254 lines
31 KiB
XML
254 lines
31 KiB
XML
<document id="F219826BB4735AA632F47C942BBC858C" ID-DOI="10.21248/contrib.entomol.69.1.033-070" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742766" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638269617685" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2019" docId="03EE87A5FFC23B44FCBFFC1473279D1E" docLanguage="en" docName="ContribEntomol.69.1.33-70.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 69 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) shavrini Assing 2019, spec. nov." docType="treatment" docUuid="A7C7B14F-AA8A-47EB-AA6F-F7A7C2B040A4" docUuidSource="ZooBank" docVersion="3" lastPageNumber="68" masterDocId="FFD7FFDDFFE03B67FFC3FFDE77509866" masterDocTitle="On the Lomechusini fauna of the Palaearctic and Oriental regions. XXVI. New species, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="70" masterPageNumber="33" pageNumber="67" updateTime="1699437235520" updateUser="plazi">
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<mods:titleInfo id="45A72DA8C2AAD4336A3198FF466D312E">
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<mods:title id="E78332475DD70F5DD51D79DA25347EAC">On the Lomechusini fauna of the Palaearctic and Oriental regions. XXVI. New species, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="14EF1F382BB2FE28E1C0715EE3BF9008" type="personal">
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<mods:namePart id="FDBAF822E74042FC522EAE8DE5509BE1">Assing, Volker</mods:namePart>
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<mods:typeOfResource id="AA0FEE1CE5BB90DB15754A52888C2202">text</mods:typeOfResource>
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<mods:title id="97B3DF2D4716AA76687CEC78E1DC5044">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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</mods:titleInfo>
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<mods:part id="380D226B8379B710607F2DD64988A970">
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<mods:date id="22376592EC518EF609BEE44A6CD02165">2019</mods:date>
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<mods:number id="67E9E5A65953CB6CA23177BE599735CE">2019-06-24</mods:number>
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</mods:detail>
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<mods:detail id="3534C146B6C86BBE3CB6B07FF5A804EF" type="volume">
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<mods:number id="A83127F265B17FA133058E2ADF89DA85">69</mods:number>
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<mods:start id="9E8B5DBDC490DD977A1C3285F5C6FD71">33</mods:start>
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<mods:end id="80F1C107C5B1C18CD7F7CADFCFA89687">70</mods:end>
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</mods:extent>
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<mods:classification id="098D438E18B27B40110F185685C855CD">journal article</mods:classification>
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<mods:identifier id="F8DB1D148B30DA07BDF5334B9C986E31" type="DOI">10.21248/contrib.entomol.69.1.033-070</mods:identifier>
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<mods:identifier id="44D29AE23F72384BCA331885D0D1591F" type="ISSN">0005-805X</mods:identifier>
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<mods:identifier id="2FF88DCB2D8924254234E845D36DA473" type="Zenodo-Dep">5742766</mods:identifier>
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<treatment id="03EE87A5FFC23B44FCBFFC1473279D1E" LSID="urn:lsid:zoobank.org:act:A7C7B14F-AA8A-47EB-AA6F-F7A7C2B040A4" httpUri="http://treatment.plazi.org/id/03EE87A5FFC23B44FCBFFC1473279D1E" lastPageId="35" lastPageNumber="68" pageId="34" pageNumber="67">
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<subSubSection id="C35D6538FFC23B45FCBFFC1472DC9C63" pageId="34" pageNumber="67" type="nomenclature">
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<paragraph id="8BF836B3FFC23B45FCBFFC1473939B80" blockId="34.[892,1420,970,1061]" box="[892,1219,970,998]" pageId="34" pageNumber="67">
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<heading id="D0B081DFFFC23B45FCBFFC1473939B80" box="[892,1219,970,998]" fontSize="12" level="2" pageId="34" pageNumber="67" reason="2">
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<taxonomicName id="4C474D30FFC23B45FCBFFC1473049B80" ID-CoL="8YCH4" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" box="[892,1108,970,998]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="67" phylum="Arthropoda" rank="species" species="shavrini" status="sp. nov." subGenus="Zyras">
