGURES 10–12. Cybaeus hesper, male from Skyline Boulevard at King’s Mountain, California, right palpus. 10 Patella and tibia, dorsal. 11–12 Genital bulb (11 ventral, 12 retrolateral). PA—proximal arm of tegular apophysis, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752546" httpUri="https://zenodo.org/record/4752546/files/figure.png" pageId="2" pageNumber="403">Fig. 11</figureCitation>
). Atrium convex medially, concave laterally (
<figureCitation id="13192A501901BE2DFAE76A6BFA5499CF" box="[1354,1424,1406,1428]" captionStart="FIGURES 13–17" captionStartId="8.[151,264,1664,1688]" captionTargetBox="[164,1435,184,1637]" captionTargetId="figure-27@8.[151,1436,181,1640]" captionTargetPageId="8" captionText="FIGURES 13–17. Cybaeus spp., females, copulatory organ (13–15 C. hesper, syntype of C. minor; 16–17 C. consocius, holotype of C. marinensis). 13, 16 Epigynum, ventral; 14–15, 17 Vulva (14, 17 ventral, 15 dorsal). CD—copulatory duct. Unlabelled arrows indicate anterior margin of atrium." figureDoi="http://doi.org/10.5281/zenodo.4752550" httpUri="https://zenodo.org/record/4752550/files/figure.png" pageId="2" pageNumber="403">Fig. 13</figureCitation>
). Copulatory ducts attached to anterolateral corners of atrium and extending anteriorly from the atrium a distance equal to about 1/4 to 2/5 the distance from epigastric groove to anterior margin of atrium (
<figureCitation id="13192A501901BE2DFC186AACFBE69994" box="[949,1058,1465,1487]" captionStart="FIGURES 13–17" captionStartId="8.[151,264,1664,1688]" captionTargetBox="[164,1435,184,1637]" captionTargetId="figure-27@8.[151,1436,181,1640]" captionTargetPageId="8" captionText="FIGURES 13–17. Cybaeus spp., females, copulatory organ (13–15 C. hesper, syntype of C. minor; 16–17 C. consocius, holotype of C. marinensis). 13, 16 Epigynum, ventral; 14–15, 17 Vulva (14, 17 ventral, 15 dorsal). CD—copulatory duct. Unlabelled arrows indicate anterior margin of atrium." figureDoi="http://doi.org/10.5281/zenodo.4752550" httpUri="https://zenodo.org/record/4752550/files/figure.png" pageId="2" pageNumber="403">Figs 14–15</figureCitation>
).........
<taxonomicName id="4C224D561901BE2DFB0C6AACFA589994" ID-CoL="6BSVG" authority="Chamberlin &amp; Ivie" authorityName="Chamberlin &amp; Ivie" authorityYear="1932" box="[1185,1436,1465,1487]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="hesper">
<emphasis id="B956EAC71901BE2DFB0C6AACFB259995" box="[1185,1249,1465,1486]" italics="true" pageId="2" pageNumber="403">hesper</emphasis>
Chamberlin &amp; Ivie
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9D36D51901BE2DFF3A6AC2FA5F9A51" blockId="2.[151,1437,1084,2015]" pageId="2" pageNumber="403">
<keyLead id="30D395E01901BE2DFF3A6AC2FA5F9A51" pageId="2" pageNumber="403">
- Proximal arm of tegular apophysis with right-angled fork (
<figureCitation id="13192A501901BE2DFCBD6AC3FC8E99B7" box="[784,842,1494,1516]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1863,1887]" captionTargetBox="[252,1338,241,1831]" captionTargetId="figure-62@4.[246,1341,234,1837]" captionTargetPageId="4" captionText="FIGURES 1–4. Cybaeus spp., males, left palpus (1 C. simplex from Ashland watershed, Oregon; 2–4 C. consocius from Muir Woods, California). 1 Patella, tibia, and genital bulb, ventral. 2 Patella and tibia, dorsal. 3–4 Genital bulb (3 ventral, 4 retrolateral). DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, PS—peg setae, PTA—patellar apophysis, RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752533" httpUri="https://zenodo.org/record/4752533/files/figure.png" pageId="2" pageNumber="403">Fig. 3</figureCitation>
). Atrium concave (
<figureCitation id="13192A501901BE2DFBAD6AC3FB9299B0" box="[1024,1110,1494,1516]" captionStart="FIGURES 5–9" captionStartId="5.[151,264,1417,1441]" captionTargetBox="[156,1436,180,1384]" captionTargetId="figure-17@5.[151,1436,179,1391]" captionTargetPageId="5" captionText="FIGURES 5–9. Cybaeus consocius, females from California, copulatory organ (5–7 holotype, 8 from Mount Tamalpais, 9 from near Oakville). 5 Epigynum, ventral. 6–8 Vulva (6, 8 ventral, 7 dorsal). 9 Atria, ventral. BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate anterior margin of atrium." figureDoi="http://doi.org/10.5281/zenodo.4752539" httpUri="https://zenodo.org/record/4752539/files/figure.png" pageId="2" pageNumber="403">Figs 5, 9</figureCitation>
,
<figureCitation id="13192A501901BE2DFBF26AC3FBBE99B7" box="[1119,1146,1494,1516]" captionStart="FIGURES 13–17" captionStartId="8.[151,264,1664,1688]" captionTargetBox="[164,1435,184,1637]" captionTargetId="figure-27@8.[151,1436,181,1640]" captionTargetPageId="8" captionText="FIGURES 13–17. Cybaeus spp., females, copulatory organ (13–15 C. hesper, syntype of C. minor; 16–17 C. consocius, holotype of C. marinensis). 13, 16 Epigynum, ventral; 14–15, 17 Vulva (14, 17 ventral, 15 dorsal). CD—copulatory duct. Unlabelled arrows indicate anterior margin of atrium." figureDoi="http://doi.org/10.5281/zenodo.4752550" httpUri="https://zenodo.org/record/4752550/files/figure.png" pageId="2" pageNumber="403">16</figureCitation>
). Copulatory ducts attached to lateral margins of atrium (
<figureCitation id="13192A501901BE2DFE716AE1FDF79A51" box="[476,563,1524,1546]" captionStart="FIGURES 5–9" captionStartId="5.[151,264,1417,1441]" captionTargetBox="[156,1436,180,1384]" captionTargetId="figure-17@5.[151,1436,179,1391]" captionTargetPageId="5" captionText="FIGURES 5–9. Cybaeus consocius, females from California, copulatory organ (5–7 holotype, 8 from Mount Tamalpais, 9 from near Oakville). 5 Epigynum, ventral. 6–8 Vulva (6, 8 ventral, 7 dorsal). 9 Atria, ventral. BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate anterior margin of atrium." figureDoi="http://doi.org/10.5281/zenodo.4752539" httpUri="https://zenodo.org/record/4752539/files/figure.png" pageId="2" pageNumber="403">Figs 6–8</figureCitation>
,
<figureCitation id="13192A501901BE2DFD906AE1FD939A51" box="[573,599,1524,1546]" captionStart="FIGURES 13–17" captionStartId="8.[151,264,1664,1688]" captionTargetBox="[164,1435,184,1637]" captionTargetId="figure-27@8.[151,1436,181,1640]" captionTargetPageId="8" captionText="FIGURES 13–17. Cybaeus spp., females, copulatory organ (13–15 C. hesper, syntype of C. minor; 16–17 C. consocius, holotype of C. marinensis). 13, 16 Epigynum, ventral; 14–15, 17 Vulva (14, 17 ventral, 15 dorsal). CD—copulatory duct. Unlabelled arrows indicate anterior margin of atrium." figureDoi="http://doi.org/10.5281/zenodo.4752550" httpUri="https://zenodo.org/record/4752550/files/figure.png" pageId="2" pageNumber="403">17</figureCitation>
); extending little or no distance anteriorly from atrium…
<taxonomicName id="4C224D561901BE2DFB2C6AE1FA5F9A51" ID-CoL="6C55P" authority="Chamberlin &amp; Ivie" authorityName="Chamberlin &amp; Ivie" authorityYear="1932" box="[1153,1435,1524,1546]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="consocius">
<emphasis id="B956EAC71901BE2DFB2C6AE1FB249A52" box="[1153,1248,1524,1545]" italics="true" pageId="2" pageNumber="403">consocius</emphasis>
Chamberlin &amp; Ivie
