treatments-xml/data/03/49/87/0349878BDD03A46A02FC0CCFFD09FC17.xml
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<document id="D7E7A15646F2E076FD45D4FDA693CB54" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629691827126" checkinUser="felipe" docAuthor="Chamorro, Maria Lourdes &amp; Holzenthal, Ralph W." docDate="2010" docId="0349878BDD03A46A02FC0CCFFD09FC17" docLanguage="en" docName="zt02582p252.pdf" docOrigin="Zootaxa 2582" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Polyplectropus clauseni , Chamorro-Lacayo &amp; Holzenthal 2004" docType="treatment" docVersion="3" lastPageNumber="85" masterDocId="FF70FFF3DD52A43E026B0979FF8EFFED" masterDocTitle="2582" masterLastPageNumber="252" masterPageNumber="1" pageNumber="82" updateTime="1699096773090" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:namePart id="826DE2263C925921B144BCC09712CC91">Chamorro, Maria Lourdes</mods:namePart>
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<paragraph id="8B5F369DDD03A46F02FC0CCFFCDBFA3D" blockId="81.[151,853,1462,1523]" box="[151,853,1462,1488]" pageId="81" pageNumber="82">
<heading id="D01781F1DD03A46F02FC0CCFFCDBFA3D" bold="true" box="[151,853,1462,1488]" fontSize="11" level="1" pageId="81" pageNumber="82" reason="1">
<taxonomicName id="4CE04D1EDD03A46F02FC0CCFFCDBFA3D" authority="Chamorro-Lacayo &amp; Holzenthal" authorityName=", Chamorro-Lacayo &amp; Holzenthal" authorityYear="2004" box="[151,853,1462,1488]" class="Insecta" family="Polycentropodidae" genus="Polyplectropus" kingdom="Animalia" order="Trichoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="clauseni">
<emphasis id="B994EA8FDD03A46F02FC0CCFFCDBFA3D" bold="true" box="[151,853,1462,1488]" pageId="81" pageNumber="82">
<emphasis id="B994EA8FDD03A46F02FC0CCFFE38FA3D" bold="true" box="[151,438,1462,1488]" italics="true" pageId="81" pageNumber="82">Polyplectropus clauseni</emphasis>
Chamorro-Lacayo &amp; Holzenthal
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="C3FA6516DD03A46F02FC0CA0FE57FA1E" box="[151,473,1497,1523]" pageId="81" pageNumber="82" type="description">
<paragraph id="8B5F369DDD03A46F02FC0CA0FE57FA1E" blockId="81.[151,853,1462,1523]" box="[151,473,1497,1523]" pageId="81" pageNumber="82">
<figureCitation id="13DB2A18DD03A46F02FC0CA0FE87FA1E" box="[151,265,1497,1523]" captionStart="FIGURE 15" captionStartId="36.[151,255,1803,1827]" captionTargetBox="[216,1367,187,1774]" captionTargetId="figure-62@36.[216,1370,187,1774]" captionTargetPageId="36" captionText="FIGURE 15. Right fore and hind wings of New World Polyplectropus spp., dorsal views: A—P. andinensis; B—P. colombianus; C—P. charlesi; D—P. dubitatus; E—P. clauseni; F—Polycentropus chilensis; G—P. bravoae; H—P. profaupar." pageId="81" pageNumber="82">Figs. 15E</figureCitation>
,
<figureCitation id="13DB2A18DD03A46F037D0CA0FEC7FA1E" box="[278,329,1497,1523]" captionStart="FIGURE 38" captionStartId="81.[151,255,1283,1307]" captionTargetBox="[237,1347,186,1260]" captionTargetId="figure-213@81.[237,1349,186,1267]" captionTargetPageId="81" captionText="FIGURE 38. Female genitalia of New World Polyplectropus spp.: AB—Polyplectropus bredini Flint, 1968 (allotype): A—left lateral view; B—ventral view; C—Polyplectropus clauseni ChamorroLacayo &amp; Holzenthal, 2004, ventral view." pageId="81" pageNumber="82">38C</figureCitation>
,
<figureCitation id="13DB2A18DD03A46F033E0CA0FEFBFA1E" box="[341,373,1497,1523]" captionStart="FIGURE 39" captionStartId="82.[151,255,1607,1631]" captionTargetBox="[227,1361,198,1573]" captionTargetId="figure-147@82.[226,1361,197,1580]" captionTargetPageId="82" captionText="FIGURE 39. Male genitalia of Polyplectropus clauseni ChamorroLacayo &amp; Holzenthal, 2004 (holotype): A—left lateral view; B—dorsal view; C—caudal view; D—ventral view; E—phallus, left lateral view; F—phallus apex, dorsal view; G—left inferior appendage, caudal view." pageId="81" pageNumber="82">39</figureCitation>
,