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<emphasis id="B933EAA1FFC23B45FCBFFC1474E29B80" box="[892,946,970,998]" italics="true" pageId="34" pageNumber="67">Zyras</emphasis>
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(
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<emphasis id="B933EAA1FFC23B45FC03FC1474A69B80" box="[960,1014,970,998]" italics="true" pageId="34" pageNumber="67">Zyras</emphasis>
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)
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<emphasis id="B933EAA1FFC23B45FBC7FC1473049B80" box="[1028,1108,970,998]" italics="true" pageId="34" pageNumber="67">shavrini</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20057DAFFC23B45FB99FC1473939B80" box="[1114,1219,970,998]" pageId="34" pageNumber="67" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BF836B3FFC23B45FCBFFC2F72DC9C63" blockId="34.[892,1420,970,1061]" box="[892,1420,1009,1029]" pageId="34" pageNumber="67">
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<uri id="FFD63AB1FFC23B45FCBFFC2F72DC9C63" box="[892,1420,1009,1029]" pageId="34" pageNumber="67">
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urn:lsid:zoobank.org:act:
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<uuid id="FFE10C66FFC23B45FBE8FC2F72DC9C63" box="[1067,1420,1009,1029]" pageId="34" pageNumber="67">A7C7B14F-AA8A-47EB-AA6F-F7A7C2B040A4</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D6538FFC23B45FCBFFBCF73759C43" box="[892,1061,1041,1061]" pageId="34" pageNumber="67" type="description">
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<paragraph id="8BF836B3FFC23B45FCBFFBCF73759C43" blockId="34.[892,1420,970,1061]" box="[892,1061,1041,1061]" pageId="34" pageNumber="67">(Figs 75–78, 163–165)</paragraph>
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</subSubSection>
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<subSubSection id="C35D6538FFC23B45FC95FB9E72099CBE" pageId="34" pageNumber="67" type="materials_examined">
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<paragraph id="8BF836B3FFC23B45FC95FB9E72099CBE" blockId="34.[854,1465,1087,1240]" pageId="34" pageNumber="67">
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<emphasis id="B933EAA1FFC23B45FC95FB9E74A09C31" bold="true" box="[854,1008,1087,1112]" pageId="34" pageNumber="67">Type material</emphasis>
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:
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<materialsCitation id="3B2F3CEEFFC23B45FBC3FB9E72099CBE" collectingDate="2018-03-26" collectingDateMax="2018-03-28" collectingDateMin="2018-03-26" collectionCode="PHILIP-, PINES, V" collectorName="Shavrin" latitude="7.75" longLatPrecision="1303" longitude="125.38333" pageId="34" pageNumber="67" specimenCount="2" typeStatus="holotype">
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<typeStatus id="54FC8811FFC23B45FBC3FB9E73399C3E" box="[1024,1129,1088,1112]" pageId="34" pageNumber="67" type="holotype">Holotype</typeStatus>
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[slightly teneral]: “
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<collectionCode id="ED56AE76FFC23B45FA94FB9E74F19C1E" pageId="34" pageNumber="67">PHILIP- PINES</collectionCode>
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: Mindanao, Barangay Baganihan, Marilog D., Eagles Ridge,
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<geoCoordinate id="EE735074FFC23B45FC3BFB5E73129CFE" box="[1016,1090,1152,1176]" degrees="7" direction="north" minutes="45" orientation="latitude" pageId="34" pageNumber="67" precision="925" value="7.75">7°45'N</geoCoordinate>
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,
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<geoCoordinate id="EE735074FFC23B45FB90FB5E73E29CFE" box="[1107,1202,1152,1176]" degrees="125" direction="east" minutes="23" orientation="longitude" pageId="34" pageNumber="67" precision="925" value="125.38333">125°23'E</geoCoordinate>
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, secondary forest, wet litter sifted,
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<date id="FFF91073FFC23B45FC1EFB7E732B9CDE" box="[989,1147,1184,1208]" pageId="34" pageNumber="67" value="2018-03-26" valueMax="2018-03-28" valueMin="2018-03-26">
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<collectingDate id="EFBDE99BFFC23B45FC1EFB7E732B9CDE" box="[989,1147,1184,1208]" pageId="34" pageNumber="67" value="2018-03-26" valueMax="2018-03-28" valueMin="2018-03-26">26–28.III.2018</collectingDate>
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</date>
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, leg.