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D62E701901BE2DFF3A6904FA589B4A" pageId="2" pageNumber="403">
<paragraph id="8B9D36D51901BE2DFF3A6904FA589AC0" blockId="2.[151,1437,1084,2015]" pageId="2" pageNumber="403">
<keyLead id="30D395E01901BE2DFF3A6904FA589AC0" pageId="2" pageNumber="403">
4(1). Patellar apophysis with small dorsobasal prominence bearing several peg setae (
<figureCitation id="13192A501901BE2DFC4F6904FB959A7C" box="[994,1105,1553,1575]" captionStart="FIGURES 68–71" captionStartId="24.[151,264,1762,1786]" captionTargetBox="[151,1418,186,1738]" captionTargetId="figure-27@24.[151,1436,181,1738]" captionTargetPageId="24" captionText="FIGURES 68–71. Cybaeus ubicki spec. nov., males from California, left palpus (68–69, 71 from Novato, 70 from Anchor Bay). 68–69 Patella and tibia (68 retrolateral, 69 dorsal). 70–71 Genital bulb (70 ventral, 71 retrolateral). E—embolus, PA—proximal arm of tegular apophysis. RTA—retrolateral tibial apophysis. Unlabelled arrows indicate dorsal tibial ridge and dorsal prominence of patellar apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752607" httpUri="https://zenodo.org/record/4752607/files/figure.png" pageId="2" pageNumber="403">Figs 68–69</figureCitation>
,
<figureCitation id="13192A501901BE2DFBF76904FBB19A7C" box="[1114,1141,1553,1575]" captionStart="FIGURES 72–73" captionStartId="25.[151,264,977,1001]" captionTargetBox="[151,1436,181,953]" captionTargetId="figure-17@25.[151,1436,181,953]" captionTargetPageId="25" captionText="FIGURES 72–73. Cybaeus ubicki spec. nov., male from Novato, California, left palpus. 72 Patellar apophysis, ventral. 73 Proximal arm of tegular apophysis, ventral. Arrows indicate diagnostic characters (72 dorsal prominence of patellar apophysis, 73 ventral longitudinal groove of proximal arm of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.4752611" httpUri="https://zenodo.org/record/4752611/files/figure.png" pageId="2" pageNumber="403">72</figureCitation>
). Tibia with small ridge dorsomedially above retrolateral tibial apophysis (
<figureCitation id="13192A501901BE2DFD27693BFD3D9A1F" box="[650,761,1582,1604]" captionStart="FIGURES 68–71" captionStartId="24.[151,264,1762,1786]" captionTargetBox="[151,1418,186,1738]" captionTargetId="figure-27@24.[151,1436,181,1738]" captionTargetPageId="24" captionText="FIGURES 68–71. Cybaeus ubicki spec. nov., males from California, left palpus (68–69, 71 from Novato, 70 from Anchor Bay). 68–69 Patella and tibia (68 retrolateral, 69 dorsal). 70–71 Genital bulb (70 ventral, 71 retrolateral). E—embolus, PA—proximal arm of tegular apophysis. RTA—retrolateral tibial apophysis. Unlabelled arrows indicate dorsal tibial ridge and dorsal prominence of patellar apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752607" httpUri="https://zenodo.org/record/4752607/files/figure.png" pageId="2" pageNumber="403">Figs 68–69</figureCitation>
). Medial portion of embolus flattened, ribbon-like (
<figureCitation id="13192A501901BE2DFB4C693BFA959A1F" box="[1249,1361,1582,1604]" captionStart="FIGURES 68–71" captionStartId="24.[151,264,1762,1786]" captionTargetBox="[151,1418,186,1738]" captionTargetId="figure-27@24.[151,1436,181,1738]" captionTargetPageId="24" captionText="FIGURES 68–71. Cybaeus ubicki spec. nov., males from California, left palpus (68–69, 71 from Novato, 70 from Anchor Bay). 68–69 Patella and tibia (68 retrolateral, 69 dorsal). 70–71 Genital bulb (70 ventral, 71 retrolateral). E—embolus, PA—proximal arm of tegular apophysis. RTA—retrolateral tibial apophysis. Unlabelled arrows indicate dorsal tibial ridge and dorsal prominence of patellar apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752607" httpUri="https://zenodo.org/record/4752607/files/figure.png" pageId="2" pageNumber="403">Figs 70–71</figureCitation>
). Proximal arm of tegular apophysis blunt with narrow longitudinal trough ventrally (
<figureCitation id="13192A501901BE2DFC726959FB8B9A3A" box="[991,1103,1612,1634]" captionStart="FIGURES 68–71" captionStartId="24.[151,264,1762,1786]" captionTargetBox="[151,1418,186,1738]" captionTargetId="figure-27@24.[151,1436,181,1738]" captionTargetPageId="24" captionText="FIGURES 68–71. Cybaeus ubicki spec. nov., males from California, left palpus (68–69, 71 from Novato, 70 from Anchor Bay). 68–69 Patella and tibia (68 retrolateral, 69 dorsal). 70–71 Genital bulb (70 ventral, 71 retrolateral). E—embolus, PA—proximal arm of tegular apophysis. RTA—retrolateral tibial apophysis. Unlabelled arrows indicate dorsal tibial ridge and dorsal prominence of patellar apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752607" httpUri="https://zenodo.org/record/4752607/files/figure.png" pageId="2" pageNumber="403">Figs 70–71</figureCitation>
,
<figureCitation id="13192A501901BE2DFBF76959FBB19A39" box="[1114,1141,1612,1634]" captionStart="FIGURES 72–73" captionStartId="25.[151,264,977,1001]" captionTargetBox="[151,1436,181,953]" captionTargetId="figure-17@25.[151,1436,181,953]" captionTargetPageId="25" captionText="FIGURES 72–73. Cybaeus ubicki spec. nov., male from Novato, California, left palpus. 72 Patellar apophysis, ventral. 73 Proximal arm of tegular apophysis, ventral. Arrows indicate diagnostic characters (72 dorsal prominence of patellar apophysis, 73 ventral longitudinal groove of proximal arm of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.4752611" httpUri="https://zenodo.org/record/4752611/files/figure.png" pageId="2" pageNumber="403">73</figureCitation>
). Atrium large, nearly as wide as vulva (
<figureCitation id="13192A501901BE2DFEED697CFE769A25" box="[320,434,1641,1663]" captionStart="FIGURES 74–79" captionStartId="26.[151,264,1772,1796]" captionTargetBox="[162,1436,186,1747]" captionTargetId="figure-27@26.[151,1436,181,1748]" captionTargetPageId="26" captionText="FIGURES 74–79. Cybaeus ubicki spec. nov., females from California, copulatory organ (74–76 from Point Reyes Station, 77– 79 from Anchor Bay). 74, 77 Epigynum, ventral. 75–76, 78–79 Vulva (75, 78, ventral, 76, 79 dorsal). CD—copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.4752616" httpUri="https://zenodo.org/record/4752616/files/figure.png" pageId="2" pageNumber="403">Figs 74, 77</figureCitation>
). Copulatory ducts broad, prominent; converging from atrium to midline then diverging before turning posteriorly at anterior margin of vulva (
<figureCitation id="13192A501901BE2DFDF06993FCD79AC7" box="[605,787,1670,1692]" captionStart="FIGURES 74–79" captionStartId="26.[151,264,1772,1796]" captionTargetBox="[162,1436,186,1747]" captionTargetId="figure-27@26.[151,1436,181,1748]" captionTargetPageId="26" captionText="FIGURES 74–79. Cybaeus ubicki spec. nov., females from California, copulatory organ (74–76 from Point Reyes Station, 77– 79 from Anchor Bay). 74, 77 Epigynum, ventral. 75–76, 78–79 Vulva (75, 78, ventral, 76, 79 dorsal). CD—copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.4752616" httpUri="https://zenodo.org/record/4752616/files/figure.png" pageId="2" pageNumber="403">Figs 75–76, 78–79</figureCitation>
)................................
