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<document id="16F1083EFD457CAA4B476DF31765F183" ID-DOI="10.5281/zenodo.273851" ID-GBIF-Dataset="91e318c7-d881-4379-bced-1d8b63355d41" ID-ISSN="1175-5326" ID-Zenodo-Dep="273851" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459869041990" checkinUser="plazi" docAuthor="Stuke, Jens-Hermann &amp; Skevington, Jeffrey H." docDate="2007" docId="03FE02361A23FFD8FF71D64BFE01F927" docLanguage="en" docName="zt01528p040.pdf" docOrigin="Zootaxa 1528" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Physocephala herrerai Stuke &amp; Skevington, 2007, spec. nov." docType="treatment" docVersion="6" lastPageNumber="10" masterDocId="FFC77A4E1A24FFD1FFE6D171FFB4FFD5" masterDocTitle="The Conopidae of Costa Rica (Diptera) (Part 1: Conopinae Conopini &amp; Tropidomyiini)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="8" updateTime="1698226465273" updateUser="plazi">
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<mods:title id="53F7C33105689969B89DCE660D1F74DD">The Conopidae of Costa Rica (Diptera) (Part 1: Conopinae Conopini &amp; Tropidomyiini)</mods:title>
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<mods:namePart id="110EF3EB76A2057EF31522779A2E88E1">Stuke, Jens-Hermann</mods:namePart>
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<mods:namePart id="231A8DC133DE82477BD127251FE6061C">Skevington, Jeffrey H.</mods:namePart>
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<mods:date id="2813685DBAE21815A5B8D6800E6A8D2F">2007</mods:date>
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<mods:number id="EDDAE1B4A09F601BA13205B9016D3E0B">1528</mods:number>
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<treatment id="03FE02361A23FFD8FF71D64BFE01F927" ID-DOI="http://doi.org/10.5281/zenodo.6251531" ID-GBIF-Taxon="119369141" ID-Zenodo-Dep="6251531" LSID="urn:lsid:plazi:treatment:03FE02361A23FFD8FF71D64BFE01F927" httpUri="http://treatment.plazi.org/id/03FE02361A23FFD8FF71D64BFE01F927" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
<subSubSection id="C34DE0AB1A23FFD6FF71D64BFD5CF881" box="[151,744,1850,1876]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BE8B3201A23FFD6FF71D64BFD5CF881" blockId="7.[151,744,1850,1876]" box="[151,744,1850,1876]" pageId="7" pageNumber="8">
<heading id="D0A0044C1A23FFD6FF71D64BFD5CF881" bold="true" box="[151,744,1850,1876]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B9236F321A23FFD6FF71D64BFD5CF881" bold="true" box="[151,744,1850,1876]" pageId="7" pageNumber="8">
<taxonomicName id="4C57C8A31A23FFD6FF71D64BFE1DF881" ID-CoL="9HCBX" box="[151,425,1850,1876]" class="Insecta" family="Conopidae" genus="Physocephala" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="herrerai" status="sp. nov.">
<emphasis id="B9236F321A23FFD6FF71D64BFE1DF881" bold="true" box="[151,425,1850,1876]" italics="true" pageId="7" pageNumber="8">Physocephala herrerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A210D2491A23FFD6FE56D64BFD9CF881" box="[432,552,1850,1876]" pageId="7" pageNumber="8" rank="species">spec. nov.</taxonomicNameLabel>
(figures 1621)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34DE0AB1A23FFD8FF71D60BFB07FE27" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8BE8B3201A23FFD6FF71D60BFA8DF831" blockId="7.[151,1436,1914,2020]" pageId="7" pageNumber="8">
<emphasis id="B9236F321A23FFD6FF71D60BFEBDF841" bold="true" box="[151,265,1914,1940]" pageId="7" pageNumber="8">
<typeStatus id="54EC0D821A23FFD6FF71D60BFEBDF841" box="[151,265,1914,1940]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
</emphasis>
Ψ: &quot;Sect. San Ramón de dos Rios, Prov. / Alaju,
<collectingCountry id="F340F3B01A23FFD6FCA1D60BFC41F841" box="[839,1013,1914,1940]" name="Costa Rica" pageId="7" pageNumber="8">COSTA RICA</collectingCountry>
,
<specimenCount id="9D5178A91A23FFD6FBE6D60BFBFEF841" box="[1024,1098,1914,1940]" pageId="7" pageNumber="8" type="generic">
<quantity id="4CAF1EC51A23FFD6FBE6D60BFBF2F841" box="[1024,1094,1914,1940]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="7" pageNumber="8" unit="m" value="620.0">620m</quantity>
.