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<collectorName id="26B25365FFC23B45FB74FB7E72599CDE" box="[1207,1289,1184,1208]" pageId="34" pageNumber="67">Shavrin</collectorName>
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/
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<typeStatus id="54FC8811FFC23B45FAE6FB7E72CA9CDE" box="[1317,1434,1184,1208]" pageId="34" pageNumber="67" type="holotype">Holotypus</typeStatus>
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<taxonomicName id="4C474D30FFC23B45FC95FB1E74A19CBE" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" box="[854,1009,1216,1240]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="67" phylum="Arthropoda" rank="species" species="shavrini" status="sp. nov.">
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<emphasis id="B933EAA1FFC23B45FC95FB1E74A19CBE" box="[854,1009,1216,1240]" italics="true" pageId="34" pageNumber="67">Zyras shavrini</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20057DAFFC23B45FC32FB1E73799CB1" box="[1009,1065,1216,1239]" pageId="34" pageNumber="67" rank="species">sp. n.</taxonomicNameLabel>
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, det.
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<collectionCode id="ED56AE76FFC23B45FBA1FB1E73249CB1" box="[1122,1140,1216,1239]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="34" pageNumber="67" type="Museum">V</collectionCode>
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. Assing 2018” (cAss).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D6538FFC23B45FC95FADE732B9D3E" pageId="34" pageNumber="67" type="etymology">
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<paragraph id="8BF836B3FFC23B45FC95FADE732B9D3E" blockId="34.[854,1465,1280,1368]" pageId="34" pageNumber="67">
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<emphasis id="B933EAA1FFC23B45FC95FADE749D9D7E" bold="true" box="[854,973,1280,1304]" pageId="34" pageNumber="67">Etymology</emphasis>
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: This species is dedicated to Alexey Shavrin (Daugavpils), specialist of
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<taxonomicName id="4C474D30FFC23B45FB49FAFE72E49D5E" authorityName="MacLeay" authorityYear="1825" box="[1162,1460,1312,1336]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="67" phylum="Arthropoda" rank="subFamily" subFamily="Omaliinae">Omaliinae (Staphylinidae)</taxonomicName>
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, who collected the
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<typeStatus id="54FC8811FFC23B45FBD4FA9E73279D3E" box="[1047,1143,1344,1368]" pageId="34" pageNumber="67" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D6538FFC23B44FC95FAA173A39CBE" lastPageId="35" lastPageNumber="68" pageId="34" pageNumber="67" type="description">
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<paragraph id="8BF836B3FFC23B45FC95FAA174B39E3E" blockId="34.[854,1465,1407,2104]" pageId="34" pageNumber="67">
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<emphasis id="B933EAA1FFC23B45FC95FAA174889DF1" bold="true" box="[854,984,1407,1431]" pageId="34" pageNumber="67">Description</emphasis>
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: Rather larger species: body length
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<quantity id="4CBF9B56FFC23B45FA98FA5E72FF9DF1" box="[1371,1455,1408,1432]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="34" pageNumber="67" unit="mm" value="8.2">8.2 mm</quantity>
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; length of forebody
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<quantity id="4CBF9B56FFC23B45FBE3FA7E73249DD1" box="[1056,1140,1440,1464]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="34" pageNumber="67" unit="mm" value="3.7">3.7 mm</quantity>
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. Habitus as in Fig. 75. Coloration: body pale-reddish, except for the slightly infuscate median portion of tergite VI; legs yellow; antennae with antennomeres I–II reddish, III–IV brown, V yellowishbrown, and VI–XI pale–yellowish; maxillary palpi pale-reddish.