<taxonomicName id="4C224D561901BE2DFB046993FAF19AC7" ID-CoL="98YZS" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1193,1333,1670,1692]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="ubicki" status="sp. nov.">
<emphasis id="B956EAC71901BE2DFB046993FB209AC0" box="[1193,1252,1670,1691]" italics="true" pageId="2" pageNumber="403">ubicki</emphasis>
Bennett
</taxonomicName>
<emphasis id="B956EAC71901BE2DFA976993FA589AC0" bold="true" box="[1338,1436,1670,1691]" pageId="2" pageNumber="403">
<taxonomicNameLabel id="A26557BC1901BE2DFA976993FA589AC0" box="[1338,1436,1670,1691]" pageId="2" pageNumber="403" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9D36D51901BE2DFF3A69B1FA589B4A" blockId="2.[151,1437,1084,2015]" pageId="2" pageNumber="403">
<keyLead id="30D395E01901BE2DFF3A69B1FA589B4A" pageId="2" pageNumber="403">
- Patellar apophysis without dorsobasal prominence (
<figureCitation id="13192A501901BE2DFD7969B1FCDB9AE2" box="[724,799,1700,1722]" captionStart="FIGURES 27–29" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[162,1415,184,1917]" captionTargetId="figure-27@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURES 27–29. Cybaeus opulentus spec. nov., male from Riverton, California, left palpus. 27 Patella and tibia, dorsal. 28–29 Genital bulb (28 ventral, 29 retrolateral). PA—proximal arm of tegular apophysis. Unlabelled arrows indicate compound curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752568" httpUri="https://zenodo.org/record/4752568/files/figure.png" pageId="2" pageNumber="403">Figs 27</figureCitation>
,
<figureCitation id="13192A501901BE2DFC8469B1FC879AE2" box="[809,835,1700,1721]" captionStart="FIGURES 33–36" captionStartId="14.[151,264,1933,1957]" captionTargetBox="[207,1372,273,1901]" captionTargetId="figure-71@14.[207,1381,273,1909]" captionTargetPageId="14" captionText="FIGURES 33–36. Cybaeus pan spec. nov., male from Calpine, California, left palpus. 33–34 Patella and tibia (33 retrolateral, 34 dorsal). 35–36 Genital bulb (35 ventral, 36 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752576" httpUri="https://zenodo.org/record/4752576/files/figure.png" pageId="2" pageNumber="403">34</figureCitation>
,
<figureCitation id="13192A501901BE2DFCE369B1FCAC9AE2" box="[846,872,1700,1721]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1806,1830]" captionTargetBox="[151,1419,155,1775]" captionTargetId="figure-17@17.[151,1436,151,1783]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus penedentatus, male from Holy City, California, left palpus. 42–43 Patella and tibia (42 retrolateral, 43 dorsal). 44–45 Genital bulb (44 ventral, 45 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate ventral keel of proximal arm of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752582" httpUri="https://zenodo.org/record/4752582/files/figure.png" pageId="2" pageNumber="403">43</figureCitation>
,
<figureCitation id="13192A501901BE2DFCDF69B1FC489AE1" box="[882,908,1700,1722]" captionStart="FIGURES 59–63" captionStartId="21.[151,264,1872,1896]" captionTargetBox="[176,1407,183,1835]" captionTargetId="figure-17@21.[176,1410,181,1849]" captionTargetPageId="21" captionText="FIGURES 59–63. Cybaeus simplex, males from Oregon, left palpus (59 from Ashland watershed, 60–63 from H.J. Andrews Experimental Forest). 59 Tibia, retrolateral. 60 Patella and tibia, dorsal. 61–62 Genital bulb (61 ventral, 62 retrolateral). 63 Tip of proximal arm of tegular apophysis, ventral. Arrows indicate diagnostic characters (59 distal extension of retrolateral tibial apophysis, 61 strongly twisted tip and prominent basal projection of proximal arm of tegular apophysis, 62 compound curve of dorsal margin of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.4752600" httpUri="https://zenodo.org/record/4752600/files/figure.png" pageId="2" pageNumber="403">60</figureCitation>
,
<figureCitation id="13192A501901BE2DFC3A69B1FC759AE1" box="[919,945,1700,1722]" captionStart="FIGURES 80–83" captionStartId="27.[151,264,1843,1867]" captionTargetBox="[151,1420,199,1819]" captionTargetId="figure-17@27.[151,1436,181,1819]" captionTargetPageId="27" captionText="FIGURES 80–83. Cybaeus vulpinus, male from Columbia, California, left palpus. 80–81 Patella and tibia (80 retrolateral, 81 dorsal). 82–83 Genital bulb (82 ventral, 83 retrolateral). Arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752620" httpUri="https://zenodo.org/record/4752620/files/figure.png" pageId="2" pageNumber="403">81</figureCitation>
). Tibia without dorso-medial ridge. Embolus “normal” (
<figureCitation id="13192A501901BE2DFEB369D4FEAD9A8C" box="[286,361,1729,1751]" captionStart="FIGURES 33–36" captionStartId="14.[151,264,1933,1957]" captionTargetBox="[207,1372,273,1901]" captionTargetId="figure-71@14.[207,1381,273,1909]" captionTargetPageId="14" captionText="FIGURES 33–36. Cybaeus pan spec. nov., male from Calpine, California, left palpus. 33–34 Patella and tibia (33 retrolateral, 34 dorsal). 35–36 Genital bulb (35 ventral, 36 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752576" httpUri="https://zenodo.org/record/4752576/files/figure.png" pageId="2" pageNumber="403">Figs 35</figureCitation>
,
<figureCitation id="13192A501901BE2DFED969D4FE4A9A8D" box="[372,398,1729,1750]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1806,1830]" captionTargetBox="[151,1419,155,1775]" captionTargetId="figure-17@17.[151,1436,151,1783]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus penedentatus, male from Holy City, California, left palpus. 42–43 Patella and tibia (42 retrolateral, 43 dorsal). 44–45 Genital bulb (44 ventral, 45 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate ventral keel of proximal arm of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752582" httpUri="https://zenodo.org/record/4752582/files/figure.png" pageId="2" pageNumber="403">44</figureCitation>
,
<figureCitation id="13192A501901BE2DFE3569D4FE779A8C" box="[408,435,1729,1751]" captionStart="FIGURES 59–63" captionStartId="21.[151,264,1872,1896]" captionTargetBox="[176,1407,183,1835]" captionTargetId="figure-17@21.[176,1410,181,1849]" captionTargetPageId="21" captionText="FIGURES 59–63. Cybaeus simplex, males from Oregon, left palpus (59 from Ashland watershed, 60–63 from H.J. Andrews Experimental Forest). 59 Tibia, retrolateral. 60 Patella and tibia, dorsal. 61–62 Genital bulb (61 ventral, 62 retrolateral). 63 Tip of proximal arm of tegular apophysis, ventral. Arrows indicate diagnostic characters (59 distal extension of retrolateral tibial apophysis, 61 strongly twisted tip and prominent basal projection of proximal arm of tegular apophysis, 62 compound curve of dorsal margin of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.4752600" httpUri="https://zenodo.org/record/4752600/files/figure.png" pageId="2" pageNumber="403">61</figureCitation>
). Tip of proximal arm of tegular apophysis acuminate (
<figureCitation id="13192A501901BE2DFC6469D4FBD19A8C" box="[969,1045,1729,1751]" captionStart="FIGURES 33–36" captionStartId="14.[151,264,1933,1957]" captionTargetBox="[207,1372,273,1901]" captionTargetId="figure-71@14.[207,1381,273,1909]" captionTargetPageId="14" captionText="FIGURES 33–36. Cybaeus pan spec. nov., male from Calpine, California, left palpus. 33–34 Patella and tibia (33 retrolateral, 34 dorsal). 35–36 Genital bulb (35 ventral, 36 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752576" httpUri="https://zenodo.org/record/4752576/files/figure.png" pageId="2" pageNumber="403">Figs 35</figureCitation>
,
<figureCitation id="13192A501901BE2DFBB269D4FBFD9A8D" box="[1055,1081,1729,1750]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1806,1830]" captionTargetBox="[151,1419,155,1775]" captionTargetId="figure-17@17.[151,1436,151,1783]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus penedentatus, male from Holy City, California, left palpus. 42–43 Patella and tibia (42 retrolateral, 43 dorsal). 44–45 Genital bulb (44 ventral, 45 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate ventral keel of proximal arm of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752582" httpUri="https://zenodo.org/record/4752582/files/figure.png" pageId="2" pageNumber="403">44</figureCitation>
,
<figureCitation id="13192A501901BE2DFBE969D4FB479A8C" box="[1092,1155,1729,1751]" captionStart="FIGURES 59–63" captionStartId="21.[151,264,1872,1896]" captionTargetBox="[176,1407,183,1835]" captionTargetId="figure-17@21.[176,1410,181,1849]" captionTargetPageId="21" captionText="FIGURES 59–63. Cybaeus simplex, males from Oregon, left palpus (59 from Ashland watershed, 60–63 from H.J. Andrews Experimental Forest). 59 Tibia, retrolateral. 60 Patella and tibia, dorsal. 61–62 Genital bulb (61 ventral, 62 retrolateral). 63 Tip of proximal arm of tegular apophysis, ventral. Arrows indicate diagnostic characters (59 distal extension of retrolateral tibial apophysis, 61 strongly twisted tip and prominent basal projection of proximal arm of tegular apophysis, 62 compound curve of dorsal margin of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.4752600" httpUri="https://zenodo.org/record/4752600/files/figure.png" pageId="2" pageNumber="403">61, 63</figureCitation>
). Atrium large but not nearly as wide as vulva (
<figureCitation id="13192A501901BE2DFE2769CBFE109AA8" box="[394,468,1758,1780]" captionStart="FIGURES 30–32" captionStartId="13.[151,264,1111,1135]" captionTargetBox="[156,1436,186,1087]" captionTargetId="figure-17@13.[151,1436,181,1087]" captionTargetPageId="13" captionText="FIGURES 30–32. Cybaeus opulentus spec. nov., female from Riverton, California, copulatory organ. 30 Epigynum, ventral. 31–32 Vulva (31 ventral, 32 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4752572" httpUri="https://zenodo.org/record/4752572/files/figure.png" pageId="2" pageNumber="403">Figs 30</figureCitation>