<figureCitation id="13DB2A18DD03A46F03EA0CA0FE2FFA1E" box="[385,417,1497,1523]" captionStart="FIGURE 40" captionStartId="83.[151,255,1609,1633]" captionTargetBox="[218,1368,186,1576]" captionTargetId="figure-146@83.[218,1369,186,1582]" captionTargetPageId="83" captionText="FIGURE 40. Male genitalia of Polyplectropus clauseni ChamorroLacayo &amp; Holzenthal, 2004 (Panama): A—left lateral view; B—dorsal view; C—caudal view; D—ventral view; E—phallus, left lateral view." pageId="81" pageNumber="82">40</figureCitation>
,
<figureCitation id="13DB2A18DD03A46F03C60CA0FE57FA1E" box="[429,473,1497,1523]" captionStart="FIGURE 127" captionStartId="235.[151,255,1010,1034]" captionTargetBox="[169,1413,196,988]" captionTargetId="figure-57@235.[169,1413,170,988]" captionTargetPageId="235" captionText="FIGURE 127. Distributions of Polyplectropus denticulus Bueno, 1990 and Polyplectropus clauseni ChamorroLacayo &amp; Holzenthal, 2004." pageId="81" pageNumber="82">127</figureCitation>
</paragraph>
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<subSubSection id="C3FA6516DD03A46F02FC0F50FE33F880" pageId="81" pageNumber="82" type="reference_group">
<paragraph id="8B5F369DDD03A46F02FC0F50FC75F98D" blockId="81.[151,1436,1577,1632]" pageId="81" pageNumber="82">
<treatmentCitationGroup id="ABF011B3DD03A46F02FC0F50FC75F98D" pageId="81" pageNumber="82">
<treatmentCitation id="0A41108CDD03A46F02FC0F50FCF0F9AD" author="Chamorro-Lacayo, M. L. &amp; Holzenthal, R. W." box="[151,894,1577,1600]" page="202" pageId="81" pageNumber="82" year="2004">
<taxonomicName id="4CE04D1EDD03A46F02FC0F50FCF0F9AD" authority=", Chamorro-Lacayo &amp; Holzenthal, 2004: 202" authorityName=", Chamorro-Lacayo &amp; Holzenthal" authorityPageNumber="202" authorityYear="2004" box="[151,894,1577,1600]" class="Insecta" family="Polycentropodidae" genus="Polyplectropus" kingdom="Animalia" order="Trichoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="clauseni">
<emphasis id="B994EA8FDD03A46F02FC0F50FE16F9AD" box="[151,408,1577,1600]" italics="true" pageId="81" pageNumber="82">Polyplectropus clauseni</emphasis>
,
<materialsCitation id="3B883CC0DD03A46F03CE0F50FCF0F9AD" box="[421,894,1577,1600]" collectionCode="UMSP" collectorName="NW Dos Rios" country="Costa Rica" location="Rio Bochinche" pageId="81" pageNumber="82" specimenCount="1" stateProvince="Alajuela">
<bibRefCitation id="EF714B6CDD03A46F03CE0F50FCF0F9AD" author="Chamorro-Lacayo, M. L. &amp; Holzenthal, R. W." box="[421,894,1577,1600]" pageId="81" pageNumber="82" pagination="202 - 216" refId="ref122934" refString="Chamorro-Lacayo, M. L. &amp; Holzenthal, R. W. (2004) Seven new species of Polyplectropus Ulmer (Trichoptera: Polycentropodidae) from Costa Rica. Proceedings of the Entomological Society of Washington, 106, 202 - 216." type="journal article" year="2004">Chamorro-Lacayo &amp; Holzenthal, 2004: 202</bibRefCitation>
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. [
<typeStatus id="545B883FDD03A46F01FD0F50FC40F9AD" box="[918,974,1577,1600]" pageId="81" pageNumber="82">Type</typeStatus>
locality:
<collectingCountry id="F3F7760DDD03A46F06570F50FB53F9AD" box="[1084,1245,1577,1600]" name="Costa Rica" pageId="81" pageNumber="82">COSTA RICA</collectingCountry>
:
<collectingRegion id="4924F87FDD03A46F06840F50FAC3F9AD" box="[1263,1357,1577,1600]" country="Costa Rica" name="Alajuela" pageId="81" pageNumber="82">Alajuela</collectingRegion>
: Cerro Campana,
<location id="8E3F6046DD03A46F03520F30FE55F98D" LSID="urn:lsid:plazi:treatment:0349878BDD03A46A02FC0CCFFD09FC17:8E3F6046DD03A46F03520F30FE55F98D" box="[313,475,1609,1632]" country="Costa Rica" name="Rio Bochinche" pageId="81" pageNumber="82" stateProvince="Alajuela">Rio Bochinche</location>
, tributary,
<quantity id="4C189B78DD03A46F00260F30FD0CF98D" box="[589,642,1609,1632]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="81" pageNumber="82" unit="km" value="6.0">6 km</quantity>
(air)
<collectorName id="2615534BDD03A46F00D60F30FCD8F98D" box="[701,854,1609,1632]" pageId="81" pageNumber="82">NW Dos Rios</collectorName>
; UMSP; male.]