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</paragraph>
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<paragraph id="8BF836B3FFC23B45FC95F9BE73729F11" blockId="34.[854,1465,1407,2104]" pageId="34" pageNumber="67">
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Head (Fig. 76) 1.22 times as broad as long; punctation moderately coarse and dense, median portion impunctate. Eyes large, approximately twice as long as postocular region in dorsal view. Antenna (Fig. 77)
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<quantity id="4CBF9B56FFC23B45FAE2F91E72239EB1" box="[1313,1395,1728,1752]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="34" pageNumber="67" unit="mm" value="2.5">2.5 mm</quantity>
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long; antennomere IV weakly transverse; antennomeres V–X transverse, of gradually increasing width, and increasingly transverse, X slightly less than twice as broad as long, and XI of conical shape, shorter than the combined length of IX and X.
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</paragraph>
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<paragraph id="8BF836B3FFC23B44FC95F85E76019931" blockId="34.[854,1465,1407,2104]" lastBlockId="35.[174,785,256,1304]" lastPageId="35" lastPageNumber="68" pageId="34" pageNumber="67">Pronotum (Fig. 76) approximately 1.1 times as broad as long and 1.2 times as broad as head, broadest near anterior angles, weakly convex in cross-section; lateral margins very weakly sinuate in posterior two-thirds in dorsal view; posterior angles obtusely marked; punctation relatively fine and very dense; impunctate median band narrow; pubescence long, black, and suberect; near lateral and anterior margins with numerous long black and erect setae.</paragraph>
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<paragraph id="8BF836B3FFC33B44FF6DFEBE76929991" blockId="35.[174,785,256,1304]" pageId="35" pageNumber="68">Elytra (Fig. 76) 0.8 times as long as pronotum; punctation rather fine and very dense; pubescence long, black, and suberect. Hind wings fully developed. Legs moderately long and slender; metatarsomere I nearly as long as the combined length of II–IV.</paragraph>
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<paragraph id="8BF836B3FFC33B44FF6DFDDE76E09C31" blockId="35.[174,785,256,1304]" pageId="35" pageNumber="68">Abdomen (Fig. 78) broad, slightly narrower than elytra, and somewhat wedge-shaped; with rather deep anterior impressions on tergites III–V; all sternites and paratergites with long, dense, and suberect to erect pubescence; anterior impressions of tergites III–V each with a transverse row of non-setiferous punctures, with two transverse series of setiferous punctures in posterior portion, and with scattered micropunctation; tergite VI with a transverse band of dense non-setiferous punctures anteriorly, with sparse non-setiferous punctures across the middle, and with setiferous punctures near posterior margin and postero-laterally; tergite VII with a broad transverse band of dense non-setiferous punctures anteriorly and with setiferous punctures in posterior half, this setiferous punctation much denser in postero-lateral portions than in the middle, posterior margin with palisade fringe; tergite VIII with rather dense setiferous punctation in posterior half, posterior margin broadly convex.</paragraph>
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<paragraph id="8BF836B3FFC33B44FF00FBBE76C99C9E" blockId="35.[174,785,256,1304]" pageId="35" pageNumber="68">
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: posterior margin convex, in the middle nearly truncate; median lobe of aedeagus approximately