,
<figureCitation id="13192A501901BE2DFE7369CBFE3C9AA8" box="[478,504,1758,1779]" captionStart="FIGURES 37–41" captionStartId="15.[151,264,1667,1691]" captionTargetBox="[151,1431,185,1639]" captionTargetId="figure-17@15.[151,1436,181,1643]" captionTargetPageId="15" captionText="FIGURES 37–41. Cybaeus spp., females from California, copulatory organ (37–38 C. pan spec. nov. from near Calpine, 39 C. pan spec. nov. from Sierra City, 40–41 C. vulpinus from Big Trees State Park). 37 Epigynum, ventral. 38–41 Vulva (38, 40 ventral; 39, 41 dorsal). CD—copulatory duct, SS—spermathecal stalk. Unlabelled arrow indicates epigynal notch." figureDoi="http://doi.org/10.5281/zenodo.4752580" httpUri="https://zenodo.org/record/4752580/files/figure.png" pageId="2" pageNumber="403">37</figureCitation>
,
<figureCitation id="13192A501901BE2DFDAC69CBFDDE9AAF" box="[513,538,1758,1780]" captionStart="FIGURES 64–67" captionStartId="22.[151,264,1249,1273]" captionTargetBox="[153,1436,184,1225]" captionTargetId="figure-27@22.[151,1436,181,1226]" captionTargetPageId="22" captionText="FIGURES 64–67. Cybaeus simplex, females from Cascadia, Oregon, copulatory organ. 64 Epigynum, ventral. 65–67 Vulva (65 ventral, 66–67 dorsal). CD—copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.4752604" httpUri="https://zenodo.org/record/4752604/files/figure.png" pageId="2" pageNumber="403">64</figureCitation>
) or small (
<figureCitation id="13192A501901BE2DFD2D69CBFD2A9AAF" box="[640,750,1758,1780]" captionStart="FIGURES 46–51" captionStartId="18.[151,264,1787,1811]" captionTargetBox="[151,1433,181,1759]" captionTargetId="figure-27@18.[151,1437,181,1763]" captionTargetPageId="18" captionText="FIGURES 46–51. Cybaeus penedentatus, females from California, copulatory organ (46–48 from Ben Lomond, 49 from Pescadero Creek, 50–51 from Mount Madonna). 46 Epigynum, ventral. 47–51 Vulva (47, 49–50 ventral; 48, 51 dorsal). AT—atrium." figureDoi="http://doi.org/10.5281/zenodo.4752586" httpUri="https://zenodo.org/record/4752586/files/figure.png" pageId="2" pageNumber="403">Figs 46–47</figureCitation>
,
<figureCitation id="13192A501901BE2DFD5569CBFCD79AAF" box="[760,787,1758,1780]" captionStart="FIGURES 52–54" captionStartId="19.[151,264,1076,1100]" captionTargetBox="[176,1401,188,1050]" captionTargetId="figure-17@19.[176,1411,181,1052]" captionTargetPageId="19" captionText="FIGURES 52–54. Cybaeus penedentatus, female from Brookdale, California, copulatory organ. 52 Epigynum, ventral. 53–54 Vulva (53 ventral, 54 dorsal). AT—atrium." figureDoi="http://doi.org/10.5281/zenodo.4752592" httpUri="https://zenodo.org/record/4752592/files/figure.png" pageId="2" pageNumber="403">52</figureCitation>
). Copulatory ducts narrower (or, if broad, then membranous and not prominent); not following trajectory of converging then diverging before turning posteriorly (
<figureCitation id="13192A501901BE2DFBCE69E9FB699B4A" box="[1123,1197,1788,1810]" captionStart="FIGURES 30–32" captionStartId="13.[151,264,1111,1135]" captionTargetBox="[156,1436,186,1087]" captionTargetId="figure-17@13.[151,1436,181,1087]" captionTargetPageId="13" captionText="FIGURES 30–32. Cybaeus opulentus spec. nov., female from Riverton, California, copulatory organ. 30 Epigynum, ventral. 31–32 Vulva (31 ventral, 32 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4752572" httpUri="https://zenodo.org/record/4752572/files/figure.png" pageId="2" pageNumber="403">Figs 31</figureCitation>
,
<figureCitation id="13192A501901BE2DFB1A69E9FB159B4A" box="[1207,1233,1788,1809]" captionStart="FIGURES 37–41" captionStartId="15.[151,264,1667,1691]" captionTargetBox="[151,1431,185,1639]" captionTargetId="figure-17@15.[151,1436,181,1643]" captionTargetPageId="15" captionText="FIGURES 37–41. Cybaeus spp., females from California, copulatory organ (37–38 C. pan spec. nov. from near Calpine, 39 C. pan spec. nov. from Sierra City, 40–41 C. vulpinus from Big Trees State Park). 37 Epigynum, ventral. 38–41 Vulva (38, 40 ventral; 39, 41 dorsal). CD—copulatory duct, SS—spermathecal stalk. Unlabelled arrow indicates epigynal notch." figureDoi="http://doi.org/10.5281/zenodo.4752580" httpUri="https://zenodo.org/record/4752580/files/figure.png" pageId="2" pageNumber="403">40</figureCitation>
,
<figureCitation id="13192A501901BE2DFB7669E9FB319B4A" box="[1243,1269,1788,1809]" captionStart="FIGURES 46–51" captionStartId="18.[151,264,1787,1811]" captionTargetBox="[151,1433,181,1759]" captionTargetId="figure-27@18.[151,1437,181,1763]" captionTargetPageId="18" captionText="FIGURES 46–51. Cybaeus penedentatus, females from California, copulatory organ (46–48 from Ben Lomond, 49 from Pescadero Creek, 50–51 from Mount Madonna). 46 Epigynum, ventral. 47–51 Vulva (47, 49–50 ventral; 48, 51 dorsal). AT—atrium." figureDoi="http://doi.org/10.5281/zenodo.4752586" httpUri="https://zenodo.org/record/4752586/files/figure.png" pageId="2" pageNumber="403">47</figureCitation>
,
<figureCitation id="13192A501901BE2DFB5269EEFADC9B4A" box="[1279,1304,1787,1809]" captionStart="FIGURES 64–67" captionStartId="22.[151,264,1249,1273]" captionTargetBox="[153,1436,184,1225]" captionTargetId="figure-27@22.[151,1436,181,1226]" captionTargetPageId="22" captionText="FIGURES 64–67. Cybaeus simplex, females from Cascadia, Oregon, copulatory organ. 64 Epigynum, ventral. 65–67 Vulva (65 ventral, 66–67 dorsal). CD—copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.4752604" httpUri="https://zenodo.org/record/4752604/files/figure.png" pageId="2" pageNumber="403">65</figureCitation>
)......... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D62E701901BE2DFF3A680CFA589B99" pageId="2" pageNumber="403">
<paragraph id="8B9D36D51901BE2DFF3A680CFA599B32" blockId="2.[151,1437,1084,2015]" pageId="2" pageNumber="403">
<keyLead id="30D395E01901BE2DFF3A680CFA599B32" pageId="2" pageNumber="403">
5(4). Patellar apophysis with two to six peg setae (
<figureCitation id="13192A501901BE2DFD38680CFD199B74" box="[661,733,1817,1839]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1806,1830]" captionTargetBox="[151,1419,155,1775]" captionTargetId="figure-17@17.[151,1436,151,1783]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus penedentatus, male from Holy City, California, left palpus. 42–43 Patella and tibia (42 retrolateral, 43 dorsal). 44–45 Genital bulb (44 ventral, 45 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate ventral keel of proximal arm of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752582" httpUri="https://zenodo.org/record/4752582/files/figure.png" pageId="2" pageNumber="403">Fig. 43</figureCitation>
). Proximal arm of tegular apophysis with prominent ventral keel and terminating in short, slightly twisted tip (
<figureCitation id="13192A501901BE2DFDFE6823FD079B17" box="[595,707,1846,1868]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1806,1830]" captionTargetBox="[151,1419,155,1775]" captionTargetId="figure-17@17.[151,1436,151,1783]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus penedentatus, male from Holy City, California, left palpus. 42–43 Patella and tibia (42 retrolateral, 43 dorsal). 44–45 Genital bulb (44 ventral, 45 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate ventral keel of proximal arm of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752582" httpUri="https://zenodo.org/record/4752582/files/figure.png" pageId="2" pageNumber="403">Figs 44–45</figureCitation>
,
<figureCitation id="13192A501901BE2DFD606823FD2C9B17" box="[717,744,1846,1868]" captionStart="FIGURES 55–58" captionStartId="20.[151,264,1895,1919]" captionTargetBox="[177,1409,381,1872]" captionTargetId="figure-139@20.[177,1411,381,1872]" captionTargetPageId="20" captionText="FIGURES 55–58. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis (55 C. opulentus spec. nov. from Riverton, 56 C. penedentatus from Holy City, 57 C. vulpinus from Columbia, 58 C. pan from Calpine) (55–56 ventral, 57–58 prolateral). Arrows indicate diagnostic characters (55 slightly twisted tip and prominent basal protrusion; 56 acuminate slightly twisted tip and longitudinal ventral keel; 57 smoothly curved, twisted tip; 58 triangular keel dorsally on tip)." figureDoi="http://doi.org/10.5281/zenodo.4752596" httpUri="https://zenodo.org/record/4752596/files/figure.png" pageId="2" pageNumber="403">56</figureCitation>
). Atrium small, inconspicuous, its width equal to about 1/3 of the width of the vulva (
<figureCitation id="13192A501901BE2DFECC6841FE149B31" box="[353,464,1876,1898]" captionStart="FIGURES 46–51" captionStartId="18.[151,264,1787,1811]" captionTargetBox="[151,1433,181,1759]" captionTargetId="figure-27@18.[151,1437,181,1763]" captionTargetPageId="18" captionText="FIGURES 46–51. Cybaeus penedentatus, females from California, copulatory organ (46–48 from Ben Lomond, 49 from Pescadero Creek, 50–51 from Mount Madonna). 46 Epigynum, ventral. 47–51 Vulva (47, 49–50 ventral; 48, 51 dorsal). AT—atrium." figureDoi="http://doi.org/10.5281/zenodo.4752586" httpUri="https://zenodo.org/record/4752586/files/figure.png" pageId="2" pageNumber="403">Figs 46–47</figureCitation>
,
<figureCitation id="13192A501901BE2DFE776846FE379B32" box="[474,499,1875,1897]" captionStart="FIGURES 52–54" captionStartId="19.[151,264,1076,1100]" captionTargetBox="[176,1401,188,1050]" captionTargetId="figure-17@19.[176,1411,181,1052]" captionTargetPageId="19" captionText="FIGURES 52–54. Cybaeus penedentatus, female from Brookdale, California, copulatory organ. 52 Epigynum, ventral. 53–54 Vulva (53 ventral, 54 dorsal). AT—atrium." figureDoi="http://doi.org/10.5281/zenodo.4752592" httpUri="https://zenodo.org/record/4752592/files/figure.png" pageId="2" pageNumber="403">52</figureCitation>
)............................................................
