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<document id="D0AC227D3E1DAD43A3F182FE701E90D5" ID-CLB-Dataset="39090" ID-DOI="10.11646/zootaxa.4109.5.3" ID-GBIF-Dataset="9da062a3-576b-4c79-8535-6ff59ccaf5b4" ID-ISSN="1175-5326" ID-Zenodo-Dep="264020" ID-ZooBank="FA042AA5-A18F-467C-AE31-1CAF47E807F1" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1462950601769" checkinUser="plazi" docAuthor="Komai, Tomoyuki &amp; Higashiji, Takuo" docDate="2016" docId="038B879CFFBE6113EAC7FDAAEAA9B38E" docLanguage="en" docName="zootaxa.4109.5.3.pdf.imf" docOrigin="Zootaxa 4109 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Munida pollioculus Komai &amp; Higashiji, 2016, n. sp." docType="treatment" docVersion="10" lastPageNumber="547" masterDocId="FFB2FFE4FFBF6116EA50FFD7EB48B516" masterDocTitle="New records of the squat lobster genus Munida Leach, 1820 (Crustacea: Decapoda: Anomura: Munididae) from deep-water off Okinawa Islands, Ryukyu Islands, Japan, with description of a new species" masterLastPageNumber="554" masterPageNumber="542" pageNumber="543" updateTime="1698662464318" updateUser="ExternalLinkService">
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<mods:title id="FF71EC4FD8DF21A4B522ED14FED2E843">New records of the squat lobster genus Munida Leach, 1820 (Crustacea: Decapoda: Anomura: Munididae) from deep-water off Okinawa Islands, Ryukyu Islands, Japan, with description of a new species</mods:title>
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<mods:namePart id="212075404D3B6C7F869D2DC43A1B4EF6">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="51511A8EF7DD57E068EE6D1C6EC094DD">Higashiji, Takuo</mods:namePart>
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<mods:title id="E75D82D06A03AC8ACC0F8556921648B0">Zootaxa</mods:title>
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<mods:date id="0F25B43663ADD86D34270E58779D1784">2016</mods:date>
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<treatment id="038B879CFFBE6113EAC7FDAAEAA9B38E" ID-DOI="http://doi.org/10.5281/zenodo.6087277" ID-GBIF-Taxon="120677326" ID-Zenodo-Dep="6087277" LSID="urn:lsid:plazi:treatment:038B879CFFBE6113EAC7FDAAEAA9B38E" httpUri="http://treatment.plazi.org/id/038B879CFFBE6113EAC7FDAAEAA9B38E" lastPageId="5" lastPageNumber="547" pageId="1" pageNumber="543">
<subSubSection id="C3386501FFBE6117EAC7FDAAEA43B7CA" pageId="1" pageNumber="543" type="nomenclature">
<paragraph id="8B9D368AFFBE6117EAC7FDAAEA85B781" blockId="1.[151,691,637,733]" box="[151,461,637,663]" pageId="1" pageNumber="543">
<heading id="D0D581E6FFBE6117EAC7FDAAEA85B781" bold="true" box="[151,461,637,663]" fontSize="11" level="1" pageId="1" pageNumber="543" reason="1">
<emphasis id="B956EA98FFBE6117EAC7FDAAEA85B781" bold="true" box="[151,461,637,663]" pageId="1" pageNumber="543">
<taxonomicName id="4C224D09FFBE6117EAC7FDAAEACBB781" box="[151,387,637,663]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="543" phylum="Arthropoda" rank="species" species="pollioculus" status="sp. nov.">
<emphasis id="B956EA98FFBE6117EAC7FDAAEACBB781" bold="true" box="[151,387,637,663]" italics="true" pageId="1" pageNumber="543">Munida pollioculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557E3FFBE6117EBDBFDAAEA85B781" box="[395,461,637,663]" pageId="1" pageNumber="543" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8B9D368AFFBE6117EAC7FD77EA43B7CA" blockId="1.[151,691,637,733]" pageId="1" pageNumber="543">
[New Japanese name: Chihiro-chu-koshiori-ebi] (
<figureCitation id="13192A0FFFBE6117EACFFD13EBADB7CA" box="[159,229,708,733]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="1" pageNumber="543">Figs 1</figureCitation>
,
<figureCitation id="13192A0FFFBE6117EAA3FD13EA4AB7CA" box="[243,258,708,732]" captionStart="FIGURE 2" captionStartId="3.[151,250,1602,1624]" captionTargetBox="[166,1425,206,1572]" captionTargetId="figure@3.[151,1436,193,1581]" captionTargetPageId="3" captionText="FIGURE 2. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, left third maxilliped, lateral view (setae omitted); right cheliped, merus, ventral view (setae omitted); B, cheliped ischium and merus, ventral view (setae omitted); C, same, carpus and chela, ventral view; D, right second pereopod, lateral view; E, same, dactylus and distal part of propodus, lateral view; F, right third pereopod, lateral view (setae omitted); G, right fourth pereopod, lateral view (setae omitted). Scale bars: B D, F, G = 5 mm; A = 2 mm; E = 1 mm." httpUri="https://zenodo.org/record/264022/files/figure.png" pageId="1" pageNumber="543">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3386501FFBE6117EAC7FCDBEA1EB67A" pageId="1" pageNumber="543" type="materials_examined">
<paragraph id="8B9D368AFFBE6117EAC7FCDBEA1EB67A" blockId="1.[151,1437,780,2029]" pageId="1" pageNumber="543">
<emphasis id="B956EA98FFBE6117EAC7FCDBEA32B633" bold="true" box="[151,378,780,805]" pageId="1" pageNumber="543">Material examined</emphasis>
.
<typeStatus id="54998828FFBE6117EBD9FCDBEABEB633" box="[393,502,780,805]" pageId="1" pageNumber="543" type="holotype">Holotype</typeStatus>
: East of Kunigami Village, Okinawa Island,
<geoCoordinate id="EE16504DFFBE6117E9A2FCDBEFCFB633" box="[1010,1159,780,805]" direction="north" orientation="latitude" pageId="1" pageNumber="543" precision="1" value="26.782583">26°46.955N</geoCoordinate>
,
<geoCoordinate id="EE16504DFFBE6117EEC3FCDBEE7BB632" box="[1171,1331,780,804]" direction="east" orientation="longitude" pageId="1" pageNumber="543" precision="1" value="128.51569">128°30.941E</geoCoordinate>
,
<quantity id="4CDA9B6FFFBE6117EF6FFCDBEEDFB632" box="[1343,1431,780,804]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="543" unit="m" value="1000.0">1000 m</quantity>
,
<date id="FF9C104AFFBE6117EAC7FCF8EA3FB65E" box="[151,375,815,840]" pageId="1" pageNumber="543" value="2005-09-15">15 September 2005</date>
, bait trap, coll. A. Kaneko &amp; T. Higashiji,
<specimenCount id="9D24FD03FFBE6117E932FCE7E88DB65E" box="[866,965,815,840]" pageId="1" pageNumber="543" type="female">1 female</specimenCount>
(cl
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), infested by rhizocephalan, CBM-ZC 11348.
</paragraph>
</subSubSection>
<subSubSection id="C3386501FFBE6113EA97FCA0E96EB72B" lastPageId="5" lastPageNumber="547" pageId="1" pageNumber="543" type="description">
<paragraph id="8B9D368AFFBE6117EA97FCA0EA01B073" blockId="1.[151,1437,780,2029]" pageId="1" pageNumber="543">
<emphasis id="B956EA98FFBE6117EA97FCA0EA19B686" bold="true" box="[199,337,887,912]" pageId="1" pageNumber="543">Description</emphasis>
. Carapace (excluding rostrum) (
<figureCitation id="13192A0FFFBE6117E8EFFCA0E9B6B686" box="[703,766,887,912]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="1" pageNumber="543">Fig. 1</figureCitation>
A) 1.2 times as long as wide. Dorsal surface gently convex transversely; main transverse ridges mostly uninterrupted medially; few secondary transverse striae intervening main ridges on anterior part (anterior to cervical groove); most ridges and striae with row of short setae. Gastric region slightly elevated, with 5 pairs of epigastric spines (second pair just behind supraocular spines strongest, 2 lateral pairs obsolescent) plus 1 scale-like median tubercle; postrostral carina low, obsolescent. Cervical groove distinct. Parahepatic, branchial dorsal and postcervical spines present. Anterior part of branchial region between cervical groove and transverse groove with some short, scale-like ridges; posterior part of branchial region with 6 main transverse ridges (excluding posterodorsal ridge) and a few short secondary striae laterally between main ridges. Cardiac region with 3 main transverse ridges. Intestinal region with pair of short striae, but medially without ridges or stria; posterodorsal ridge distinct, slightly convex medially, with 1 secondary stria. Frontal margins slightly oblique. Anterolateral spines each located at anterolateral angle, strong, slightly overreaching level of sinus between rostrum and supraocular spines. Second marginal spine anterior to cervical groove also strong, though distinctly shorter than anterolateral spine. Branchial margins each with 5 small spines decreasing in size posteriorly.
</paragraph>
<paragraph id="8B9D368AFFBE6117EA97FAA6EFA6B0C6" blockId="1.[151,1437,780,2029]" pageId="1" pageNumber="543">
Rostrum (
<figureCitation id="13192A0FFFBE6117EB6BFAB8EA35B09E" box="[315,381,1391,1416]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="1" pageNumber="543">Fig. 1</figureCitation>
A) spiniform, about 0.4 times as long as carapace, slightly sinuous in lateral view; lateral margins with minute denticles. Supraocular spines about 0.3 length of rostrum, slightly diverging distally and slightly ascending in lateral view. Inner orbital spine curved laterally, clearly visible in dorsal view.
</paragraph>
<paragraph id="8B9D368AFFBE6117EA97FA0BE818B30E" blockId="1.[151,1437,780,2029]" pageId="1" pageNumber="543">
Pterygostomial flap bluntly pointed anteriorly with minute granules (
<figureCitation id="13192A0FFFBE6117E9AFFA0BEF0DB0E2" box="[1023,1093,1500,1525]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="1" pageNumber="543">Fig. 1</figureCitation>
E); anterior margin minutely granulate: lateral face rugose with obliquely transverse ridges.
</paragraph>
<paragraph id="8B9D368AFFBE6117EA97F9F3E895B3DA" blockId="1.[151,1437,780,2029]" pageId="1" pageNumber="543">
Thoracic sternite 3 (
<figureCitation id="13192A0FFFBE6117EBFCF9F3EAA4B32A" box="[428,492,1572,1597]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="1" pageNumber="543">Fig. 1</figureCitation>
B) 1.3 times wider than long, distinctly separated from sternite 4, much wider than anterior width of sternite 4; anterior margin faintly granulate, with shallow, V-shaped median notch; anterolateral angles pointed, directed anterolaterally. Sternite 4 with transverse ridge adjacent to anterior margin; ventral surface shallowly concave medially, with paired short scale-like ridge adjacent to midline. Sternites 57 smooth. Transverse ridges dividing sternites nearly smooth, with sparse short setae.
</paragraph>
<paragraph id="8B9D368AFFBE6117EA97F900E919B2FB" blockId="1.[151,1437,780,2029]" pageId="1" pageNumber="543">
Pleomere 2 (
<figureCitation id="13192A0FFFBE6117EB08F900EAD0B3E6" box="[344,408,1751,1776]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="1" pageNumber="543">Fig. 1</figureCitation>
A) with 1 distinct transverse ridge on tergum; anterior ridge with 4 pairs of strong spines. Pleomere 3 (
<figureCitation id="13192A0FFFBE6117EB74F92BEA2AB202" box="[292,354,1788,1813]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="1" pageNumber="543">Fig. 1</figureCitation>
A) with 1 medially interrupted stria and 1 transverse ridge on tergum, unarmed on anterior ridge. Fourth pleomere with 1 medially interrupted stria and 1 transverse ridge on tergum, anterior ridge, unarmed. Pleura of pleomeres 24 each with scale-like or short ridges. Pleomere 5 with 2 transverse ridges on tergum. Pleomere 6 (
<figureCitation id="13192A0FFFBE6117EACFF8B0EBA8B296" box="[159,224,1895,1920]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="1" pageNumber="543">Fig. 1</figureCitation>
C) with 1 laterally interrupted stria anterior to midlength and broadly W-shaped groove on posterior half, otherwise smooth; posterior margin tri-lobed. All ridges or striae on pleomeres with fringe of short setae. Telson (
<figureCitation id="13192A0FFFBE6117EACFF867EBACB2DE" box="[159,228,1968,1993]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="1" pageNumber="543">Fig. 1</figureCitation>
D) 1.2 times as wide as long, incompletely divided in 7 plates; several scale-like ridges present on posterolateral and posteromesial plates.
</paragraph>
<caption id="DF5D6602FFBD6114EAC7F8F5E80CB2A4" httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="2" pageNumber="544" targetBox="[166,1415,209,1791]" targetPageId="2">
<paragraph id="8B9D368AFFBD6114EAC7F8F5E80CB2A4" blockId="2.[151,1436,1826,1970]" pageId="2" pageNumber="544">
<emphasis id="B956EA98FFBD6114EAC7F8F5EA58B221" bold="true" box="[151,272,1826,1848]" pageId="2" pageNumber="544">FIGURE 1</emphasis>
.
<taxonomicName id="4C224D09FFBD6114EB70F8F5EAA2B22E" box="[288,490,1826,1848]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="544" phylum="Arthropoda" rank="species" species="pollioculus" status="sp. nov.">
<emphasis id="B956EA98FFBD6114EB70F8F5EAA2B22E" box="[288,490,1826,1848]" italics="true" pageId="2" pageNumber="544">Munida pollioculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557E3FFBD6114EBA3F8F5E978B221" box="[499,560,1826,1847]" pageId="2" pageNumber="544" rank="species">
<emphasis id="B956EA98FFBD6114EBA3F8F5E978B221" bold="true" box="[499,560,1826,1847]" pageId="2" pageNumber="544">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; BE = 2 mm.
</paragraph>
</caption>
<caption id="DF5D6602FFBC6115EAC7F995E851B3C4" httpUri="https://zenodo.org/record/264022/files/figure.png" pageId="3" pageNumber="545" targetBox="[166,1425,206,1572]" targetPageId="3">
<paragraph id="8B9D368AFFBC6115EAC7F995E851B3C4" blockId="3.[151,1436,1602,1746]" pageId="3" pageNumber="545">
<emphasis id="B956EA98FFBC6115EAC7F995EA46B341" bold="true" box="[151,270,1602,1624]" pageId="3" pageNumber="545">FIGURE 2</emphasis>
.
<taxonomicName id="4C224D09FFBC6115EB4BF995EAAAB34E" box="[283,482,1602,1624]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="545" phylum="Arthropoda" rank="species" species="pollioculus" status="sp. nov.">
<emphasis id="B956EA98FFBC6115EB4BF995EAAAB34E" box="[283,482,1602,1624]" italics="true" pageId="3" pageNumber="545">Munida pollioculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557E3FFBC6115EBB9F995E96BB341" box="[489,547,1602,1623]" pageId="3" pageNumber="545" rank="species">
<emphasis id="B956EA98FFBC6115EBB9F995E96BB341" bold="true" box="[489,547,1602,1623]" pageId="3" pageNumber="545">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, female (cl 16.9 mm), CBM-ZC 11348. A, left third maxilliped, lateral view (setae omitted); right cheliped, merus, ventral view (setae omitted); B, cheliped ischium and merus, ventral view (setae omitted); C, same, carpus and chela, ventral view; D, right second pereopod, lateral view; E, same, dactylus and distal part of propodus, lateral view; F, right third pereopod, lateral view (setae omitted); G, right fourth pereopod, lateral view (setae omitted). Scale bars: BD, F, G = 5 mm; A = 2 mm; E = 1 mm.
</paragraph>
</caption>
<paragraph id="8B9D368AFFBC6115EA97F92CEE5DB220" blockId="3.[151,1436,1786,2027]" pageId="3" pageNumber="545">
Eyes (
<figureCitation id="13192A0FFFBC6115EB40F92DEA1AB204" box="[272,338,1786,1811]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="3" pageNumber="545">Fig. 1</figureCitation>
A) small. Cornea slightly longer than eyestalk, not dilated, corneal width slightly greater than width between rostrum and supraocular spine and 0.13 cl. Eyestalk widened distally; eyelashes absent.
</paragraph>
<paragraph id="8B9D368AFFBC6115EA97F895E98FB2FC" blockId="3.[151,1436,1786,2027]" pageId="3" pageNumber="545">
Basal article of antennular peduncle (
<figureCitation id="13192A0FFFBC6115E839F895E9E0B24C" box="[617,680,1858,1883]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="3" pageNumber="545">Fig. 1</figureCitation>
A, E) moderately stout, length excluding distal spines 1.8 of width; distal spines unequal with mesial spine much shorter than lateral; 2 lateral spines present, distal spine located proximal to base of distolateral spine, directed somewhat upward, extending as far as distolateral spine, proximal spine located slightly distal to midlength of article; statocyst lobe weakly inflated, ventral surface rounded, with scattered minute tubercles and tufts of short setae.
</paragraph>
<paragraph id="8B9D368AFFBB6112EA97FF40EE00B456" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="546">
Antennal peduncle (
<figureCitation id="13192A0FFFBB6112EBE1FF40EABCB5A6" box="[433,500,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="4" pageNumber="546">Fig. 1</figureCitation>
A, E) moderately stout, overreaching distal corneal margins by length of fourth article. First article with moderately long distomesial spine overreaching midlength of second article; distolateral angle unarmed. Second article with distomesial spine reaching distal end of fourth article, distolateral spine much shorter, slightly falling short of distal end of third article; mesial and lateral margins with few minute tubercles. Third article with minute spine at distomesial angle, unarmed on distolateral angle. Fourth article unarmed.
</paragraph>
<paragraph id="8B9D368AFFBB6112EA97FE9BEA4CB70E" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="546">
Third maxilliped (
<figureCitation id="13192A0FFFBB6112EBC9FE9BEA97B472" box="[409,479,332,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1602,1624]" captionTargetBox="[166,1425,206,1572]" captionTargetId="figure@3.[151,1436,193,1581]" captionTargetPageId="3" captionText="FIGURE 2. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, left third maxilliped, lateral view (setae omitted); right cheliped, merus, ventral view (setae omitted); B, cheliped ischium and merus, ventral view (setae omitted); C, same, carpus and chela, ventral view; D, right second pereopod, lateral view; E, same, dactylus and distal part of propodus, lateral view; F, right third pereopod, lateral view (setae omitted); G, right fourth pereopod, lateral view (setae omitted). Scale bars: B D, F, G = 5 mm; A = 2 mm; E = 1 mm." httpUri="https://zenodo.org/record/264022/files/figure.png" pageId="4" pageNumber="546">Fig. 2</figureCitation>
A) moderately slender. Ischium distinctly longer than merus, with tiny ventrodistal spine, extensor distal angle unarmed; lateral surface with some scale-like tubercles. Merus not markedly narrowed distally, flexor margin with 2 greatly unequal spines, distal spine tiny, proximal spine prominent, arising slightly proximal to midlength; extensor distal margin unarmed; extensor margin and lateral face with several small tubercles. Exopod narrowing distally, distinctly overreaching distal margin of merus; lateral surface with minute tubercles.
</paragraph>
<paragraph id="8B9D368AFFBB6112EA97FDF3EED8B01A" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="546">
Cheliped (first pereopod) (
<figureCitation id="13192A0FFFBB6112EBA1FDF3E97EB72A" box="[497,566,548,573]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="4" pageNumber="546">Figs 1</figureCitation>
F; 2B, C) about 2.4 times as long as carapace. Merus with row of 8 spines on dorsal surface laterally (spines increasing in size distally, distolateral spine diverging) and 2 prominent spines distomesially (distal spine stronger than distolateral spine); lateral face with numerous scale-like tubercles becoming larger ventrally and with strong spine at ventrodistal angle; mesial face with 3 well-spaced strong spines adjacent to ventral margin (distalmost spine at ventromesial distal angle) and scattered tiny tubercles; ventral surface with scattered tiny to small scale-like tubercles, distal edge of these tubercles usually multidenticulate, bearing short setae. Carpus 1.2 times as long as palm, 2.3 times longer than distal width; dorsolateral margin with 4 spines noticeably increasing in size distally (distalmost spine located at dorsolateral distal angle), dorsomesial margin with 5 spines (second spine strongest, somewhat diverging); lateral face with several scale-like tubercles (distal edges multidenticulate) and 1 small subterminal spine; mesial face with scattered scale-like tubercles and 1 strong subdistal spine; ventral surface also with scattered scale-like tubercles and small spine at ventrodistal angle. Palm 2.6 times longer than wide; dorsal surface with row of 3 moderately strong spines on proximal 0.6 of midline, dorsolateral margin 4 spines increasing in size distally, dorsomesial margin with 3 spines also increasing in size distally, articular condyle with small spine; mesial face without conspicuous spines; surfaces with scale-like tubercles of various size. Fixed finger straight, terminating in sharp claw; lateral margin with 2 small subdistal spines, 1 moderately strong spine somewhat proximal to midlength and row of tiny, low protuberances; cutting edge straight, with row of minute, rounded denticles interspersing tiny triangular teeth. Dactylus 1.1 times as long as palm, without conspicuous spines, terminating in sharp, curved claw crossing tip of fixed finger; mesial margin with row of low tubercles; cutting edge with row of minute, rounded or triangular denticles over entire length, no hiatus between dactylus and fixed finger. Setae primarily plumose (no iridescent setae present), most arising from tubercles or bases of spines, length variable, particularly long and numerous on mesial faces of merus and carpus.
</paragraph>
<paragraph id="8B9D368AFFBB6113EA97FAC0EE03B456" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1437,151,1688]" lastPageId="5" lastPageNumber="547" pageId="4" pageNumber="546">
Ambulatory legs (second to fourth pereopods) moderately long and slender, second and third pereopods subequal in length, fourth pereopod shorter than preceding pereopods. Second pereopod (
<figureCitation id="13192A0FFFBB6112EE2EFAEBEF89B042" box="[1150,1217,1340,1365]" captionStart="FIGURE 2" captionStartId="3.[151,250,1602,1624]" captionTargetBox="[166,1425,206,1572]" captionTargetId="figure@3.[151,1436,193,1581]" captionTargetPageId="3" captionText="FIGURE 2. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, left third maxilliped, lateral view (setae omitted); right cheliped, merus, ventral view (setae omitted); B, cheliped ischium and merus, ventral view (setae omitted); C, same, carpus and chela, ventral view; D, right second pereopod, lateral view; E, same, dactylus and distal part of propodus, lateral view; F, right third pereopod, lateral view (setae omitted); G, right fourth pereopod, lateral view (setae omitted). Scale bars: B D, F, G = 5 mm; A = 2 mm; E = 1 mm." httpUri="https://zenodo.org/record/264022/files/figure.png" pageId="4" pageNumber="546">Fig. 2</figureCitation>
D), when anteriorly extended, falling slightly short of tip of rostrum by tip of mero-carpal articulation, about twice of carapace length; merus 1.4 times as long as carapace, extensor margin with row of numerous setae and row of 10 (left) or 13 (right) spines noticeably increasing in length distally, flexor margin with 1 strong distolateral spine followed by 3 smaller spines and scale-like tubercles, lateral face with scattered scale-like tubercles and short setae, flexor distal margin with 1 small subterminal spine on mesial side; carpus with prominent extensor distal spine followed by 1 or 2 small spines on extensor margin, flexor distal margin produced in spine, lateral face with weak longitudinal ridge dorsally and few small scale-like tubercles ventrally; propodus with sparse tufts or individual long setae on extensor margin, lateral surface with some low, scale-like tubercles ventrally; mesial surface with row of numerous plumose setae adjacent to extensor margin, flexor margin with row of 9 (left) or 10 (right) evenly spaced movable spines, all but distalmost spine arising from low protuberances, distalmost spine supported by contiguous acute fixed spine, ventromesial distal margin with 1 movable spine; dactylus (
<figureCitation id="13192A0FFFBB6112E9EEF910EF48B3F6" box="[958,1024,1735,1760]" captionStart="FIGURE 2" captionStartId="3.[151,250,1602,1624]" captionTargetBox="[166,1425,206,1572]" captionTargetId="figure@3.[151,1436,193,1581]" captionTargetPageId="3" captionText="FIGURE 2. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, left third maxilliped, lateral view (setae omitted); right cheliped, merus, ventral view (setae omitted); B, cheliped ischium and merus, ventral view (setae omitted); C, same, carpus and chela, ventral view; D, right second pereopod, lateral view; E, same, dactylus and distal part of propodus, lateral view; F, right third pereopod, lateral view (setae omitted); G, right fourth pereopod, lateral view (setae omitted). Scale bars: B D, F, G = 5 mm; A = 2 mm; E = 1 mm." httpUri="https://zenodo.org/record/264022/files/figure.png" pageId="4" pageNumber="546">Fig. 2</figureCitation>
E) 0.6 times as long as propodus and 5 times longer than greatest width, slightly curved distally, bearing sparse short to long stiff setae, flexor margin mostly straight, with 9 or 10 slender corneous spinules, increasing in length distally, along entire length and 1 very slender subterminal spinule contiguous with unguis. Third pereopod (
<figureCitation id="13192A0FFFBB6112E9FAF8E3E8A5B25A" box="[938,1005,1844,1869]" captionStart="FIGURE 2" captionStartId="3.[151,250,1602,1624]" captionTargetBox="[166,1425,206,1572]" captionTargetId="figure@3.[151,1436,193,1581]" captionTargetPageId="3" captionText="FIGURE 2. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, left third maxilliped, lateral view (setae omitted); right cheliped, merus, ventral view (setae omitted); B, cheliped ischium and merus, ventral view (setae omitted); C, same, carpus and chela, ventral view; D, right second pereopod, lateral view; E, same, dactylus and distal part of propodus, lateral view; F, right third pereopod, lateral view (setae omitted); G, right fourth pereopod, lateral view (setae omitted). Scale bars: B D, F, G = 5 mm; A = 2 mm; E = 1 mm." httpUri="https://zenodo.org/record/264022/files/figure.png" pageId="4" pageNumber="546">Fig. 2</figureCitation>
F) similar to second pereopod, though shorter; merus bearing row of 8 (right) or 10 (left) spines on extensor margin, flexor margin with 1 moderately strong distolateral spine followed by 3 or 4 smaller spine and scale-like tubercles or ridges; carpus with 2 spines on extensor margin; propodus with 8 or 9 movable spines on flexor margin; dactylus with 9 spinules on flexor margin (except for subterminal contiguous spinule). Fourth pereopod (
<figureCitation id="13192A0FFFBB6112E928F813E8F7B2CA" box="[888,959,1988,2013]" captionStart="FIGURE 2" captionStartId="3.[151,250,1602,1624]" captionTargetBox="[166,1425,206,1572]" captionTargetId="figure@3.[151,1436,193,1581]" captionTargetPageId="3" captionText="FIGURE 2. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, left third maxilliped, lateral view (setae omitted); right cheliped, merus, ventral view (setae omitted); B, cheliped ischium and merus, ventral view (setae omitted); C, same, carpus and chela, ventral view; D, right second pereopod, lateral view; E, same, dactylus and distal part of propodus, lateral view; F, right third pereopod, lateral view (setae omitted); G, right fourth pereopod, lateral view (setae omitted). Scale bars: B D, F, G = 5 mm; A = 2 mm; E = 1 mm." httpUri="https://zenodo.org/record/264022/files/figure.png" pageId="4" pageNumber="546">Fig. 2</figureCitation>
G) falling short of anterolateral angle of carapace by mero-carpal articulation; merus 0.8 length of that of second pereopod, extensor margin with small distal spine and some tiny spines or spinulose tubercles proximally, flexor margin with strong distolateral spine following by 1 small spine and scale-like ridges or tubercles, flexor mesial margin with 1 small subdistal spine; carpus with extensor distal spine and flexor distal spine, distinctly smaller than those of preceding pereopods; propodus with 6 or 7 movable spines on flexor margin; dactylus with 9 corneous spinules on flexor margin.
</paragraph>
<paragraph id="8B9D368AFFBA6113EA97FE9AE807B473" blockId="5.[151,1437,151,1688]" box="[199,847,332,357]" pageId="5" pageNumber="547">Merus of fifth pereopod slightly rugose on lateral surface.</paragraph>
<paragraph id="8B9D368AFFBA6113EA97FEB8EFB5B70E" blockId="5.[151,1437,151,1688]" pageId="5" pageNumber="547">
Uropodal exopod with row of minute movable spinules on nearly straight lateral margin, distal margin truncate, also with row of closely spaced movable spinules; outer surface with sets of 14 minute movable spinules laterally. Endopod (
<figureCitation id="13192A0FFFBA6113EB27FE60EAFFB4C6" box="[375,439,439,464]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="5" pageNumber="547">Fig. 1</figureCitation>
D) with row of scale-like tubercles and distal row of minute movable spinules distally, extending onto truncate distal margin; outer surface with scale-like ridges or tubercles, each bearing 14 minute movable spines on distal edge. Uropodal protopod (
<figureCitation id="13192A0FFFBA6113E88AFE28E850B70E" box="[730,792,511,536]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="5" pageNumber="547">Fig. 1</figureCitation>
D) with small spine on inner distal margin.
</paragraph>
<paragraph id="8B9D368AFFBA6113EA97FDF3E96EB72B" blockId="5.[151,1437,151,1688]" box="[199,550,548,573]" pageId="5" pageNumber="547">
<emphasis id="B956EA98FFBA6113EA97FDF3EADAB72B" bold="true" box="[199,402,548,573]" pageId="5" pageNumber="547">Coloration in life</emphasis>
. Not known.
</paragraph>
</subSubSection>
<subSubSection id="C3386501FFBA6113EA97FD90EFF0B749" box="[199,1208,583,608]" pageId="5" pageNumber="547" type="distribution">
<paragraph id="8B9D368AFFBA6113EA97FD90EFF0B749" blockId="5.[151,1437,151,1688]" box="[199,1208,583,608]" pageId="5" pageNumber="547">
<emphasis id="B956EA98FFBA6113EA97FD90EA11B776" bold="true" box="[199,345,583,608]" pageId="5" pageNumber="547">Distribution</emphasis>
. Known only from the
<typeStatus id="54998828FFBA6113E830FD9FE9D8B776" box="[608,656,584,608]" pageId="5" pageNumber="547">type</typeStatus>
locality, off Okinawa Island, at depth of
<quantity id="4CDA9B6FFFBA6113EE0CFD9FEFFCB749" box="[1116,1204,584,608]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="547" unit="m" value="1000.0">1000 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3386501FFBA6113EA97FDBBE885B346" pageId="5" pageNumber="547" type="discussion">
<paragraph id="8B9D368AFFBA6113EA97FDBBE88FB1D2" blockId="5.[151,1437,151,1688]" pageId="5" pageNumber="547">
<emphasis id="B956EA98FFBA6113EA97FDBBEA7CB793" bold="true" box="[199,308,620,645]" pageId="5" pageNumber="547">Remarks</emphasis>
. The present new species resembles closely
<taxonomicName id="4C224D09FFBA6113E913FDBAE8E7B792" box="[835,943,620,644]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="clevai">
<emphasis id="B956EA98FFBA6113E913FDBAE8E7B792" box="[835,943,620,644]" italics="true" pageId="5" pageNumber="547">M. clevai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C224D09FFBA6113E9BDFDBAEF38B792" box="[1005,1136,621,644]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="microps">
<emphasis id="B956EA98FFBA6113E9BDFDBAEF38B792" box="[1005,1136,621,644]" italics="true" pageId="5" pageNumber="547">M. microps</emphasis>
</taxonomicName>
, sharing many diagnostic characters, e.g., (1) branchial margin bearing five spines, (2) pleomere 2 with row of spines on anterior ridge while pleomere 3 unarmed, (3) no granules on thoracic sternites 6 and 7, (4) eye small, (5) basal article of antennular peduncle with distomesial spine much shorter than distolateral spine, (6) distomesial spine of second article of antennal peduncle reaching distal end of fourth article, (7) fixed finger of cheliped armed with one or more spines on lateral margin in addition to subterminal spines, and (8) dactylus of second pereopod with accessory spines along entire length of flexor margin.
<taxonomicName id="4C224D09FFBA6113E86EFC93E9BEB64A" box="[574,758,836,860]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="microps">
<emphasis id="B956EA98FFBA6113E86EFC93E9BEB64A" box="[574,758,836,860]" italics="true" pageId="5" pageNumber="547">Munida microps</emphasis>
</taxonomicName>
has been recorded from rather wide areas in the Indo-West Pacific (e.g.,
<bibRefCitation id="EFB34B7BFFBA6113EB7BFCB0EAF5B696" author="Alcock" box="[299,445,871,896]" pageId="5" pageNumber="547" refString="Alcock, A. (1894) Natural history notes from H. M. Royal Indian Marine Survey Steamer &quot; Investigator &quot;, commander R. F. Hoskin, R. N., commanding. - Series II, No. 1. On the results of deep-sea dredging during the season of 1890 - 91 (continued). Annals and Magazine of Natural History, Series 6, 13, 321 - 334." type="journal article" year="1894">Alcock 1894</bibRefCitation>
,
<bibRefCitation id="EFB34B7BFFBA6113EB98FCBFE94BB696" author="Alcock" box="[456,515,872,896]" pageId="5" pageNumber="547" refString="Alcock, A. (1901) A descriptive catalogue of the Indian deep-sea Crustacea Decapoda Macrura and Anomala, in the Indian Museum. Being a revised account of the deep-sea species collected by the Royal Indian Marine Survey Ship Investigator. Trustees of the Indian Museum, Calcutta, 286 pp., 3 pls." type="book" year="1901">1901</bibRefCitation>
;
<bibRefCitation id="EFB34B7BFFBA6113E85EFCBFE84CB696" author="Kemp" box="[526,772,871,896]" pageId="5" pageNumber="547" refString="Kemp, S. W. &amp; Sewell, R. B. S. (1912) Notes on Decapoda in the Indian Museum. III. The species obtained by R. I. M. S. S. ' Investigator' during the survey season 1910 - 11. Records of the Indian Museum, 7, 15 - 32, pl. 1." type="journal article" year="1912">Kemp &amp; Sewell 1912</bibRefCitation>
;
<bibRefCitation id="EFB34B7BFFBA6113E95FFCB0E8EDB696" author="Tirmizi" box="[783,933,871,896]" pageId="5" pageNumber="547" refString="Tirmizi, N. M. (1966) Crustacea: Galatheidae. The John Murray Expedition 1933 - 34. Scientific Reports, 11, 167 - 234." type="journal article" year="1966">Tirmizi 1966</bibRefCitation>
;
<bibRefCitation id="EFB34B7BFFBA6113E9FFFCB0EF33B696" author="Macpherson" box="[943,1147,871,896]" pageId="5" pageNumber="547" refString="Macpherson, E. (1994) Crustacea Decapoda: Studies on the genus Munida Leach, 1820 (Galatheidae) in New Caledonia and adjacent waters with descriptions of 56 new species. In: Crosnier, A. (Ed.), Resultats des Campagnes MUSORSTOM. Vol. 12. Memoires du Museum National d'Histoire Naturelle, Paris, 161, pp. 421 - 569." type="journal article" year="1994">Macpherson 1994</bibRefCitation>
,
<bibRefCitation id="EFB34B7BFFBA6113EED6FCB0EF89B696" author="Macpherson" box="[1158,1217,871,896]" pageId="5" pageNumber="547" refString="Macpherson, E. (1996) Crustacea Decapoda: species of the genera Munida Leach, 1820 and Paramunida Baba, 1988 (Galatheidae) from the seas around the Wallis and Futuna Islands. In: Crosnier, A. (Ed.), Resultats des Campagnes MUSORSTOM. Vol. 15. Memoires du Museum National d'Histoire Naturelle, Paris, 168, pp. 387 - 421." type="journal article" year="1996">1996</bibRefCitation>
,
<bibRefCitation id="EFB34B7BFFBA6113EE9BFCBFEE4EB696" author="Macpherson" box="[1227,1286,872,896]" pageId="5" pageNumber="547" refString="Macpherson, E. (1997) Crustacea Decapoda: species of the genera Agononida Baba &amp; de Saint Laurent, 1996 and Munida Leach, 1820 (Galatheidae) from the KARUBAR cruise. In: Crosnier, A. &amp; Bouchet, P. (Eds.), Resultats des Campagnes MUSORSTOM. Vol. 16. Memoires du Museum National d'Histoire Naturelle, Paris, 172, pp. 597 - 612." type="journal article" year="1997">1997</bibRefCitation>
; Macpherson &amp; de
<bibRefCitation id="EFB34B7BFFBA6113EAB6FC5BEA9AB6B2" author="Saint" box="[230,466,908,933]" pageId="5" pageNumber="547" refString="Macpherson, E. &amp; Saint Laurent, M. de (2002) On the genus Munida Leach, 1820 (Decapoda, Galatheidae) from the western and southern Indian Ocean, with the description of four new species. Crustaceana, 75, 465 - 484. http: // dx. doi. org / 10.1163 / 156854002760095525" type="journal article" year="2002">Saint Laurent 2002</bibRefCitation>
; Macpherson 2004;
<bibRefCitation id="EFB34B7BFFBA6113E89FFC5BE81CB6B3" author="Baba" box="[719,852,908,933]" pageId="5" pageNumber="547" refString="Baba, K. (2005) Deep-sea chirostylid and galatheid crustaceans (Decapoda: Anomura) from the Indo-West Pacific, with a list of species. Galathea Reports, 20, 1 - 317." type="journal article" year="2005">Baba 2005</bibRefCitation>
), but some workers pointed out possibilities of misidentifications in literature or the presence of closely related species under the name (
<bibRefCitation id="EFB34B7BFFBA6113EE93FC78EEDFB6DE" author="Macpherson" box="[1219,1431,943,968]" pageId="5" pageNumber="547" refString="Macpherson, E. (1994) Crustacea Decapoda: Studies on the genus Munida Leach, 1820 (Galatheidae) in New Caledonia and adjacent waters with descriptions of 56 new species. In: Crosnier, A. (Ed.), Resultats des Campagnes MUSORSTOM. Vol. 12. Memoires du Museum National d'Histoire Naturelle, Paris, 161, pp. 421 - 569." type="journal article" year="1994">Macpherson 1994</bibRefCitation>
;
<bibRefCitation id="EFB34B7BFFBA6113EAC7FC03EAD0B6FA" author="Ahyong" box="[151,408,980,1005]" pageId="5" pageNumber="547" refString="Ahyong, S. T. &amp; Poore, G. C. G. (2004) Deep-water Galatheidae (Crustacea: Decapoda: Anomura) from southern and eastern Australia. Zootaxa, 472, 3 - 76." type="journal article" year="2004">Ahyong &amp; Poore 2004</bibRefCitation>
;
<bibRefCitation id="EFB34B7BFFBA6113EBF4FC03E96AB6FB" author="Baba" box="[420,546,980,1005]" pageId="5" pageNumber="547" refString="Baba, K. (2005) Deep-sea chirostylid and galatheid crustaceans (Decapoda: Anomura) from the Indo-West Pacific, with a list of species. Galathea Reports, 20, 1 - 317." type="journal article" year="2005">Baba 2005</bibRefCitation>
). Therefore, we refer to
<bibRefCitation id="EFB34B7BFFBA6113E961FC03E8BFB6FB" author="Baba" box="[817,1015,980,1005]" pageId="5" pageNumber="547" refString="Baba, K. (2005) Deep-sea chirostylid and galatheid crustaceans (Decapoda: Anomura) from the Indo-West Pacific, with a list of species. Galathea Reports, 20, 1 - 317." type="journal article" year="2005">Baba (2005: 116)</bibRefCitation>
, who examined a topotypic specimen from the Andaman Sea, for comparison. The new species differs from
<taxonomicName id="4C224D09FFBA6113E9FAFC2EEF5AB106" box="[938,1042,1016,1040]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="clevai">
<emphasis id="B956EA98FFBA6113E9FAFC2EEF5AB106" box="[938,1042,1016,1040]" italics="true" pageId="5" pageNumber="547">M. clevai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C224D09FFBA6113EE18FC2EEF8EB119" box="[1096,1222,1016,1040]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="microps">
<emphasis id="B956EA98FFBA6113EE18FC2EEF8EB119" box="[1096,1222,1016,1040]" italics="true" pageId="5" pageNumber="547">M. microps</emphasis>
</taxonomicName>
in the absence of a proximal spine on the mesial margin of the cheliped dactylus and the clearly stouter dactylus of the second pereopod (5 times as long as wide versus 10 times as long) (
<bibRefCitation id="EFB34B7BFFBA6113E934FBE8E8ADB14E" author="Baba" box="[868,997,1087,1112]" pageId="5" pageNumber="547" refString="Baba, K. (2005) Deep-sea chirostylid and galatheid crustaceans (Decapoda: Anomura) from the Indo-West Pacific, with a list of species. Galathea Reports, 20, 1 - 317." type="journal article" year="2005">Baba 2005</bibRefCitation>
; Macpherson 2009). Furthermore, the frontal margin of the carapace is more strongly oblique in
<taxonomicName id="4C224D09FFBA6113E970FBB2E8C0B16A" box="[800,904,1124,1148]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="clevai">
<emphasis id="B956EA98FFBA6113E970FBB2E8C0B16A" box="[800,904,1124,1148]" italics="true" pageId="5" pageNumber="547">M. clevai</emphasis>
</taxonomicName>
(cf. Macpherson 2009, fig. 2A) than in the new species (cf.
<figureCitation id="13192A0FFFBA6113EB4BFB50EA1FB1B6" box="[283,343,1159,1184]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="5" pageNumber="547">Fig. 1</figureCitation>
A) and
<taxonomicName id="4C224D09FFBA6113EBFEFB5EE964B189" box="[430,556,1160,1184]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="microps">
<emphasis id="B956EA98FFBA6113EBFEFB5EE964B189" box="[430,556,1160,1184]" italics="true" pageId="5" pageNumber="547">M. microps</emphasis>
</taxonomicName>
. The fixed finger of the cheliped has two or three spines in
<taxonomicName id="4C224D09FFBA6113EE91FB5EEE77B189" box="[1217,1343,1160,1184]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="microps">
<emphasis id="B956EA98FFBA6113EE91FB5EEE77B189" box="[1217,1343,1160,1184]" italics="true" pageId="5" pageNumber="547">M. microps</emphasis>
</taxonomicName>
, instead of only one relatively strong spine in
<taxonomicName id="4C224D09FFBA6113E86AFB7AE993B1D2" box="[570,731,1196,1220]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="pollioculus" status="sp. nov.">
<emphasis id="B956EA98FFBA6113E86AFB7AE993B1D2" box="[570,731,1196,1220]" italics="true" pageId="5" pageNumber="547">M. pollioculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557E3FFBA6113E8B3FB7BE86AB1D3" box="[739,802,1196,1221]" pageId="5" pageNumber="547" rank="species">
<emphasis id="B956EA98FFBA6113E8B3FB7BE86AB1D3" bold="true" box="[739,802,1196,1221]" pageId="5" pageNumber="547">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C224D09FFBA6113E908FB7AE888B1D2" box="[856,960,1196,1220]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="clevai">
<emphasis id="B956EA98FFBA6113E908FB7AE888B1D2" box="[856,960,1196,1220]" italics="true" pageId="5" pageNumber="547">M. clevai</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9D368AFFBA6113EA97FB18E885B346" blockId="5.[151,1437,151,1688]" pageId="5" pageNumber="547">
The new species is also substantially similar to
<taxonomicName id="4C224D09FFBA6113E945FB06EFB5B1FE" authority="Ahyong &amp; Poore, 2004" authorityName="Ahyong &amp; Poore" authorityYear="2004" box="[789,1277,1231,1256]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="endeavourae">
<emphasis id="B956EA98FFBA6113E945FB06E89AB1FE" box="[789,978,1232,1256]" italics="true" pageId="5" pageNumber="547">M. endeavourae</emphasis>
<bibRefCitation id="EFB34B7BFFBA6113E9B1FB18EFB5B1FE" author="Ahyong" box="[993,1277,1231,1256]" pageId="5" pageNumber="547" refString="Ahyong, S. T. &amp; Poore, G. C. G. (2004) Deep-water Galatheidae (Crustacea: Decapoda: Anomura) from southern and eastern Australia. Zootaxa, 472, 3 - 76." type="journal article" year="2004">Ahyong &amp; Poore, 2004</bibRefCitation>
</taxonomicName>
known from southeastern
<collectingCountry id="F335761AFFBA6113EB7CFB23EADEB01B" box="[300,406,1268,1293]" name="Australia" pageId="5" pageNumber="547">Australia</collectingCountry>
, Tasmania,
<collectingCountry id="F335761AFFBA6113E84EFB23E93FB01B" box="[542,631,1268,1293]" name="Norfolk Island" pageId="5" pageNumber="547">Norfolk</collectingCountry>
Ridge, Kermadec Ridge and Bay of Plenty,
<collectingCountry id="F335761AFFBA6113EE28FB22EE58B01B" box="[1144,1296,1268,1293]" name="New Zealand" pageId="5" pageNumber="547">New Zealand</collectingCountry>
(
<bibRefCitation id="EFB34B7BFFBA6113EF71FB23EA53B026" author="Ahyong" pageId="5" pageNumber="547" refString="Ahyong, S. T. &amp; Poore, G. C. G. (2004) Deep-water Galatheidae (Crustacea: Decapoda: Anomura) from southern and eastern Australia. Zootaxa, 472, 3 - 76." type="journal article" year="2004">Ahyong &amp; Poore 2004</bibRefCitation>
;
<bibRefCitation id="EFB34B7BFFBA6113EB76FAC0EA8BB026" author="Ahyong" box="[294,451,1303,1328]" pageId="5" pageNumber="547" refString="Ahyong, S. T. (2007) Decapod Crustacea collected by the NORFANZ Expedition: Galatheidae and Polychelidae. Zootaxa, 1593, 1 - 54." type="journal article" year="2007">Ahyong 2007</bibRefCitation>
; with
<taxonomicName id="4C224D09FFBA6113E857FACEE82FB026" authority="Vereshchaka, 2005" authorityName="Vereshchaka" authorityYear="2005" box="[519,871,1303,1328]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="grieveae">
<emphasis id="B956EA98FFBA6113E857FACEE9C4B039" box="[519,652,1304,1328]" italics="true" pageId="5" pageNumber="547">M. grieveae</emphasis>
Vereshchaka, 2005
</taxonomicName>
as a junior subjective synonym) in the small eyes, armature and ornamentation of the carapace, pleon, antennae and pereopods.
<taxonomicName id="4C224D09FFBA6113EE43FAEBEE4DB042" box="[1043,1285,1340,1364]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="endeavourae">
<emphasis id="B956EA98FFBA6113EE43FAEBEE4DB042" box="[1043,1285,1340,1364]" italics="true" pageId="5" pageNumber="547">Munida endeavourae</emphasis>
</taxonomicName>
was actually misidentified with
<taxonomicName id="4C224D09FFBA6113EB3EFAB6EAA6B061" box="[366,494,1376,1400]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="microps">
<emphasis id="B956EA98FFBA6113EB3EFAB6EAA6B061" box="[366,494,1376,1400]" italics="true" pageId="5" pageNumber="547">M. microps</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB34B7BFFBA6113E84CFA88E9EEB06E" author="Haig" box="[540,678,1375,1400]" pageId="5" pageNumber="547" refString="Haig, J. (1973) Galatheidea (Crustacea, Decapoda, Anomura) collected by the F. I. S. Endeavour. Records of the Australian Museum, 28, 269 - 289. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.28.1973.411" type="journal article" year="1973">Haig (1973)</bibRefCitation>
(
<bibRefCitation id="EFB34B7BFFBA6113E8E7FA88E889B06E" author="Ahyong" box="[695,961,1375,1400]" pageId="5" pageNumber="547" refString="Ahyong, S. T. &amp; Poore, G. C. G. (2004) Deep-water Galatheidae (Crustacea: Decapoda: Anomura) from southern and eastern Australia. Zootaxa, 472, 3 - 76." type="journal article" year="2004">Ahyong &amp; Poore 2004</bibRefCitation>
). Nevertheless,
<taxonomicName id="4C224D09FFBA6113EE26FAB7EE1BB06E" box="[1142,1363,1376,1400]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="pollioculus" status="sp. nov.">
<emphasis id="B956EA98FFBA6113EE26FAB7EE1BB06E" box="[1142,1363,1376,1400]" italics="true" pageId="5" pageNumber="547">Munida pollioculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557E3FFBA6113EF0BFA88EED4B06E" box="[1371,1436,1375,1400]" pageId="5" pageNumber="547" rank="species">
<emphasis id="B956EA98FFBA6113EF0BFA88EED4B06E" bold="true" box="[1371,1436,1375,1400]" pageId="5" pageNumber="547">n. sp.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C224D09FFBA6113EB76FA52EA96B08A" box="[294,478,1412,1436]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="endeavourae">
<emphasis id="B956EA98FFBA6113EB76FA52EA96B08A" box="[294,478,1412,1436]" italics="true" pageId="5" pageNumber="547">M. endeavourae</emphasis>
</taxonomicName>
in many characters, such as relatively stout basal article of antennular peduncle, the shorter distomesial spine on the second article of the antennal peduncle (reaching to the distal end of the fourth article of the antennal peduncle versus clearly overreaching it), the presence of a proximal spine on the cheliped dactylus and the presence of a single strong spine on the lateral margin of the fixed finger of the cheliped, instead of two small spines
<taxonomicName id="4C224D09FFBA6113EB09F9C2E947B33A" box="[345,527,1556,1580]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="endeavourae">
<emphasis id="B956EA98FFBA6113EB09F9C2E947B33A" box="[345,527,1556,1580]" italics="true" pageId="5" pageNumber="547">M. endeavourae</emphasis>
</taxonomicName>
. The cheliped palm seems to have more numerous spines in
<taxonomicName id="4C224D09FFBA6113EEE9F9C2EE27B33A" box="[1209,1391,1556,1580]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="endeavourae">
<emphasis id="B956EA98FFBA6113EEE9F9C2EE27B33A" box="[1209,1391,1556,1580]" italics="true" pageId="5" pageNumber="547">M. endeavourae</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFB34B7BFFBA6113EAC7F9E0EAD1B346" author="Ahyong" box="[151,409,1591,1616]" pageId="5" pageNumber="547" refString="Ahyong, S. T. &amp; Poore, G. C. G. (2004) Deep-water Galatheidae (Crustacea: Decapoda: Anomura) from southern and eastern Australia. Zootaxa, 472, 3 - 76." type="journal article" year="2004">Ahyong &amp; Poore 2004</bibRefCitation>
: fig. 5e) than in
<taxonomicName id="4C224D09FFBA6113E800F9EEE9BAB346" box="[592,754,1592,1616]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="547" phylum="Arthropoda" rank="species" species="pollioculus" status="sp. nov.">
<emphasis id="B956EA98FFBA6113E800F9EEE9BAB346" box="[592,754,1592,1616]" italics="true" pageId="5" pageNumber="547">M. pollioculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557E3FFBA6113E8A8F9E0E87FB346" box="[760,823,1591,1616]" pageId="5" pageNumber="547" rank="species">
<emphasis id="B956EA98FFBA6113E8A8F9E0E87FB346" bold="true" box="[760,823,1591,1616]" pageId="5" pageNumber="547">n. sp.</emphasis>
</taxonomicNameLabel>
(cf.
<figureCitation id="13192A0FFFBA6113E93CF9E0E8E2B346" box="[876,938,1591,1616]" captionStart="FIGURE 1" captionStartId="2.[151,250,1826,1848]" captionTargetBox="[166,1415,209,1791]" captionTargetId="figure@2.[151,1436,193,1805]" captionTargetPageId="2" captionText="FIGURE 1. Munida pollioculus n. sp., holotype, female (cl 16.9 mm), CBM-ZC 11348. A, carapace, pleon and cephalic appendages, dorsal view (setae on carapace ridges and striae omitted; distal 2 articles of antennular peduncles omitted); B, thoracic sternum, ventral view; C, pleomere 6, outer view; D, telson and right uropod, outer view (setae omitted); E, basal article of left antennular peduncle, antennal peduncle and anterior part of pterygostomial flap, ventral view; F, right cheliped, dorsal view (setae omitted). Scale bars: A, F = 5 mm; B E = 2 mm." httpUri="https://zenodo.org/record/264021/files/figure.png" pageId="5" pageNumber="547">Fig. 1</figureCitation>
F).
</paragraph>
</subSubSection>
<subSubSection id="C3386501FFBA6113EA97F98BEAA9B38E" pageId="5" pageNumber="547" type="etymology">
<paragraph id="8B9D368AFFBA6113EA97F98BEAA9B38E" blockId="5.[151,1437,151,1688]" pageId="5" pageNumber="547">
<emphasis id="B956EA98FFBA6113EA97F98BEA0EB363" bold="true" box="[199,326,1628,1653]" pageId="5" pageNumber="547">Etymology</emphasis>
. From the Latin
<emphasis id="B956EA98FFBA6113E855F98BE902B362" box="[517,586,1628,1652]" italics="true" pageId="5" pageNumber="547">pollus</emphasis>
(= small) and
<emphasis id="B956EA98FFBA6113E8BEF98BE87FB362" box="[750,823,1628,1652]" italics="true" pageId="5" pageNumber="547">oculus</emphasis>
(= eye), referring to the small eye in this new species. Used as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>