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<quantity id="4CBF9B56FFC33B44FD45FB5E75879CF1" box="[646,727,1152,1176]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="35" pageNumber="68" unit="mm" value="0.9">0.9 mm</quantity>
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long and shaped as in
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<figureCitation id="137C2A36FFC33B44FEB1FB7E75509CDE" box="[370,512,1184,1208]" captionStart="Figs 145–165" captionStartId="13.[174,214,2001,2023]" captionTargetBox="[176,1415,260,1989]" captionTargetId="figure-118@13.[173,1421,255,1995]" captionTargetPageId="13" captionText="Figs 145–165:Drusilla spiniventris(145–147), D. breviuter(148–152), Pheidologitonetes biplicatus (153–154), Zyras conlectus (155– 157), Z. hetzeli (158–159), Z. reductus(160–162), andZ. shavrini(163–165): male tergite VIII (145, 148); median lobe of aedeagus in lateral and in ventral view (146–147, 149–150, 153–156, 158–161, 163–164); female tergite VIII (151); spermatheca (152); paramere (157, 162, 165). Scale bars: 145–151, 153–165: 0.2 mm; 152: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742777" httpUri="https://zenodo.org/record/5742777/files/figure.png" pageId="35" pageNumber="68">Figs 163–164</figureCitation>
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, ventral process apically long and acute; paramere (
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<figureCitation id="137C2A36FFC33B44FE1BFB1E75609CBE" box="[472,560,1216,1240]" captionStart="Figs 145–165" captionStartId="13.[174,214,2001,2023]" captionTargetBox="[176,1415,260,1989]" captionTargetId="figure-118@13.[173,1421,255,1995]" captionTargetPageId="13" captionText="Figs 145–165:Drusilla spiniventris(145–147), D. breviuter(148–152), Pheidologitonetes biplicatus (153–154), Zyras conlectus (155– 157), Z. hetzeli (158–159), Z. reductus(160–162), andZ. shavrini(163–165): male tergite VIII (145, 148); median lobe of aedeagus in lateral and in ventral view (146–147, 149–150, 153–156, 158–161, 163–164); female tergite VIII (151); spermatheca (152); paramere (157, 162, 165). Scale bars: 145–151, 153–165: 0.2 mm; 152: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742777" httpUri="https://zenodo.org/record/5742777/files/figure.png" pageId="35" pageNumber="68">Fig. 165</figureCitation>
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)
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<quantity id="4CBF9B56FFC33B44FD82FB1E75F09CB1" box="[577,672,1216,1240]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="35" pageNumber="68" unit="mm" value="1.05">1.05 mm</quantity>
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long and with short apical lobe.
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</paragraph>
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<paragraph id="8BF836B3FFC33B44FF00FADE766C9D7E" blockId="35.[174,785,256,1304]" box="[195,316,1280,1304]" pageId="35" pageNumber="68">: unknown.</paragraph>
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<paragraph id="8BF836B3FFC33B44FF6DFAE176F89E9E" blockId="35.[174,785,1343,2104]" pageId="35" pageNumber="68">
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<emphasis id="B933EAA1FFC33B44FF6DFAE176D59D3E" bold="true" box="[174,389,1343,1368]" pageId="35" pageNumber="68">Comparative notes</emphasis>
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: As can be inferred from the long and dense pubescence of the whole body, the morphology of the antennae (antennomere XI short and of conical shape), the robust habitus with a broad and wedge-shaped abdomen, the long and dense pale pubescence of the abdomen, and the morphology of the aedeagus (shape of the ventral process; apical lobe of the paramere very short),
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<taxonomicName id="4C474D30FFC33B44FE32F9FE75359E5E" authorityName="Assing" authorityYear="2019" box="[497,613,1568,1592]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="shavrini">