<taxonomicName id="4C224D561901BE2DFB606841FA599B32" ID-CoL="32RGT" authority="Bennett" authorityName="Bennett" authorityYear="2009" box="[1229,1437,1876,1897]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="penedentatus">
<emphasis id="B956EAC71901BE2DFB606841FA889B32" box="[1229,1356,1876,1897]" italics="true" pageId="2" pageNumber="403">penedentatus</emphasis>
Bennett
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</paragraph>
<paragraph id="8B9D36D51901BE2DFF3A6864FA589B99" blockId="2.[151,1437,1084,2015]" pageId="2" pageNumber="403">
<keyLead id="30D395E01901BE2DFF3A6864FA589B99" pageId="2" pageNumber="403">
- Patellar apophysis with 13 to 40 peg setae (
<figureCitation id="13192A501901BE2DFD216864FD1C9BDD" box="[652,728,1905,1927]" captionStart="FIGURES 27–29" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[162,1415,184,1917]" captionTargetId="figure-27@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURES 27–29. Cybaeus opulentus spec. nov., male from Riverton, California, left palpus. 27 Patella and tibia, dorsal. 28–29 Genital bulb (28 ventral, 29 retrolateral). PA—proximal arm of tegular apophysis. Unlabelled arrows indicate compound curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752568" httpUri="https://zenodo.org/record/4752568/files/figure.png" pageId="2" pageNumber="403">Figs 27</figureCitation>
,
<figureCitation id="13192A501901BE2DFD4E6864FD399BDD" box="[739,765,1905,1926]" captionStart="FIGURES 33–36" captionStartId="14.[151,264,1933,1957]" captionTargetBox="[207,1372,273,1901]" captionTargetId="figure-71@14.[207,1381,273,1909]" captionTargetPageId="14" captionText="FIGURES 33–36. Cybaeus pan spec. nov., male from Calpine, California, left palpus. 33–34 Patella and tibia (33 retrolateral, 34 dorsal). 35–36 Genital bulb (35 ventral, 36 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752576" httpUri="https://zenodo.org/record/4752576/files/figure.png" pageId="2" pageNumber="403">34</figureCitation>
,
<figureCitation id="13192A501901BE2DFCA56864FCE69BDC" box="[776,802,1905,1927]" captionStart="FIGURES 59–63" captionStartId="21.[151,264,1872,1896]" captionTargetBox="[176,1407,183,1835]" captionTargetId="figure-17@21.[176,1410,181,1849]" captionTargetPageId="21" captionText="FIGURES 59–63. Cybaeus simplex, males from Oregon, left palpus (59 from Ashland watershed, 60–63 from H.J. Andrews Experimental Forest). 59 Tibia, retrolateral. 60 Patella and tibia, dorsal. 61–62 Genital bulb (61 ventral, 62 retrolateral). 63 Tip of proximal arm of tegular apophysis, ventral. Arrows indicate diagnostic characters (59 distal extension of retrolateral tibial apophysis, 61 strongly twisted tip and prominent basal projection of proximal arm of tegular apophysis, 62 compound curve of dorsal margin of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.4752600" httpUri="https://zenodo.org/record/4752600/files/figure.png" pageId="2" pageNumber="403">60</figureCitation>
,
<figureCitation id="13192A501901BE2DFC806864FC839BDC" box="[813,839,1905,1927]" captionStart="FIGURES 80–83" captionStartId="27.[151,264,1843,1867]" captionTargetBox="[151,1420,199,1819]" captionTargetId="figure-17@27.[151,1436,181,1819]" captionTargetPageId="27" captionText="FIGURES 80–83. Cybaeus vulpinus, male from Columbia, California, left palpus. 80–81 Patella and tibia (80 retrolateral, 81 dorsal). 82–83 Genital bulb (82 ventral, 83 retrolateral). Arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752620" httpUri="https://zenodo.org/record/4752620/files/figure.png" pageId="2" pageNumber="403">81</figureCitation>
). Proximal arm of tegular apophysis without ventral keel; tip twisted or not (
<figureCitation id="13192A501901BE2DFEDC689BFE7F9BF8" box="[369,443,1934,1956]" captionStart="FIGURES 27–29" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[162,1415,184,1917]" captionTargetId="figure-27@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURES 27–29. Cybaeus opulentus spec. nov., male from Riverton, California, left palpus. 27 Patella and tibia, dorsal. 28–29 Genital bulb (28 ventral, 29 retrolateral). PA—proximal arm of tegular apophysis. Unlabelled arrows indicate compound curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752568" httpUri="https://zenodo.org/record/4752568/files/figure.png" pageId="2" pageNumber="403">Figs 28</figureCitation>
,
<figureCitation id="13192A501901BE2DFE68689BFE1B9BFF" box="[453,479,1934,1956]" captionStart="FIGURES 33–36" captionStartId="14.[151,264,1933,1957]" captionTargetBox="[207,1372,273,1901]" captionTargetId="figure-71@14.[207,1381,273,1909]" captionTargetPageId="14" captionText="FIGURES 33–36. Cybaeus pan spec. nov., male from Calpine, California, left palpus. 33–34 Patella and tibia (33 retrolateral, 34 dorsal). 35–36 Genital bulb (35 ventral, 36 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752576" httpUri="https://zenodo.org/record/4752576/files/figure.png" pageId="2" pageNumber="403">35</figureCitation>
,
<figureCitation id="13192A501901BE2DFE45689BFDC69BFF" box="[488,514,1934,1956]" captionStart="FIGURES 55–58" captionStartId="20.[151,264,1895,1919]" captionTargetBox="[177,1409,381,1872]" captionTargetId="figure-139@20.[177,1411,381,1872]" captionTargetPageId="20" captionText="FIGURES 55–58. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis (55 C. opulentus spec. nov. from Riverton, 56 C. penedentatus from Holy City, 57 C. vulpinus from Columbia, 58 C. pan from Calpine) (55–56 ventral, 57–58 prolateral). Arrows indicate diagnostic characters (55 slightly twisted tip and prominent basal protrusion; 56 acuminate slightly twisted tip and longitudinal ventral keel; 57 smoothly curved, twisted tip; 58 triangular keel dorsally on tip)." figureDoi="http://doi.org/10.5281/zenodo.4752596" httpUri="https://zenodo.org/record/4752596/files/figure.png" pageId="2" pageNumber="403">57</figureCitation>
,
<figureCitation id="13192A501901BE2DFDA1689BFD8F9BFF" box="[524,587,1934,1956]" captionStart="FIGURES 59–63" captionStartId="21.[151,264,1872,1896]" captionTargetBox="[176,1407,183,1835]" captionTargetId="figure-17@21.[176,1410,181,1849]" captionTargetPageId="21" captionText="FIGURES 59–63. Cybaeus simplex, males from Oregon, left palpus (59 from Ashland watershed, 60–63 from H.J. Andrews Experimental Forest). 59 Tibia, retrolateral. 60 Patella and tibia, dorsal. 61–62 Genital bulb (61 ventral, 62 retrolateral). 63 Tip of proximal arm of tegular apophysis, ventral. Arrows indicate diagnostic characters (59 distal extension of retrolateral tibial apophysis, 61 strongly twisted tip and prominent basal projection of proximal arm of tegular apophysis, 62 compound curve of dorsal margin of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.4752600" httpUri="https://zenodo.org/record/4752600/files/figure.png" pageId="2" pageNumber="403">61, 63</figureCitation>
). Atrium larger, more conspicuous, its width about 1/2 to 2/3 width of the vulva (
<figureCitation id="13192A501901BE2DFAE3689BFA5C9BF8" box="[1358,1432,1934,1956]" captionStart="FIGURES 30–32" captionStartId="13.[151,264,1111,1135]" captionTargetBox="[156,1436,186,1087]" captionTargetId="figure-17@13.[151,1436,181,1087]" captionTargetPageId="13" captionText="FIGURES 30–32. Cybaeus opulentus spec. nov., female from Riverton, California, copulatory organ. 30 Epigynum, ventral. 31–32 Vulva (31 ventral, 32 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4752572" httpUri="https://zenodo.org/record/4752572/files/figure.png" pageId="2" pageNumber="403">Figs 30</figureCitation>
,