</specimenCount>
<date id="FFE995E01A23FFD6FBB7D60BFAC8F841" box="[1105,1404,1914,1940]" pageId="7" pageNumber="8" value="1995-03-18" valueMax="1995-04-13" valueMin="1995-03-18">
18 /
<collectionCode id="ED462BE51A23FFD6FB63D60BFB7FF841" box="[1157,1227,1914,1940]" httpUri="http://grbio.org/cool/p96j-ip08" name="Grasslands Rhizobium Collection" pageId="7" pageNumber="8">MAR</collectionCode>
13 ABR 1995
</date>
. F. A. Quesada, / L_N_318100_381900 #5274&quot;; (2) &quot;
<collectingCountry id="F340F3B01A23FFD6FD05D6D3FC27F869" box="[739,915,1954,1980]" name="Costa Rica" pageId="7" pageNumber="8">COSTA RICA</collectingCountry>
<collectionCode id="ED462BE51A23FFD6FC7DD6D3FC59F869" box="[923,1005,1954,1980]" pageId="7" pageNumber="8">INBIO</collectionCode>
/
<collectionCode id="ED462BE51A23FFD6FBE3D6D3FB82F869" LSID="urn:lsid:biocol.org:col:14323" box="[1029,1078,1954,1980]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="7" pageNumber="8">CRI</collectionCode>
002 / 246242&quot; [label with barcode]; (3) &quot;
<typeStatus id="54EC0D821A23FFD6FEFFD6BBFE22F831" box="[281,406,1994,2020]" pageId="7" pageNumber="8" type="holotype">Holotypus</typeStatus>
/
<taxonomicName id="4C57C8A31A23FFD6FE4BD6BAFD0EF831" box="[429,698,1995,2020]" class="Insecta" family="Conopidae" genus="Physocephala" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="herrerai">
<emphasis id="B9236F321A23FFD6FE4BD6BAFD0EF831" box="[429,698,1995,2020]" italics="true" pageId="7" pageNumber="8">Physocephala herrerai</emphasis>
</taxonomicName>
/ spec. nov. / det. Stuke &amp; Skevington, 2005&quot; (
<collectionCode id="ED462BE51A23FFD6FB06D6BBFA98F831" box="[1248,1324,1994,2020]" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="7" pageNumber="8">INBC</collectionCode>
).
</paragraph>
<caption id="DF28E3A81A2CFFD9FF71D6B0FC05F80C" box="[151,945,1985,2009]" httpUri="https://zenodo.org/record/273856/files/figure.png" pageId="8" pageNumber="9" targetBox="[263,1323,181,1970]" targetPageId="8">
<paragraph id="8BE8B3201A2CFFD9FF71D6B0FC05F80C" blockId="8.[151,945,1985,2009]" box="[151,945,1985,2009]" pageId="8" pageNumber="9">
<emphasis id="B9236F321A2CFFD9FF71D6B0FEE2F80C" bold="true" box="[151,342,1985,2009]" pageId="8" pageNumber="9">FIGURES 1621</emphasis>
:
<taxonomicName id="4C57C8A31A2CFFD9FE82D6B3FDEDF80C" box="[356,601,1986,2009]" class="Insecta" family="Conopidae" genus="Physocephala" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="herrerai" status="sp. nov.">
<emphasis id="B9236F321A2CFFD9FE82D6B3FDEDF80C" box="[356,601,1986,2009]" italics="true" pageId="8" pageNumber="9">Physocephala herrerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A210D2491A2CFFD9FDB9D6B3FD7EF80C" box="[607,714,1986,2009]" pageId="8" pageNumber="9" rank="species">spec. nov.</taxonomicNameLabel>
[specimen holotype].
</paragraph>
</caption>
<paragraph id="8BE8B3201A2DFFD8FF71D1E9FDF7FF67" blockId="9.[151,1437,152,1258]" box="[151,579,152,178]" pageId="9" pageNumber="10">
Description of the
<typeStatus id="54EC0D821A2DFFD8FE94D1E9FE6DFF67" box="[370,473,152,178]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
(female)
</paragraph>
<paragraph id="8BE8B3201A2DFFD8FF20D1CEFB07FE27" blockId="9.[151,1437,152,1258]" pageId="9" pageNumber="10">
<emphasis id="B9236F321A2DFFD8FF20D1CEFEBDFF0C" bold="true" box="[198,265,191,217]" pageId="9" pageNumber="10">Head</emphasis>
3.5 mm high. Eye brown, facets a little bit larger in the centre near the face.