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<emphasis id="B933EAA1FFC33B44FE32F9FE75359E5E" box="[497,613,1568,1592]" italics="true" pageId="35" pageNumber="68">Z. shavrini</emphasis>
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</taxonomicName>
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belongs to the
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<taxonomicName id="4C474D30FFC33B44FF6DF99E765A9E3E" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[174,266,1600,1624]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="hirtus">
|
||
<emphasis id="B933EAA1FFC33B44FF6DF99E765A9E3E" box="[174,266,1600,1624]" italics="true" pageId="35" pageNumber="68">Z. hirtus</emphasis>
|
||
</taxonomicName>
|
||
group. The new species is distinguished from other similarly coloured species of this group by the shape of the median lobe of the aedeagus (particularly the long and acute apex of the ventral process), by the conspicuously dense punctation of the forebody, and additionally as follows:
|
||
</paragraph>
|
||
<paragraph id="8BF836B3FFC33B44FF6DF8DE750D9F3E" blockId="35.[174,785,1343,2104]" pageId="35" pageNumber="68">
|
||
from
|
||
<taxonomicName id="4C474D30FFC33B44FF2FF8DE76AA9F7E" authority="(KRAATZ, 1859)" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[236,506,1792,1816]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="hirtus">
|
||
<emphasis id="B933EAA1FFC33B44FF2FF8DE76169F7E" box="[236,326,1792,1816]" italics="true" pageId="35" pageNumber="68">Z. hirtus</emphasis>
|
||
(KRAATZ, 1859)
|
||
</taxonomicName>
|
||
(South
|
||
<collectingCountry id="F3507623FFC33B44FD97F8DE75DE9F7E" box="[596,654,1792,1816]" name="India" pageId="35" pageNumber="68">India</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3507623FFC33B44FD5FF8DE74589F7E" box="[668,776,1792,1816]" name="Sri Lanka" pageId="35" pageNumber="68">Sri Lanka</collectingCountry>
|
||
) by slightly larger size, the coloration of the antennae, and the blackish pubescence of the forebody;
|
||
</paragraph>
|
||
<paragraph id="8BF836B3FFC33B44FF6DF8BE767C9051" blockId="35.[174,785,1343,2104]" pageId="35" pageNumber="68">
|
||
from
|
||
<taxonomicName id="4C474D30FFC33B44FF32F8BE76B49F1E" authority="ASSING, 2017" authorityName="ASSING" authorityYear="2017" box="[241,484,1888,1912]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="titan">
|
||
<emphasis id="B933EAA1FFC33B44FF32F8BE766F9F11" box="[241,319,1888,1911]" italics="true" pageId="35" pageNumber="68">Z. titan</emphasis>
|
||
ASSING, 2017
|
||
</taxonomicName>
|
||
(Sulawesi; largest species of the subgenus), by smaller size, the coloration of the antennae, a more transverse pronotum, blackish pubescence of the forebody, shorter and less slender legs, by the punctation pattern of the abdomen (
|
||
<taxonomicName id="4C474D30FFC33B44FD85F83E75C49F91" authorityName="ASSING" authorityYear="2017" box="[582,660,2016,2039]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="titan">
|
||
<emphasis id="B933EAA1FFC33B44FD85F83E75C49F91" box="[582,660,2016,2039]" italics="true" pageId="35" pageNumber="68">Z. titan</emphasis>
|
||
</taxonomicName>
|
||
: setiferous punctures much sparser), and by the shape of the male tergite VIII;
|
||
</paragraph>
|
||
<paragraph id="8BF836B3FFC33B44FCF3FEDE74B299BE" blockId="35.[816,1428,256,1112]" pageId="35" pageNumber="68">
|
||
from
|
||
<taxonomicName id="4C474D30FFC33B44FCBAFEDE7231997E" authority="CAMERON, 1943" authorityName="CAMERON" authorityYear="1943" box="[889,1377,256,280]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="matangensis">
|
||
<emphasis id="B933EAA1FFC33B44FCBAFEDE734A9971" box="[889,1050,256,279]" italics="true" pageId="35" pageNumber="68">Z. matangensis</emphasis>