<figureCitation id="13192A501901BE2DFF4F68B9FF389B9A" box="[226,252,1964,1985]" captionStart="FIGURES 37–41" captionStartId="15.[151,264,1667,1691]" captionTargetBox="[151,1431,185,1639]" captionTargetId="figure-17@15.[151,1436,181,1643]" captionTargetPageId="15" captionText="FIGURES 37–41. Cybaeus spp., females from California, copulatory organ (37–38 C. pan spec. nov. from near Calpine, 39 C. pan spec. nov. from Sierra City, 40–41 C. vulpinus from Big Trees State Park). 37 Epigynum, ventral. 38–41 Vulva (38, 40 ventral; 39, 41 dorsal). CD—copulatory duct, SS—spermathecal stalk. Unlabelled arrow indicates epigynal notch." figureDoi="http://doi.org/10.5281/zenodo.4752580" httpUri="https://zenodo.org/record/4752580/files/figure.png" pageId="2" pageNumber="403">37</figureCitation>
,
<figureCitation id="13192A501901BE2DFEAB68B9FEDB9B99" box="[262,287,1964,1986]" captionStart="FIGURES 64–67" captionStartId="22.[151,264,1249,1273]" captionTargetBox="[153,1436,184,1225]" captionTargetId="figure-27@22.[151,1436,181,1226]" captionTargetPageId="22" captionText="FIGURES 64–67. Cybaeus simplex, females from Cascadia, Oregon, copulatory organ. 64 Epigynum, ventral. 65–67 Vulva (65 ventral, 66–67 dorsal). CD—copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.4752604" httpUri="https://zenodo.org/record/4752604/files/figure.png" pageId="2" pageNumber="403">64</figureCitation>
).............................................................................................. 6
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<keyStep id="30D62E701901BE2CFF3A68DCFA589D5E" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403">
<paragraph id="8B9D36D51901BE2CFF3A68DCFA589CF6" blockId="2.[151,1437,1084,2015]" lastBlockId="3.[151,1437,151,614]" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403">
<keyLead id="30D395E01901BE2CFF3A68DCFA589CF6" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403">
6(5). In retrolateral view, dorsal margin of tegular apophysis a simple curve, lacking a proximal projection (
<figureCitation id="13192A501901BE2DFB1C68DCFB3F9B84" box="[1201,1275,1993,2015]" captionStart="FIGURES 33–36" captionStartId="14.[151,264,1933,1957]" captionTargetBox="[207,1372,273,1901]" captionTargetId="figure-71@14.[207,1381,273,1909]" captionTargetPageId="14" captionText="FIGURES 33–36. Cybaeus pan spec. nov., male from Calpine, California, left palpus. 33–34 Patella and tibia (33 retrolateral, 34 dorsal). 35–36 Genital bulb (35 ventral, 36 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752576" httpUri="https://zenodo.org/record/4752576/files/figure.png" pageId="2" pageNumber="403">Figs 36</figureCitation>
,
<figureCitation id="13192A501901BE2DFAA868DCFAE49B84" box="[1285,1312,1993,2015]" captionStart="FIGURES 80–83" captionStartId="27.[151,264,1843,1867]" captionTargetBox="[151,1420,199,1819]" captionTargetId="figure-17@27.[151,1436,181,1819]" captionTargetPageId="27" captionText="FIGURES 80–83. Cybaeus vulpinus, male from Columbia, California, left palpus. 80–81 Patella and tibia (80 retrolateral, 81 dorsal). 82–83 Genital bulb (82 ventral, 83 retrolateral). Arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752620" httpUri="https://zenodo.org/record/4752620/files/figure.png" pageId="2" pageNumber="403">83</figureCitation>
). Copulatory ducts usually only contiguous at atrium (
<figureCitation id="13192A501900BE2CFDC46F82FD1D9CF5" box="[617,729,151,174]" captionStart="FIGURES 37–41" captionStartId="15.[151,264,1667,1691]" captionTargetBox="[151,1431,185,1639]" captionTargetId="figure-17@15.[151,1436,181,1643]" captionTargetPageId="15" captionText="FIGURES 37–41. Cybaeus spp., females from California, copulatory organ (37–38 C. pan spec. nov. from near Calpine, 39 C. pan spec. nov. from Sierra City, 40–41 C. vulpinus from Big Trees State Park). 37 Epigynum, ventral. 38–41 Vulva (38, 40 ventral; 39, 41 dorsal). CD—copulatory duct, SS—spermathecal stalk. Unlabelled arrow indicates epigynal notch." figureDoi="http://doi.org/10.5281/zenodo.4752580" httpUri="https://zenodo.org/record/4752580/files/figure.png" pageId="3" pageNumber="404">Figs 38, 40</figureCitation>
). Femora usually banded..................................... 7
</keyLead>
</paragraph>
<paragraph id="8B9D36D51900BE2CFF3A6FA0FA589D5E" blockId="3.[151,1437,151,614]" pageId="3" pageNumber="404">
<keyLead id="30D395E01900BE2CFF3A6FA0FA589D5E" pageId="3" pageNumber="404">
- In retrolateral view, dorsal margin of tegular apophysis a compound curve with a prominent proximally directed projection (
<figureCitation id="13192A501900BE2CFF446FC7FEF79CBC" box="[233,307,210,232]" captionStart="FIGURES 27–29" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[162,1415,184,1917]" captionTargetId="figure-27@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURES 27–29. Cybaeus opulentus spec. nov., male from Riverton, California, left palpus. 27 Patella and tibia, dorsal. 28–29 Genital bulb (28 ventral, 29 retrolateral). PA—proximal arm of tegular apophysis. Unlabelled arrows indicate compound curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752568" httpUri="https://zenodo.org/record/4752568/files/figure.png" pageId="3" pageNumber="404">Figs 29</figureCitation>
,
<figureCitation id="13192A501900BE2CFE906FC7FEB99CB3" box="[317,381,210,232]" captionStart="FIGURES 59–63" captionStartId="21.[151,264,1872,1896]" captionTargetBox="[176,1407,183,1835]" captionTargetId="figure-17@21.[176,1410,181,1849]" captionTargetPageId="21" captionText="FIGURES 59–63. Cybaeus simplex, males from Oregon, left palpus (59 from Ashland watershed, 60–63 from H.J. Andrews Experimental Forest). 59 Tibia, retrolateral. 60 Patella and tibia, dorsal. 61–62 Genital bulb (61 ventral, 62 retrolateral). 63 Tip of proximal arm of tegular apophysis, ventral. Arrows indicate diagnostic characters (59 distal extension of retrolateral tibial apophysis, 61 strongly twisted tip and prominent basal projection of proximal arm of tegular apophysis, 62 compound curve of dorsal margin of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.4752600" httpUri="https://zenodo.org/record/4752600/files/figure.png" pageId="3" pageNumber="404">61–62</figureCitation>
). Copulatory ducts usually contiguous at atrium and medially (
<figureCitation id="13192A501900BE2CFC756FC7FB829CBC" box="[984,1094,210,232]" captionStart="FIGURES 30–32" captionStartId="13.[151,264,1111,1135]" captionTargetBox="[156,1436,186,1087]" captionTargetId="figure-17@13.[151,1436,181,1087]" captionTargetPageId="13" captionText="FIGURES 30–32. Cybaeus opulentus spec. nov., female from Riverton, California, copulatory organ. 30 Epigynum, ventral. 31–32 Vulva (31 ventral, 32 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4752572" httpUri="https://zenodo.org/record/4752572/files/figure.png" pageId="3" pageNumber="404">Figs 31–32</figureCitation>
,
<figureCitation id="13192A501900BE2CFBFC6FC7FB549CB3" box="[1105,1168,210,232]" captionStart="FIGURES 64–67" captionStartId="22.[151,264,1249,1273]" captionTargetBox="[153,1436,184,1225]" captionTargetId="figure-27@22.[151,1436,181,1226]" captionTargetPageId="22" captionText="FIGURES 64–67. Cybaeus simplex, females from Cascadia, Oregon, copulatory organ. 64 Epigynum, ventral. 65–67 Vulva (65 ventral, 66–67 dorsal). CD—copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.4752604" httpUri="https://zenodo.org/record/4752604/files/figure.png" pageId="3" pageNumber="404">65–67</figureCitation>
). Femora usually unbanded ................................................................................................... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D62E701900BE2CFF3A6E18FA589DEE" pageId="3" pageNumber="404">
<paragraph id="8B9D36D51900BE2CFF3A6E18FA599D06" blockId="3.[151,1437,151,614]" pageId="3" pageNumber="404">
<keyLead id="30D395E01900BE2CFF3A6E18FA599D06" pageId="3" pageNumber="404">