<collectingCountry id="F340F3B01A2DFFD8FB57D1B1FB58FF0F" box="[1201,1260,192,218]" name="India" pageId="9" pageNumber="10">Hind</collectingCountry>
margin of eye slightly concave in the middle. Antenna completely orange, with black bristles at the top of the scape and mainly dorsally on the pedicel. Ventral flagellum membrane-like. Second segment of the style as long as the third segment, the third segment slightly blackish at the tip. Head mainly yellow, gena black with clear yellow centre, upper parts of frons black, vertex and ptilinal fissure light brown. Obvious pubescence only bordering the eyes with the exception of the frons, pubescence golden at the postcranium and grey at the gena. A less obvious pubescence can be seen when looking at a suitable angle at the gena and the postcranium. Vertex and postcranium (with the exception of the postgena) with long black hairs. Proboscis orange.
</paragraph>
</subSubSection>
<subSubSection id="C34DE0AB1A2DFFD8FF20D08EFE01F927" pageId="9" pageNumber="10" type="description">
<paragraph id="8BE8B3201A2DFFD8FF20D08EFEE7FD47" blockId="9.[151,1437,152,1258]" pageId="9" pageNumber="10">
<emphasis id="B9236F321A2DFFD8FF20D08EFE97FDCC" bold="true" box="[198,291,511,537]" pageId="9" pageNumber="10">Thorax</emphasis>
black. Only a few hard-to-find black hairs at the dorsal posterior part of the katepisternum and the whole mesoscutum with scutellum and postpronotum covered with short black hairs. Thorax completely covered with a less obvious grey pubescence, the dorsal posterior part of the katepisternum with more obvious silver pubescence.
</paragraph>
<paragraph id="8BE8B3201A2DFFD8FF20D3D1FBE0FC8F" blockId="9.[151,1437,152,1258]" pageId="9" pageNumber="10">Legs orange brown with the exception of the hind femur, the apical part of the hind tibia and the top of the claws that are all blackish. Legs covered with black hairs with these exceptions: anteroventral side of fore tibia with short dense golden hairs, fore and middle tibia posterior at the apical part with dense golden pubescence, hind tibia apically covered with dense golden hairs, ventral and posterior part of the femora without hairs. Coxa densely silver to golden pubescent, femora less obvious silver dusted.</paragraph>
<paragraph id="8BE8B3201A2DFFD8FF20D219FF5EFBF7" blockId="9.[151,1437,152,1258]" pageId="9" pageNumber="10">
Length of wing
<quantity id="4CAF1EC51A2DFFD8FE62D219FE63FC57" box="[388,471,872,898]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="10" unit="mm" value="12.0">12 mm</quantity>
(measured from the top of the wing to the tegula). Wing completely covered with microtrichia with the exception of the alula, the anal lobe and the basal part of the posterior cubital cell
<emphasis id="B9236F321A2DFFD8FA8CD2E1FA20FC7C" box="[1386,1428,912,937]" italics="true" pageId="9" pageNumber="10">cup</emphasis>
. Venation and colouration as shown on figure 21. Haltere yellow with a black basis and a black apex; 56 black bristles at the stem. Calypters yellow brown with a slightly darker border and small golden hairs on the thicker border.
</paragraph>
<paragraph id="8BE8B3201A2DFFD8FF20D55EFB5BFB3F" blockId="9.[151,1437,152,1258]" pageId="9" pageNumber="10">
<emphasis id="B9236F321A2DFFD8FF20D55EFEF5FB9C" bold="true" box="[198,321,1071,1097]" pageId="9" pageNumber="10">Abdomen</emphasis>
completely black to dark brown; covered with silver grey pubescence with the exception of tergite 5, syntergite 6, and partly the lateral margins of tergite 2. Abdomen completely covered with black hairs that are smaller and less dense at the base of tergite 1. Form of the tergites as shown in figures 16 and 18. The elongated syntergite 6 and the rounded tergite 5 are distinctive. Theca very broad and completely covered with black bristles that are arranged in small regular lines with about 612 bristles in each line.
</paragraph>
<paragraph id="8BE8B3201A2DFFD8FF71D451FEADFAEF" blockId="9.[151,1436,1312,1418]" box="[151,281,1312,1338]" pageId="9" pageNumber="10">Etymology</paragraph>
<paragraph id="8BE8B3201A2DFFD8FF20D439FD0EFA5F" blockId="9.[151,1436,1312,1418]" pageId="9" pageNumber="10">
The species is named after Alvaro Herrera (INBIO; Santo
<collectingCountry id="F340F3B01A2DFFD8FC89D439FC55FAB7" box="[879,993,1352,1378]" name="Dominican Republic" pageId="9" pageNumber="10">Domingo</collectingCountry>
, Heredia) who was very helpful when we visited the INBIO collection in
<collectingCountry id="F340F3B01A2DFFD8FDD5D401FD02FA5F" box="[563,694,1392,1418]" name="Costa Rica" pageId="9" pageNumber="10">Costa Rica</collectingCountry>
.
</paragraph>
<paragraph id="8BE8B3201A2DFFD8FF71D4B1FEB8FA0F" blockId="9.[151,1436,1472,1778]" box="[151,268,1472,1498]" pageId="9" pageNumber="10">Diagnosis</paragraph>
<paragraph id="8BE8B3201A2DFFD8FF20D499FE01F927" blockId="9.[151,1436,1472,1778]" pageId="9" pageNumber="10">
The first couplet in
<bibRefCitation id="EFC6CED11A2DFFD8FE57D499FDD4F9D7" author="Camras" box="[433,608,1512,1538]" pageId="9" pageNumber="39" refString="Camras, S. (1996). New information on the New World Physocephala (Diptera: Conopidae). Entomological News, 107, 104 - 112." type="journal article" year="1996">Camras (1996)</bibRefCitation>
doesnt work when identifying this species because the frons is about half black and yellow. Following couplet 1 &quot;Frons mainly or entirely dark&quot; will lead to couplet 14 with the species
<taxonomicName id="4C57C8A31A2DFFD8FF15D749FD97F984" box="[243,547,1592,1617]" class="Insecta" family="Conopidae" genus="Physocephala" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="carbonaria">
<emphasis id="B9236F321A2DFFD8FF15D749FD97F984" box="[243,547,1592,1617]" italics="true" pageId="9" pageNumber="10">Physocephala carbonaria</emphasis>
</taxonomicName>
and
<taxonomicName id="4C57C8A31A2DFFD8FDBBD749FCC1F984" box="[605,885,1592,1617]" class="Insecta" family="Conopidae" genus="Physocephala" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aurifrons">
<emphasis id="B9236F321A2DFFD8FDBBD749FCC1F984" box="[605,885,1592,1617]" italics="true" pageId="9" pageNumber="10">Physocephala aurifrons</emphasis>
</taxonomicName>
. The distinction of
<taxonomicName id="4C57C8A31A2DFFD8FBBBD749FADEF984" box="[1117,1386,1592,1617]" class="Insecta" family="Conopidae" genus="Physocephala" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="herrerai">
<emphasis id="B9236F321A2DFFD8FBBBD749FADEF984" box="[1117,1386,1592,1617]" italics="true" pageId="9" pageNumber="10">Physocephala herrerai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C57C8A31A2DFFD8FF71D711FE7CF9AC" box="[151,456,1632,1657]" class="Insecta" family="Conopidae" genus="Physocephala" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="carbonaria">
<emphasis id="B9236F321A2DFFD8FF71D711FE7CF9AC" box="[151,456,1632,1657]" italics="true" pageId="9" pageNumber="10">Physocephala carbonaria</emphasis>
</taxonomicName>
is given in our key.
<collectingCountry id="F340F3B01A2DFFD8FD5DD711FD68F9AF" box="[699,732,1632,1658]" name="American Samoa" pageId="9" pageNumber="10">As</collectingCountry>
stated above, the status of
<taxonomicName id="4C57C8A31A2DFFD8FBF8D711FA83F9AC" box="[1054,1335,1632,1657]" class="Insecta" family="Conopidae" genus="Physocephala" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aurifrons">
<emphasis id="B9236F321A2DFFD8FBF8D711FA83F9AC" box="[1054,1335,1632,1657]" italics="true" pageId="9" pageNumber="10">Physocephala aurifrons</emphasis>
</taxonomicName>
remains doubtful and cannot be separated. Following couplet 1' &quot;Frons mainly or entirely pale&quot; will lead to couplet 30 where both species offered don´t fit
<taxonomicName id="4C57C8A31A2DFFD8FDA5D7C1FCE4F91C" box="[579,848,1712,1737]" class="Insecta" family="Conopidae" genus="Physocephala" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="herrerai">
<emphasis id="B9236F321A2DFFD8FDA5D7C1FCE4F91C" box="[579,848,1712,1737]" italics="true" pageId="9" pageNumber="10">Physocephala herrerai</emphasis>
</taxonomicName>
(i.e. it is not a rufous species and does not have a pollinose pleural stripe).
</paragraph>
</subSubSection>
</treatment>
</document>