|
||
CAMERON, 1943 (Borneo)
|
||
</taxonomicName>
|
||
by larger size, the coloration of the antennae, a more transverse pronotum, and a broader, more distinctly wedge-shaped abdomen with denser setiferous punctation, and with distinct non-setiferous punctation in the anterior impressions of tergite III–V (absent in
|
||
<taxonomicName id="4C474D30FFC33B44FCF3FE1E748199B1" authorityName="CAMERON" authorityYear="1943" box="[816,977,448,471]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="matangensis">
|
||
<emphasis id="B933EAA1FFC33B44FCF3FE1E748199B1" box="[816,977,448,471]" italics="true" pageId="35" pageNumber="68">Z. matangensis</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
</paragraph>
|
||
<paragraph id="8BF836B3FFC33B44FCF3FE3E73F39ADE" blockId="35.[816,1428,256,1112]" pageId="35" pageNumber="68">
|
||
from
|
||
<taxonomicName id="4C474D30FFC33B44FCB6FE3E73D9999E" authority="ASSING, 2017" authorityName="ASSING" authorityYear="2017" box="[885,1161,480,504]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="lunatus">
|
||
<emphasis id="B933EAA1FFC33B44FCB6FE3E74B0999E" box="[885,992,480,504]" italics="true" pageId="35" pageNumber="68">Z. lunatus</emphasis>
|
||
ASSING, 2017
|
||
</taxonomicName>
|
||
(Peninsular
|
||
<collectingCountry id="F3507623FFC33B44FAE7FE3E72DC999E" box="[1316,1420,480,504]" name="Malaysia" pageId="35" pageNumber="68">Malaysia</collectingCountry>
|
||
) by a larger and more robust body, the coloration of the antennae, a more transverse pronotum, and an abdomen with distinct non-setiferous punctation in the anterior impressions of tergites III–V, with coarser nonsetiferous punctation on the remaining tergites, and with denser setiferous punctation;
|
||
</paragraph>
|
||
<paragraph id="8BF836B3FFC33B44FCF3FD1E73089BFE" blockId="35.[816,1428,256,1112]" pageId="35" pageNumber="68">
|
||
from
|
||
<taxonomicName id="4C474D30FFC33B44FCA8FD1E73BD9ABE" authority="ASSING, 2017" authorityName="ASSING" authorityYear="2017" box="[875,1261,704,728]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="parahirtus">
|
||
<emphasis id="B933EAA1FFC33B44FCA8FD1E74A49ABE" box="[875,1012,704,728]" italics="true" pageId="35" pageNumber="68">Z. parahirtus</emphasis>
|
||
ASSING, 2017 (Borneo)
|
||
</taxonomicName>
|
||
by paler coloration, a larger and more robust body, the coloration of the abdomen, a less convex (cross-section) and more transverse pronotum, the presence of non-setiferous punctation in the anterior impressions of tergites III–V (indistinct in
|
||
<taxonomicName id="4C474D30FFC33B44FC13FCBE73099B1E" authorityName="ASSING" authorityYear="2017" box="[976,1113,864,888]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="parahirtus">
|
||
<emphasis id="B933EAA1FFC33B44FC13FCBE73099B1E" box="[976,1113,864,888]" italics="true" pageId="35" pageNumber="68">Z. parahirtus</emphasis>
|
||
</taxonomicName>
|
||
), and by denser setiferous punctation of the abdomen;
|
||
</paragraph>
|
||
<paragraph id="8BF836B3FFC33B44FCF3FC7E74FE9C3E" blockId="35.[816,1428,256,1112]" pageId="35" pageNumber="68">
|
||
from
|
||
<taxonomicName id="4C474D30FFC33B44FCADFC7E73E09BDE" authority="CAMERON, 1939" authorityName="CAMERON" authorityYear="1939" box="[878,1200,928,952]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="flavorufus">
|
||
<emphasis id="B933EAA1FFC33B44FCADFC7E74A79BDE" box="[878,1015,928,952]" italics="true" pageId="35" pageNumber="68">Z. flavorufus</emphasis>
|
||
CAMERON, 1939
|
||
</taxonomicName>
|
||
by larger body size, the coloration of the antennae, a less convex pronotum (cross-section), and an abdomen with distinct non-setiferous punctation in the anterior impressions of tergites III–V and with much denser setiferous punctation.