7(6). About 13 peg setae around tip of short, anteriorly directed patellar apophysis (
<figureCitation id="13192A501900BE2CFC676E18FBD49D78" box="[970,1040,269,291]" captionStart="FIGURES 80–83" captionStartId="27.[151,264,1843,1867]" captionTargetBox="[151,1420,199,1819]" captionTargetId="figure-17@27.[151,1436,181,1819]" captionTargetPageId="27" captionText="FIGURES 80–83. Cybaeus vulpinus, male from Columbia, California, left palpus. 80–81 Patella and tibia (80 retrolateral, 81 dorsal). 82–83 Genital bulb (82 ventral, 83 retrolateral). Arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752620" httpUri="https://zenodo.org/record/4752620/files/figure.png" pageId="3" pageNumber="404">Fig. 81</figureCitation>
). Proximal arm of tegular apophysis with simple moderately corkscrewed tip lacking a dorsal process (
<figureCitation id="13192A501900BE2CFC826E3FFCBE9D1B" box="[815,890,298,320]" captionStart="FIGURES 55–58" captionStartId="20.[151,264,1895,1919]" captionTargetBox="[177,1409,381,1872]" captionTargetId="figure-139@20.[177,1411,381,1872]" captionTargetPageId="20" captionText="FIGURES 55–58. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis (55 C. opulentus spec. nov. from Riverton, 56 C. penedentatus from Holy City, 57 C. vulpinus from Columbia, 58 C. pan from Calpine) (55–56 ventral, 57–58 prolateral). Arrows indicate diagnostic characters (55 slightly twisted tip and prominent basal protrusion; 56 acuminate slightly twisted tip and longitudinal ventral keel; 57 smoothly curved, twisted tip; 58 triangular keel dorsally on tip)." figureDoi="http://doi.org/10.5281/zenodo.4752596" httpUri="https://zenodo.org/record/4752596/files/figure.png" pageId="3" pageNumber="404">Figs 57</figureCitation>
,
<figureCitation id="13192A501900BE2CFC296E3FFC5B9D1B" box="[900,927,298,320]" captionStart="FIGURES 80–83" captionStartId="27.[151,264,1843,1867]" captionTargetBox="[151,1420,199,1819]" captionTargetId="figure-17@27.[151,1436,181,1819]" captionTargetPageId="27" captionText="FIGURES 80–83. Cybaeus vulpinus, male from Columbia, California, left palpus. 80–81 Patella and tibia (80 retrolateral, 81 dorsal). 82–83 Genital bulb (82 ventral, 83 retrolateral). Arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752620" httpUri="https://zenodo.org/record/4752620/files/figure.png" pageId="3" pageNumber="404">83</figureCitation>
). Spermathecal stalks slightly sinuous, describing an obtuse angle of about 120
<emphasis id="B956EAC71900BE2CFE776E50FE219D07" bold="true" box="[474,485,325,348]" pageId="3" pageNumber="404">⁰</emphasis>
–130
<emphasis id="B956EAC71900BE2CFDB86E50FDE49D07" bold="true" box="[533,544,325,348]" pageId="3" pageNumber="404">⁰</emphasis>
near Bennett’s glands (
<figureCitation id="13192A501900BE2CFCAF6E52FC8C9D05" box="[770,840,327,350]" captionStart="FIGURES 37–41" captionStartId="15.[151,264,1667,1691]" captionTargetBox="[151,1431,185,1639]" captionTargetId="figure-17@15.[151,1436,181,1643]" captionTargetPageId="15" captionText="FIGURES 37–41. Cybaeus spp., females from California, copulatory organ (37–38 C. pan spec. nov. from near Calpine, 39 C. pan spec. nov. from Sierra City, 40–41 C. vulpinus from Big Trees State Park). 37 Epigynum, ventral. 38–41 Vulva (38, 40 ventral; 39, 41 dorsal). CD—copulatory duct, SS—spermathecal stalk. Unlabelled arrow indicates epigynal notch." figureDoi="http://doi.org/10.5281/zenodo.4752580" httpUri="https://zenodo.org/record/4752580/files/figure.png" pageId="3" pageNumber="404">Fig. 41</figureCitation>
)...................................
<taxonomicName id="4C224D561900BE2CFB576E52FA599D06" ID-CoL="32RJ9" authority="Bennett" authorityName="Bennett. Eight" authorityYear="2009" box="[1274,1437,327,349]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="vulpinus">
<emphasis id="B956EAC71900BE2CFB576E52FA8F9D07" box="[1274,1355,327,348]" italics="true" pageId="3" pageNumber="404">vulpinus</emphasis>
Bennett
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9D36D51900BE2CFF3A6E70FA589DEE" blockId="3.[151,1437,151,614]" pageId="3" pageNumber="404">
<keyLead id="30D395E01900BE2CFF3A6E70FA589DEE" pageId="3" pageNumber="404">
- About 35 peg setae dorsally on long, anterolaterally directed patellar apophysis (
<figureCitation id="13192A501900BE2CFC5F6E70FBFF9D20" box="[1010,1083,357,379]" captionStart="FIGURES 33–36" captionStartId="14.[151,264,1933,1957]" captionTargetBox="[207,1372,273,1901]" captionTargetId="figure-71@14.[207,1381,273,1909]" captionTargetPageId="14" captionText="FIGURES 33–36. Cybaeus pan spec. nov., male from Calpine, California, left palpus. 33–34 Patella and tibia (33 retrolateral, 34 dorsal). 35–36 Genital bulb (35 ventral, 36 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752576" httpUri="https://zenodo.org/record/4752576/files/figure.png" pageId="3" pageNumber="404">Fig. 34</figureCitation>
). Proximal arm of tegular apophysis with simple untwisted tip; tip with small, triangular dorsal process (
<figureCitation id="13192A501900BE2CFCD96E97FC049DC3" box="[884,960,386,408]" captionStart="FIGURES 33–36" captionStartId="14.[151,264,1933,1957]" captionTargetBox="[207,1372,273,1901]" captionTargetId="figure-71@14.[207,1381,273,1909]" captionTargetPageId="14" captionText="FIGURES 33–36. Cybaeus pan spec. nov., male from Calpine, California, left palpus. 33–34 Patella and tibia (33 retrolateral, 34 dorsal). 35–36 Genital bulb (35 ventral, 36 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate simple curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752576" httpUri="https://zenodo.org/record/4752576/files/figure.png" pageId="3" pageNumber="404">Figs 35</figureCitation>
,
<figureCitation id="13192A501900BE2CFC666E97FC229DC3" box="[971,998,386,408]" captionStart="FIGURES 55–58" captionStartId="20.[151,264,1895,1919]" captionTargetBox="[177,1409,381,1872]" captionTargetId="figure-139@20.[177,1411,381,1872]" captionTargetPageId="20" captionText="FIGURES 55–58. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis (55 C. opulentus spec. nov. from Riverton, 56 C. penedentatus from Holy City, 57 C. vulpinus from Columbia, 58 C. pan from Calpine) (55–56 ventral, 57–58 prolateral). Arrows indicate diagnostic characters (55 slightly twisted tip and prominent basal protrusion; 56 acuminate slightly twisted tip and longitudinal ventral keel; 57 smoothly curved, twisted tip; 58 triangular keel dorsally on tip)." figureDoi="http://doi.org/10.5281/zenodo.4752596" httpUri="https://zenodo.org/record/4752596/files/figure.png" pageId="3" pageNumber="404">58</figureCitation>
). Spermathecal stalks more strongly sinuous, describing an acute angle of about 55
<emphasis id="B956EAC71900BE2CFDE56E88FD979DEF" bold="true" box="[584,595,413,436]" pageId="3" pageNumber="404">⁰</emphasis>
–65
<emphasis id="B956EAC71900BE2CFDD56E88FD479DEF" bold="true" box="[632,643,413,436]" pageId="3" pageNumber="404">⁰</emphasis>
near Bennett’s gland (
<figureCitation id="13192A501900BE2CFCF66E8AFC659DED" box="[859,929,415,438]" captionStart="FIGURES 37–41" captionStartId="15.[151,264,1667,1691]" captionTargetBox="[151,1431,185,1639]" captionTargetId="figure-17@15.[151,1436,181,1643]" captionTargetPageId="15" captionText="FIGURES 37–41. Cybaeus spp., females from California, copulatory organ (37–38 C. pan spec. nov. from near Calpine, 39 C. pan spec. nov. from Sierra City, 40–41 C. vulpinus from Big Trees State Park). 37 Epigynum, ventral. 38–41 Vulva (38, 40 ventral; 39, 41 dorsal). CD—copulatory duct, SS—spermathecal stalk. Unlabelled arrow indicates epigynal notch." figureDoi="http://doi.org/10.5281/zenodo.4752580" httpUri="https://zenodo.org/record/4752580/files/figure.png" pageId="3" pageNumber="404">Fig. 39</figureCitation>
)......................
<taxonomicName id="4C224D561900BE2CFB6D6EB5FAF19DEE" ID-CoL="98YZW" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1216,1333,416,437]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="pan" status="sp. nov.">
<emphasis id="B956EAC71900BE2CFB6D6EB5FB209DEE" box="[1216,1252,416,437]" italics="true" pageId="3" pageNumber="404">pan</emphasis>
Bennett
</taxonomicName>
<emphasis id="B956EAC71900BE2CFA976EB5FA589DEE" bold="true" box="[1338,1436,416,437]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="A26557BC1900BE2CFA976EB5FA589DEE" box="[1338,1436,416,437]" pageId="3" pageNumber="404" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D62E701900BE2CFF3A6EA8FA589E3E" pageId="3" pageNumber="404">
<paragraph id="8B9D36D51900BE2CFF3A6EA8FA599E56" blockId="3.[151,1437,151,614]" pageId="3" pageNumber="404">
<keyLead id="30D395E01900BE2CFF3A6EA8FA599E56" pageId="3" pageNumber="404">
8(6). Patellar apophysis long, about as long as width of patella, with 15–25 peg setae (
<figureCitation id="13192A501900BE2CFC5E6EA8FBF89D88" box="[1011,1084,445,467]" captionStart="FIGURES 59–63" captionStartId="21.[151,264,1872,1896]" captionTargetBox="[176,1407,183,1835]" captionTargetId="figure-17@21.[176,1410,181,1849]" captionTargetPageId="21" captionText="FIGURES 59–63. Cybaeus simplex, males from Oregon, left palpus (59 from Ashland watershed, 60–63 from H.J. Andrews Experimental Forest). 59 Tibia, retrolateral. 60 Patella and tibia, dorsal. 61–62 Genital bulb (61 ventral, 62 retrolateral). 63 Tip of proximal arm of tegular apophysis, ventral. Arrows indicate diagnostic characters (59 distal extension of retrolateral tibial apophysis, 61 strongly twisted tip and prominent basal projection of proximal arm of tegular apophysis, 62 compound curve of dorsal margin of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.4752600" httpUri="https://zenodo.org/record/4752600/files/figure.png" pageId="3" pageNumber="404">Fig. 60</figureCitation>
). Proximal arm of tegular apophysis with prominent basal projection and strongly twisted tip (
<figureCitation id="13192A501900BE2CFCAE6ECFFCB79DAB" box="[771,883,474,496]" captionStart="FIGURES 59–63" captionStartId="21.[151,264,1872,1896]" captionTargetBox="[176,1407,183,1835]" captionTargetId="figure-17@21.[176,1410,181,1849]" captionTargetPageId="21" captionText="FIGURES 59–63. Cybaeus simplex, males from Oregon, left palpus (59 from Ashland watershed, 60–63 from H.J. Andrews Experimental Forest). 59 Tibia, retrolateral. 60 Patella and tibia, dorsal. 61–62 Genital bulb (61 ventral, 62 retrolateral). 63 Tip of proximal arm of tegular apophysis, ventral. Arrows indicate diagnostic characters (59 distal extension of retrolateral tibial apophysis, 61 strongly twisted tip and prominent basal projection of proximal arm of tegular apophysis, 62 compound curve of dorsal margin of tegular apophysis)." figureDoi="http://doi.org/10.5281/zenodo.4752600" httpUri="https://zenodo.org/record/4752600/files/figure.png" pageId="3" pageNumber="404">Figs 61–63</figureCitation>
). Copulatory ducts and spermathecal stalks elongate (
<figureCitation id="13192A501900BE2CFADF6ECFFEDB9E56" captionStart="FIGURES 64–67" captionStartId="22.[151,264,1249,1273]" captionTargetBox="[153,1436,184,1225]" captionTargetId="figure-27@22.[151,1436,181,1226]" captionTargetPageId="22" captionText="FIGURES 64–67. Cybaeus simplex, females from Cascadia, Oregon, copulatory organ. 64 Epigynum, ventral. 65–67 Vulva (65 ventral, 66–67 dorsal). CD—copulatory duct." figureDoi="http://doi.org/10.5281/zenodo.4752604" httpUri="https://zenodo.org/record/4752604/files/figure.png" pageId="3" pageNumber="404">Figs 65–67</figureCitation>
).....................................................................................
<taxonomicName id="4C224D561900BE2CFAB36EE2FA599E56" ID-CoL="32RHK" authority="Roth" authorityName="Roth" authorityYear="1952" box="[1310,1437,503,525]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B956EAC71900BE2CFAB36EE2FAA39E57" box="[1310,1383,503,524]" italics="true" pageId="3" pageNumber="404">simplex</emphasis>
Roth
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9D36D51900BE2CFF3A6D00FA589E3E" blockId="3.[151,1437,151,614]" pageId="3" pageNumber="404">
<keyLead id="30D395E01900BE2CFF3A6D00FA589E3E" pageId="3" pageNumber="404">
- Patellar apophysis shorter, length about 2/3 patellar width, with 30–40 peg setae (
<figureCitation id="13192A501900BE2CFC586D00FBF99E70" box="[1013,1085,533,555]" captionStart="FIGURES 27–29" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[162,1415,184,1917]" captionTargetId="figure-27@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURES 27–29. Cybaeus opulentus spec. nov., male from Riverton, California, left palpus. 27 Patella and tibia, dorsal. 28–29 Genital bulb (28 ventral, 29 retrolateral). PA—proximal arm of tegular apophysis. Unlabelled arrows indicate compound curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752568" httpUri="https://zenodo.org/record/4752568/files/figure.png" pageId="3" pageNumber="404">Fig. 27</figureCitation>
). Proximal arm of tegular apophysis with much less prominent basal projection and a weakly twisted tip (
<figureCitation id="13192A501900BE2CFC286D27FC329E13" box="[901,1014,562,584]" captionStart="FIGURES 27–29" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[162,1415,184,1917]" captionTargetId="figure-27@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURES 27–29. Cybaeus opulentus spec. nov., male from Riverton, California, left palpus. 27 Patella and tibia, dorsal. 28–29 Genital bulb (28 ventral, 29 retrolateral). PA—proximal arm of tegular apophysis. Unlabelled arrows indicate compound curve of dorsal margin of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752568" httpUri="https://zenodo.org/record/4752568/files/figure.png" pageId="3" pageNumber="404">Figs 28–29</figureCitation>
). Copulatory ducts and spermathecal stalks short (
<figureCitation id="13192A501900BE2CFEB26D5AFE489E3D" box="[287,396,591,614]" captionStart="FIGURES 30–32" captionStartId="13.[151,264,1111,1135]" captionTargetBox="[156,1436,186,1087]" captionTargetId="figure-17@13.[151,1436,181,1087]" captionTargetPageId="13" captionText="FIGURES 30–32. Cybaeus opulentus spec. nov., female from Riverton, California, copulatory organ. 30 Epigynum, ventral. 31–32 Vulva (31 ventral, 32 dorsal). BG—Bennett’s gland, CD—copulatory duct, HS—head of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4752572" httpUri="https://zenodo.org/record/4752572/files/figure.png" pageId="3" pageNumber="404">Figs 31–32</figureCitation>
)...............................................................
<taxonomicName id="4C224D561900BE2CFB2B6D5AFAF19E3E" ID-CoL="98YZT" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2021" box="[1158,1333,591,613]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="opulentus" status="sp. nov.">
<emphasis id="B956EAC71900BE2CFB2B6D5AFB279E3F" box="[1158,1251,591,612]" italics="true" pageId="3" pageNumber="404">opulentus</emphasis>
Bennett
</taxonomicName>
<emphasis id="B956EAC71900BE2CFA976D45FA589E3E" bold="true" box="[1338,1436,592,613]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="A26557BC1900BE2CFA976D45FA589E3E" box="[1338,1436,592,613]" pageId="3" pageNumber="404" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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