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</paragraph>
<paragraph id="8B5F369DDD03A46F02FC0FF2FE33F880" blockId="81.[151,1437,1675,2021]" pageId="81" pageNumber="82">
This species is most similar to
<taxonomicName id="4CE04D1EDD03A46F03950FF5FD14F948" authority="Chamorro &amp; Holzenthal, 2010" authorityName="Chamorro &amp; Holzenthal" authorityYear="2010" box="[510,666,1676,1701]" class="Insecta" family="Polycentropodidae" genus="Polyplectropus" kingdom="Animalia" order="Trichoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="brasilensis" status="sp. nov.">
<emphasis id="B994EA8FDD03A46F03950FF5FD14F948" box="[510,666,1676,1701]" italics="true" pageId="81" pageNumber="82">P. brasilensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2A757F4DD03A46F00CC0FF2FCBDF948" box="[679,819,1675,1701]" pageId="81" pageNumber="82" rank="species">new species</taxonomicNameLabel>
in the overall shape of the inferior appendage, in the presence of a dorsolateral process of the preanal appendage, and in the overall shape of the phallus.
<taxonomicName id="4CE04D1EDD03A46F02FC0FA5FE3AF918" authorityName=", Chamorro-Lacayo &amp; Holzenthal" authorityYear="2004" box="[151,436,1756,1781]" class="Insecta" family="Polycentropodidae" genus="Polyplectropus" kingdom="Animalia" order="Trichoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="clauseni">
<emphasis id="B994EA8FDD03A46F02FC0FA5FE3AF918" box="[151,436,1756,1781]" italics="true" pageId="81" pageNumber="82">Polyplectropus clauseni</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4CE04D1EDD03A46F01680FA5FC2CF918" authorityName="Chamorro &amp; Holzenthal" authorityYear="2010" box="[771,930,1756,1781]" class="Insecta" family="Polycentropodidae" genus="Polyplectropus" kingdom="Animalia" order="Trichoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="brasilensis">
<emphasis id="B994EA8FDD03A46F01680FA5FC2CF918" box="[771,930,1756,1781]" italics="true" pageId="81" pageNumber="82">P. brasilensis</emphasis>
</taxonomicName>
and all other species in the group by the broad and largely membranous intermediate appendage, by the subquadrate dorsolaterally produced mesoventral processes of the preanal appendage bearing apical peg-like setae, and by the presence of a prominent papillate lobe.
</paragraph>
</subSubSection>
<subSubSection id="C3FA6516DD03A46A02AD0E05FD09FC17" lastPageId="84" lastPageNumber="85" pageId="81" pageNumber="82" type="description">
<paragraph id="8B5F369DDD03A46F02AD0E05FCEAF808" blockId="81.[151,1437,1675,2021]" pageId="81" pageNumber="82">
<emphasis id="B994EA8FDD03A46F02AD0E05FE83F878" box="[198,269,1916,1941]" italics="true" pageId="81" pageNumber="82">Adult.</emphasis>
Length of forewing
<quantity id="4C189B78DD03A46F00630E02FDEBF878" box="[520,613,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="81" pageNumber="82" unit="mm" value="5.1">5.1 mm</quantity>
, n = 4. Color of head, thorax, and legs yellowish-brown; tegulae with long, golden brown and grey setae; setae on wings golden brown along costal margin with patches of grey setae interspersed along margin and on most of wing surface.
</paragraph>
<caption id="DF9F6615DD00A46C02FC0F3EFD03F94E" pageId="82" pageNumber="83" startId="82.[151,255,1607,1631]" targetBox="[227,1361,198,1573]" targetPageId="82">
<paragraph id="8B5F369DDD00A46C02FC0F3EFD03F94E" blockId="82.[151,1436,1607,1699]" pageId="82" pageNumber="83">
<emphasis id="B994EA8FDD00A46C02FC0F3EFEA5F9B2" bold="true" box="[151,299,1607,1631]" pageId="82" pageNumber="83">FIGURE 39.</emphasis>
Male genitalia of
<taxonomicName id="4CE04D1EDD00A46C006B0F3EFCF2F9B3" authority="Chamorro" authorityName=", Chamorro-Lacayo &amp; Holzenthal" authorityYear="2004" box="[512,892,1607,1630]" class="Insecta" family="Polycentropodidae" genus="Polyplectropus" kingdom="Animalia" order="Trichoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="clauseni">
<emphasis id="B994EA8FDD00A46C006B0F3EFC8AF9B3" box="[512,772,1607,1630]" italics="true" pageId="82" pageNumber="83">Polyplectropus clauseni</emphasis>
Chamorro
</taxonomicName>
Lacayo &amp;
<bibRefCitation id="EF714B6CDD00A46C06690F3EFB30F9B3" author="Holzenthal, R. W. &amp; Andersen, T." box="[1026,1214,1607,1630]" pageId="82" pageNumber="83" pagination="1 - 80" refId="ref124231" refString="Holzenthal, R. W. &amp; Andersen, T. (2004) The caddisfly genus Triaenodes in the Neotropics (Trichoptera: Leptoceridae). Zootaxa, 511, 1 - 80." type="journal article" year="2004">Holzenthal, 2004</bibRefCitation>
(holotype): A—left lateral view; B—dorsal view; C—caudal view; D—ventral view; E—phallus, left lateral view; F—phallus apex, dorsal view; G—left inferior appendage, caudal view.
</paragraph>
</caption>
<paragraph id="8B5F369DDD00A46A02AD0FAEFD29FE27" blockId="82.[151,1436,1751,2017]" lastBlockId="84.[151,1437,152,1018]" lastPageId="84" lastPageNumber="85" pageId="82" pageNumber="83">
<emphasis id="B994EA8FDD00A46C02AD0FAEFEF2F91D" box="[198,380,1751,1776]" italics="true" pageId="82" pageNumber="83">Male genitalia.</emphasis>
Sternum IX in lateral view deltoid, anterior margin sinuate and supramedially convex, posterior margin submedially produced and becoming concave ventrally; in ventral view rectangular, anterior margin entire to sinuate, posterior margin with median flange and submedially deeply concave. Tergum X membranous, oblong, bearing dorsal microsetae. Intermediate appendage not longer than inferior appendage, broad, membranous and with ventral digitate projection and long setae basally and apically; in dorsal view digitate to clavate, elongate; in caudal view clavate, produced posteroventrad and entire appendage produced posterad and ventromesally; median papillate lobe flanking phallocrypt laterally, bearing small apical setae intermediate appendage apparently fused to mesoventral process of preanal appendage. Preanal appendage tripartite; dorsolateral process elongate, originating from dorsum of mesolateral process, directed anterad, recurved posteromesad then posterolaterad, tapering mesally into acute apex; mesolateral process setose; in dorsal view sickle-shaped, in lateral view orbicular; mesoventral process setose; in lateral view hook-like or subquadrate, mesoventral process dorsally fused to intermediate appendage with dorsolateral expanded lobe with narrowed apex, process with ventrad-directed sclerotized apex, anteroventral margin angled to concave, posterior margin angled to concave; in caudal view fused basomesally, processes separated apically, ventral margin of process undulate, with many sclerotized, peg-like setae. Inferior appendage bipartite with anterior basal plate not extending anterad beyond sternum IX when observed in lateral view; dorsal branch setose, in lateral view oblong, directed posterodorsad; in ventral view broad, lateral margin convex, entire, posterior margin acute, mesal margin entire, concave, shallowly fused to ventral branch; ventral branch setose, short and broad, bearing robust, sclerotized spines apically; in lateral view oblong and almost forming a continuous structure with dorsal branch, rounded posterad; in ventral view deltoid, mesal margin slightly concave, gradually converging mesally. Phallus short; dorsal phallic sclerite in lateral view sinuate, apex oblong; lateral projection of phallobase in lateral view cylindrical, with truncate apex; endothecal membrane without embedded spines, membranous basodorsally.
</paragraph>
<caption id="DF9F6615DD01A46D02FC0F30FB0BF96E" pageId="83" pageNumber="84" startId="83.[151,255,1609,1633]" targetBox="[218,1368,186,1576]" targetPageId="83">
<paragraph id="8B5F369DDD01A46D02FC0F30FB0BF96E" blockId="83.[151,1436,1609,1667]" pageId="83" pageNumber="84">
<emphasis id="B994EA8FDD01A46D02FC0F30FEA5F98C" bold="true" box="[151,299,1609,1633]" pageId="83" pageNumber="84">FIGURE 40.</emphasis>
Male genitalia of
<taxonomicName id="4CE04D1EDD01A46D006F0F33FC0FF98D" authority="Chamorro" authorityName=", Chamorro-Lacayo &amp; Holzenthal" authorityYear="2004" box="[516,897,1609,1633]" class="Insecta" family="Polycentropodidae" genus="Polyplectropus" kingdom="Animalia" order="Trichoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="clauseni">
<emphasis id="B994EA8FDD01A46D006F0F33FC87F98C" box="[516,777,1610,1633]" italics="true" pageId="83" pageNumber="84">Polyplectropus clauseni</emphasis>
Chamorro
</taxonomicName>
Lacayo &amp;
<bibRefCitation id="EF714B6CDD01A46D06630F30FB4BF98D" author="Holzenthal, R. W. &amp; Andersen, T." box="[1032,1221,1609,1632]" pageId="83" pageNumber="84" pagination="1 - 80" refId="ref124231" refString="Holzenthal, R. W. &amp; Andersen, T. (2004) The caddisfly genus Triaenodes in the Neotropics (Trichoptera: Leptoceridae). Zootaxa, 511, 1 - 80." type="journal article" year="2004">Holzenthal, 2004</bibRefCitation>
(Panama): A—left lateral view; B—dorsal view; C—caudal view; D—ventral view; E—phallus, left lateral view.
</paragraph>
</caption>
<paragraph id="8B5F369DDD06A46A02AD08A1FB89FD7F" blockId="84.[151,1437,152,1018]" pageId="84" pageNumber="85">
<emphasis id="B994EA8FDD06A46A02AD08A1FE1BFE1C" box="[198,405,472,497]" italics="true" pageId="84" pageNumber="85">Female genitalia.</emphasis>
Ventral plates of segment VIII setose, in ventral view narrowly oblong, narrowly fused to sternite VIII; external part of gonopod of segment VIII in ventral view with posterior margin rounded. Tergum IX devoid of setae, lightly sclerotized. Segment X setose, apically bearing a pair of sclerotized cerci surrounded by 2 pairs of membranous digitate processes. Genital chamber opening to processus spermathecae through sclerotized circular opening of ductus spermathecae.
</paragraph>
<paragraph id="8B5F369DDD06A46A02AD0BD9FDFAFC3F" blockId="84.[151,1437,152,1018]" pageId="84" pageNumber="85">
<materialsCitation id="3B883CC0DD06A46A02AD0BD9FE06FCE7" collectingDate="1987-07-22" collectingDateMax="1987-07-23" collectingDateMin="1987-07-22" collectionCode="UMSP" collectorName="Morse &amp; Clausen" country="Costa Rica" county="Material Examined." elevation="640" latitude="10.9" location="Rio Bochinche" longLatPrecision="21" longitude="-85.4" municipality="Cerro Campana" pageId="84" pageNumber="85" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela" typeStatus="holotype">
<collectingCounty id="623E4E11DD06A46A02AD0BD9FE3CFD57" box="[198,434,672,698]" pageId="84" pageNumber="85">Material Examined.</collectingCounty>
<emphasis id="B994EA8FDD06A46A03AF0BE6FC82FD54" bold="true" box="[452,780,671,697]" pageId="84" pageNumber="85">
<collectingCountry id="F3F7760DDD06A46A03AF0BE6FD0EFD54" box="[452,640,671,697]" name="Costa Rica" pageId="84" pageNumber="85">COSTA RICA</collectingCountry>
:
<collectingRegion id="4924F87FDD06A46A00F20BE6FC89FD54" box="[665,775,671,697]" country="Costa Rica" name="Alajuela" pageId="84" pageNumber="85">Alajuela</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6B3BACE7DD06A46A014B0BD9FC50FD57" box="[800,990,672,698]" pageId="84" pageNumber="85">Cerro Campana</collectingMunicipality>
,
<location id="8E3F6046DD06A46A01870BD9FB2EFD57" LSID="urn:lsid:plazi:treatment:0349878BDD03A46A02FC0CCFFD09FC17:8E3F6046DD06A46A01870BD9FB2EFD57" box="[1004,1184,672,698]" country="Costa Rica" county="Material Examined." latitude="10.9" longLatPrecision="21" longitude="-85.4" municipality="Cerro Campana" name="Rio Bochinche" pageId="84" pageNumber="85" stateProvince="Alajuela">Rio Bochinche</location>
, tributary,
<quantity id="4C189B78DD06A46A07490BD9FAD1FD57" box="[1314,1375,672,698]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="84" pageNumber="85" unit="km" value="6.0">6 km</quantity>
(air) NW
<location id="8E3F6046DD06A46A02BF0BB1FEC9FD0F" LSID="urn:lsid:plazi:treatment:0349878BDD03A46A02FC0CCFFD09FC17:8E3F6046DD06A46A02BF0BB1FEC9FD0F" box="[212,327,712,738]" country="Costa Rica" county="Material Examined." latitude="10.9" longLatPrecision="21" longitude="-85.4" municipality="Cerro Campana" name="Dos Rios" pageId="84" pageNumber="85" stateProvince="Alajuela">Dos Rios</location>
,
<geoCoordinate id="EED4505ADD06A46A03330BB1FE6BFD0F" box="[344,485,712,738]" degrees="10" direction="north" minutes="54" orientation="latitude" pageId="84" pageNumber="85" precision="15" seconds="00" value="10.9">10°54'00&quot;N</geoCoordinate>
,
<geoCoordinate id="EED4505ADD06A46A039D0BB1FD08FD0F" box="[502,646,712,738]" degrees="85" direction="west" minutes="24" orientation="longitude" pageId="84" pageNumber="85" precision="15" seconds="00" value="-85.4">85°24'00&quot;W</geoCoordinate>
,
<quantity id="4C189B78DD06A46A00FC0BB1FD66FD0F" box="[663,744,712,738]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="84" pageNumber="85" unit="m" value="640.0">
<elevation id="00CDD1AEDD06A46A00FC0BB1FD66FD0F" box="[663,744,712,738]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="84" pageNumber="85" unit="m" value="640.0">640 m</elevation>
</quantity>
,
<date id="FF5E105DDD06A46A00910BB1FC23FD0F" box="[762,941,712,738]" pageId="84" pageNumber="85" value="1987-07-22" valueMax="1987-07-23" valueMin="1987-07-22">
<collectingDate id="EF1AE9B5DD06A46A00910BB1FC23FD0F" box="[762,941,712,738]" pageId="84" pageNumber="85" value="1987-07-22" valueMax="1987-07-23" valueMin="1987-07-22">2223.vii.1987</collectingDate>
</date>
, Holzenthal,
<collectorName id="2615534BDD06A46A06390BB1FB13FD0F" box="[1106,1181,712,738]" pageId="84" pageNumber="85">Morse</collectorName>
&amp;
<collectorName id="2615534BDD06A46A06A70BB1FAA0FD0F" box="[1228,1326,712,738]" pageId="84" pageNumber="85">Clausen</collectorName>
<specimenCount id="9DE6FD14DD06A46A07250BB1FA12FD0F" box="[1358,1436,712,738]" pageId="84" pageNumber="85" type="male">1 male</specimenCount>
(
<collectionCode id="EDF1AE58DD06A46A02F40B89FF7CFCE7" box="[159,242,752,778]" country="USA" name="University of Minnesota Insect Collection" pageId="84" pageNumber="85" type="University or college">UMSP</collectionCode>
) [
<typeStatus id="545B883FDD06A46A037A0B89FE0FFCE7" box="[273,385,752,778]" pageId="84" pageNumber="85" type="holotype">holotype</typeStatus>
]
</materialsCitation>
;
<materialsCitation id="3B883CC0DD06A46A03F70B96FBDEFCDF" collectingDate="1986-03-07" collectionCode="NMNH" collectorName="Holzenthal &amp; Fasth" country="Costa Rica" elevation="470" latitude="10.865001" location="Rio Los Ahogados" longLatPrecision="21" longitude="-85.42305" pageId="84" pageNumber="85" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<emphasis id="B994EA8FDD06A46A03F70B96FDB6FCE4" bold="true" box="[412,568,751,777]" pageId="84" pageNumber="85">
<collectingRegion id="4924F87FDD06A46A03F70B96FDBDFCE4" box="[412,563,751,777]" country="Costa Rica" name="Guanacaste" pageId="84" pageNumber="85">Guanacaste</collectingRegion>
:
</emphasis>
<location id="8E3F6046DD06A46A00330B89FCC6FCE7" LSID="urn:lsid:plazi:treatment:0349878BDD03A46A02FC0CCFFD09FC17:8E3F6046DD06A46A00330B89FCC6FCE7" box="[600,840,752,778]" country="Costa Rica" latitude="10.865001" longLatPrecision="21" longitude="-85.42305" name="Rio Los Ahogados" pageId="84" pageNumber="85" stateProvince="Guanacaste">Río Los Ahogados</location>
,
<locationDeviation id="9AED2FBFDD06A46A01370B89FB78FCE7" box="[860,1270,752,778]" location="Quebrada Grande" pageId="84" pageNumber="85">
<quantity id="4C189B78DD06A46A01370B89FC4CFCE7" box="[860,962,752,778]" metricMagnitude="4" metricUnit="m" metricValue="1.13" pageId="84" pageNumber="85" unit="km" value="11.3">11.3 km</quantity>
ENE Quebrada Grande
</locationDeviation>
,
<geoCoordinate id="EED4505ADD06A46A07600B89FA19FCE7" box="[1291,1431,752,778]" degrees="10" direction="north" minutes="51" orientation="latitude" pageId="84" pageNumber="85" precision="15" seconds="54" value="10.865001">10°51'54&quot;N</geoCoordinate>
,
<geoCoordinate id="EED4505ADD06A46A02FC0A61FEA8FCDF" box="[151,294,792,818]" degrees="85" direction="west" minutes="25" orientation="longitude" pageId="84" pageNumber="85" precision="15" seconds="23" value="-85.42305">85°25'23&quot;W</geoCoordinate>
,
<quantity id="4C189B78DD06A46A03580A61FEF1FCDF" box="[307,383,792,818]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="84" pageNumber="85" unit="m" value="470.0">
<elevation id="00CDD1AEDD06A46A03580A61FEF1FCDF" box="[307,383,792,818]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="84" pageNumber="85" unit="m" value="470.0">470 m</elevation>
</quantity>
,
<date id="FF5E105DDD06A46A03E00A61FE70FCDF" box="[395,510,792,818]" pageId="84" pageNumber="85" value="1986-03-07">
<collectingDate id="EF1AE9B5DD06A46A03E00A61FE70FCDF" box="[395,510,792,818]" pageId="84" pageNumber="85" value="1986-03-07">7.iii.1986</collectingDate>
</date>
,
<collectorName id="2615534BDD06A46A00620A61FD07FCDF" box="[521,649,792,818]" pageId="84" pageNumber="85">Holzenthal</collectorName>
&amp;
<collectorName id="2615534BDD06A46A00C40A61FD7CFCDF" box="[687,754,792,818]" pageId="84" pageNumber="85">Fasth</collectorName>
<specimenCount id="9DE6FD14DD06A46A017A0A61FCD4FCDF" box="[785,858,792,818]" pageId="84" pageNumber="85" type="male">1 male</specimenCount>
(
<collectionCode id="EDF1AE58DD06A46A01030A61FC4BFCDF" box="[872,965,792,818]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="84" pageNumber="85" type="Museum">NMNH</collectionCode>
) [
<typeStatus id="545B883FDD06A46A01B70A61FBC4FCDF" box="[988,1098,792,818]" pageId="84" pageNumber="85" type="paratype">paratype</typeStatus>
]
</materialsCitation>
;
<materialsCitation id="3B883CC0DD06A46A06370A61FDEDFC6F" collectingDate="1986-06-25" collectionCode="UMSP" collectorName="Holzenthal &amp; Heyn &amp; Armitage" country="Costa Rica" elevation="795" latitude="10.795" location="Quebrada" longLatPrecision="21" longitude="-85.29195" municipality="Parque Nacional Rincon de la Vieja" pageId="84" pageNumber="85" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<collectingMunicipality id="6B3BACE7DD06A46A06370A61FF75FCB7" pageId="84" pageNumber="85">Parque Nacional Rincón de la Vieja</collectingMunicipality>
,
<location id="8E3F6046DD06A46A03670A39FEF0FCB7" LSID="urn:lsid:plazi:treatment:0349878BDD03A46A02FC0CCFFD09FC17:8E3F6046DD06A46A03670A39FEF0FCB7" box="[268,382,832,858]" country="Costa Rica" latitude="10.795" longLatPrecision="21" longitude="-85.29195" municipality="Parque Nacional Rincon de la Vieja" name="Quebrada" pageId="84" pageNumber="85" stateProvince="Guanacaste">Quebrada</location>
Agua Apinolada,
<geoCoordinate id="EED4505ADD06A46A00030A39FD7BFCB7" box="[616,757,832,858]" degrees="10" direction="north" minutes="47" orientation="latitude" pageId="84" pageNumber="85" precision="15" seconds="42" value="10.795">10°47'42&quot;N</geoCoordinate>
,
<geoCoordinate id="EED4505ADD06A46A016C0A39FC19FCB7" box="[775,919,832,858]" degrees="85" direction="west" minutes="17" orientation="longitude" pageId="84" pageNumber="85" precision="15" seconds="31" value="-85.29195">85°17'31&quot;W</geoCoordinate>
,
<quantity id="4C189B78DD06A46A01C10A39FC70FCB7" box="[938,1022,832,858]" metricMagnitude="2" metricUnit="m" metricValue="7.95" pageId="84" pageNumber="85" unit="m" value="795.0">
<elevation id="00CDD1AEDD06A46A01C10A39FC70FCB7" box="[938,1022,832,858]" metricMagnitude="2" metricUnit="m" metricValue="7.95" pageId="84" pageNumber="85" unit="m" value="795.0">795 m</elevation>
</quantity>
,
<date id="FF5E105DDD06A46A067B0A39FB1EFCB7" box="[1040,1168,832,858]" pageId="84" pageNumber="85" value="1986-06-25">
<collectingDate id="EF1AE9B5DD06A46A067B0A39FB1EFCB7" box="[1040,1168,832,858]" pageId="84" pageNumber="85" value="1986-06-25">25.vi.1986</collectingDate>
</date>
,
<collectorName id="2615534BDD06A46A06C90A39FAA8FCB7" box="[1186,1318,832,858]" pageId="84" pageNumber="85">Holzenthal</collectorName>
,
<collectorName id="2615534BDD06A46A07530A39FAF9FCB7" box="[1336,1399,832,858]" pageId="84" pageNumber="85">Heyn</collectorName>
&amp;
<collectorName id="2615534BDD06A46A02FC0A11FE84FC6F" box="[151,266,872,898]" pageId="84" pageNumber="85">Armitage</collectorName>
<specimenCount id="9DE6FD14DD06A46A03430A11FEFDFC6F" box="[296,371,872,898]" pageId="84" pageNumber="85" type="male">1 male</specimenCount>
(
<collectionCode id="EDF1AE58DD06A46A03E80A11FE58FC6F" box="[387,470,872,898]" country="USA" name="University of Minnesota Insect Collection" pageId="84" pageNumber="85" type="University or college">UMSP</collectionCode>
) [
<typeStatus id="545B883FDD06A46A03860A11FDD5FC6F" box="[493,603,872,898]" pageId="84" pageNumber="85" type="paratype">paratype</typeStatus>
]
</materialsCitation>
;
<materialsCitation id="3B883CC0DD06A46A00040A1EFCA6FC47" collectingDate="1988-08-03" collectingDateMax="1988-08-09" collectingDateMin="1988-08-03" collectionCode="NMNH" collectorName="H. Wolda" country="Panama" location="Snyder-Molino Trail" municipality="Barro Colorado Island" pageId="84" pageNumber="85" specimenCount="1" specimenCount-male="1" stateProvince="Panama" typeStatus="paratype">
<emphasis id="B994EA8FDD06A46A00040A1EFCF4FC6C" bold="true" box="[623,890,871,897]" pageId="84" pageNumber="85">
<collectingCountry id="F3F7760DDD06A46A00040A1EFD7AFC6C" box="[623,756,871,897]" name="Panama" pageId="84" pageNumber="85">PANAMA</collectingCountry>
:
<collectingRegion id="4924F87FDD06A46A01610A1EFCFBFC6C" box="[778,885,871,897]" country="Panama" name="Panama" pageId="84" pageNumber="85">Panama</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6B3BACE7DD06A46A01E10A11FB1BFC6F" box="[906,1173,872,898]" pageId="84" pageNumber="85">Barro Colorado Island</collectingMunicipality>
,
<location id="8E3F6046DD06A46A06C90A11FA19FC6F" LSID="urn:lsid:plazi:treatment:0349878BDD03A46A02FC0CCFFD09FC17:8E3F6046DD06A46A06C90A11FA19FC6F" box="[1186,1431,872,898]" country="Panama" municipality="Barro Colorado Island" name="Snyder-Molino Trail" pageId="84" pageNumber="85" stateProvince="Panama">Snyder-Molino Trail</location>
, marker 3,
<date id="FF5E105DDD06A46A03730AE9FE39FC47" box="[280,439,912,938]" pageId="84" pageNumber="85" value="1988-08-03" valueMax="1988-08-09" valueMin="1988-08-03">
<collectingDate id="EF1AE9B5DD06A46A03730AE9FE39FC47" box="[280,439,912,938]" pageId="84" pageNumber="85" value="1988-08-03" valueMax="1988-08-09" valueMin="1988-08-03">39.viii.1988</collectingDate>
</date>
,
<collectorName id="2615534BDD06A46A03AC0AE9FDCEFC47" box="[455,576,912,938]" pageId="84" pageNumber="85">H. Wolda</collectorName>
<specimenCount id="9DE6FD14DD06A46A00340AE9FD22FC47" box="[607,684,912,938]" pageId="84" pageNumber="85" type="male">1 male</specimenCount>
(
<collectionCode id="EDF1AE58DD06A46A00AB0AE9FCAEFC47" box="[704,800,912,938]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="84" pageNumber="85" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B883CC0DD06A46A01520AE9FAD1FC47" box="[825,1375,912,938]" collectingDate="1989-01-25" collectingDateMax="1989-01-31" collectingDateMin="1989-01-25" collectionCode="NMNH" collectorName="Wolda" country="Panama" pageId="84" pageNumber="85" specimenCount="1" specimenCount-female="1" stateProvince="Panama" typeStatus="paratype">
<date id="FF5E105DDD06A46A01520AE9FC59FC47" box="[825,983,912,938]" pageId="84" pageNumber="85" value="1989-01-25" valueMax="1989-01-31" valueMin="1989-01-25">
<collectingDate id="EF1AE9B5DD06A46A01520AE9FC59FC47" box="[825,983,912,938]" pageId="84" pageNumber="85" value="1989-01-25" valueMax="1989-01-31" valueMin="1989-01-25">2531.i.1989</collectingDate>
</date>
, H.
<collectorName id="2615534BDD06A46A067B0AE9FBEEFC47" box="[1040,1120,912,938]" pageId="84" pageNumber="85">Wolda</collectorName>
<specimenCount id="9DE6FD14DD06A46A06140AE9FB6DFC47" box="[1151,1251,912,938]" pageId="84" pageNumber="85" type="female">1 female</specimenCount>
(
<collectionCode id="EDF1AE58DD06A46A069C0AE9FAD9FC47" box="[1271,1367,912,938]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="84" pageNumber="85" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B883CC0DD06A46A071B0AE9FDFEFC3F" collectingDate="1991-02-13" collectingDateMax="1991-02-19" collectingDateMin="1991-02-13" collectionCode="NMNH" collectorName="Wolda" country="Panama" pageId="84" pageNumber="85" specimenCount="1" specimenCount-male="1" stateProvince="Panama" typeStatus="paratype">
<date id="FF5E105DDD06A46A071B0AE9FE9EFC3F" pageId="84" pageNumber="85" value="1991-02-13" valueMax="1991-02-19" valueMin="1991-02-13">
<collectingDate id="EF1AE9B5DD06A46A071B0AE9FE9EFC3F" pageId="84" pageNumber="85" value="1991-02-13" valueMax="1991-02-19" valueMin="1991-02-13">13 19.ii.1991</collectingDate>
</date>
, H.
<collectorName id="2615534BDD06A46A032B0AC1FE1EFC3F" box="[320,400,952,978]" pageId="84" pageNumber="85">Wolda</collectorName>
<specimenCount id="9DE6FD14DD06A46A03C40AC1FE76FC3F" box="[431,504,952,978]" pageId="84" pageNumber="85" type="male">1 male</specimenCount>
(
<collectionCode id="EDF1AE58DD06A46A006C0AC1FDE6FC3F" box="[519,616,952,978]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="84" pageNumber="85" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B5F369DDD06A46A02AD0A99FD09FC17" blockId="84.[151,1437,152,1018]" box="[198,647,992,1018]" pageId="84" pageNumber="85">
Distribution.
<collectingCountry id="F3F7760DDD06A46A03010A99FE64FC17" box="[362,490,992,1018]" name="Costa Rica" pageId="84" pageNumber="85">Costa Rica</collectingCountry>
and
<collectingCountry id="F3F7760DDD06A46A00490A99FD0DFC17" box="[546,643,992,1018]" name="Panama" pageId="84" pageNumber="85">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>