|
||
</paragraph>
|
||
<paragraph id="8BF836B3FFC33B44FCF3FB5E73A39CBE" blockId="35.[816,1427,1152,1240]" pageId="35" pageNumber="68">
|
||
For illustrations and detailed descriptions of
|
||
<taxonomicName id="4C474D30FFC33B44FAEEFB5E72D79CFE" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[1325,1415,1152,1176]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="hirtus">
|
||
<emphasis id="B933EAA1FFC33B44FAEEFB5E72D79CFE" box="[1325,1415,1152,1176]" italics="true" pageId="35" pageNumber="68">Z. hirtus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D30FFC33B44FCF3FB7E742E9CD1" authorityName="ASSING" authorityYear="2017" box="[816,894,1184,1207]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="titan">
|
||
<emphasis id="B933EAA1FFC33B44FCF3FB7E742E9CD1" box="[816,894,1184,1207]" italics="true" pageId="35" pageNumber="68">Z. titan</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D30FFC33B44FC57FB7E73619CD1" authorityName="CAMERON" authorityYear="1943" box="[916,1073,1184,1207]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="matangensis">
|
||
<emphasis id="B933EAA1FFC33B44FC57FB7E73619CD1" box="[916,1073,1184,1207]" italics="true" pageId="35" pageNumber="68">Z. matangensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D30FFC33B44FB85FB7E73E09CDE" authorityName="ASSING" authorityYear="2017" box="[1094,1200,1184,1208]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="lunatus">
|
||
<emphasis id="B933EAA1FFC33B44FB85FB7E73E09CDE" box="[1094,1200,1184,1208]" italics="true" pageId="35" pageNumber="68">Z. lunatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D30FFC33B44FB05FB7E721F9CDE" authorityName="ASSING" authorityYear="2017" box="[1222,1359,1184,1208]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="parahirtus">
|
||
<emphasis id="B933EAA1FFC33B44FB05FB7E721F9CDE" box="[1222,1359,1184,1208]" italics="true" pageId="35" pageNumber="68">Z. parahirtus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C474D30FFC33B44FCF3FB1E74E69CBE" authorityName="CAMERON" authorityYear="1939" box="[816,950,1216,1240]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="68" phylum="Arthropoda" rank="species" species="flavorufus">
|
||
<emphasis id="B933EAA1FFC33B44FCF3FB1E74E69CBE" box="[816,950,1216,1240]" italics="true" pageId="35" pageNumber="68">Z. flavorufus</emphasis>
|
||
</taxonomicName>
|
||
see
|
||
<bibRefCitation id="EFD64B42FFC33B44FC20FB1E73D29CBE" author="ASSING, V." box="[995,1154,1216,1240]" pageId="35" pageNumber="68" pagination="87 - 175" refId="ref29569" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175. - https: // www. zobodat. at / pdf / Stuttgarter-Beitraege-Naturkunde _ NS _ 9 _ A _ 0087 - 0175. pdf." type="journal article" year="2016">ASSING (2016a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD64B42FFC33B44FB4FFB1E739E9CBE" author="ASSING, V." box="[1164,1230,1216,1240]" pageId="35" pageNumber="68" pagination="117 - 192" refId="ref29844" refString="ASSING, V. 2017 b: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192. - https: // www. contributions-to-entomology. org / article / view / 1920 / 1923." type="journal article" year="2017">2017b</bibRefCitation>
|
||
, c).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D6538FFC33B44FCF3FB2173279D1E" pageId="35" pageNumber="68" type="distribution">
|
||
<paragraph id="8BF836B3FFC33B44FCF3FB2173279D1E" blockId="35.[816,1427,1279,1400]" pageId="35" pageNumber="68">
|
||
<emphasis id="B933EAA1FFC33B44FCF3FB2174E99D71" bold="true" box="[816,953,1279,1303]" pageId="35" pageNumber="68">Distribution</emphasis>
|
||
: The type locality is situated in Central Mindanao, South
|
||
<collectingCountry id="F3507623FFC33B44FC37FAFE733F9D5E" box="[1012,1135,1312,1336]" name="Philippines" pageId="35" pageNumber="68">Philippines</collectingCountry>
|
||
. The slightly teneral
|
||
<typeStatus id="54FC8811FFC33B44FA97FAFE740E9D3E" pageId="35" pageNumber="68" type="holotype">holotype</typeStatus>
|
||
was sifted from wet litter near small rocks in a secondary broad-leaved forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |